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<mods:title id="E4F62F2C2F093FED227041BFE06981B5">Revision of the deep-water spider crab genus, Scyramathia A. Milne-Edwards, 1880, with the description of a new species from the Mediterranean and notes on Rochinia A. Milne-Edwards, 1875, and Anamathia Smith, 1885 (Crustacea, Decapoda, Brachyura, Epialtidae)</mods:title>
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<mods:namePart id="2958B7C8F23B9B029AA2FAC4ADAC9472">Lee, Bee Yan</mods:namePart>
<mods:affiliation id="AF29973FD974F0A02B2ABAA938FCB259">Tropical Marine Science Institute, National University of Singapore, 18 Kent Ridge Road, Singapore 119227, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<mods:namePart id="3E8F3F389CDD454856B45F8E1410A543">Richer De Forges, Bertrand</mods:namePart>
<mods:affiliation id="4B7D6CB85AFD81C9F55690CB7A5DA6C5">c / o Museum National d'Histoire Naturelle, Institut de Systematique, Evolution, Biodiversite, ISYEB - UMR 7205 - CNRS, MNHN, UPMC, EPHE, Departement Systematique et Evolution, 57 rue Cuvier, CP 26, F- 75005, Paris, France</mods:affiliation>
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<treatment id="89D4A77EFCA05B3FA2B9FBA3D6952549" ID-GBIF-Taxon="167407581" LSID="urn:lsid:plazi:treatment:89D4A77EFCA05B3FA2B9FBA3D6952549" httpUri="http://treatment.plazi.org/id/89D4A77EFCA05B3FA2B9FBA3D6952549" lastPageNumber="537" pageId="0" pageNumber="537" scope_family="Epialtidae" scope_infraOrder="Brachyura" scope_order="Decapoda" scope_superFamily="Majoidea">
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<taxonomicName id="F87CD0DD458BB0DAE746025AE00D2ED3" ID-CoL="7YZ87" LSID="89D4A77E-FCA0-5B3F-A2B9-FBA3D6952549" authority="(Norman, in Thomson, 1873)" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">Scyramathia carpenteri (Norman, in Thomson, 1873)</taxonomicName>
<figureCitation id="6F7BE4897053290D828A6E14323ADB2B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">Figs 3</figureCitation>
<figureCitation id="70A582C7985AE83FEFA4A1DC603AB5F4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">, 4A-C</figureCitation>
<figureCitation id="89F891AA0879D9F9F8BDB8CBE29F88EA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">, 5A-C</figureCitation>
<figureCitation id="7753CD17131791198A0264960A5823FD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">, 6A-F</figureCitation>
<figureCitation id="E1DA3C13C8390526ED5E908F7FA451C5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">, 7A-D</figureCitation>
<figureCitation id="65A47D7025AE03290D180E390F7A1075" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, C-H. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); B. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, B. lateral view of carapace; C. ventral view of G 1; D. ventral view of distal portion of G 1; E. dorsal view of G 1; F. dorsal view of distal portion of G 1; G. ventral view of G 1; H. dorsal view of G 2. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48041.figure8" httpUri="https://binary.pensoft.net/fig/447925" pageId="0" pageNumber="537">, 8A-H</figureCitation>
<figureCitation id="8DA448FEF702DB3F42756C41CB8E874B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">, 9A-I</figureCitation>
<figureCitation id="F3E6883757F55A4D7C6F6E378AED1494" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dorsal view of P 5. A, B. Scyramathia carpenteri (Norman, in Thomson 1873); A. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; B. ovigerous female (35.8 x 29.0 mm) (NHM 1978.99), Atlantic Ocean; C, D. S. hertwigi Doflein, in Chun 1900; C. male (34.6 x 27.3 mm) (NHM 1924.7.19.1 - 3), South Africa; D. female (24.5 x 18.4 mm) (SAM-A 47872), South Africa; E, S. umbonata (Stimpson, 1871), female (36.7 x 27.9 mm) (NSMT-Cr 7178), off Suriname; F-J. S. tenuipes sp. nov.; F. holotype male (33.9 x 26.3 mm) (ICMD 288), Mediterranean Sea; G. paratype male (29.7 x 22.0 mm) (ICMD 000707), Mediterranean Sea; H. paratype male (32.0 x 25.4 mm) (ZRC 2019.1636), Mediterranean Sea,; I. paratype ovigerous female (31.2 x 24.6 mm) (ICMD 000708), Mediterranean Sea,; J. paratype ovigerous female (32.2 x 25.8 mm) (ZRC 2019.1635), Mediterranean Sea." figureDoi="10.3897/zse.96.48041.figure14" httpUri="https://binary.pensoft.net/fig/447931" pageId="0" pageNumber="537">, 14A, B</figureCitation>
</paragraph>
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<paragraph id="D64B81AEB29560AE157103EC17E7F894" pageId="0" pageNumber="537">
<taxonomicName id="989B87934665EF23BCE167B3DD64B9C3" class="Malacostraca" family="Epialtidae" genus="Amathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">Amathia carpenteri</taxonomicName>
Norman, in
<bibRefCitation id="227ADC4C55427B4EEEC92C262A3C5060" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
: 175 (fig.), 176, fig. 35 (type locality: &quot;sandy chalk of the Holtenia ground&quot; [= between North Scotland and the Faeroe Islands]).
</paragraph>
<paragraph id="F8CB17DE9F8DAA23F627C57B7761D57F" pageId="0" pageNumber="537">
<taxonomicName id="14431F5780533FF30813187DE6BB95E1" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia Carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="Carpenteri">Scyramathia Carpenteri</taxonomicName>
: A.
<bibRefCitation id="7B0260910E24935D1C8FEAE596353F21" author="Milne-Edwards, A" journalOrPublisher="The European Zoological Journal" pageId="0" pageNumber="537" refId="B72" refString="Milne-Edwards, A, 1880a. Compte rendu sommaire d'une exploration zoologique faite dans le golfe de Gascogne a bord du navire de l'Etat le Travailleur. Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Paris 91(7): 355-360. [23 August 1880 (p. 424 of issue 8 of same volume implies that issue 7 had already been published)]." title="Compte rendu sommaire d'une exploration zoologique faite dans le golfe de Gascogne a bord du navire de l'Etat le Travailleur. Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Paris 91 (7): 355 - 360. [23 August 1880 (p. 424 of issue 8 of same volume implies that issue 7 had already been published)]." year="1880 a">Milne-Edwards 1880a</bibRefCitation>
: 356 [new combination]; A.
<bibRefCitation id="F78DB42DA28BBE4C2040BB4DFC4EB629" author="Milne-Edwards, A" journalOrPublisher="The European Zoological Journal" pageId="0" pageNumber="537" refId="B73" refString="Milne-Edwards, A, 1880b. Rapport sur l'exploration scientifique du Travailleur. Le Naturaliste, Journal des echanges et des nouvelles 2(35): 275-277. [1 September 1880 (p. 273 of same issue)]." title="Rapport sur l'exploration scientifique du Travailleur. Le Naturaliste, Journal des echanges et des nouvelles 2 (35): 275 - 277. [1 September 1880 (p. 273 of same issue)]." year="1880 b">Milne-Edwards 1880b</bibRefCitation>
: 277;
<bibRefCitation id="B8B8DB058E6B37F94F5C4CBE7E25EC97" author="Filhol, H" journalOrPublisher="Biodiversity. Texas A &amp; M University Press, College Station, Texas" pageId="0" pageNumber="537" refId="B37" refString="Filhol, H, 1885. La vie au fond des mers. Les explorations sous-marines et les voyages du Travailleur et du Talisman. G. Masson, Paris, viii + 303 pp. [pls. 1-8]" title="La vie au fond des mers. Les explorations sous-marines et les voyages du Travailleur et du Talisman. G. Masson, Paris, viii + 303 pp. [pls. 1 - 8]" year="1885">Filhol 1885</bibRefCitation>
: 123, 154, fig. 38;
<bibRefCitation id="04DB44B12A1D4DCDAC1978FC800A32C8" author="Sars, GO" journalOrPublisher="Zoologi" pageId="0" pageNumber="537" pagination="1 - 210" refId="B112" refString="Sars, GO, 1885. Crustacea I. Den Norske Nordhavs-Expedition (= The Norwegian North-Atlantic Expedition) 1876-1878. Zoologi 6: 1 - 210" title="Crustacea I. Den Norske Nordhavs-Expedition (= The Norwegian North-Atlantic Expedition) 1876 - 1878." volume="6" year="1885">Sars 1885</bibRefCitation>
: 6-11, 274, pl. 1 figs 1-7;
<bibRefCitation id="1FF54275526DA6FAC1AA29A630F009D1" author="Perrier, E" journalOrPublisher="Hachette, Paris" pageId="0" pageNumber="537" refId="B93" refString="Perrier, E, 1886. Les explorations sous-marines. Hachette, Paris" title="Les explorations sous-marines." year="1886">Perrier 1886</bibRefCitation>
: 208, fig. 217;
<bibRefCitation id="2D56366AFFB5ACC68AFFA9CF28B9DBE0" author="Smith, SI" journalOrPublisher="Crustaceana" pageId="0" pageNumber="537" refId="B117" refString="Smith, SI, 1886. Report on the decapod Crustacea of the Albatross dredgings off the east coast of the United States during the summer and autumn of 1884. Annual Report of the Commissioner of Fish and Fisheries for 1885. Report of the Commission on Fish and Fisheries 13: 605-698 [1-94]. [pls. 1-6]" title="Report on the decapod Crustacea of the Albatross dredgings off the east coast of the United States during the summer and autumn of 1884. Annual Report of the Commissioner of Fish and Fisheries for 1885. Report of the Commission on Fish and Fisheries 13: 605 - 698 [1 - 94]. [pls. 1 - 6]" year="1886">Smith 1886</bibRefCitation>
: 625[21], 626[22];
<bibRefCitation id="B7F8E9C7CDC27636B82F29B52661EF14" DOI="https://doi.org/10.1017/S0025315400072167" author="Bourne, GC" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="537" pagination="306 - 323" refId="B10" refString="Bourne, GC, 1890. Report of a trawling cruise in H.M.S. &quot;Research&quot; off the South-west coast of Ireland. Journal of the Marine Biological Association of the United Kingdom 1 (3): 306 - 323, DOI: https://doi.org/10.1017/S0025315400072167" title="Report of a trawling cruise in H. M. S. &quot; Research &quot; off the South-west coast of Ireland." url="https://doi.org/10.1017/S0025315400072167" volume="1" year="1890">Bourne 1890</bibRefCitation>
: 308; A.
<bibRefCitation id="3804ABB903CBE91A4F2491ADE0873962" author="Milne-Edwards, A" journalOrPublisher="Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" pageId="0" pageNumber="537" pagination="3 - 112" refId="B75" refString="Milne-Edwards, A, Bouvier, E-L, 1894. Crustaces decapodes provenant des campagnes du yacht l'Hirondelle (1886, 1887, 1888). I. Brachyures et Anomoures. Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 7: 3 - 112" title="Crustaces decapodes provenant des campagnes du yacht l'Hirondelle (1886, 1887, 1888). I. Brachyures et Anomoures." volume="7" year="1894">Milne-Edwards and Bouvier 1894</bibRefCitation>
: 13, 14 (in part); A.
<bibRefCitation id="7A3CEB4284105596D671A65EF6F54341" author="Milne-Edwards, A" journalOrPublisher="Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" pageId="0" pageNumber="537" pagination="1 - 106" refId="B76" refString="Milne-Edwards, A, Bouvier, E-L, 1899. Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891-1897). Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 13: 1 - 106" title="Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891 - 1897)." volume="13" year="1899">Milne-Edwards and Bouvier 1899</bibRefCitation>
: 43 (in part);
<bibRefCitation id="E45DF818AAF28C9A7365CDE1782E7633" author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein 1904</bibRefCitation>
: 81 (list); A.
