266 lines
33 KiB
XML
266 lines
33 KiB
XML
<document id="9381FD8FEAC64213C76D5703C02B75C8" ID="10.11646/zootaxa.4144.1.7" ID-DOI="10.11646/zootaxa.4144.1.7" ID-GBIF-Dataset="acb70f7b-aa2e-4729-bc9a-4c794f63e119" ID-ISSN="1175-5326" ID-Zenodo-Dep="257699" ID-ZooBank="C9DCB2BE-6DE6-416D-A0A7-076AA2716C6D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469515202917" checkinUser="plazi" docAuthor="Bukejs, Andris & Alekseev, Vitalii I." docDate="2016" docId="895A87DEFFDDFFAA35EEFC181D2646F7" docLanguage="en" docName="zootaxa.4144.1.7.pdf" docOrigin="Zootaxa 4144 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Passandra septentrionaria Bukejs & Alekseev, 2016, sp. nov." docType="treatment" docUuid="930B731B-7CB0-4E7D-A6DA-9153BFD70062" docUuidSource="ZooBank" docVersion="8" lastPageNumber="121" masterDocId="7563FFA6FFDCFFAE3579FFDA19114257" masterDocTitle="Passandra septentrionaria sp. nov.: the first described species of Passandridae (Coleoptera: Cucujoidea) from Eocene Baltic amber" masterLastPageNumber="123" masterPageNumber="117" pageNumber="118" updateTime="1698366181931" updateUser="plazi">
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<mods:mods id="709497F96B96289D980529BBC00DB7BF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="135D43AE5B2C9A0C3E99EEFAC798E31C">
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<mods:title id="E21A1C5A3AEC80BDF72F2657A96BC4D2">Passandra septentrionaria sp. nov.: the first described species of Passandridae (Coleoptera: Cucujoidea) from Eocene Baltic amber</mods:title>
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<mods:name id="D19019DD54FCB2B6F0E1E78060CFD2BE" type="personal">
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<mods:roleTerm id="C9674EEE3CAA1092B2A26CC537FFC23D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E97246B20CD4E576BE481EC5A62161C">Bukejs, Andris</mods:namePart>
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</mods:name>
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<mods:name id="8C512B5C1D3E4EAD299BA0F20803ED11" type="personal">
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<mods:role id="9EF825A6A4044445D1AFBD1BC5CC96DF">
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<mods:roleTerm id="C2ED5A299E0C491EB3CD0B4F53296512">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BFC0DC19D5A8AE43DA3E7C60C56F2818">Alekseev, Vitalii I.</mods:namePart>
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<mods:typeOfResource id="6844043F129927A715BD5E2F25776949">text</mods:typeOfResource>
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<mods:titleInfo id="6BFFC75599C099836CFDCCA125595C2B">
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<mods:title id="E0FAEB101CFE2D267174300DF5C3F3EB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A8C871588100D34D241268DD77CDFA34">
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<mods:date id="52E27809F9FCAB4288C1C9119B8158FC">2016</mods:date>
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<mods:detail id="5BABA95D61032011E4AB447F3CFCB1ED" type="volume">
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<mods:number id="1231CF3D3A4B163A07C72E123A095BB3">4144</mods:number>
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</mods:detail>
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<mods:detail id="899AEF0BA85A1683C52B4B81AA463E49" type="issue">
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<mods:number id="D8CDD7EA5424DE384A5096897872F8A8">1</mods:number>
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<mods:classification id="975D4B8D9FB2B16847C19F0914BFE361">journal article</mods:classification>
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<mods:identifier id="CEE62E3DE7613AD772090FF6870B787E" type="DOI">10.11646/zootaxa.4144.1.7</mods:identifier>
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<mods:identifier id="F73DE939EE44F9D083FA516153F3462D" type="GBIF-Dataset">acb70f7b-aa2e-4729-bc9a-4c794f63e119</mods:identifier>
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<mods:identifier id="70BFE0514A7165F268648A9DF8AC1353" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D91B2F10F86F4D75DF4D3A220AA6A171" type="Zenodo-Dep">257699</mods:identifier>
