160 lines
18 KiB
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160 lines
18 KiB
XML
<document ID-CLB-Dataset="9290" ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.60525" ID-GBIF-Dataset="217bea89-5bd1-4ae7-bfce-fb945a933fc4" ID-Pensoft-Pub="2535-0730-2-119" ID-Pensoft-UUID="6301B37BBB7D51A185FB132ED24E4E81" ID-ZooBank="8BD62DBF438C46508C4037C837FD573F" ModsDocID="2535-0730-2-119" checkinTime="1608157809646" checkinUser="pensoft" docAuthor="Ingrisch, Sigfrid" docDate="2020" docId="8B066D77C1C05840BF8F9ECC83471BBB" docLanguage="en" docName="EvolutSyst 4(2): 119-132" docOrigin="Evolutionary Systematics 4 (2)" docPubDate="2020-12-15" docSource="http://dx.doi.org/10.3897/evolsyst.4.60525" docTitle="Pseudosubria assamensis Ingrisch 2020, sp. nov." docType="treatment" docUuid="61582288-A0A6-45DB-990A-79E54E7ED294" docUuidSource="ZooBank" docVersion="5" id="6301B37BBB7D51A185FB132ED24E4E81" lastPageNumber="119" masterDocId="6301B37BBB7D51A185FB132ED24E4E81" masterDocTitle="New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections" masterLastPageNumber="132" masterPageNumber="119" pageNumber="119" updateTime="1732741141311" updateUser="ExternalLinkService">
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<mods:mods id="CC06BAE0D25828FBF33EB86AA44B1C1F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="64B8384055930BB04E2C5FB64F2E3612">
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<mods:title id="517C4EA0BAF10AA047F3B8D353A5C6DB">New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections</mods:title>
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</mods:titleInfo>
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<mods:name id="77B9F2C1F2A469C746125D92F798584E" type="personal">
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<mods:role id="1355BBE9D35C6ABADB171E6717B59263">
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<mods:roleTerm id="97B91C8B195120BFF4C77F6B3A66936D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="039F5F168872D4DDE1E0AC7D4996240D">Ingrisch, Sigfrid</mods:namePart>
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<mods:nameIdentifier id="4D61DF9B1BB768B03E53DFE03B86095A" type="ORCID">https://orcid.org/0000-0002-8624-0472</mods:nameIdentifier>
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<mods:affiliation id="D1B88A6EA9E468271CF77A0B14F8CCE5">Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="3B26DF016E2FA383D61C748ED95FCFB5" type="email">s.ingrisch@leibniz-zfmk.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="2C69958171903C29BA7C6D3F2FFD6250">text</mods:typeOfResource>
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<mods:relatedItem id="14C48D05E7CD082A8A684208BEFBE965" type="host">
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<mods:titleInfo id="F8FBD74F3166119EF94619177FF9CBC8">
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<mods:title id="A83A8E7C59FFC01FB5A0BC419B9B03B8">Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part id="C85DA01198FDEC05666E61DFAC12B2AB">
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<mods:date id="6149E0509053B446CE5C586BA0A9041E">2020</mods:date>
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<mods:detail id="134387BFC3CD70FFCE1D8DB0DECA011B" type="pubDate">
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<mods:number id="F07EB442095986184F32954CA714B375">2020-12-15</mods:number>
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</mods:detail>
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<mods:detail id="0650A64A81042A8A3899A5DA37691CF4" type="volume">
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<mods:number id="FEBA39DBF9B872286F4BFF049D0D13E5">4</mods:number>
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</mods:detail>
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<mods:detail id="E7418A878604090D8C9B4E49AC4B0BD5" type="issue">
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<mods:number id="163DC8906A5718F764B5F3491D262E59">2</mods:number>
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</mods:detail>
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<mods:extent id="5C1C04D07C8BC23BAFE977D35F15ED40" unit="page">
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<mods:start id="2A3C202182B36B2129802B144FD76483">119</mods:start>
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<mods:end id="1D631A581D72E9C63F32928CA3CA77CC">132</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="65F3B421505935DDFC9C4EC978699479">