<bibRefCitation id="2FA8778C8D9554EBA6D17A6A1A1A0B70" DOI="https://doi.org/10.5962/bhl.title.11668" author="Milne-Edwards, A" editor="Milne-Edwards, A" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="537" pagination="1 - 396" refId="B77" refString="Milne-Edwards, A, Bouvier, E-L, 1900. Crustaces decapodes. Premiere partie. Brachyures et Anomoures. In: Milne-Edwards, A, Ed., Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883. Masson, Paris: 1 - 396, DOI: https://doi.org/10.5962/bhl.title.11668" title="Crustaces decapodes. Premiere partie. Brachyures et Anomoures." url="https://doi.org/10.5962/bhl.title.11668" volumeTitle="Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883." year="1900">Milne-Edwards and Bouvier 1900</bibRefCitation>
: 133-138, pl. 20 fig. 1-10;
<bibRefCitation id="5925E3DB38602DB29E185C83395B4E3C" author="Hansen, HJ" journalOrPublisher="Danish Ingolf-Expedition" pageId="0" pageNumber="537" pagination="1 - 120" refId="B47" refString="Hansen, HJ, 1908. Crustacea Malacostraca. I. Danish Ingolf-Expedition 3 (2): 1 - 120" title="Crustacea Malacostraca. I." volume="3" year="1908">Hansen 1908</bibRefCitation>
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<bibRefCitation id="355C1D7C85ED80AE60670408EEEE0CB9" author="Bouvier, E-L" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="537" refId="B11" refString="Bouvier, E-L, 1922. Observations complementaires sur les Crustaces decapodes (Abstraction faite des Carides) provenant des Campagnes de S.A.S. le Prince de Monaco. In: Resultats des campagnes scientifiques accomplies sur son yacht par Albert Ier, Prince Souverain de Monaco 62: 106 + [1] pp. [pls. 1-4] [In French]" title="Observations complementaires sur les Crustaces decapodes (Abstraction faite des Carides) provenant des Campagnes de S. A. S. le Prince de Monaco. In: Resultats des campagnes scientifiques accomplies sur son yacht par Albert Ier, Prince Souverain de Monaco 62: 106 + [1] pp. [pls. 1 - 4] [In French]" year="1922">Bouvier 1922</bibRefCitation>
: 81 (in part);
<bibRefCitation id="4B76AF50E738C1847A43846A100DE130" author="Grieg, JA" journalOrPublisher="Bergens Museums Aarbok Naturvidenskabelig raekke" pageId="0" pageNumber="537" pagination="1 - 53" refId="B43" refString="Grieg, JA, 1927. Decapoda Crustacea from the West Coast of Norway and the North Atlantic. Bergens Museums Aarbok Naturvidenskabelig raekke 7: 1 - 53" title="Decapoda Crustacea from the West Coast of Norway and the North Atlantic." volume="7" year="1927">Grieg 1927</bibRefCitation>
: 44-45.
</paragraph>
<paragraph id="5A2DE7BEFE207ABD2BC905E8B8877531" pageId="0" pageNumber="537">
<taxonomicName id="2EE3522A3B95F47F8837EABD13FEFF00" class="Malacostraca" family="Epialtidae" genus="Anamathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anamathia Carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="Carpenteri">Anamathia Carpenteri</taxonomicName>
:
<bibRefCitation id="1A3C31B8C8752C2DAEB87C30C46D1163" author="Smith, SI" journalOrPublisher="Crustaceana" pageId="0" pageNumber="537" refId="B117" refString="Smith, SI, 1886. Report on the decapod Crustacea of the Albatross dredgings off the east coast of the United States during the summer and autumn of 1884. Annual Report of the Commissioner of Fish and Fisheries for 1885. Report of the Commission on Fish and Fisheries 13: 605-698 [1-94]. [pls. 1-6]" title="Report on the decapod Crustacea of the Albatross dredgings off the east coast of the United States during the summer and autumn of 1884. Annual Report of the Commissioner of Fish and Fisheries for 1885. Report of the Commission on Fish and Fisheries 13: 605 - 698 [1 - 94]. [pls. 1 - 6]" year="1886">Smith 1886</bibRefCitation>
: 626[22]; Pocock 1889: 425, 426;
<bibRefCitation id="78B5DCCB56F326CD547447173DB89305" DOI="https://doi.org/10.1017/S0025315400072167" author="Bourne, GC" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="537" pagination="306 - 323" refId="B10" refString="Bourne, GC, 1890. Report of a trawling cruise in H.M.S. &quot;Research&quot; off the South-west coast of Ireland. Journal of the Marine Biological Association of the United Kingdom 1 (3): 306 - 323, DOI: https://doi.org/10.1017/S0025315400072167" title="Report of a trawling cruise in H. M. S. &quot; Research &quot; off the South-west coast of Ireland." url="https://doi.org/10.1017/S0025315400072167" volume="1" year="1890">Bourne 1890</bibRefCitation>
: 314, 315.
</paragraph>
<paragraph id="41BFF255EBE6DC179E6CCDE124646809" pageId="0" pageNumber="537">
<taxonomicName id="C40A547ACD4FAF006EA362F3F67AD677" class="Malacostraca" family="Epialtidae" genus="Anamathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anamathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">Anamathia carpenteri</taxonomicName>
:
<bibRefCitation id="63A5EE9A8E43B974406AC533080EB5B9" author="Faxon, W" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B35" refString="Faxon, W, 1895. The Stalk-Eyed Crustacea. In: Reports on an exploration off the west coasts of Mexico, Central and South America, and off the Galapagos Islands, in charge of Alexander Agassiz, by the U.S. Fish Commission steamer &quot;Albatross,&quot; during 1891, Lieut.-Commander Z. L. Tanner, U.S.N. commanding. XV. Memoirs of the Museum of Comparative Zooelogy at Harvard College XVIII: 1-292. [pls. A-K + I-LVII] [Apr.1895]." title="The Stalk-Eyed Crustacea. In: Reports on an exploration off the west coasts of Mexico, Central and South America, and off the Galapagos Islands, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; during 1891, Lieut. - Commander Z. L. Tanner, U. S. N. commanding. XV. Memoirs of the Museum of Comparative Zooelogy at Harvard College XVIII: 1 - 292. [pls. A-K + I-LVII] [Apr. 1895]." year="1895">Faxon 1895</bibRefCitation>
: 10.
</paragraph>
<paragraph id="00D1BAB256DA8E88FAE0F0684058A1D8" pageId="0" pageNumber="537">
<taxonomicName id="C945734A11CA73B7AF6C2AD3CB3FB31D" baseAuthorityName="C. W. Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Rochinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rochinia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">Rochinia carpenteri</taxonomicName>
:
<bibRefCitation id="8DC5EC2D80071B61C0D25FB366A09BA3" DOI="https://doi.org/10.5479/si.03629236.129.i" author="Rathbun, MJ" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="537" pagination="1 - 613" refId="B101" refString="Rathbun, MJ, 1925. The spider crabs of America. United States National Museum Bulletin 129: 1 - 613, DOI: https://doi.org/10.5479/si.03629236.129.i" title="The spider crabs of America." url="https://doi.org/10.5479/si.03629236.129.i" volume="129" year="1925">Rathbun 1925</bibRefCitation>
: 204 (list) [new combination]; Miranda y Rivera 1933: 41;
<bibRefCitation id="5DAF857DB4F2E562DDBBB0CCEF8B912F" author="Sivertsen, E" journalOrPublisher="Crustaceana" pageId="0" pageNumber="537" refId="B115" refString="Sivertsen, E, Holthuis, LB, 1956. CrustaceaDecapoda (The Penaeidae and Stenopodidae excepted). Report on the Scientific results of the &quot;Michael Sars&quot; North Atlantic Deep-Sea Expedition (1910) 5(12): 1-54. [pls. 1-4]" title="CrustaceaDecapoda (The Penaeidae and Stenopodidae excepted). Report on the Scientific results of the &quot; Michael Sars &quot; North Atlantic Deep-Sea Expedition (1910) 5 (12): 1 - 54. [pls. 1 - 4]" year="1956">Sivertsen and Holthuis 1956</bibRefCitation>
: 49;
<bibRefCitation id="D3A5649961801BBBD90D135C2A6DC8EC" author="Allen, JA" journalOrPublisher="Scientia Marina 66 (supplement 2)" pageId="0" pageNumber="537" refId="B4" refString="Allen, JA, 1967. Crustacea: Euphausiacea and Decapoda with an illustrated key to the Bristish species. The Fauna of the Clyde Sea Area, Bell and Bain Ltd., Glasgow, 1-116 pp." title="Crustacea: Euphausiacea and Decapoda with an illustrated key to the Bristish species. The Fauna of the Clyde Sea Area, Bell and Bain Ltd., Glasgow, 1 - 116 pp." year="1967">Allen 1967</bibRefCitation>
: 35 (list), 64 (key), 97 (fig.);
<bibRefCitation id="E0F0C4A7CC6F1A6D3786DD5D472FD148" DOI="https://doi.org/10.1080/00364827.1968.10411109" author="Sankarankutty, C" journalOrPublisher="Sarsia" pageId="0" pageNumber="537" pagination="43 - 56" refId="B111" refString="Sankarankutty, C, 1968. The first male pleopod in Norwegian Decapoda Brachyura. Sarsia 31: 43 - 56, DOI: https://doi.org/10.1080/00364827.1968.10411109" title="The first male pleopod in Norwegian Decapoda Brachyura." url="https://doi.org/10.1080/00364827.1968.10411109" volume="31" year="1968">Sankarankutty 1968</bibRefCitation>
: 51, figs 5D-F;
<bibRefCitation id="43EC5AB6D30537C357AF46C017703B9E" author="Christiansen, ME" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B18" refString="Christiansen, ME, 1969. CrustaceaDecapodaBrachyura. Marine Invertebrates of Scandinacia number 2. Universitetsforlaget Oslo, Denmark" title="CrustaceaDecapodaBrachyura. Marine Invertebrates of Scandinacia number 2." year="1969">Christiansen 1969</bibRefCitation>
: 20, 122-124, 123 (map), fig. 50;
<bibRefCitation id="9A8768602BD5149E2F5F65C1FBF626DA" author="Ingle, RW" journalOrPublisher="Bulletin of British Museum of Natural History (Zoology)" pageId="0" pageNumber="537" pagination="47 - 66" refId="B51" refString="Ingle, RW, 1979. The larval development of the spider crab Rochinia carpenteri (Thomson) [Oxyrhyncha: Majidae] with a review of majid subfamilial larval features. Bulletin of British Museum of Natural History (Zoology) 37 (1): 47 - 66" title="The larval development of the spider crab Rochinia carpenteri (Thomson) [Oxyrhyncha: Majidae] with a review of majid subfamilial larval features." volume="37" year="1979">Ingle 1979</bibRefCitation>
: 47-53, 59, figs 1-7;
<bibRefCitation id="B18A083E7C110F38E7B3A949E66451C0" author="Ingle, RW" journalOrPublisher="Bulletin of British Museum of Natural History (Zoology)" pageId="0" pageNumber="537" refId="B52" refString="Ingle, RW, 1980. British Crabs. British Museum (Natural History), London, [i] + 222 pp." title="British Crabs. British Museum (Natural History), London, [i] + 222 pp." year="1980">Ingle 1980</bibRefCitation>