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<mods:identifier id="7C5691C975CDA0EA4610A7D1EF93A539" type="ZooBank">C9DCB2BE-6DE6-416D-A0A7-076AA2716C6D</mods:identifier>
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</mods:mods>
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<treatment id="895A87DEFFDDFFAA35EEFC181D2646F7" ID-DOI="http://doi.org/10.5281/zenodo.6067071" ID-GBIF-Taxon="123207640" ID-Zenodo-Dep="6067071" LSID="urn:lsid:zoobank.org:act:930B731B-7CB0-4E7D-A6DA-9153BFD70062" httpUri="http://treatment.plazi.org/id/895A87DEFFDDFFAA35EEFC181D2646F7" lastPageId="4" lastPageNumber="121" pageId="1" pageNumber="118">
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<subSubSection id="49E96543FFDDFFAF35EEFC18181A41A9" pageId="1" pageNumber="118" type="nomenclature">
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<paragraph id="014C36C8FFDDFFAF35EEFC181B2B418B" blockId="1.[151,570,962,1022]" box="[151,570,962,988]" pageId="1" pageNumber="118">
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<heading id="5A0481A4FFDDFFAF35EEFC181B2B418B" bold="true" box="[151,570,962,988]" fontSize="11" level="1" pageId="1" pageNumber="118" reason="1">
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<emphasis id="3387EADAFFDDFFAF35EEFC181B2B418B" bold="true" box="[151,570,962,988]" pageId="1" pageNumber="118">
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<taxonomicName id="C6F34D4BFFDDFFAF35EEFC1818C4418B" box="[151,469,962,988]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionaria" status="sp. nov.">
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<emphasis id="3387EADAFFDDFFAF35EEFC1818C4418B" bold="true" box="[151,469,962,988]" italics="true" pageId="1" pageNumber="118">Passandra septentrionaria</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="28B457A1FFDDFFAF34A5FC181B2B418B" box="[476,570,962,988]" pageId="1" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="014C36C8FFDDFFAF35EEFC3F181A41A9" blockId="1.[151,570,962,1022]" box="[151,267,997,1022]" pageId="1" pageNumber="118">
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(
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<figureCitation id="99C82A4DFFDDFFAF35E6FC3F181341A9" box="[159,258,997,1022]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 7" captionStartId-0="2.[151,264,1631,1654]" captionStartId-1="3.[151,263,1638,1661]" captionTargetBox-0="[151,1436,195,1607]" captionTargetBox-1="[151,1436,265,1616]" captionTargetId-0="figure@2.[151,1436,195,1607]" captionTargetId-1="figure@3.[151,1436,265,1617]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 3. Passandra septentrionaria sp. nov., holotype: 1 — habitus, dorsal view; 2 — habitus, ventro-lateral view, 3 — habitus, dorso-lateral view. Scale bar = 1 mm." captionText-1="FIGURES 4 – 7. Passandra septentrionaria sp. nov.: 4 — habitus, dorsal view, paratype; 5 — details of forebody, dorso-lateral view, holotype; 6 — apical antennomere, paratype; 7 — right antenna of holotype, the arrows indicate narrow grooves. Not reproduced to the same scale." httpUri-0="https://zenodo.org/record/257700/files/figure.png" httpUri-1="https://zenodo.org/record/257701/files/figure.png" pageId="1" pageNumber="118">Figs 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="49E96543FFDDFFAF35EEFBF71BAF44FE" pageId="1" pageNumber="118" type="materials_examined">
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<paragraph id="014C36C8FFDDFFAF35EEFBF71A6747DD" blockId="1.[151,1437,1069,1994]" pageId="1" pageNumber="118">
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<materialsCitation id="B19B3C95FFDDFFAF35EEFBF71A2946D9" ID-GBIF-Occurrence="1288049026" collectionCode="RSM" collectorName="Bukejs A. & Alekseev V. I. & McKellar R. C." country="Russia" location="Yantarny village" pageId="1" pageNumber="118" specimenCount="3" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="3387EADAFFDDFFAF35EEFBF718584611" bold="true" box="[151,329,1069,1094]" pageId="1" pageNumber="118">
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<typeStatus id="DE48886AFFDDFFAF35EEFBF719C34611" box="[151,210,1069,1094]" pageId="1" pageNumber="118">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="DE48886AFFDDFFAF342DFBF718D04611" box="[340,449,1069,1094]" pageId="1" pageNumber="118" type="holotype">Holotype</typeStatus>
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: Nr.