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<mods:url id="DC50B632C919E0E8362E24F83BC96666">http://dx.doi.org/10.3897/evolsyst.4.60525</mods:url>
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</mods:location>
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<mods:classification id="2C3F37BE9C7F0CCB177D19CACE449AE6">journal article</mods:classification>
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<mods:identifier id="660AED114E7BF9C02100ED4585A902B2" type="DOI">http://dx.doi.org/10.3897/evolsyst.4.60525</mods:identifier>
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<mods:identifier id="70BFDEB7A0C68E1F5181D284839B00CA" type="Pensoft-Pub">2535-0730-2-119</mods:identifier>
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<mods:identifier id="4CE8E8B2D1D56B81AFE5F16DA2FD3973" type="ZooBank">8BD62DBF438C46508C4037C837FD573F</mods:identifier>
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<mods:identifier id="D3710880AC1F3E1DAA161BEE1A87E1D9" type="Pensoft-UUID">6301B37BBB7D51A185FB132ED24E4E81</mods:identifier>
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</mods:mods>
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<treatment id="8B066D77C1C05840BF8F9ECC83471BBB" ID-GBIF-Taxon="176088081" LSID="urn:lsid:zoobank.org:act:61582288-A0A6-45DB-990A-79E54E7ED294" httpUri="http://treatment.plazi.org/id/8B066D77C1C05840BF8F9ECC83471BBB" lastPageNumber="119" pageId="0" pageNumber="119" scope_family="Tettigoniidae" scope_order="Orthoptera" scope_subFamily="Conocephalinae" scope_tribe="Agraeciini">
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<subSubSection id="BCC8FD75EB6B9D46D06A91EB1D9A7433" pageId="0" pageNumber="119" type="nomenclature">
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<paragraph id="9550CA9822E41E579E4109C1FECB9D66" pageId="0" pageNumber="119">
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<taxonomicName id="DC08BE08E4A4E1AEC6BC6D6E5A5C6F26" ID-CoL="8QHWG" LSID="http://zoobank.org/61582288-A0A6-45DB-990A-79E54E7ED294" authority="Ingrisch, 2020" authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Pseudosubria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosubria assamensis" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="assamensis" status="sp. nov.">Pseudosubria assamensis</taxonomicName>
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<taxonomicNameLabel id="84EB86C30052D9EF198673E554C988E3" pageId="0" pageNumber="119">sp. nov.</taxonomicNameLabel>
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<figureCitation id="8787A236E1C3E72F6A23B3405C7A15E0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudosubria assamensis sp. nov. A-E female: A habitus lateral view; B head and pronotum; C ovipositor; D, E subgenital plate in ventral (D) and lateral view (E). - F-N male holotype: F left tegmen; G stridulatory file; H base of left tegmen; I mirror of right tegmen; J subgenital plate in lateral view; K, L cercus in external (K) and internal (L) view; M, N phallus with titillators. Abbeviations: 7 seventh sternite, m membrane, ov base of ovipositor, sg lateral lobes of subgenital plate." httpUri="https://binary.pensoft.net/fig/487063" pageId="0" pageNumber="119">Fig. 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFEC808D64A257FB374C11582B7C33BE" pageId="0" pageNumber="119" type="holotype">
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<paragraph id="317C402E21600E5ABB7E470BFA891759" pageId="0" pageNumber="119">Holotype</paragraph>
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<paragraph id="BDE3B1969052869BD8FD1762B4A42377" pageId="0" pageNumber="119">
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<emphasis id="BD6B01A23639EBDC07421915D589DB1E" bold="true" pageId="0" pageNumber="119">(male).</emphasis>
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India: Assam, Imphal (
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<geoCoordinate id="A9EF27ECB36922A66D77452BDB7DA659" degrees="24" direction="north" minutes="51" orientation="latitude" precision="925" value="24.85">24°51'N</geoCoordinate>
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,
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<geoCoordinate id="F23576DF4F24B5BD291F84F3ED70C94E" degrees="93" direction="east" minutes="54" orientation="longitude" precision="925" value="93.9">93°54'E</geoCoordinate>
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), 22.iii.1946, coll. T.J. Lawrence - depository: BMNH (Lo037S001, B.M.1946-228).