: 15 (list), 19 (table), 20 (table), 140, fig. 108, pl. 33 fig. b;
<bibRefCitation id="349E39D27DFBA87BB568D676C4F91D91" DOI="https://doi.org/10.5479/si.00810282.306" author="Manning, RB" journalOrPublisher="Fauna Ryukyuana" pageId="0" pageNumber="537" refId="B63" refString="Manning, RB, Holthuis, LB, 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology 306: i-xii, 1-379. https://doi.org/10.5479/si.00810282.306" title="West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology 306: i-xii, 1 - 379." url="https://doi.org/10.5479/si.00810282.306" year="1981">Manning and Holthuis 1981</bibRefCitation>
: 254;
<bibRefCitation id="D8E98BBBF745C76065B73BDAF23A792A" DOI="https://doi.org/10.1163/156854082X00768" author="Shelton, RGJ" journalOrPublisher="Crustaceana" pageId="0" pageNumber="537" pagination="108 - 110" refId="B114" refString="Shelton, RGJ, Dooley, HD, 1982. New records of the geryonid crab, Geryon affinis Milne Edwards &amp; Bouvier, and other deep water Brachyura from the North East Atlantic. Crustaceana 42 (1): 108 - 110, DOI: https://doi.org/10.1163/156854082X00768" title="New records of the geryonid crab, Geryon affinis Milne Edwards &amp; Bouvier, and other deep water Brachyura from the North East Atlantic." url="https://doi.org/10.1163/156854082X00768" volume="42" year="1982">Shelton and Dooley 1982</bibRefCitation>
: 109;
<bibRefCitation id="9A96844E867508F73A0DE22CCEC4E3DA" DOI="https://doi.org/10.5962/bhl.title.14876" author="Clark, PF" journalOrPublisher="Marine Conservation Society, Englang, U. K." pageId="0" pageNumber="537" refId="B21" refString="Clark, PF, 1986. North-East Atlantic Crabs. An Atlas of Distribution. Marine Conservation Society, Englang, U.K., DOI: https://doi.org/10.5962/bhl.title.14876" title="North-East Atlantic Crabs. An Atlas of Distribution." url="https://doi.org/10.5962/bhl.title.14876" year="1986">Clark 1986</bibRefCitation>
: 190, 191 (map);
<bibRefCitation id="ABCA11EB5EA1B70DD749FF497FB39675" author="Griffin, DJG" journalOrPublisher="Bergens Museums Aarbok Naturvidenskabelig raekke" pageId="0" pageNumber="537" refId="B44" refString="Griffin, DJG, Tranter, HA, 1986. The DecapodaBrachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie 39: [i]-[vii], 1-335." title="The DecapodaBrachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie 39: [i] - [vii], 1 - 335." year="1986">Griffin and Tranter 1986</bibRefCitation>
: 180 (list);
<bibRefCitation id="41C85DB3E10F928D3F62D5EDE84F6888" author="Rice, AL" journalOrPublisher="Bulletin of the British Museum of Natural History (History Series)" pageId="0" pageNumber="537" pagination="1 - 23" refId="B104" refString="Rice, AL, 1990. The nomenclature of crabs collected during the cruises of HMS Porcupine in 1869 and 1870, with notes on the Norman collection of invertebrata and its acquisition by the British Museum (Natural History). Bulletin of the British Museum of Natural History (History Series) 18 (1): 1 - 23" title="The nomenclature of crabs collected during the cruises of HMS Porcupine in 1869 and 1870, with notes on the Norman collection of invertebrata and its acquisition by the British Museum (Natural History)." volume="18" year="1990">Rice 1990</bibRefCitation>
: 10, fig. 7;
<bibRefCitation id="833C546BA48C5A140690796E59F83690" author="Tavares, MS" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="537" refId="B124" refString="Tavares, MS, 1991. Redefinition des genres Rochinia A. Milne Edwards, Sphenocarcinus A. Milne Edwards et Oxypleurodon Miers, et etablissement du genre Nasutocarcinus gen. nov. (Crustacea, Brachyura, Majidae). Bulletin de Museum national d'Histoire naturelle, Section A (Zoology) (Series 4) 13 (1-2): 159-179. [figs 1-6, pl. 1] [In French with and English abstract]." title="Redefinition des genres Rochinia A. Milne Edwards, Sphenocarcinus A. Milne Edwards et Oxypleurodon Miers, et etablissement du genre Nasutocarcinus gen. nov. (Crustacea, Brachyura, Majidae). Bulletin de Museum national d'Histoire naturelle, Section A (Zoology) (Series 4) 13 (1 - 2): 159 - 179. [figs 1 - 6, pl. 1] [In French with and English abstract]." year="1991">Tavares 1991</bibRefCitation>
: 161 (list), 172;
<bibRefCitation id="78FD1AAEE296D8533763A258217A65A8" author="D'Udekem d'Acoz, C" journalOrPublisher="Marine Conservation Society, Englang, U. K." pageId="0" pageNumber="537" refId="B23" refString="D'Udekem d'Acoz, C, 1999. Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25°N. Patrimoines naturels, Museum national d'Histoire naturelle, Paris 40: i-x, 1-394." title="Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Patrimoines naturels, Museum national d'Histoire naturelle, Paris 40: i-x, 1 - 394." year="1999">
<normalizedToken id="A6A8315497E783E86660645A67E6C057" originalValue="dUdekem">d'Udekem</normalizedToken>
<normalizedToken id="FEBD2436804425C7367F136B5631AEAC" originalValue="dAcoz">d'Acoz</normalizedToken>
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</bibRefCitation>
: 194 (in part);
<bibRefCitation id="08BF6AC672965138A360DDCA0D3DA014" DOI="https://doi.org/10.2992/0097-4463(2005)74[151:MFDCBF]2.0.CO;2" author="Casadio, S" journalOrPublisher="Proceedings of the Royal Society of London" pageId="0" pageNumber="537" refId="B16" refString="Casadio, S, Feldmann, RM, Parras, A, Schweitzer, CE, 2005. Miocene fossil Decapoda (Crustacea: Brachyura) from Patagonia, Argentina, and their paleoecological setting. Annals of Carnegie Museum 74(3): 151-188. https://doi.org/10.2992/0097-4463(2005)74[151:MFDCBF]2.0.CO;2" title="Miocene fossil Decapoda (Crustacea: Brachyura) from Patagonia, Argentina, and their paleoecological setting. Annals of Carnegie Museum 74 (3): 151 - 188." url="https://doi.org/10.2992/0097-4463(2005)74[151:MFDCBF]2.0.CO;2" year="2005">
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: 158 (list);
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: 63 (list);
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: 105 (list);
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: 362, fig. 5B.
</paragraph>
<paragraph id="96E4953DCA24363C1565AC4E2E94F7D9" pageId="0" pageNumber="537">
<taxonomicName id="B1BC4C08CF2EC1BB59DBBB3365D6E5F1" baseAuthorityName="C. W. Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Rochinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rochinia Carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="Carpenteri">Rochinia Carpenteri</taxonomicName>
:
<bibRefCitation id="77974D9DDDDFBC4E398241A2D0B25300" author="Bouvier, E-L" journalOrPublisher="Faune de France" pageId="0" pageNumber="537" pagination="1 - 404" refId="B12" refString="Bouvier, E-L, 1940. Decapodes Marcheurs. Faune de France 37: 1 - 404" title="Decapodes Marcheurs." volume="37" year="1940">Bouvier 1940</bibRefCitation>
: 344, 345 (in part).
</paragraph>
<paragraph id="AA92ACB5C44040BE75F388C0BF04E3FC" pageId="0" pageNumber="537">
<taxonomicName id="25BC7692519F057D71BF45118BC4C141" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">Scyramathia carpenteri</taxonomicName>
:
<bibRefCitation id="73FF913BCB88C968CC982C1DEE9EFAFC" DOI="https://doi.org/10.11646/zootaxa.4418.3.1" author="Tavares, MS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="201 - 227" refId="B126" refString="Tavares, MS, Santana, W, 2018. Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae). Zootaxa 4418 (3): 201 - 227, DOI: https://doi.org/10.11646/zootaxa.4418.3.1" title="Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae)." url="https://doi.org/10.11646/zootaxa.4418.3.1" volume="4418" year="2018">Tavares and Santana 2018</bibRefCitation>
: 204 (in part).
</paragraph>
</subSubSection>
<subSubSection id="C4E1FC2F8AB1636316023954408D31FF" pageId="0" pageNumber="537" type="materials_examined">
<paragraph id="8D059546E47D1F2BE818FF121FA73E31" pageId="0" pageNumber="537">Material examined.</paragraph>
<paragraph id="54271149647DD6DC02B42CD7B147061F" pageId="0" pageNumber="537">
<emphasis id="D30676084491ABF016290FF1EC48D44D" bold="true" italics="true" pageId="0" pageNumber="537">Lectotype</emphasis>
: ♂ (21.6
<normalizedToken id="A14465124C28161171719194DBEED421" originalValue="×">x</normalizedToken>
15.9 mm) (NHM 1907.8.28.3), stn 47, 'Holtenia
<normalizedToken id="AC0E025E5C799DDF457244676FD882DC" originalValue="ground">ground'</normalizedToken>
,
<geoCoordinate id="86E26266B68E29F69487B651D36A34B4" degrees="59" direction="north" minutes="34" orientation="latitude" precision="925" value="59.566666">59°34'N</geoCoordinate>
,
<geoCoordinate id="7010D7FB9633AA0DFD0ED38B8E3FCF8D" degrees="7" direction="west" minutes="18" orientation="longitude" precision="925" value="-7.3">7°18'W</geoCoordinate>
, 958 m, coll. H.M.S.
<normalizedToken id="9EE83F5AC95497FCC39C89B3AB76CD9A" originalValue="“Porcupine”">&quot;Porcupine&quot;</normalizedToken>
, 1869. Paralectotypes: 1 ♀ (13.0
<normalizedToken id="7B50835D2FE925D6104BE3FDD08B0058" originalValue="×">x</normalizedToken>
8.4 mm) (NHM 1911.11.8.377), same locality and collection data as lectotype; 1 damaged juvenile ♀ (5.4
<normalizedToken id="5704B5275CE66D2ADBC8D1FD598F55E5" originalValue="×">x</normalizedToken>
3.0 mm) (NHM 1910.2.4.213), stn 48, 'Holtenia
<normalizedToken id="1E531A6B86EE597D6FB89B396341D225" originalValue="ground">ground'</normalizedToken>
<geoCoordinate id="087B15945533D01600C575151326EE1B" degrees="59" direction="north" minutes="32" orientation="latitude" precision="925" value="59.533333">59°32'N</geoCoordinate>
,
<geoCoordinate id="CA6103114516EFBC3C062CFEB14911B9" degrees="6" direction="west" minutes="59" orientation="longitude" precision="925" value="-6.9833336">6°59'W</geoCoordinate>
, coll. H.M.S.
<normalizedToken id="217D0D6380BD3225BCA16F576D4EA894" originalValue="“Porcupine”">&quot;Porcupine&quot;</normalizedToken>
, 1869.