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<collectionCode id="67E2AE0DFFDDFFAF3487FBF41B284611" box="[510,569,1070,1094]" country="Canada" httpUri="http://grbio.org/cool/7sv4-pazk" name="Royal Saskatchewan Museum" pageId="1" pageNumber="118">RSM</collectionCode>
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P3300.013 [
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<collectionCode id="67E2AE0DFFDDFFAF37B6FBF41A034611" box="[719,786,1070,1094]" country="Canada" httpUri="http://grbio.org/cool/7sv4-pazk" name="Royal Saskatchewan Museum" pageId="1" pageNumber="118">RSM</collectionCode>
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], “
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<typeStatus id="DE48886AFFDDFFAF364BFBF71A8C4611" box="[818,925,1069,1094]" pageId="1" pageNumber="118" type="holotype">Holotype</typeStatus>
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/
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<taxonomicName id="C6F34D4BFFDDFFAF36C5FBF71DFD4612" box="[956,1260,1069,1093]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionaria" status="sp. nov.">
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<emphasis id="3387EADAFFDDFFAF36C5FBF71DFD4612" box="[956,1260,1069,1093]" italics="true" pageId="1" pageNumber="118">Passandra septentrionaria</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="28B457A1FFDDFFAF318EFBF41C434612" box="[1271,1362,1070,1093]" pageId="1" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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/ des.
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<collectorName id="AC06531EFFDDFFAF35EEFB8B181F463D" box="[151,270,1105,1130]" pageId="1" pageNumber="118">Bukejs A.</collectorName>
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,
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<collectorName id="AC06531EFFDDFFAF3466FB8B18AD463E" box="[287,444,1105,1130]" pageId="1" pageNumber="118">Alekseev V.I.</collectorName>
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&
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<collectorName id="AC06531EFFDDFFAF3493FB8B1B87463D" box="[490,662,1105,1130]" pageId="1" pageNumber="118">McKellar R.C.</collectorName>
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” [red printed label];
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<location id="042C6013FFDDFFAF36EEFB881D44463D" LSID="urn:lsid:plazi:treatment:895A87DEFFDDFFAA35EEFC181D2646F7:042C6013FFDDFFAF36EEFB881D44463D" box="[919,1109,1105,1130]" country="Russia" name="Yantarny village" pageId="1" pageNumber="118" stateProvince="Kaliningrad">Yantarny village</location>
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[formerly Palmnicken], the Sambian [Samland] peninsula,
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<collectingRegion id="C337F82AFFDDFFAF3779FBAF1B9746D9" box="[512,646,1141,1166]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="118">Kaliningrad</collectingRegion>
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region,
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<collectingCountry id="79E47658FFDDFFAF3793FBAF1A2946D9" box="[746,824,1141,1166]" name="Russia" pageId="1" pageNumber="118">Russia</collectingCountry>
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</materialsCitation>
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; sex unknown. The complete beetle is included in a piece of amber that has been polished into a rhombic shape in order to create viewing planes (maximum length
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<quantity id="C60B9B2DFFDDFFAF30F9FB4019D44682" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="118" unit="mm" value="10.0">10 mm</quantity>
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, width
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<quantity id="C60B9B2DFFDDFFAF3462FB6718714682" box="[283,352,1213,1238]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="118" unit="mm" value="6.0">6 mm</quantity>
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, and thickness
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<quantity id="C60B9B2DFFDDFFAF3776FB641B474681" box="[527,598,1214,1238]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="118" unit="mm" value="4.0">4 mm</quantity>
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), and preserved without supplementary fixation. Ventral details are partly obscured by prominent drying line within amber that contains many particulates; thin veil of “milky” amber (created by decay products interacting with resin) is pervasive on ventral surface, but most anatomical details remain visible; much of the cuticle has carbonized and pulled away from the surface of the surrounding amber, creating silvery areas dorsally. Syninclusions: spider web (silk) fragments, and a few plant trichomes, separated during preparation and catalogued as specimen RSM P 3300.014.
