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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.54.475" ID-GBIF-Dataset="74dea868-e511-4276-ba0a-facd3de94406" ID-PMC="PMC3088039" ID-Pensoft-Pub="1313-2970-54-1" ID-PubMed="21594134" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-54-1" ModsDocOrigin="ZooKeys 54" ModsDocTitle="Revision of the Agathidinae (Hymenoptera, Braconidae) of Vietnam, with the description of forty-two new species and three new genera" checkinTime="1451251028578" checkinUser="pensoft" docAuthor="van Achterberg, Cornelis &amp; Long, Khuat Dang" docDate="2010" docId="8A368BCA138BE14653996C571016A68D" docLanguage="en" docName="ZooKeys 54: 1-184" docOrigin="ZooKeys 54" docSource="http://dx.doi.org/10.3897/zookeys.54.475" docTitle="Therophilus contrastus Achterberg &amp; Long, 2010, sp. n." docType="treatment" docVersion="7" lastPageNumber="81" masterDocId="FFCCFF9CFFF5FFE2FF809D194441FFD8" masterDocTitle="Revision of the Agathidinae (Hymenoptera, Braconidae) of Vietnam, with the description of forty-two new species and three new genera" masterLastPageNumber="184" masterPageNumber="1" pageNumber="79" updateTime="1668162270754" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Agathidinae (Hymenoptera, Braconidae) of Vietnam, with the description of forty-two new species and three new genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>van Achterberg, Cornelis</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Long, Khuat Dang</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>184</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.54.475</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.54.475</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-54-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159359717" LSID="urn:lsid:zoobank.org:act:D8721AA6-7169-45A9-B605-97DDFB9B8E58" httpUri="http://treatment.plazi.org/id/8A368BCA138BE14653996C571016A68D" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79">
<subSubSection pageId="78" pageNumber="79" type="nomenclature">
<paragraph pageId="78" pageNumber="79">
<taxonomicName LSID="urn:lsid:zoobank.org:act:D8721AA6-7169-45A9-B605-97DDFB9B8E58" class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus contrastus" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="contrastus">Therophilus contrastus</taxonomicName>
<taxonomicNameLabel pageId="78" pageNumber="79">sp. n.</taxonomicNameLabel>
Figs 238-246
</paragraph>
</subSubSection>
<subSubSection lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79" type="type material">
<paragraph pageId="78" pageNumber="79">Type material.</paragraph>
<paragraph lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
Holotype, ♀ (RMNH), &quot;S. Vietnam: Dong Nai, Cat Tien N.P., c 100 m, 13-19.v.2007, Crocodile tr[ail], Mal. traps, C. v. Achterberg &amp; R. de Vries,
<normalizedToken originalValue="RMNH07”">RMNH'07&quot;</normalizedToken>
. Paratypes (3 ♀): 1 ♀ (IEBR), id. but
<taxonomicName class="Gastropoda" family="Ficidae" genus="Ficus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ficus" order="Littorinimorpha" pageId="78" pageNumber="79" phylum="Mollusca" rank="genus">Ficus</taxonomicName>
trail, 9-30.iv.2007; 1 ♀ (RMNH), &quot;Museum Leiden, Vietnam (Dong Nai Prov.), Cat Tien N.P., Ben Cu trail, 14-20.v.2007, C. van Achterberg, R. de Vries &amp; E.
<normalizedToken originalValue="Gassó">Gasso</normalizedToken>
Miracle&quot;, &quot;mixed bamboo and wood [= deciduous]
<pageBreakToken pageId="79" pageNumber="80" start="start">forest</pageBreakToken>
, by hand, 220 m, 11°26'54.8N; 107°26'30.9E; 1 ♀ (IEBR), Aga. 092, &quot;N. Vietnam: Ha Noi, Gia Lam, orchard, MT, 20-30.iv.2001, K.D. Long&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="diagnosis">
<paragraph pageId="79" pageNumber="80">Diagnosis.</paragraph>
<paragraph pageId="79" pageNumber="80">
The new species is similar to
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus lienhoachihensis" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="lienhoachihensis">Therophilus lienhoachihensis</taxonomicName>
(Chou &amp; Sharkey, 1989), but differs by having the notauli fused posteriorly, forming a large crenulate area (
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus lienhoachihensis" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="lienhoachihensis">Therophilus lienhoachihensis</taxonomicName>
:rugose); the scutellum with sparse fine punctures (lienhoachihensis:rugose-punctate) and the mesonotum reddish yellow (lienhoachihensis:black).