<emphasis id="93AD296706C7E9D0954959759437D162" bold="true" italics="true" pageId="0" pageNumber="537">Other material</emphasis>
: Scotland/North Sea • 1 ♂ (35.0
<normalizedToken id="06E16FF16B35715E4FDAFAE5510CEE0C" originalValue="×">x</normalizedToken>
27.1 mm) (NHM 1981.108), stn 18, Sula Sgeir,
<geoCoordinate id="8ABBD8293B5ABCD1CD0A16E4785EDACA" degrees="59" direction="north" minutes="33.7" orientation="latitude" precision="92" value="59.56167">59°33.7'N</geoCoordinate>
,
<geoCoordinate id="3C303545C38F41271C723026B0E36DE8" degrees="06" direction="west" minutes="49" orientation="longitude" precision="925" value="-6.8166666">06°49'W</geoCoordinate>
to
<geoCoordinate id="40AE5F2225AC32941B95AE7FEBC2C1E4" degrees="59" direction="north" minutes="29.5" orientation="latitude" precision="92" value="59.491665">59°29.5'N</geoCoordinate>
,
<geoCoordinate id="D09049E11A0E7848E98FF15F4A0A2E6C" degrees="06" direction="west" minutes="43.3" orientation="longitude" precision="92" value="-6.721667">06°43.3'W</geoCoordinate>
, 915-880 m, coll. A Wheeler, 16 July 1973; 2 ♂♂ (41.3
<normalizedToken id="2A00BE413A870C7897D324213291C117" originalValue="×">x</normalizedToken>
34.0 mm, 38.8
<normalizedToken id="5E550155036915C6217C55539754B9A8" originalValue="×">x</normalizedToken>
31.6 mm) (NHM 1983.449),
<geoCoordinate id="54395955E502A9DA4A1F375E41AF0253" degrees="56" direction="north" minutes="23" orientation="latitude" precision="925" value="56.383335">56°23'N</geoCoordinate>
,
<geoCoordinate id="6B71DDB43BF475C4DBC63697108EFFCC" degrees="09" direction="west" minutes="18.2" orientation="longitude" precision="92" value="-9.303333">09°18.2'W</geoCoordinate>
, 1010-1030 m, coll. W Wales, 20 October 1977; 7 ♂♂ (37.1
<normalizedToken id="3C797B5B245D2E12A07B7CD45BD3A6D0" originalValue="×">x</normalizedToken>
30.5 mm, 35.0
<normalizedToken id="FF1F4B2622D2B4AADCA9511ED8A486F3" originalValue="×">x</normalizedToken>
27.6 mm, 30.5
<normalizedToken id="F73B69E1CA495FBB7E1A13A1541914D9" originalValue="×">x</normalizedToken>
23.0 mm, 28.2
<normalizedToken id="84C2EF5E0BC6B1D2BB6BEA4A92018311" originalValue="×">x</normalizedToken>
22.7 mm, 24.6
<normalizedToken id="EFD5FA94D41AB4BB00272D0F4F1AE689" originalValue="×">x</normalizedToken>
17.2 mm), 3 ♀♀ (37.1
<normalizedToken id="744FF52A8FE67D28BB6FB1E1BA406046" originalValue="×">x</normalizedToken>
30.6 mm, 33.5
<normalizedToken id="4FFB2A235C5E3DBEA94436CB89784DA6" originalValue="×">x</normalizedToken>
27.5 mm, 22.4
<normalizedToken id="A9F9D10EF41E1B7F131087371D58A59E" originalValue="×">x</normalizedToken>
17.2 mm), 1 ovigerous ♀ (35.8
<normalizedToken id="B80AC04D3FA888B4B65BDE8D417430C6" originalValue="×">x</normalizedToken>
29.0 mm) (NHM 1978.99), Atlantic Ocean,
<geoCoordinate id="C4095CFFC67A282584B913E343185C28" degrees="56" direction="north" minutes="33" orientation="latitude" precision="925" value="56.55">56°33'N</geoCoordinate>
,
<geoCoordinate id="CF81AAB345EEFE04209CFA24306047D6" degrees="09" direction="west" minutes="13" orientation="longitude" precision="925" value="-9.216666">09°13'W</geoCoordinate>
to
<geoCoordinate id="DDADA8B46E518C711312C025117D12B2" degrees="56" direction="north" minutes="30" orientation="latitude" precision="925" value="56.5">56°30'N</geoCoordinate>
,
<geoCoordinate id="20B34D6D4392463A0590A0BF6668B7F5" degrees="09" direction="west" minutes="14" orientation="longitude" precision="925" value="-9.233334">09°14'W</geoCoordinate>
, 750 m, coll. RW Ingle, 23 October 1977. Porcupine Seabight • 3 ♂♂ (25.0
<normalizedToken id="9CD1F2501161A6C63AC257B37CC37948" originalValue="×">x</normalizedToken>
18.9 mm, 15.5
<normalizedToken id="183DEAE0FD0F12A5629BEE14913326E2" originalValue="×">x</normalizedToken>
11.1 mm, 9.0
<normalizedToken id="49AAD2C96B398AB76BE2B23ABDDAFAA7" originalValue="×">x</normalizedToken>
6.2 mm), 1 ♀ (34.8
<normalizedToken id="7178D18E00F9106F695A59309378C526" originalValue="×">x</normalizedToken>
28.2 mm), 1 ♀ (with bopyrid; 15.6
<normalizedToken id="D617C5F5350882F4B5F03B20AF3A811A" originalValue="×">x</normalizedToken>
11.6 mm) (National Oceanography Centre), stn 50601,
<geoCoordinate id="A4C4349C5E881613C11F064AAEDFFC43" degrees="51" direction="north" minutes="19.2" orientation="latitude" precision="92" value="51.32">51°19.2'N</geoCoordinate>
,
<geoCoordinate id="546A9152D53A1661C6D0CF8319DF4FAE" degrees="11" direction="west" minutes="41.1" orientation="longitude" precision="92" value="-11.685">11°41.1'W</geoCoordinate>
-
<geoCoordinate id="B862D106A23CAB14042B4176B2C354AD" degrees="51" direction="north" minutes="21.1" orientation="latitude" precision="92" value="51.351665">51°21.1'N</geoCoordinate>
,
<geoCoordinate id="4046A0D64F32A52FC2DB0F0BBF2C4458" degrees="11" direction="west" minutes="42.9" orientation="longitude" precision="92" value="-11.715">11°42.9'W</geoCoordinate>
, 927-770 m, coll. H.M.S.
<normalizedToken id="AEC5FF8A7326338906E02AEB7BEF33E9" originalValue="“Discovery”">&quot;Discovery&quot;</normalizedToken>
Collection, 1 July 1979; 1 ♂ (23.9
<normalizedToken id="3C97046A7F1D9F9C0B224DB92E2A546E" originalValue="×">x</normalizedToken>
18.4 mm), 3 ♀♀ (17.1
<normalizedToken id="086AC006E1054FA7F9103EB68BF43FE6" originalValue="×">x</normalizedToken>
13.8 mm, 13.6
<normalizedToken id="92E2B0970486D54EE5655875B50BB40C" originalValue="×">x</normalizedToken>
10.6 mm, 12.5
<normalizedToken id="A1A5CA815D42066140F5F2B80CE15C9D" originalValue="×">x</normalizedToken>
9.4 mm) (National Oceanography Centre), stn 9752 #1,
<geoCoordinate id="878B4F4F56D8A2FBBD477C5075286773" degrees="51" direction="north" minutes="16.3" orientation="latitude" precision="92" value="51.271667">51°16.3'N</geoCoordinate>
,
<geoCoordinate id="3895B65A2B1692F81401A89CED468309" degrees="11" direction="west" minutes="42.5" orientation="longitude" precision="92" value="-11.708333">11°42.5'W</geoCoordinate>
-
<geoCoordinate id="E5DAF21E0BF51B0CBA1B781B389166AE" degrees="51" direction="north" minutes="18.6" orientation="latitude" precision="92" value="51.31">51°18.6'N</geoCoordinate>
,
<geoCoordinate id="6F6B2AF8E487A888D0D06A6CCE9AC1AE" degrees="11" direction="west" minutes="42.8" orientation="longitude" precision="92" value="-11.713333">11°42.8'W</geoCoordinate>
, 1010-1045 m, coll. H.M.S.
<normalizedToken id="BB56D1DC52A1DDAD64BEC0969BC241DF" originalValue="“Discovery”">&quot;Discovery&quot;</normalizedToken>
Collection, 7 April 1978.
</paragraph>
</subSubSection>
<subSubSection id="6F6F6EBB91A5426F5DA0952208D95AB4" pageId="0" pageNumber="537" type="diagnosis">
<paragraph id="48E1EBB0323AE6B8E93D2CE49ED7CDF7" pageId="0" pageNumber="537">Diagnosis.</paragraph>
<paragraph id="AB35221DE03D50B58FC03D3419DF649B" pageId="0" pageNumber="537">
Carapace pyriform, strong diverging pseudorostral spines. Postorbital and hepatic spine fused, forming distinct L-shape plate on larger specimens. Carapace with 1 oblong mesogastric plate, 1 oblong cardiac plate, 2 metabranchial plates, 2 horseshoe-shaped epibranchial plates (Figs
<figureCitation id="EBF70196C430B8565C92DE9E61DD41C8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
,
<figureCitation id="FC58C2C4824709E59AE033BE28A731A5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="E0B22B5065D27F4031034170BF1CA9DC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="C67EC3168027CEC239C58728B3462DC1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A, C</figureCitation>
,
<figureCitation id="F41A81B59D295D9D48DBF62CE8B77474" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
). Lateral branchial spine plate-like on specimens with strong carapace plates, present as strong spine on specimens with weak carapace plates (Figs
<figureCitation id="33C584D25FBC2E85523C43B1109A6E76" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
,
<figureCitation id="49A650B410B8B8C6BE365701E320AF8C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="7ADAF5FFAB326F51CE62C23E403FD0F3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="29C438C5791C824B19D7C96B69F41729" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A, C</figureCitation>
,
<figureCitation id="12307E2623A75115914878B3DE8105DF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
). Male thoracic sternum deeply concave with distinct grooves (Figs
<figureCitation id="BDFDD73472E615F92DE0B03BF54D93DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4B</figureCitation>
,
<figureCitation id="615AF8E718898FE12808D8E2455E5FD0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5B</figureCitation>
,
<figureCitation id="6BCB523CC9C7D1C455762AB308A59E78" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6D</figureCitation>
). G1 straight, with flattened, sharp tip; straight top margin (Fig.
<figureCitation id="C18DBAE19CC0EAB3182B0B0F2382797E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, C-H. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); B. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, B. lateral view of carapace; C. ventral view of G 1; D. ventral view of distal portion of G 1; E. dorsal view of G 1; F. dorsal view of distal portion of G 1; G. ventral view of G 1; H. dorsal view of G 2. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48041.figure8" httpUri="https://binary.pensoft.net/fig/447925" pageId="0" pageNumber="537">8C-F</figureCitation>
).
</paragraph>
<caption id="DE45DDED9295B3B7EC8B910A124908A4" doi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537" start="Figure 6" startId="F6">
<paragraph id="BF0465FB15F3CCB01A3D0DAD34383DFE" pageId="0" pageNumber="537">
<emphasis id="14F6CF32C4D351A980C9C3E571E394B8" bold="true" pageId="0" pageNumber="537">Figure 6.</emphasis>
<emphasis id="C70B51A8C7642EF9617F1ED9F3CD729D" bold="true" pageId="0" pageNumber="537">A.</emphasis>
<taxonomicName id="43616D5C980E448E518435F821E9F7AC" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="791D30E1E5EE03517CDE0246C696318F" italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
(Norman, in
<bibRefCitation id="50BACE451B270B3FC9C90CD1302FC59B" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
), male (35.0
<normalizedToken id="5216DBE2EDA4DDC8A63A402DB8DF4CEF" originalValue="×">x</normalizedToken>
27.1 mm) (NHM 1981.108), North Sea;
<emphasis id="3B2C1CC4EE0DF03BB9B4A6F07B9CA55F" bold="true" pageId="0" pageNumber="537">B-F.</emphasis>
<taxonomicName id="82CC1CF07176825DA875E4653B68BCA2" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="4F999D291D2F80E4C753E9F4E62B746E" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
(Norman, in
<bibRefCitation id="3BE7CB9E0A467AE0E91337F23A87A537" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
), male (41.3
<normalizedToken id="928D82F4437F6D45080D5A77932D25DF" originalValue="×">x</normalizedToken>
34.0 mm) (NHM 1983.449), North Sea.
<emphasis id="255E5D969902A6A0E481205485920D9A" bold="true" pageId="0" pageNumber="537">A, B.</emphasis>
lateral view of carapace;
<emphasis id="6FCDFCB845DFF4E508BCF60E4E2D31CD" bold="true" pageId="0" pageNumber="537">C.</emphasis>
ventral frontal surface of carapace;
<emphasis id="CB0F71B5035A21029FCA696E37316C98" bold="true" pageId="0" pageNumber="537">D.</emphasis>
male thoracic sternum view;
<emphasis id="230C382EF8EC436B41DD270D79F315AF" bold="true" pageId="0" pageNumber="537">E.</emphasis>
dorsal surface of male left chela;
<emphasis id="1132F2C3CA7C6F2C1E9A4380E71ED4C9" bold="true" pageId="0" pageNumber="537">F.</emphasis>
posterior ventral view of male pleon.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="CBA491D0CFBD3400595518EC45F8FEA1" pageId="0" pageNumber="537" type="description">
<paragraph id="372A3AA1FD6437562B666810DEE0AAD9" pageId="0" pageNumber="537">Description.</paragraph>
<paragraph id="9F231E51EA2C3A0501F28364EA54490E" pageId="0" pageNumber="537">
Carapace covered by a thick tomentum of setae; with several plates and spines. Pseudorostral spines sharp, straight, diverging, slightly deflected. Supraorbital eave with sharp preorbital spine; sharp angle preorbital spine on juvenile specimens. Hepatic spine pointing upwards, fused with postorbital plate; proportionally longer on juvenile specimens (Figs
<figureCitation id="FCF67897CA0F2B8956DCA85DD1E13AB2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4C</figureCitation>
,
<figureCitation id="A6C8E027723A5F0096A17007ACAC5E60" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5C</figureCitation>
,
<figureCitation id="B380EB8078929148FF696D67259F88CE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6A, B</figureCitation>
,
<figureCitation id="6047EE3E49E432E80786A57405439E90" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, C-H. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); B. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, B. lateral view of carapace; C. ventral view of G 1; D. ventral view of distal portion of G 1; E. dorsal view of G 1; F. dorsal view of distal portion of G 1; G. ventral view of G 1; H. dorsal view of G 2. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48041.figure8" httpUri="https://binary.pensoft.net/fig/447925" pageId="0" pageNumber="537">8A, B</figureCitation>
). Carapace with plate-like structures: 1 oblong mesogastric plate; 1 oblong cardiac plate; 2 metabranchial plates; 2 horseshoe-shaped epibranchial plates (Figs
<figureCitation id="4A1C3C49A1FC53B884248988EDE56142" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
,
<figureCitation id="5C33F182294EA1627AF7829A749609E7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="EA1EB5C11BF425364416E8A7A6087976" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="8B0F563FC61BD0F0635B7715327A1FF6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A, C</figureCitation>
,
<figureCitation id="F19F56C6522675604AF1EBFCF021B99A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
). One long sharp branchial spine pointing laterally; proportionately longer in juveniles and specimens with weak carapace. One large intestinal granule on posterior border of carapace (Figs
<figureCitation id="29FDCE4ADD2371190AEF859A1FBDA8BF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
,
<figureCitation id="5E5DFBB606664098CC217E64E35602C6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="8016010F37531C83D6697D77A34B5D16" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="F9C51C48B0A7FDF5CD482E52B22106D7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A, C</figureCitation>
,
<figureCitation id="DD62866202E6B3FBBF87D82C5BD8A32D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
).