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</paragraph>
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</subSubSection>
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<subSubSection id="CA681301C21FBB7ABDDE09AFB0F0FAAA" pageId="0" pageNumber="119" type="paratype">
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<paragraph id="78DE51683B8E23FA6ED6EA67E5E3BE82" pageId="0" pageNumber="119">Paratype</paragraph>
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<paragraph id="4BB3CBA269D369A58F3FBF277B099483" pageId="0" pageNumber="119">
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<emphasis id="195464C01A419E0284C25114AC859DD5" bold="true" pageId="0" pageNumber="119">(1 female).</emphasis>
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India: Assam, Imphal, 15.iii.1946, coll. T.J. Lawrence - depository: BMNH (Lo037S002, B.M.1946-228).
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</paragraph>
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</subSubSection>
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<subSubSection id="0C93FC20A225386CB38CEDEEA457684F" pageId="0" pageNumber="119" type="etymology">
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<paragraph id="0BA2B06F331AD65F37E273488FBFA35B" pageId="0" pageNumber="119">Etymology.</paragraph>
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<paragraph id="2A1EC6C73F96CEA8240A38115DC44C4A" pageId="0" pageNumber="119">The name of the new species refers to the type locality.</paragraph>
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</subSubSection>
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<subSubSection id="C44916751F906D6B60D1026523EF3EC2" pageId="0" pageNumber="119" type="diagnosis">
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<paragraph id="C3DBAF147180A6591525210DF6894140" pageId="0" pageNumber="119">Diagnosis.</paragraph>
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<paragraph id="78F8ABFF4824A99BC792915399F4F8BA" pageId="0" pageNumber="119">
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The shape of the male cerci is similar to that in
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<taxonomicName id="49657BE649AC3C0EA331AD2843A851F5" lsidName="P. bispinosa subsp. obtusa" pageId="0" pageNumber="119" rank="subSpecies" species="bispinosa" subSpecies="obtusa">
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<emphasis id="6450CD5011BB0F3A429BD2EBD2E6A9F0" italics="true" pageId="0" pageNumber="119">P. bispinosa obtusa</emphasis>
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</taxonomicName>
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Ingrisch, 1998 but has the ventral branch very narrow, regularly upcurved throughout instead of substraight behind the basal bent and reaches the top of the cercus stem (Fig.
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<figureCitation id="BE707B8D5F888FA6A87D75B13DEAB0CA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudosubria assamensis sp. nov. A-E female: A habitus lateral view; B head and pronotum; C ovipositor; D, E subgenital plate in ventral (D) and lateral view (E). - F-N male holotype: F left tegmen; G stridulatory file; H base of left tegmen; I mirror of right tegmen; J subgenital plate in lateral view; K, L cercus in external (K) and internal (L) view; M, N phallus with titillators. Abbeviations: 7 seventh sternite, m membrane, ov base of ovipositor, sg lateral lobes of subgenital plate." httpUri="https://binary.pensoft.net/fig/487063" pageId="0" pageNumber="119">6K-L</figureCitation>
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). The titillators have a rather long fused central area, also as in
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<taxonomicName id="5DE92BEC051AB30DC0E52F2F2CD2AE44" lsidName="P. b. subsp. obtusa" pageId="0" pageNumber="119" rank="subSpecies" species="b." subSpecies="obtusa">
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<emphasis id="3088D7A0D39873E15802E9A8596E9364" italics="true" pageId="0" pageNumber="119">P. b. obtusa</emphasis>
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</taxonomicName>
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, but the free arms are narrower than in the latter species (Fig.
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<figureCitation id="A91F1980932AD5706CABA0399CE3957C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudosubria assamensis sp. nov. A-E female: A habitus lateral view; B head and pronotum; C ovipositor; D, E subgenital plate in ventral (D) and lateral view (E). - F-N male holotype: F left tegmen; G stridulatory file; H base of left tegmen; I mirror of right tegmen; J subgenital plate in lateral view; K, L cercus in external (K) and internal (L) view; M, N phallus with titillators. Abbeviations: 7 seventh sternite, m membrane, ov base of ovipositor, sg lateral lobes of subgenital plate." httpUri="https://binary.pensoft.net/fig/487063" pageId="0" pageNumber="119">6M</figureCitation>
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). The female differs from that of the other species by the shape of the ovipositor that is only little dorso-ventrally widened around mid-length and by the shape of the subgenital plate that is split to the base allowing the base of the ovipositor to project between the lateral lobes (Fig.