<taxonomicName class="Insecta" family="Braconidae" genus="Bassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassus albifasciatus" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="albifasciatus">Bassus albifasciatus</taxonomicName>
(Watanabe, 1934) is similar, but
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus contrastus" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="contrastus">Therophilus contrastus</taxonomicName>
differs by having the tarsal claws with a large lobe (
<taxonomicName class="Insecta" family="Braconidae" genus="Bassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassus albifasciatus" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="albifasciatus">Bassus albifasciatus</taxonomicName>
: claws without a lobe); outer side of the middle tibia with a row of 4-5 pegs (albifasciatus: 4-10 pegs); the pronotum crenulate anteriorly (albifasciatus: with two carinae) and the head entirely black (albifasciatus: reddish-brown). The new species is also close to
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus festivus" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="festivus">Therophilus festivus</taxonomicName>
(Muesebeck, 1953), but differs by having the first tergite about 1.7 times as long as its apical width (
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus festivus" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="festivus">Therophilus festivus</taxonomicName>
: 1.8-1.9 times); pronotum and mesonotum reddish-yellow (festivus: black)and basal half of hind tibia without a dark brown basal ring (festivus: basal ring usually present).
</paragraph>
</subSubSection>
<subSubSection lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="description">
<paragraph pageId="79" pageNumber="80">Description.</paragraph>
<paragraph pageId="79" pageNumber="80">Holotype, ♀, length of body 5.2 mm, of fore wing 3.6 mm, ovipositor sheath 3.0 mm.</paragraph>
<subSection lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="head">
<paragraph pageId="79" pageNumber="80">Head.</paragraph>
<paragraph lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
Antennal segments 36, length of third segment 1.2 times fourth segment, length of third, fourth and penultimate segments 3.7, 3.1 and 2.0 times their width, respectively; length of apical antennal segment 1.3 times as long as penultimate segment; maxillary palp 2.2 times height of head; malar space 2.7 times as long as basal width of mandible; in dorsal view length of eye 3.5 times temple; temple directly narrowed posteriorly (Fig. 246); POL:OD:OOL= 9:6:11; face with densely punctate
<pageBreakToken pageId="80" pageNumber="81" start="start">;</pageBreakToken>
frons rather dull and distinctly punctate laterally (Fig. 246); vertex and temple with sparse fine punctures.