</paragraph>
<caption id="8C79893319D6D77DC87D6814CF0FD462" doi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537" start="Figure 7" startId="F7">
<paragraph id="D720DAA9F7A530AAA116DA49B19906F6" pageId="0" pageNumber="537">
<emphasis id="F4DF60F4FB6261E31391BCF60AC48DDE" bold="true" pageId="0" pageNumber="537">Figure 7.</emphasis>
Illustration of
<taxonomicName id="1215C9FB6A5D881368B5E5D39EE67ACC" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="A7C3AF060E98E225694545227F41C134" italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
(Norman, in
<bibRefCitation id="9CA5D5A035E2A56D4C55897466E416A5" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
).
<emphasis id="3FFACF3C1F418194235AE40DDD8A08E1" bold="true" pageId="0" pageNumber="537">A, B.</emphasis>
lectotype male (21.6
<normalizedToken id="CBBB1973602666ED3D7BE6D43B3E66ED" originalValue="×">x</normalizedToken>
15.9 mm) (NHM 1907.8.28.3), &quot;Holtenia ground&quot; (= between north Scotland and the Faeroe Islands);
<emphasis id="406A719125E164E321E2785CE4B16295" bold="true" pageId="0" pageNumber="537">C, D.</emphasis>
paralectotype female (13.0
<normalizedToken id="E146061D1EA6DCC865B52A223E54F913" originalValue="×">x</normalizedToken>
8.4 mm) (NHM 1911.11.8.377), same locality as lectotype.
<emphasis id="00F5A6729467265231EEF26729073F83" bold="true" pageId="0" pageNumber="537">A, C.</emphasis>
dorsal view of carapace;
<emphasis id="58B967A703E775C4F5AF9FB93532BC27" bold="true" pageId="0" pageNumber="537">B, D.</emphasis>
ventral frontal surface of carapace.
</paragraph>
</caption>
<paragraph id="C9E1C7449917FFAEC359F2472F3A21F2" pageId="0" pageNumber="537">
Antennal flagellum shorter than pseudorostral spines, about half or nearly equal in length. Basal antennal article fused to carapace; longer than broad; straight to gently convex outer margin; blunt distal angle of article. Pterygostomial region with several granules on outer margin (Figs
<figureCitation id="135529EFACDFA38EE4AB1D53496E824D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4B</figureCitation>
,
<figureCitation id="E5A2DE7E31830900A8C51343BB22A0EA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5B</figureCitation>
,
<figureCitation id="51D68F753E12B2A5493750A27213D540" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6C</figureCitation>
,
<figureCitation id="DF69444E09CEA94B1D666DD2514AFA25" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7B, D</figureCitation>
). Buccal frame square, covered by third maxiliped; strong, blunt distal angle of buccal frame (Figs
<figureCitation id="145EC99849BEFCF419AE6616BEC199E3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4B</figureCitation>
,
<figureCitation id="A123605652A2A4B321DBE00075354442" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5B</figureCitation>
,
<figureCitation id="5002862499EDB97F48E088750BDA6360" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6C</figureCitation>
,
<figureCitation id="4E0B8115308DA36DB1F3BEB869D8F355" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7B, D</figureCitation>
).
</paragraph>
<paragraph id="61CF1B499A9493F36963654F67DB37F3" pageId="0" pageNumber="537">
Chelipeds long, slender; propodus longer than dactylus; dactylus curved, serrulated along entire length (Fig.
<figureCitation id="2A64A3AAF71E932635D6A89D360BEB2E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6E</figureCitation>
); carpus short with granules on margin, dorsal margin carinated; merus with 1 tooth on distal upper border. Ambulatory legs slender; merus with sharp angle on distal end; male P2 merus length between 0.6-1.3 times carapace length, female P2 merus length between 0.8-1.0 times carapace length for female, male P2 merus length 8.2-14.0 times width, female P2 merus length 10.5-13.4 times width; male P5 merus length 0.4-0.6 times carapace length, female P5 merus length 0.3-0.5 times carapace length, male P5 merus length 4.2-7.3 times width, female P5 merus length 4.6-5.8 times width (Figs
<figureCitation id="8F72DA64C3F1791C14CFF183B93365ED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
,
<figureCitation id="2155E8E7A8C767D5CF93077AF606F459" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="4687E69F9E32BD5DD7E19AD7CDA3DDED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="B9E7B4B27487010B660F0D48C2E4A8CF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dorsal view of P 5. A, B. Scyramathia carpenteri (Norman, in Thomson 1873); A. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; B. ovigerous female (35.8 x 29.0 mm) (NHM 1978.99), Atlantic Ocean; C, D. S. hertwigi Doflein, in Chun 1900; C. male (34.6 x 27.3 mm) (NHM 1924.7.19.1 - 3), South Africa; D. female (24.5 x 18.4 mm) (SAM-A 47872), South Africa; E, S. umbonata (Stimpson, 1871), female (36.7 x 27.9 mm) (NSMT-Cr 7178), off Suriname; F-J. S. tenuipes sp. nov.; F. holotype male (33.9 x 26.3 mm) (ICMD 288), Mediterranean Sea; G. paratype male (29.7 x 22.0 mm) (ICMD 000707), Mediterranean Sea; H. paratype male (32.0 x 25.4 mm) (ZRC 2019.1636), Mediterranean Sea,; I. paratype ovigerous female (31.2 x 24.6 mm) (ICMD 000708), Mediterranean Sea,; J. paratype ovigerous female (32.2 x 25.8 mm) (ZRC 2019.1635), Mediterranean Sea." figureDoi="10.3897/zse.96.48041.figure14" httpUri="https://binary.pensoft.net/fig/447931" pageId="0" pageNumber="537">14A, B</figureCitation>
).
</paragraph>
<paragraph id="0B8A2C5B3759020EA47953A83C1AC748" pageId="0" pageNumber="537">
Male thoracic sternum with sternites 1-4 deep concave; sternites 1, 2 wider than long; sternites 3, 4 widest at base, lateral margin slightly constricted; sternites strongly defined (Figs
<figureCitation id="8C2CB4C52A4E38AD8E5C39C4B9242D55" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4B</figureCitation>
,
<figureCitation id="4971567A246F7EDD0A8525ABC22343A7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5B</figureCitation>
,
<figureCitation id="87874BBEDF33602DB472B59CE72F4C98" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6D, F</figureCitation>
,). Male pleon with triangular telson and 6 somites, all free; widest on second and third somites. Adult female with round pleon, with all somites distinct. G1 straight, with flattened, sharp distal tip; straight top margin; slightly constricted at distal region (Fig.
<figureCitation id="72E1DE7866F09417B8CA38C105C796DC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, C-H. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); B. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, B. lateral view of carapace; C. ventral view of G 1; D. ventral view of distal portion of G 1; E. dorsal view of G 1; F. dorsal view of distal portion of G 1; G. ventral view of G 1; H. dorsal view of G 2. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48041.figure8" httpUri="https://binary.pensoft.net/fig/447925" pageId="0" pageNumber="537">8C-F</figureCitation>
).
</paragraph>
<caption id="260F9455CF81CAE706927AE4E50027E3" doi="10.3897/zse.96.48041.figure8" httpUri="https://binary.pensoft.net/fig/447925" pageId="0" pageNumber="537" start="Figure 8" startId="F8">
<paragraph id="9B141D7D3A62B130244B102D2CE8C8FB" pageId="0" pageNumber="537">
<emphasis id="913CB9DC23D276A5D4622CD95536FA12" bold="true" pageId="0" pageNumber="537">Figure 8.</emphasis>
Illustration of
<taxonomicName id="2F69A2BECDC4F35912A06CEDF29D0ADC" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="44413B55B6E8A89287435625BBA71024" italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
(Norman, in
<bibRefCitation id="94AD9087C7AB79E29E1E3B221D46EEBA" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
).
<emphasis id="C3ABE7395BCBF289AC499D9AC5555AAF" bold="true" pageId="0" pageNumber="537">A, C-H.</emphasis>
lectotype male (21.6
<normalizedToken id="5B33B6B4E611C89B5774518154B30D65" originalValue="×">x</normalizedToken>
15.9 mm) (NHM 1907.8.28.3), &quot;Holtenia ground&quot; (= between north Scotland and the Faeroe Islands);
<emphasis id="9355300337AFFEB61635F46DBA0847C7" bold="true" pageId="0" pageNumber="537">B.</emphasis>
paralectotype female (13.0
<normalizedToken id="21557899CBC9CBF1AA6DE35FEA280332" originalValue="×">x</normalizedToken>
8.4 mm) (NHM 1911.11.8.377), same locality as lectotype.
<emphasis id="16DEB9AD97CA6D4F221053A5E5196E6C" bold="true" pageId="0" pageNumber="537">A, B.</emphasis>
lateral view of carapace;
<emphasis id="FE5FBE8391AA87D78170C478CB488F5C" bold="true" pageId="0" pageNumber="537">C.</emphasis>
ventral view of G1;
<emphasis id="78B8A1B8E2715B33753B03DD6D1EBC9A" bold="true" pageId="0" pageNumber="537">D.</emphasis>
ventral view of distal portion of G1;
<emphasis id="1C7235F34EE4DC76657D665725752E6D" bold="true" pageId="0" pageNumber="537">E.</emphasis>
dorsal view of G1;
<emphasis id="E819E0786E5D1D9243D34FF9621CECED" bold="true" pageId="0" pageNumber="537">F.</emphasis>
dorsal view of distal portion of G1;
<emphasis id="38CABD6FD231991A814BD4CD6F595DF7" bold="true" pageId="0" pageNumber="537">G.</emphasis>
ventral view of G1;
<emphasis id="EC0F23365B689784DADCA33233303680" bold="true" pageId="0" pageNumber="537">H.</emphasis>
dorsal view of G2. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C03003CD38C2D388C1D74ED0A1AA65A7" pageId="0" pageNumber="537" type="remarks">
<paragraph id="6C87B4BB777BCE9BAFF57032E7894EF5" pageId="0" pageNumber="537">Remarks.</paragraph>
<paragraph id="D9A09701493CB99648DFD22DC51C10B3" pageId="0" pageNumber="537">
The identity of
<taxonomicName id="AB8D15509A18CFF2C67EB93497F34DED" class="Malacostraca" family="Epialtidae" genus="Amathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="5719F1FD222C710EC55E139F5720A247" italics="true" pageId="0" pageNumber="537">Amathia carpenteri</emphasis>
</taxonomicName>
Norman, in
<bibRefCitation id="7761F9BECB15CF758225596D05166D23" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
s. str., is critical to the status of
<taxonomicName id="AE2C25A17BFA9B1C96D001DB13CDA0BB" authorityName="A. Milne-Edwards" authorityYear="1880" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="302AE1EFD6FE92F9D35F39D88EDEC2E1" italics="true" pageId="0" pageNumber="537">Scyramathia</emphasis>
</taxonomicName>
as it is the type species of the genus. Although
<bibRefCitation id="BFDFD0ABEB681412872A21E64CF7C3E2" DOI="https://doi.org/10.11646/zootaxa.4418.3.1" author="Tavares, MS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="201 - 227" refId="B126" refString="Tavares, MS, Santana, W, 2018. Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae). Zootaxa 4418 (3): 201 - 227, DOI: https://doi.org/10.11646/zootaxa.4418.3.1" title="Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae)." url="https://doi.org/10.11646/zootaxa.4418.3.1" volume="4418" year="2018">Tavares and Santana (2018)</bibRefCitation>
clarified the identity of the genus and had material of the species, they did not examine the types for this species. The species was named by Thomson in 1873 in his famous book about the exploration of the deep Atlantic by the H.M.S. Porcupine, &quot;The Depths of the Seas&quot; (p. 175: fig. 35; reproduced here as Fig.