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<figureCitation id="493A553EFB2BC5C283C5F14C707B29EF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudosubria assamensis sp. nov. A-E female: A habitus lateral view; B head and pronotum; C ovipositor; D, E subgenital plate in ventral (D) and lateral view (E). - F-N male holotype: F left tegmen; G stridulatory file; H base of left tegmen; I mirror of right tegmen; J subgenital plate in lateral view; K, L cercus in external (K) and internal (L) view; M, N phallus with titillators. Abbeviations: 7 seventh sternite, m membrane, ov base of ovipositor, sg lateral lobes of subgenital plate." httpUri="https://binary.pensoft.net/fig/487063" pageId="0" pageNumber="119">6D</figureCitation>
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).
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</paragraph>
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<caption id="96783FA74096DE4D5CE709A3CED52828" httpUri="https://binary.pensoft.net/fig/487063" pageId="0" pageNumber="119" start="Figure 6" startId="F6">
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<paragraph id="B954635794198588E9E886AE47F47EB8" pageId="0" pageNumber="119">
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<emphasis id="44E397838164319CDA60CD8F34C23126" bold="true" pageId="0" pageNumber="119">Figure 6.</emphasis>
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<taxonomicName id="B3C43F90ED153A1BC668D923CF280627" authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Pseudosubria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosubria assamensis" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="assamensis">
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<emphasis id="6F81F675DEBA4BA887119DEE6FD79A7F" italics="true" pageId="0" pageNumber="119">Pseudosubria assamensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="EED4306E7595557097ACE2B370806A93" bold="true" pageId="0" pageNumber="119">A-E</emphasis>
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female:
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<emphasis id="C1144EA2FFC37FC4389A21142B6156BB" bold="true" pageId="0" pageNumber="119">A</emphasis>
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habitus lateral view;
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<emphasis id="476DADF7E68CC3879F62A28033E09E03" bold="true" pageId="0" pageNumber="119">B</emphasis>
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head and pronotum;
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<emphasis id="7BDDFD516223C8F1E63A044CACBA0BF0" bold="true" pageId="0" pageNumber="119">C</emphasis>
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ovipositor;
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<emphasis id="F6FE619967E9CF91C43D25B7385ADF2F" bold="true" pageId="0" pageNumber="119">D, E</emphasis>
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subgenital plate in ventral (
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<emphasis id="ED5FAF854BA199D03A508F4EC4BD95E9" bold="true" pageId="0" pageNumber="119">D</emphasis>
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) and lateral view (
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<emphasis id="12A553987FE8C22B0ED3D7566544CB2D" bold="true" pageId="0" pageNumber="119">E</emphasis>
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). -
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<emphasis id="1C1E7E40ED71C1C489E5BEDE77231BB2" bold="true" pageId="0" pageNumber="119">F-N</emphasis>
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male holotype:
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<emphasis id="6E9C2F6914AF97009A40DC458D24E340" bold="true" pageId="0" pageNumber="119">F</emphasis>
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left tegmen;
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<emphasis id="5DC129E331E14A54C8F0A959A7E4A77F" bold="true" pageId="0" pageNumber="119">G</emphasis>
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stridulatory file;
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<emphasis id="4520AC70E7B286692862B4650FD7300D" bold="true" pageId="0" pageNumber="119">H</emphasis>
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base of left tegmen;
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<emphasis id="ADCC0B5788E21848B99184C9BEA09EFE" bold="true" pageId="0" pageNumber="119">I</emphasis>
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mirror of right tegmen;
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<emphasis id="A540EAD6F59F733B984E7CF4A58D5844" bold="true" pageId="0" pageNumber="119">J</emphasis>
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subgenital plate in lateral view;
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<emphasis id="64BEE7CD259543759DF0A4DA475A2993" bold="true" pageId="0" pageNumber="119">K, L</emphasis>
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cercus in external (
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<emphasis id="6A87F176E3CCB1D40FDDD1430B77EFBA" bold="true" pageId="0" pageNumber="119">K</emphasis>
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) and internal (