</paragraph>
</subSection>
<subSection pageId="80" pageNumber="81" type="mesosoma">
<paragraph pageId="80" pageNumber="81">Mesosoma.</paragraph>
<paragraph pageId="80" pageNumber="81">Length of mesosoma 1.5 times its height; subpronope shallow; pronotal trough largely smooth, crenulate anteriorly, dorsally with sparse fine punctures; area near lateral carina of mesoscutum crenulate; mesoscutum with very sparse fine punctures; notauli complete and crenulate; scutellar sulcus 0.6 times as long as dorsal face of scutellum and with 3 carinae; scutellum with sparse punctures, subposterior crest short (Fig. 240); precoxal sulcus complete and largely crenulate (Fig. 239); mesopleuron with sparse distinct punctures; mesopleuron rugose-punctate anteriorly, reticulate-rugose posteriorly; propodeum largely reticulate-rugose; propodeal spiracle small, as long as wide.</paragraph>
</subSection>
<subSection pageId="80" pageNumber="81" type="wings">
<paragraph pageId="80" pageNumber="81">Wings.</paragraph>
<paragraph pageId="80" pageNumber="81">Fore wing: second submarginal cell small (Fig. 242); vein SR1 straight; r:3-SR+SR1 = 3:62. Hind wing: vein M+CU 0.8 times as long as vein 1-M.</paragraph>
</subSection>
<subSection pageId="80" pageNumber="81" type="legs">
<paragraph pageId="80" pageNumber="81">Legs.</paragraph>
<paragraph pageId="80" pageNumber="81">Length of hind femur, tibia and basitarsus 3.8, 6.1 and 9.2 times their width, respectively; hind femur (as remainder of legs) with short setae (Fig. 243); length of outer and inner spur of middle tibia 0.4 and 0.7 times middle basitarsus, respectively; outer side of middle tibia with a row of 3 pegs; length of outer and inner spur of hind tibia 0.3 and 0.5 times hind basitarsus, respectively; outer apex of hind tibia with a cluster of 7 pegs; tarsal claws with a distinct lobe.</paragraph>
</subSection>
<subSection pageId="80" pageNumber="81" type="metasoma">
<paragraph pageId="80" pageNumber="81">Metasoma.</paragraph>
<paragraph pageId="80" pageNumber="81">First tergite longitudinally striate without dorsal carinae; length of first tergite 1.7 times its apical width (Fig. 241); second tergite with weak transverse groove, slightly striate (Fig. 241); remainder of metasoma shiny and smooth; ovipositor sheath 0.8 times as long as fore wing.</paragraph>
</subSection>
<subSection pageId="80" pageNumber="81" type="colour">
<paragraph pageId="80" pageNumber="81">Colour.</paragraph>
<paragraph pageId="80" pageNumber="81">Black; mouthparts, fore and middle legs, hind trochantellus and basal half of hind tibia pale yellow; pronotum, mesonotum, mesopleuron above precoxal sulcus orange brown; second tergite basally, first-second tergites laterally ivory; pterostigma dark brown; wing membrane subhyaline.</paragraph>
</subSection>
<subSection pageId="80" pageNumber="81" type="variation">
<paragraph pageId="80" pageNumber="81">Variation.</paragraph>
<paragraph pageId="80" pageNumber="81">Antenna with 33-36 segments; first tergite 1.7-1.9 times as long as its apical width; second tergite weakly longitudinally striate or smooth; ovipositor sheath 0.8-0.9 times as long as fore wing; length of body 4.9-5.2 mm and of fore wing 3.6-3.8 mm.</paragraph>
</subSection>
</subSubSection>
<subSubSection pageId="80" pageNumber="81" type="distribution">
<paragraph pageId="80" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="80" pageNumber="81">NE Vietnam: Ha Noi and S Vietnam: Dong Nai.</paragraph>
<caption pageId="80" pageNumber="81">
<paragraph pageId="80" pageNumber="81">
Figure 238.
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus contrastus" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="contrastus">Therophilus contrastus</taxonomicName>
sp. n., female, holotype. Habitus lateral.
</paragraph>
</caption>
<caption pageId="80" pageNumber="81">
<paragraph pageId="80" pageNumber="81">
Figures 239-246.
<taxonomicName class="Insecta" family="Braconidae" genus="Therophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Therophilus contrastus" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="contrastus">Therophilus contrastus</taxonomicName>
sp. n., female, holotype. 239 mesosoma lateral 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 wings 243 hind femur lateral 244 head lateral 245 head anterior 246 head dorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="80" pageNumber="81" type="biology">
<paragraph pageId="80" pageNumber="81">Biology.</paragraph>
<paragraph pageId="80" pageNumber="81">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="81" type="etymology">
<paragraph pageId="80" pageNumber="81">Etymology.</paragraph>
<paragraph pageId="80" pageNumber="81">
From
<normalizedToken originalValue="“contra”">&quot;contra&quot;</normalizedToken>
(Latin for
<normalizedToken originalValue="“opposite”">&quot;opposite&quot;</normalizedToken>
), because of the contrasting orange and black body colour.
</paragraph>
</subSubSection>
</treatment>
</document>