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</paragraph>
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<paragraph id="014C36C8FFDDFFAF35BEFA4C1A884435" blockId="1.[151,1437,1069,1994]" pageId="1" pageNumber="118">
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<materialsCitation id="B19B3C95FFDDFFAF35BEFA4C1A884435" ID-GBIF-Occurrence="1288049025" collectionCode="CAG" collectorName="Bukejs A. & Alekseev V. I. & McKellar R. C." country="Poland" location="Poland" pageId="1" pageNumber="118" specimenCount="2" typeStatus="paratype">
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<typeStatus id="DE48886AFFDDFFAF35BEFA4C183F47F9" box="[199,302,1430,1454]" pageId="1" pageNumber="118" type="paratype">Paratype</typeStatus>
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: Nr. 1291 [
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<collectionCode id="67E2AE0DFFDDFFAF34D5FA4C18FE47F9" box="[428,495,1430,1454]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14391" name="Universit<69> degli Studi di Cagliari" pageId="1" pageNumber="118">CAG</collectionCode>
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], “
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<typeStatus id="DE48886AFFDDFFAF3775FA4C1B6047F9" box="[524,625,1430,1454]" pageId="1" pageNumber="118" type="paratype">Paratype</typeStatus>
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/
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<taxonomicName id="C6F34D4BFFDDFFAF37F0FA4F1AA647FA" box="[649,951,1429,1453]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionaria" status="sp. nov.">
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<emphasis id="3387EADAFFDDFFAF37F0FA4F1AA647FA" box="[649,951,1429,1453]" italics="true" pageId="1" pageNumber="118">Passandra septentrionaria</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="28B457A1FFDDFFAF36C6FA4C1D0647FA" box="[959,1047,1430,1453]" pageId="1" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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/ des.
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<collectorName id="AC06531EFFDDFFAF311DFA4F1DC647F9" box="[1124,1239,1429,1454]" pageId="1" pageNumber="118">Bukejs A.</collectorName>
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,
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<collectorName id="AC06531EFFDDFFAF319CFA4F1C6F47FA" box="[1253,1406,1429,1454]" pageId="1" pageNumber="118">Alekseev V.I.</collectorName>
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&
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<collectorName id="AC06531EFFDDFFAF35EEFA6318524785" box="[151,323,1465,1490]" pageId="1" pageNumber="118">McKellar R.C.</collectorName>
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” [red printed label]; Baltic Sea coast, Gdańsk,
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<collectingCountry id="79E47658FFDDFFAF3606FA631AC14785" box="[895,976,1465,1490]" name="Poland" pageId="1" pageNumber="118">Poland</collectingCountry>
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; sex unknown. The complete beetle is included in a small, elongate amber piece (length
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<quantity id="C60B9B2DFFDDFFAF379EFA041A2C47A2" box="[743,829,1502,1526]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="118" unit="mm" value="29.0">29 mm</quantity>
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, width
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||
<quantity id="C60B9B2DFFDDFFAF36E1FA041ACE47A2" box="[920,991,1502,1526]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="118" unit="mm" value="9.0">9 mm</quantity>
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, and maximum thickness
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<quantity id="C60B9B2DFFDDFFAF306AFA071C4C47A1" box="[1299,1373,1501,1526]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="118" unit="mm" value="5.0">5 mm</quantity>
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), and preserved without supplementary fixation. Elytra and anterior part of head are partly obscured by a “milky” opacity; ventral part of specimen is almost not visible because of location of the beetle in amber piece. Syninclusions: leg of unidentified insect, and few small gas vesicles.
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</materialsCitation>
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</paragraph>
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<paragraph id="014C36C8FFDDFFAF35BEF9B71D2B44D2" blockId="1.[151,1437,1069,1994]" box="[199,1082,1645,1670]" pageId="1" pageNumber="118">
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||
<materialsCitation id="B19B3C95FFDDFFAF35BEF9B71D2B44D2" ID-GBIF-Occurrence="1288049024" box="[199,1082,1645,1670]" county="Upper Eocene." location="Baltic" municipality="Blaue Erde" pageId="1" pageNumber="118" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="3387EADAFFDDFFAF35BEF9B7184444D1" bold="true" box="[199,341,1645,1670]" pageId="1" pageNumber="118">
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||
<typeStatus id="DE48886AFFDDFFAF35BEF9B7181344D1" box="[199,258,1645,1670]" pageId="1" pageNumber="118">Type</typeStatus>
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strata.