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<emphasis id="30907412F5DC3A3A5DBF0D906970E993" bold="true" pageId="0" pageNumber="119">L</emphasis>
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) view;
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<emphasis id="BE92B9D5E424DD50E95E86C18B4703F9" bold="true" pageId="0" pageNumber="119">M, N</emphasis>
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phallus with titillators. Abbeviations: 7 seventh sternite, m membrane, ov base of ovipositor, sg lateral lobes of subgenital plate.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="6F9E9476DCEA05B8505E77E84DD2AD6A" pageId="0" pageNumber="119" type="description">
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<paragraph id="65897739B8A3AFFAB671C159966C2B42" pageId="0" pageNumber="119">Description.</paragraph>
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<paragraph id="4EEBA64BF9DA43B71CA9E535CD8BFDB7" pageId="0" pageNumber="119">A rather small, slender species. Fastigium verticis conical, obtuse, with a dorsal furrow: separated by a concavity and a narrow seam from fastigium frontis. Frons shining with impressed dots, nearly subrugose. Tegmen reaching or little surpassing apex of stretched hind tibiae. Femora with the following number of spines on ventral margins: profemur 1-3 external, 2-3 internal; mesofemur 3-4 external, no internal; postfemur 5-7 external, no internal. Knee lobes of fore femur obtuse on both sides, on mid femur obtuse on external, acute on internal side, of hind femur obtuse on external, short spinose on internal side.</paragraph>
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<paragraph id="567505CDBF4B13EBE9031D26EE159B04" pageId="0" pageNumber="119">
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<emphasis id="4CD45FEF034FAB465F5F1E68017DAD62" bold="true" pageId="0" pageNumber="119">Male.</emphasis>
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Tenth abdominal tergite prolonged; apical margin in lateral areas nearly straight but converging from both sides; in middle obtuse-angularly excised; with a weak medial carina. Epiproct triangularly rounded with deep medial furrow. Paraprocts with a faint swelling on internal side. Cerci short; external surface convex, at dorso-internal margin with a small triangular expansion; at apex with a large triangular internal expansion covering apex of internal side as a cap; in apical third at ventro-medial margin with a long curved stylate projection curved dorsad behind apex of cercus and narrowing towards apex; internal surface flattened, with a weak fold. Subgenital plate long and narrow; with a medial carina fading towards apex; apical margin slightly concave; styli thin. Titillators X-shaped, fused in middle with basal branches longer than apical branches, for the greatest part hyaline; apex of apical branches darkened, separated by a narrow membranous zone from a dentate apical cap.
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</paragraph>
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<paragraph id="6082AEDB2FA2E0E6F36BEA7B10FA0ACA" pageId="0" pageNumber="119">
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<emphasis id="011B147501123107093EBC178C72D09C" bold="true" pageId="0" pageNumber="119">Female.</emphasis>
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Epiproct triangular with medial furrow. Cerci conical with a long styliform apex. Subgenital plate split to base into two conical, cap-like, lateral lobes with obtuse tip that in situ lie in wide emarginations of the base of the ventral ovipositor valves.
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</paragraph>
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</subSubSection>
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<subSubSection id="DE24AF18F2B6FC5DF0B2B939E7E26899" pageId="0" pageNumber="119" type="coloration">
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<paragraph id="4B5F67DBECB12039A5B19467B860BFDD" pageId="0" pageNumber="119">Coloration.</paragraph>
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<paragraph id="B7182C0ABEA0D0B225FAB083E9F6A3C1" pageId="0" pageNumber="119">Ochre. Head and pronotum with a dark brown medial band.</paragraph>
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</subSubSection>
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<subSubSection id="48F7C0D7022FBD6AFB2BABC6C0010BCE" pageId="0" pageNumber="119" type="description">
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<paragraph id="3256857A5E8F50A8EE056FC031D01329" pageId="0" pageNumber="119">Measurements.</paragraph>
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<paragraph id="B32A0534C9A847E6B62C0573DDA2872B" pageId="0" pageNumber="119">Body w/wings: male 37.5, female 37.5; body w/o wings: male 21, female 19.5; pronotum: male 5.8, female 5.6; tegmen: male 31.3, female 31; hind femur: male 12.5, female 12.5; antenna: female 50; ovipositor: female 12 mm.</paragraph>
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</subSubSection>
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</treatment>
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</document> |