<figureCitation id="1BC7EB2F0B251A101ABE480815B2C6D9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
), under the name &quot;
<taxonomicName id="523BF6F18B8D872F4E6ADEE0AA6E3C7B" class="Malacostraca" family="Epialtidae" genus="Amathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amathia Carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="Carpenteri">
<emphasis id="6DDFEB2EFE2A4D7482917C49667FC667" italics="true" pageId="0" pageNumber="537">Amathia Carpenteri</emphasis>
</taxonomicName>
&quot; to honour his colleague Dr. W.B. Carpenter. There was no clear indication of number of specimens examined, with the author only commenting that &quot;Another handsome new species,
<taxonomicName id="1DA1BC26BDA1E5780CCE105BF3F15434" class="Malacostraca" family="Epialtidae" genus="Amathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="FA44043952A6BFAAF383101029B80E06" italics="true" pageId="0" pageNumber="537">Amathia carpenteri</emphasis>
</taxonomicName>
, Norman (fig. 35), was common in the sandy chalkmud of the Holtenia ground.&quot; (
<bibRefCitation id="8566805914E3C9BE8EE9587191CEBF77" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
: 176). The figure shows that the cardiac region of the carapace appeared to be damaged (
<bibRefCitation id="DAB4BAB6A7F193F9694FE5A4660C589A" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
: fig. 35; reproduced here as Fig.
<figureCitation id="AFBD0315C0F2E79006F83E4A1AA8976F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
). In the NHM are three specimens that are labelled as syntypes of
<taxonomicName id="BA0208E0BF632A9F73526197F7E90F93" lsidName="A. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="41BF122684C7D1E7F385A6A5116806EF" italics="true" pageId="0" pageNumber="537">A. carpenteri</emphasis>
</taxonomicName>
, all obtained by the H.M.S. Porcupine. None of these specimens, however, match the measurements of the type figure, which was &quot;once and a half the natural size&quot; (
<bibRefCitation id="74427495AE1A6D19DF14B43F876EB01B" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
: fig. 35; reproduced here as Fig.
<figureCitation id="E548F47660489651FC771BD205B4A6F8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
). Of these, one male specimen (NHM 1907.8.28.3) agrees relatively well with the figure by
<bibRefCitation id="2C443E2A9DBE50898C776EE71DF40846" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson (1873</bibRefCitation>
: fig. 35; Fig.
<figureCitation id="F8328BB060756826411D5835A97F6203" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
), and the specimen label mentions that it was figured in
<bibRefCitation id="0541A671B45F062246749B6312FF3A4B" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson (1873)</bibRefCitation>
. This specimen, with catalogue number, NHM 1907.8.28.3, was noted by
<bibRefCitation id="29992FB3FEE2211A2022F3CCA20D55D6" author="Christiansen, ME" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B18" refString="Christiansen, ME, 1969. CrustaceaDecapodaBrachyura. Marine Invertebrates of Scandinacia number 2. Universitetsforlaget Oslo, Denmark" title="CrustaceaDecapodaBrachyura. Marine Invertebrates of Scandinacia number 2." year="1969">Christiansen (1969</bibRefCitation>
: 122) to be the lectotype for this species, and in an unpublished note, was also selected by Isabella Gordon (also unpublished data) to be the lectotype. The same specimen was also mentioned by
<bibRefCitation id="DE2B905451D1BBD734CBFC53712497C1" author="Rice, AL" journalOrPublisher="Bulletin of the British Museum of Natural History (History Series)" pageId="0" pageNumber="537" pagination="1 - 23" refId="B104" refString="Rice, AL, 1990. The nomenclature of crabs collected during the cruises of HMS Porcupine in 1869 and 1870, with notes on the Norman collection of invertebrata and its acquisition by the British Museum (Natural History). Bulletin of the British Museum of Natural History (History Series) 18 (1): 1 - 23" title="The nomenclature of crabs collected during the cruises of HMS Porcupine in 1869 and 1870, with notes on the Norman collection of invertebrata and its acquisition by the British Museum (Natural History)." volume="18" year="1990">Rice (1990</bibRefCitation>
: 10), noting that it is similar to the figure by Norman, in
<bibRefCitation id="69868ABA74BC58B74B47AEC753EBAEA7" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson (1873</bibRefCitation>
: fig. 35). The type locality of this species was stated to be on &quot;sandy chalk of the Holtenia ground&quot;. Holtenia Ground is an area between North Scotland and the Faeroe Islands that covers an area of approximately 500 km2, with a depth of 820 to 1000 m (
<bibRefCitation id="D0F02AFC179FCCACE0269E9900007A59" DOI="https://doi.org/10.1098/rspl.1869.0084" author="Carpenter, WB" journalOrPublisher="Proceedings of the Royal Society of London" pageId="0" pageNumber="537" pagination="397 - 492" refId="B14" refString="Carpenter, WB, Gwyn Jeffreys, J, Thomson, CW, 1870. Preliminary report of the scientific exploration of the deep sea in H.M. Surveying-vessel 'Porcupine,' during the summer of 1869. Proceedings of the Royal Society of London 18: 397 - 492, DOI: https://doi.org/10.1098/rspl.1869.0084" title="Preliminary report of the scientific exploration of the deep sea in H. M. Surveying-vessel ' Porcupine, ' during the summer of 1869." url="https://doi.org/10.1098/rspl.1869.0084" volume="18" year="1870">Carpenter et al. 1870</bibRefCitation>
;
<bibRefCitation id="3A4E7AB9324487A7E40B0FC91EBB74B4" DOI="https://doi.org/10.1111/j.1096-3642.1995.tb01430.x" author="Reiswig, HM" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="537" pagination="373 - 384" refId="B102" refString="Reiswig, HM, Champagne, P, 1995. The NE Atlantic glass sponges Pheronema carpenteri (Thomson) and P. grayi Kent (Porifera: Hexactinellida) are synonyms. Zoological Journal of the Linnean Society 115: 373 - 384, DOI: https://doi.org/10.1111/j.1096-3642.1995.tb01430.x" title="The NE Atlantic glass sponges Pheronema carpenteri (Thomson) and P. grayi Kent (Porifera: Hexactinellida) are synonyms." url="https://doi.org/10.1111/j.1096-3642.1995.tb01430.x" volume="115" year="1995">Reiswig and Champagne 1995</bibRefCitation>
). Specimen NHM1907.8.28.3 is here formally designated as the lectotype of
<taxonomicName id="0AEA00F08732070865B387D7DF33A7BE" class="Malacostraca" family="Epialtidae" genus="Amathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="779CEF1AEBF37BB125D670608A966F68" italics="true" pageId="0" pageNumber="537">Amathia carpenteri</emphasis>
</taxonomicName>
Norman, in
<bibRefCitation id="7010CD98AAFFE565D64A1438A1A886CD" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
, to stabilise the taxonomy of the species and the genus.
</paragraph>
<paragraph id="1FE268354F0F1E247CB7BAC8B455C93F" pageId="0" pageNumber="537">
The three type specimens differ markedly from each other in many ways although they are all from Holtenia Ground. The lectotype male (21.6
<normalizedToken id="838AA005B1942D2418E23CF3D3343275" originalValue="×">x</normalizedToken>
15.9 mm, NHM 1907.8.28.3) selected here differs markedly from the other two paralectotypes in having a smoother carapace, without the distinct raised plate-like structures on the carapace and the branchial regions are also proportionately more inflated (Figs
<figureCitation id="F81714254AB28AB17EF2FD558980D456" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="88A126FB60EAE0D925763F2AEFDA0E08" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A</figureCitation>
). The next largest type is a subadult female (13.0
<normalizedToken id="AA3F4DE5DF597DA2B689BAD6417CA5F8" originalValue="×">x</normalizedToken>
8.4 mm, NHM 1911.11.8.377), and its carapace more closely resembles typical
<taxonomicName id="B1AE83D76A1E14CD28B32B847DD18657" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="FA5596DA5A81BC0FF06AE2A464188329" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
, with the plates more developed. It is almost certainly the same specimen figured by Norman, in
<bibRefCitation id="B18C23B05C8B0B088D3441898AF48E69" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson (1873</bibRefCitation>
: fig. 35; Fig.
<figureCitation id="1A8BF7A70EC72B1E6B6B793AA8E037AC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873), &quot; sandy chalk of Holtenia ground &quot;, overall dorsal view. (After Norman, in Thomson 1873: fig. 35)." figureDoi="10.3897/zse.96.48041.figure3" httpUri="https://binary.pensoft.net/fig/447920" pageId="0" pageNumber="537">3</figureCitation>
), with the shape and structures of the spines and plates agreeing very well, althought the left pseudorostral spine of the specimen is now broken (Fig.
<figureCitation id="1F27A1EE1932C750D7060B3028B264FA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7C</figureCitation>
). The third type specimen is a small and poorly preserved juvenile female (5.4
<normalizedToken id="6ECAC3A0D6058BF3D0CCFDE929A0F6A4" originalValue="×">x</normalizedToken>
3.0 mm, NHM 1910.2.4.213) in which the dorsal surface of the carapace as well as the third maxillipeds are covered with setae but without any obvious plates. The surfaces of the third maxillipeds of the other two type specimens are finely granulated or smooth (Figs
<figureCitation id="540690F248B3C8E6D3CC0D7D28B2CF2D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4B</figureCitation>
,
<figureCitation id="EF2C258B04887D4CA87C0063D0917B33" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7B, D</figureCitation>
).
</paragraph>
<paragraph id="59F674172F29853DC66D921367A0F6AD" pageId="0" pageNumber="537">
<taxonomicName id="7B8906E92FAA1089C65D0F06EBD78F75" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="0F003C6A927E4346C189F6C245F1A171" italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
exhibits strong variation on the plate-like structures on the carapace of various sized specimens showed variation.
<bibRefCitation id="26E2B76EF9DE4FAD666EC6730D018CA1" author="D'Udekem d'Acoz, C" journalOrPublisher="Marine Conservation Society, Englang, U. K." pageId="0" pageNumber="537" refId="B23" refString="D'Udekem d'Acoz, C, 1999. Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25°N. Patrimoines naturels, Museum national d'Histoire naturelle, Paris 40: i-x, 1-394." title="Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Patrimoines naturels, Museum national d'Histoire naturelle, Paris 40: i-x, 1 - 394." year="1999">
<normalizedToken id="AC147372070473A36AAF32AF786002EA" originalValue="DUdekem">D'Udekem</normalizedToken>
<normalizedToken id="99CEC0DB7602A3FE522E04D7A42A1F51" originalValue="dAcoz">d'Acoz</normalizedToken>
(1999)
</bibRefCitation>
observed that
<normalizedToken id="BC75CE383F3D7FD11BC2B983EB432DE0" originalValue="“Lornementation">&quot;L'ornementation</normalizedToken>
de la carapace de
<taxonomicName id="BE46BD55A2980D5CFFDEC5AB124E82EC" lsidName="R. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="0820B77B0EC4FFF0E1A20E3F3BD35471" italics="true" pageId="0" pageNumber="537">R. carpenteri</emphasis>
</taxonomicName>
est
<normalizedToken id="8742228BD26CC02BECCC2DD0E2A09894" originalValue="extrêmement">extremement</normalizedToken>
variable&quot; but did not mention if it is due to the size of the specimens. Similar sized specimens showed either weak or strong plate-like structures (Fig.