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</emphasis>
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||
<location id="042C6013FFDDFFAF3425F9B7188E44D1" LSID="urn:lsid:plazi:treatment:895A87DEFFDDFFAA35EEFC181D2646F7:042C6013FFDDFFAF3425F9B7188E44D1" box="[348,415,1645,1670]" county="Upper Eocene." municipality="Blaue Erde" name="Baltic" pageId="1" pageNumber="118">Baltic</location>
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amber,
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<emphasis id="3387EADAFFDDFFAF3480F9B71B6644D2" box="[505,631,1645,1669]" italics="true" pageId="1" pageNumber="118">
|
||
<collectingMunicipality id="E128ACB2FFDDFFAF3480F9B71B6644D2" box="[505,631,1645,1669]" pageId="1" pageNumber="118">Blaue Erde</collectingMunicipality>
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||
</emphasis>
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||
deposits, mid-Eocene to
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<collectingCounty id="E82D4E44FFDDFFAF36EAF9B41D2B44D2" box="[915,1082,1646,1669]" pageId="1" pageNumber="118">Upper Eocene.</collectingCounty>
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="014C36C8FFDDFFAF35BEF94B1BAF44FE" blockId="1.[151,1437,1069,1994]" box="[199,702,1681,1706]" pageId="1" pageNumber="118">
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||
<materialsCitation id="B19B3C95FFDDFFAF35BEF94B1BAF44FE" ID-GBIF-Occurrence="1288049027" box="[199,702,1681,1706]" location="Southeastern Baltic Sea" pageId="1" pageNumber="118" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="3387EADAFFDDFFAF35BEF94B187444FD" bold="true" box="[199,357,1681,1706]" pageId="1" pageNumber="118">
|
||
<typeStatus id="DE48886AFFDDFFAF35BEF94B181344FD" box="[199,258,1681,1706]" pageId="1" pageNumber="118">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<location id="042C6013FFDDFFAF3415F94B1B6744FD" LSID="urn:lsid:plazi:treatment:895A87DEFFDDFFAA35EEFC181D2646F7:042C6013FFDDFFAF3415F94B1B6744FD" box="[364,630,1681,1706]" name="Southeastern Baltic Sea" pageId="1" pageNumber="118">Southeastern Baltic Sea</location>
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coast.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="49E96543FFDDFFAF35BEF96F1B3D44A5" pageId="1" pageNumber="118" type="etymology">
|
||
<paragraph id="014C36C8FFDDFFAF35BEF96F1B3D44A5" blockId="1.[151,1437,1069,1994]" pageId="1" pageNumber="118">
|
||
<emphasis id="3387EADAFFDDFFAF35BEF96F185F4499" bold="true" box="[199,334,1717,1742]" pageId="1" pageNumber="118">Etymology.</emphasis>
|
||
The specific epithet refers to the northern location of the specimen’s origin. It is formed after the Latin “
|
||
<emphasis id="3387EADAFFDDFFAF359CF900188D44A6" box="[229,412,1754,1777]" italics="true" pageId="1" pageNumber="118">septentrionarius</emphasis>
|
||
”—northern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49E96543FFDDFFAF35BEF92718E6459D" pageId="1" pageNumber="118" type="diagnosis">
|
||
<paragraph id="014C36C8FFDDFFAF35BEF92718E6459D" blockId="1.[151,1437,1069,1994]" pageId="1" pageNumber="118">
|
||
<emphasis id="3387EADAFFDDFFAF35BEF92718504541" bold="true" box="[199,321,1789,1814]" pageId="1" pageNumber="118">Diagnosis.</emphasis>
|
||
<taxonomicName id="C6F34D4BFFDDFFAF3431F9271B654542" box="[328,628,1789,1813]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionaria" status="sp. nov.">
|
||
<emphasis id="3387EADAFFDDFFAF3431F9271B654542" box="[328,628,1789,1813]" italics="true" pageId="1" pageNumber="118">Passandra septentrionaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3387EADAFFDDFFAF3702F9271BC54541" bold="true" box="[635,724,1789,1814]" pageId="1" pageNumber="118">
|
||
<taxonomicNameLabel id="28B457A1FFDDFFAF3702F9271BC54541" box="[635,724,1789,1814]" pageId="1" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has pronotal sublateral lines only in basal one-third of pronotal length. In all extant species of
|
||
<emphasis id="3387EADAFFDDFFAF3495F8FB1B7A456E" box="[492,619,1825,1849]" italics="true" pageId="1" pageNumber="118">
|
||
<taxonomicName id="C6F34D4BFFDDFFAF3495F8FB1B76456E" box="[492,615,1825,1849]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Passandra</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
pronotal sublateral lines are more or less complete, except for in African species
|
||
<taxonomicName id="C6F34D4BFFDDFFAF358AF89C1B234509" authority="Dalman, 1817" authorityName="Dalman" authorityYear="1817" box="[243,562,1861,1886]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="sexstriata">
|
||
<emphasis id="3387EADAFFDDFFAF358AF89C1891450A" box="[243,384,1862,1885]" italics="true" pageId="1" pageNumber="118">P. sexstriata</emphasis>
|
||
Dalman, 1817
|
||
</taxonomicName>
|
||
, which differs from the new species in (1) lacking admedian grooves and median process on head, (2) lacking pronotal sublateral lines, (3) pronotum with strongly prominent and acute anterior angles, (4) sublateral grooves on head well-defined and not joined to occipital groove, and (5) bicoloured body (rarely uniformly brown).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="558C6640FFDEFFAC35EEF9851B7844C3" httpUri="https://zenodo.org/record/257700/files/figure.png" pageId="2" pageNumber="119" targetBox="[151,1436,195,1607]" targetPageId="2">
|
||
<paragraph id="014C36C8FFDEFFAC35EEF9851B7844C3" blockId="2.[151,1435,1631,1684]" pageId="2" pageNumber="119">
|
||
<emphasis id="3387EADAFFDEFFAC35EEF985182D4422" bold="true" box="[151,316,1631,1654]" pageId="2" pageNumber="119">FIGURES 1–3.</emphasis>
|
||
<taxonomicName id="C6F34D4BFFDEFFAC343DF9BA1B454421" box="[324,596,1632,1654]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="septentrionaria" status="sp. nov.">
|
||
<emphasis id="3387EADAFFDEFFAC343DF9BA1B454421" box="[324,596,1632,1654]" italics="true" pageId="2" pageNumber="119">Passandra septentrionaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3387EADAFFDEFFAC3725F9BA1BBF4422" bold="true" box="[604,686,1632,1653]" pageId="2" pageNumber="119">
|
||
<taxonomicNameLabel id="28B457A1FFDEFFAC3725F9BA1BBF4422" box="[604,686,1632,1653]" pageId="2" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype: 1—habitus, dorsal view; 2—habitus, ventro-lateral view, 3— habitus, dorso-lateral view. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="49E96543FFDEFFAA35BEF9661C2141BA" lastPageId="4" lastPageNumber="121" pageId="2" pageNumber="119" type="description">
|
||
<paragraph id="014C36C8FFDEFFAC35BEF966184F454A" blockId="2.[151,1437,1724,2037]" pageId="2" pageNumber="119">
|
||
<emphasis id="3387EADAFFDEFFAC35BEF96618484482" bold="true" box="[199,345,1724,1749]" pageId="2" pageNumber="119">Description.</emphasis>
|
||
<typeStatus id="DE48886AFFDEFFAC341BF96618DF4482" box="[354,462,1724,1749]" pageId="2" pageNumber="119" type="holotype">Holotype</typeStatus>
|
||
. Body length
|
||
<quantity id="C60B9B2DFFDEFFAC370AF9661BCD4483" box="[627,732,1724,1749]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" pageId="2" pageNumber="119" unit="mm" value="5.45">5.45 mm</quantity>
|
||
, maximum width
|
||
<quantity id="C60B9B2DFFDEFFAC36C8F9661D0A4483" box="[945,1051,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="119" unit="mm" value="1.23">1.23 mm</quantity>
|
||
; body outline narrowly elongateoval, parallel sided, flattened dorsally, slightly convex ventrally; generally glabrous, unicolourous black, moderately shiny.
|
||
</paragraph>
|
||
<paragraph id="014C36C8FFDEFFAD35BEF8F21D3B4283" blockId="2.[151,1437,1724,2037]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="120" pageId="2" pageNumber="119">Head slightly convex, about 1.4× as wide as long, measured across compound eyes and along length of dorsal disc; covered with small and dense punctation, distance between punctures approximately 1–2× diameter of single puncture. Admedian grooves moderately deep, slightly diverging anteriorly, joined posteriorly to wide and deep occipital groove which contains shallow, broad foveae; medial process wide, slightly convex, triangular with widely rounded apex; sublateral grooves shallow and narrow but clearly impressed throughout their lengths, forming continuous lip around lateral and anterior margins of dorsal surface of head, and clearly joined to occipital groove. Clypeus narrow, slightly depressed, with emarginate anterior margin. Compound eyes small, about 1.2× as long as antennomere 1 is wide (in lateral view), subspherical, moderately convex.</paragraph>
|
||
<caption id="558C6640FFDFFFAD35EEF9BC18D344ED" httpUri="https://zenodo.org/record/257701/files/figure.png" pageId="3" pageNumber="120" targetBox="[151,1436,265,1616]" targetPageId="3">
|
||