<figureCitation id="2113CC68F75E5748B39E1AE6AA328B34" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
). These differences are also seen in the type specimens where the larger sized male lectotype specimen has weak plate-like structures on the carapace whereas the same structures on the smaller sized female paralectotype specimens are relatively stronger. The variation of the plate-like structures on the carapace are therefore, not always size dependent. Other morphological characters that show variation on
<taxonomicName id="AE70A90A27C84C34878EB81FE2EEB8B6" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="FE7118F395948813D50E33487CE60930" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
includes the divergence of the pseudorostral spines, from relatively divergent as seen on the lectotype (Fig.
<figureCitation id="6D6DFDA914F4F9C0D22DAF2A7611AABB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
) to subparallel as seen on the male specimen, with catalogue number, NHM 1983.449 (Fig.
<figureCitation id="A8D3C864D9B0ECC3D661C14CCC9B8F51" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
); the hepatic region having a hepatic spine or plate (Figs
<figureCitation id="E51191E53FD2B1BAA3E61B51EBC8B80E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="21150AB9C3B6C053D9A05ABA0D51656E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="417F19D1902261CBFA4770DF6A9E2978" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A, C</figureCitation>
,
<figureCitation id="A0E3E57572E07580A773ADA66101077D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
); and the lateral branchial spine on the carapace, from relatively straight, slightly curved or plate-like (Figs
<figureCitation id="35A05F97229468962942EB99AD51C909" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scyramathia carpenteri (Norman, in Thomson 1873), lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands). A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure4" httpUri="https://binary.pensoft.net/fig/447921" pageId="0" pageNumber="537">4A</figureCitation>
,
<figureCitation id="91DC4CC8031258AD0D34B0502F47D481" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scyramathia carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure5" httpUri="https://binary.pensoft.net/fig/447922" pageId="0" pageNumber="537">5A</figureCitation>
,
<figureCitation id="50518E822DCBACC09910B967814E1947" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Illustration of Scyramathia carpenteri (Norman, in Thomson 1873). A, B. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot; (= between north Scotland and the Faeroe Islands); C, D. paralectotype female (13.0 x 8.4 mm) (NHM 1911.11.8.377), same locality as lectotype. A, C. dorsal view of carapace; B, D. ventral frontal surface of carapace." figureDoi="10.3897/zse.96.48041.figure7" httpUri="https://binary.pensoft.net/fig/447924" pageId="0" pageNumber="537">7A, C</figureCitation>
,
<figureCitation id="56FF328DE424EAACC9137B81D5C59461" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation of plate-like structures on carapace on Scyramathia carpenteri (Norman, in Thomson 1873). A. lectotype male (21.6 x 15.9 mm) (NHM 1907.8.28.3), &quot; Holtenia ground &quot;; B, G. female (22.4 x 17.2 mm, 35.8 x 29.0 mm, respectively) (NHM 1978.99), North Sea; C. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; D, E, H, I. male (28.2 x 22.7 mm, 37.1 x 30.5 mm, 24.6 x 17.2 mm, 33.0 x 25.7 mm, respectively) (NHM 1978.99), North Sea; F. male (35.0 x 27.1 mm) (NHM 1981.108), North Sea." figureDoi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537">9A-I</figureCitation>
).
</paragraph>
<caption id="D6F618C0B551979B2F58DB9BFA6DBF25" doi="10.3897/zse.96.48041.figure9" httpUri="https://binary.pensoft.net/fig/447926" pageId="0" pageNumber="537" start="Figure 9" startId="F9">
<paragraph id="B6FF6AA7B9CA32ED92A58E03F6D987F5" pageId="0" pageNumber="537">
<emphasis id="03EE0D695DB276B64600B4B037C21F0B" bold="true" pageId="0" pageNumber="537">Figure 9.</emphasis>
Variation of plate-like structures on carapace on
<taxonomicName id="791B03CD1CBD2B924B664D10AD9BA8EA" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="7A926F4C2B7A1B36E2078E4550D654EF" italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
(Norman, in
<bibRefCitation id="4659BCC6FF24A3644FEE6A29F2985731" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
).
<emphasis id="74ADEC84266A92E0623AB9C2B70267F0" bold="true" pageId="0" pageNumber="537">A.</emphasis>
lectotype male (21.6
<normalizedToken id="87D16BAEEAFA1A149DD89354B53DF1AC" originalValue="×">x</normalizedToken>
15.9 mm) (NHM 1907.8.28.3), &quot;Holtenia ground&quot;;
<emphasis id="2ADD27E21557D4979E1B23FA978451CF" bold="true" pageId="0" pageNumber="537">B, G.</emphasis>
female (22.4
<normalizedToken id="446749D83A16C4DD5ACA1BA477D00859" originalValue="×">x</normalizedToken>
17.2 mm, 35.8
<normalizedToken id="2675BF464EC303517BD9F141E7B91825" originalValue="×">x</normalizedToken>
29.0 mm, respectively) (NHM 1978.99), North Sea;
<emphasis id="C4BFA99CC296DF27C4E8B5A8018F6A47" bold="true" pageId="0" pageNumber="537">C.</emphasis>
male (41.3
<normalizedToken id="A8DA4F6ED59B3E92C9FCBD2D0A0D041B" originalValue="×">x</normalizedToken>
34.0 mm) (NHM 1983.449), North Sea;
<emphasis id="659E869669BA8C7DDEA2D4E5E11B4891" bold="true" pageId="0" pageNumber="537">D, E, H, I.</emphasis>
male (28.2
<normalizedToken id="96F7D3A75E614519770999EA6AA9229C" originalValue="×">x</normalizedToken>
22.7 mm, 37.1
<normalizedToken id="778E64FBC092515F310CE678F15E3266" originalValue="×">x</normalizedToken>
30.5 mm, 24.6
<normalizedToken id="8D9C6A9525E50A0C322D3914D992E560" originalValue="×">x</normalizedToken>
17.2 mm, 33.0
<normalizedToken id="588132115DA34AF7AC47A3B0CD382317" originalValue="×">x</normalizedToken>
25.7 mm, respectively) (NHM 1978.99), North Sea;
<emphasis id="16B48959C1CEF9A90ACD69997813388C" bold="true" pageId="0" pageNumber="537">F.</emphasis>
male (35.0
<normalizedToken id="18B71AB4B4F17698E7858AE5CE58D33C" originalValue="×">x</normalizedToken>
27.1 mm) (NHM 1981.108), North Sea.
</paragraph>
</caption>
<paragraph id="D9D367CAA9928A5D048733F106A111FC" pageId="0" pageNumber="537">
Superficially,
<taxonomicName id="A28100FAD7390975D89BDB77D44A1325" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="67146FDDD2E3CF3E7AE45D159513BBE9" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
resembles
<taxonomicName id="DD34BBED131CD092F04F3CD2A31CE7F4" lsidName="S. umbonata" pageId="0" pageNumber="537" rank="species" species="umbonata">
<emphasis id="E17B00F306B0515F1C5620B1299454E7" italics="true" pageId="0" pageNumber="537">S. umbonata</emphasis>
</taxonomicName>
(Stimpson, 1871) from the western Atlantic, which also appears to grow to a much larger size (57 mm maximum carapace length; see Tavares et al. 2015). There are, however, consistent differences, with
<taxonomicName id="CC42CB41B765921B8296B9A151CB3647" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="7A13A04359776A893596E829E44AE7C7" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
, which has relatively longer pseudorostral spines, and the outer margin of the basal antennal article is always straight or only slightly convex. The pseudorostral spines of
<taxonomicName id="05CF03BB8CB50C01B20BC8FC21000FEE" lsidName="S. umbonata" pageId="0" pageNumber="537" rank="species" species="umbonata">
<emphasis id="266F6DE29FB11EC6DE8FF7F363F68348" italics="true" pageId="0" pageNumber="537">S. umbonata</emphasis>
</taxonomicName>
are always proportionately shorter, even when similar sized specimens are compared, and the outer margin of the basal antennal article is always distinctly convex (cf. Tavares et al. 2015: figs 1, 2, 4, 5). These differences are reliable for specimens of similar sizes. As such both species are here recognised as separate taxa.
</paragraph>
<paragraph id="9F92B775CFAE1E8DCCE035C584DB6C73" pageId="0" pageNumber="537">
It is interesting to note that some studies (A.
<bibRefCitation id="D1D8CD285B5E3A86DB6E32241E097A89" author="Milne-Edwards, A" journalOrPublisher="Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" pageId="0" pageNumber="537" pagination="1 - 106" refId="B76" refString="Milne-Edwards, A, Bouvier, E-L, 1899. Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891-1897). Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 13: 1 - 106" title="Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891 - 1897)." volume="13" year="1899">Milne-Edwards and Bouvier 1899</bibRefCitation>
,
<bibRefCitation id="80CCC1AC580DE9011D4C0D8E81D359EA" DOI="https://doi.org/10.5962/bhl.title.11668" author="Milne-Edwards, A" editor="Milne-Edwards, A" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="537" pagination="1 - 396" refId="B77" refString="Milne-Edwards, A, Bouvier, E-L, 1900. Crustaces decapodes. Premiere partie. Brachyures et Anomoures. In: Milne-Edwards, A, Ed., Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883. Masson, Paris: 1 - 396, DOI: https://doi.org/10.5962/bhl.title.11668" title="Crustaces decapodes. Premiere partie. Brachyures et Anomoures." url="https://doi.org/10.5962/bhl.title.11668" volumeTitle="Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883." year="1900">1900</bibRefCitation>
;
<bibRefCitation id="18939BC1D6B8633953028EA15A006F6A" author="Bouvier, E-L" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="537" refId="B11" refString="Bouvier, E-L, 1922. Observations complementaires sur les Crustaces decapodes (Abstraction faite des Carides) provenant des Campagnes de S.A.S. le Prince de Monaco. In: Resultats des campagnes scientifiques accomplies sur son yacht par Albert Ier, Prince Souverain de Monaco 62: 106 + [1] pp. [pls. 1-4] [In French]" title="Observations complementaires sur les Crustaces decapodes (Abstraction faite des Carides) provenant des Campagnes de S. A. S. le Prince de Monaco. In: Resultats des campagnes scientifiques accomplies sur son yacht par Albert Ier, Prince Souverain de Monaco 62: 106 + [1] pp. [pls. 1 - 4] [In French]" year="1922">Bouvier 1922</bibRefCitation>
,
<bibRefCitation id="9EF2BE061B5D71331EFF321305ADC5A3" author="Bouvier, E-L" journalOrPublisher="Faune de France" pageId="0" pageNumber="537" pagination="1 - 404" refId="B12" refString="Bouvier, E-L, 1940. Decapodes Marcheurs. Faune de France 37: 1 - 404" title="Decapodes Marcheurs." volume="37" year="1940">1940</bibRefCitation>
) mentioned that
<taxonomicName id="70A278747154F2356E75FC11FB3B62A7" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="457A097DAD7225ECE3050B3BB40A2DDA" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
does not occur in the Mediterranean, which is the region where the species,
<taxonomicName id="A29F79634660DC0491CF31E5C838FA50" lsidName="S. tenuipes" pageId="0" pageNumber="537" rank="species" species="tenuipes">
<emphasis id="272D90EC01D6CD19DB019D01BE2E3F04" italics="true" pageId="0" pageNumber="537">S. tenuipes</emphasis>
</taxonomicName>
sp. nov., is now found (see remarks for
<taxonomicName id="FC552E6DF5C75BC3D5AF7797AECC17A6" lsidName="S. tenuipes" pageId="0" pageNumber="537" rank="species" species="tenuipes">
<emphasis id="1C0DC300F057AD4882DEAB8B9ADA38DE" italics="true" pageId="0" pageNumber="537">S. tenuipes</emphasis>
</taxonomicName>
sp. nov.). Alphonse
<bibRefCitation id="9E17ACB9BEB766AB1FC4A3FFBFDF2137" author="Milne-Edwards, A" journalOrPublisher="Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" pageId="0" pageNumber="537" pagination="1 - 106" refId="B76" refString="Milne-Edwards, A, Bouvier, E-L, 1899. Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891-1897). Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 13: 1 - 106" title="Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891 - 1897)." volume="13" year="1899">Milne-Edwards and Bouvier (1899</bibRefCitation>
,
<bibRefCitation id="B433BAC9DA0432768116A8FDBC98DDC3" DOI="https://doi.org/10.5962/bhl.title.11668" author="Milne-Edwards, A" editor="Milne-Edwards, A" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="537" pagination="1 - 396" refId="B77" refString="Milne-Edwards, A, Bouvier, E-L, 1900. Crustaces decapodes. Premiere partie. Brachyures et Anomoures. In: Milne-Edwards, A, Ed., Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883. Masson, Paris: 1 - 396, DOI: https://doi.org/10.5962/bhl.title.11668" title="Crustaces decapodes. Premiere partie. Brachyures et Anomoures." url="https://doi.org/10.5962/bhl.title.11668" volumeTitle="Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883." year="1900">1900</bibRefCitation>
) and
<bibRefCitation id="59639DFEDEF7AB51F4A87E41765D5ED9" author="Bouvier, E-L" journalOrPublisher="Faune de France" pageId="0" pageNumber="537" pagination="1 - 404" refId="B12" refString="Bouvier, E-L, 1940. Decapodes Marcheurs. Faune de France 37: 1 - 404" title="Decapodes Marcheurs." volume="37" year="1940">Bouvier (1940)</bibRefCitation>
had material they referred to &quot;
<taxonomicName id="3B6E43A3E6C3FE1C2DEA2F4D4FABEB25" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="349441AAA78DB60A278EEB708CA14A10" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
&quot; from the Azores, and up north of
<normalizedToken id="9830324260AFCF467F8265C371B7D725" originalValue="São">Sao</normalizedToken>
Jorge, and the Sahara beach. Based on their figures (see A.