<paragraph id="014C36C8FFDFFFAD35EEF9BC18D344ED" blockId="3.[151,1436,1638,1722]" pageId="3" pageNumber="120">
|
||
<emphasis id="3387EADAFFDFFFAD35EEF9BC182A442B" bold="true" box="[151,315,1638,1661]" pageId="3" pageNumber="120">FIGURES 4–7.</emphasis>
|
||
<taxonomicName id="C6F34D4BFFDFFFAD343DF9BD1B45442A" box="[324,596,1639,1661]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="species" species="septentrionaria" status="sp. nov.">
|
||
<emphasis id="3387EADAFFDFFFAD343DF9BD1B45442A" box="[324,596,1639,1661]" italics="true" pageId="3" pageNumber="120">Passandra septentrionaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3387EADAFFDFFFAD3725F9BD1BBE442B" bold="true" box="[604,687,1639,1660]" pageId="3" pageNumber="120">
|
||
<taxonomicNameLabel id="28B457A1FFDFFFAD3725F9BD1BBE442B" box="[604,687,1639,1660]" pageId="3" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 4—habitus, dorsal view, paratype; 5—details of forebody, dorso-lateral view, holotype; 6—apical antennomere, paratype; 7—right antenna of holotype, the arrows indicate narrow grooves. Not reproduced to the same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="014C36C8FFDFFFAA35BEF9381B3C4368" blockId="3.[151,1437,1761,2038]" lastBlockId="4.[151,1437,151,1184]" lastPageId="4" lastPageNumber="121" pageId="3" pageNumber="120">Antennae with 11 antennomeres, moderately slender and long, extending to basal one-third of elytra; scape subcylindrical and somewhat globose, large, about 1.2× as long as wide; pedicel subquadratic, nearly as long as wide, 0.8× as long as antennomere 3; antennomeres 3–10 subcylindrical with mildly constricted bases, and markedly longer than wide, with long pale setae; antennomeres 5–6 narrowly grooved ventrally, and antennomeres 7–8 bear exceptionally faint longitudinal groove on ventral surface (other antennomeres are not clearly visible in either examined specimen); apical antennomere asymmetric, elongate-oval with broad apex, keeled, covered with setae. Setation absent from scape, but coarse setae are concentrated upon ventral and medial surfaces of subsequent antennomeres, and surround their apices; 3 or 4 short setae suberect on apex of pedicel; antennomere 3 with tuft of 20 or more suberect setae near apex, with lengths that exceed 1.5× width of antennomere; antennomere 4 with few apical setae as long as antennomere is wide, plus coarser, shorter setae dispersed along length and encircling apex of antennomere; antenommeres 5–8 bear similar setation to antennomere 4; antennomeres 9 and 10 apparently have reduced setae lengths; antennomere 11 has coarse, inclined setae basally, and encircling apex, plus dense coat of finer setae on asymmetrical apex.</paragraph>
|
||
<paragraph id="014C36C8FFD8FFAA35BEFE9718C5404F" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="121">Pronotum 1.13× as long as wide, subquadrate, nearly flat, widest in anterior one-third, distinctly narrowed posteriad; with small and dense longitudinally oval punctures (similar to punctures on head), disc with markedly sparser punctation. Sublateral lines present in basal one-third of pronotal length only. Lateral margins slightly convex, with narrow bordering; anterior margin sinuated, with faint bordering only present in medial one-third of width; posterior margin almost straight, with narrow bordering. Anterior angles obtuse, rounded, not protruding; posterior angles rectangular.</paragraph>
|
||
<paragraph id="014C36C8FFD8FFAA35BEFDFE18354142" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="121">
|
||
Scutellum moderately large, transverse, suboval. Elytra 2.5× as long as their combined width, and 2.7× as long as pronotum; base markedly wider than pronotal posterior margin. Elytral striae 1 and 6 complete: stria 1 grooved throughout entire length, stria 6 grooved in basal four-fifths of length; striae 2–5 indistinct (consisting of very fine punctures), grooved only faintly at base of elytra. Sutural interval convex; other intervals flat, smooth. Elytra of
|
||
<typeStatus id="DE48886AFFD8FFAA35EEFD6E19E8409A" box="[151,249,692,717]" pageId="4" pageNumber="121" type="holotype">holotype</typeStatus>
|
||
appear to display microsculpture and large-scale colour patterns, but these are artefacts created by carbonization (cracking) of cuticle, and by cuticle adhering to surrounding amber only in a few patches, respectively.