<bibRefCitation id="9DA4BF657FED13C82445FBEAE73CAB9A" author="Milne-Edwards, A" journalOrPublisher="Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" pageId="0" pageNumber="537" pagination="1 - 106" refId="B76" refString="Milne-Edwards, A, Bouvier, E-L, 1899. Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891-1897). Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 13: 1 - 106" title="Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891 - 1897)." volume="13" year="1899">Milne-Edwards and Bouvier 1899</bibRefCitation>
: pl. 1 fig. 4; A.
<bibRefCitation id="9E9C35F00D623BC36AF9A9EA2F7ED64A" DOI="https://doi.org/10.5962/bhl.title.11668" author="Milne-Edwards, A" editor="Milne-Edwards, A" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="537" pagination="1 - 396" refId="B77" refString="Milne-Edwards, A, Bouvier, E-L, 1900. Crustaces decapodes. Premiere partie. Brachyures et Anomoures. In: Milne-Edwards, A, Ed., Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883. Masson, Paris: 1 - 396, DOI: https://doi.org/10.5962/bhl.title.11668" title="Crustaces decapodes. Premiere partie. Brachyures et Anomoures." url="https://doi.org/10.5962/bhl.title.11668" volumeTitle="Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883." year="1900">Milne-Edwards and Bouvier 1900</bibRefCitation>
: pl. 20 Figs
<figureCitation id="767A53B02D432DA5E4AEC376442C8496" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Rochinia gracilipes A. Milne-Edwards, 1875, paralectotype male [dry] (20.0 x 15.0 mm) (MNHN-IU- 2000 - 4460 [= MNHN-B 4460]), Cap. Corrientes. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure1" httpUri="https://binary.pensoft.net/fig/447918" pageId="0" pageNumber="537">1</figureCitation>
,
<figureCitation id="C5A2A6ED8F1691472AB436E091B2259D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6</figureCitation>
;
<bibRefCitation id="75BC55720828DD78529B37A88034E333" author="Bouvier, E-L" journalOrPublisher="Faune de France" pageId="0" pageNumber="537" pagination="1 - 404" refId="B12" refString="Bouvier, E-L, 1940. Decapodes Marcheurs. Faune de France 37: 1 - 404" title="Decapodes Marcheurs." volume="37" year="1940">Bouvier 1940</bibRefCitation>
: pl. 14 fig. 1), however, the specimens they figured are more similar to
<taxonomicName id="E09AB518FA8EB3C77BF8CD5F795C0800" lsidName="S. tenuipes" pageId="0" pageNumber="537" rank="species" species="tenuipes">
<emphasis id="DFD3CF95EAAAA6C3B96E00881E973B22" italics="true" pageId="0" pageNumber="537">S. tenuipes</emphasis>
</taxonomicName>
sp. nov., with the ambulatory legs long and slender, and these records are hence referred there for now. The plates on the carapace as figured (see A.
<bibRefCitation id="9F307461A2F09DE05CF971D34E6A444B" author="Milne-Edwards, A" journalOrPublisher="Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" pageId="0" pageNumber="537" pagination="1 - 106" refId="B76" refString="Milne-Edwards, A, Bouvier, E-L, 1899. Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891-1897). Resultats des Campagnes Scientifiques accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 13: 1 - 106" title="Crustaces decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891 - 1897)." volume="13" year="1899">Milne-Edwards and Bouvier 1899</bibRefCitation>
: pl. 1 fig. 4; A.
<bibRefCitation id="4058293F7AE51EFC0D3D7D9C04749DAB" DOI="https://doi.org/10.5962/bhl.title.11668" author="Milne-Edwards, A" editor="Milne-Edwards, A" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="537" pagination="1 - 396" refId="B77" refString="Milne-Edwards, A, Bouvier, E-L, 1900. Crustaces decapodes. Premiere partie. Brachyures et Anomoures. In: Milne-Edwards, A, Ed., Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883. Masson, Paris: 1 - 396, DOI: https://doi.org/10.5962/bhl.title.11668" title="Crustaces decapodes. Premiere partie. Brachyures et Anomoures." url="https://doi.org/10.5962/bhl.title.11668" volumeTitle="Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883." year="1900">Milne-Edwards and Bouvier 1900</bibRefCitation>
: pl. 20 Figs
<figureCitation id="C2BF81E6D9A0DC7BE2E34A5E0A66EDA4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Rochinia gracilipes A. Milne-Edwards, 1875, paralectotype male [dry] (20.0 x 15.0 mm) (MNHN-IU- 2000 - 4460 [= MNHN-B 4460]), Cap. Corrientes. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure1" httpUri="https://binary.pensoft.net/fig/447918" pageId="0" pageNumber="537">1</figureCitation>
,
<figureCitation id="9303237D80182E72AA3C7F355448A0F6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A. Scyramathia carpenteri (Norman, in Thomson 1873), male (35.0 x 27.1 mm) (NHM 1981.108), North Sea; B-F. S. carpenteri (Norman, in Thomson 1873), male (41.3 x 34.0 mm) (NHM 1983.449), North Sea. A, B. lateral view of carapace; C. ventral frontal surface of carapace; D. male thoracic sternum view; E. dorsal surface of male left chela; F. posterior ventral view of male pleon." figureDoi="10.3897/zse.96.48041.figure6" httpUri="https://binary.pensoft.net/fig/447923" pageId="0" pageNumber="537">6</figureCitation>
;
<bibRefCitation id="67FE2F1910DB0D2D0FE3B528FBD114CF" author="Bouvier, E-L" journalOrPublisher="Faune de France" pageId="0" pageNumber="537" pagination="1 - 404" refId="B12" refString="Bouvier, E-L, 1940. Decapodes Marcheurs. Faune de France 37: 1 - 404" title="Decapodes Marcheurs." volume="37" year="1940">Bouvier 1940</bibRefCitation>
: pl. 14 fig. 1) appear relatively stronger than what has been observed in
<taxonomicName id="8FFAB4751BC4503DA3865F052EFB9A47" lsidName="S. tenuipes" pageId="0" pageNumber="537" rank="species" species="tenuipes">
<emphasis id="470A96EAF7047062AD61840ABAE8316F" italics="true" pageId="0" pageNumber="537">S. tenuipes</emphasis>
</taxonomicName>
sp. nov., but as discussed earlier, this character is known to vary in
<taxonomicName id="C7D0A9581CFDDD7C529CCED5EDDD0121" lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis id="7EEC533A3875153FE6C10EA9E17347B3" italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
s. str. in any case.
</paragraph>
</subSubSection>
<subSubSection id="66A4A407E5E8E1F1CAD6D795BB917CFB" pageId="0" pageNumber="537" type="distribution">
<paragraph id="85CBAF9A2C5CF1A453849627802E241E" pageId="0" pageNumber="537">Distribution.</paragraph>
<paragraph id="5D5FF0FB2F8C5917389EB003EDC45D23" pageId="0" pageNumber="537">
<taxonomicName id="75C6773C707CC9CF7CD6B803BBEE926F" baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="53FE1FEA4537BF5A3B90E0BEAE0F2F56" italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
is known from the type locality, &quot;sandy chalk of the Holtenia ground&quot; [= between North Scotland and the Faeroe Islands] (
<bibRefCitation id="9CFCD1AC2B09805F1AFB673EC010F847" DOI="https://doi.org/10.1098/rspl.1869.0084" author="Carpenter, WB" journalOrPublisher="Proceedings of the Royal Society of London" pageId="0" pageNumber="537" pagination="397 - 492" refId="B14" refString="Carpenter, WB, Gwyn Jeffreys, J, Thomson, CW, 1870. Preliminary report of the scientific exploration of the deep sea in H.M. Surveying-vessel 'Porcupine,' during the summer of 1869. Proceedings of the Royal Society of London 18: 397 - 492, DOI: https://doi.org/10.1098/rspl.1869.0084" title="Preliminary report of the scientific exploration of the deep sea in H. M. Surveying-vessel ' Porcupine, ' during the summer of 1869." url="https://doi.org/10.1098/rspl.1869.0084" volume="18" year="1870">Carpenter et al. 1870</bibRefCitation>
; Norman, in
<bibRefCitation id="8A50CF2E121BD41AB5574036D5D88765" DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
;
<bibRefCitation id="9165E7EE119238004DAC1C04C2A4DB0F" DOI="https://doi.org/10.1111/j.1096-3642.1995.tb01430.x" author="Reiswig, HM" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="537" pagination="373 - 384" refId="B102" refString="Reiswig, HM, Champagne, P, 1995. The NE Atlantic glass sponges Pheronema carpenteri (Thomson) and P. grayi Kent (Porifera: Hexactinellida) are synonyms. Zoological Journal of the Linnean Society 115: 373 - 384, DOI: https://doi.org/10.1111/j.1096-3642.1995.tb01430.x" title="The NE Atlantic glass sponges Pheronema carpenteri (Thomson) and P. grayi Kent (Porifera: Hexactinellida) are synonyms." url="https://doi.org/10.1111/j.1096-3642.1995.tb01430.x" volume="115" year="1995">Reiswig and Champagne 1995</bibRefCitation>
), South Iceland (
<bibRefCitation id="FE710CCE7F094677BCAFE8358523FA85" author="Hansen, HJ" journalOrPublisher="Danish Ingolf-Expedition" pageId="0" pageNumber="537" pagination="1 - 120" refId="B47" refString="Hansen, HJ, 1908. Crustacea Malacostraca. I. Danish Ingolf-Expedition 3 (2): 1 - 120" title="Crustacea Malacostraca. I." volume="3" year="1908">Hansen 1908</bibRefCitation>
), Southwest Faeroe Islands, South of Norway, West of British Isles (
<bibRefCitation id="F786E619C0FD61B1E26F507D62B1D041" DOI="https://doi.org/10.5962/bhl.title.14876" author="Clark, PF" journalOrPublisher="Marine Conservation Society, Englang, U. K." pageId="0" pageNumber="537" refId="B21" refString="Clark, PF, 1986. North-East Atlantic Crabs. An Atlas of Distribution. Marine Conservation Society, Englang, U.K., DOI: https://doi.org/10.5962/bhl.title.14876" title="North-East Atlantic Crabs. An Atlas of Distribution." url="https://doi.org/10.5962/bhl.title.14876" year="1986">Clark 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>