|
||
</paragraph>
|
||
<paragraph id="014C36C8FFD8FFAA35BEFCFA1DF1419F" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="121">Legs short and robust. Tarsal formula 5-5-5. Tarsomere 1 of all tarsi short, markedly shorter than tarsomere 2, partly covered by tibial apex; tarsomeres 2–4 short, subequal in length and shape; tarsomere 5 long, about as long as tarsomeres 3–4 combined; tarsomeres 2 and 3 of meso- and metatarsus with plantar comb of thick, inclined setae along apical margins, with comb most prominent at anterior and posterior corners, while tarsomeres 1 and 4 have reduced version of comb; low, broad spicules present along apex of metatibia. Claws simple, thin.</paragraph>
|
||
<paragraph id="014C36C8FFD8FFAA35BEFC0F1C2141BA" blockId="4.[151,1437,151,1184]" box="[199,1328,980,1005]" pageId="4" pageNumber="121">
|
||
Paratype. Body length
|
||
<quantity id="C60B9B2DFFD8FFAA34BEFC0E1B3141BB" box="[455,544,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="121" unit="mm" value="8.5">8.5 mm</quantity>
|
||
; otherwise similar in all visible morphological characters to holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49E96543FFD8FFAA35BEFC2D1D2646F7" pageId="4" pageNumber="121" type="discussion">
|
||
<paragraph id="014C36C8FFD8FFAA35BEFC2D1D2646F7" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="121">
|
||
<emphasis id="3387EADAFFD8FFAA35BEFC2D182C4647" bold="true" box="[199,317,1015,1040]" pageId="4" pageNumber="121">Comment</emphasis>
|
||
. Adult size varies significantly within some passandrid species, possibly due to the parasitic lifestyle of the larvae and different sizes of their hosts. For instance, adults of the extant
|
||
<taxonomicName id="C6F34D4BFFD8FFAA31D5FBC61827460F" authority="Dalman, 1817" authorityName="Dalman" authorityYear="1817" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="sexstriata">
|
||
<emphasis id="3387EADAFFD8FFAA31D5FBC61C8D4663" box="[1196,1436,1052,1076]" italics="true" pageId="4" pageNumber="121">Passandra sexstriata</emphasis>
|
||
Dalman, 1817
|
||
</taxonomicName>
|
||
can be
|
||
<quantity id="C60B9B2DFFD8FFAA34F6FBE51B184600" box="[399,521,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="3.2" metricValueMin="1.6" pageId="4" pageNumber="121" unit="mm" value="24.0" valueMax="32.0" valueMin="16.0">16–32 mm</quantity>
|
||
long; extant
|
||
<taxonomicName id="C6F34D4BFFD8FFAA37E7FB9B1AD5460F" authority="Murray, 1867" authorityName="Murray" authorityYear="1867" box="[670,964,1087,1112]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="simplex">
|
||
<emphasis id="3387EADAFFD8FFAA37E7FB9B1A01460F" box="[670,784,1088,1112]" italics="true" pageId="4" pageNumber="121">P. simplex</emphasis>
|
||
(Murray, 1867)
|
||
</taxonomicName>
|
||
is
|
||
<quantity id="C60B9B2DFFD8FFAA369FFB9A1D434600" box="[998,1106,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="4" pageNumber="121" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">7–13 mm</quantity>
|
||
in length; the body length of the extant
|
||
<taxonomicName id="C6F34D4BFFD8FFAA3475FBBF1B09462B" authority="Gray, 1832" authorityName="Gray" authorityYear="1832" box="[268,536,1124,1148]" class="Insecta" family="Passandridae" genus="Passandra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="fasciata">
|
||
<emphasis id="3387EADAFFD8FFAA3475FBBF1893462B" box="[268,386,1124,1148]" italics="true" pageId="4" pageNumber="121">P. fasciata</emphasis>
|
||
(Gray, 1832)
|
||
</taxonomicName>
|
||
varies in range
|
||
<quantity id="C60B9B2DFFD8FFAA37B7FBBE1A56462B" box="[718,839,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.8" metricValueMin="1.0" pageId="4" pageNumber="121" unit="mm" value="19.0" valueMax="28.0" valueMin="10.0">10–28 mm</quantity>
|
||
(
|
||
<bibRefCitation id="65624B39FFD8FFAA362FFBBE1AEE462B" author="Slipinski" box="[854,1023,1124,1149]" pageId="4" pageNumber="121" refString="Slipinski, S. A. (1987) A review of the Passandridae of the world (Coleoptera, Cucujoidea). I. Genus Passandra Dalman. Annali del Museo civico di Storia naturale, Genova, 86, 553 - 603." type="journal article" year="1987">Ślipiński 1987</bibRefCitation>
|
||
). As in the recent species, we do not treat body size in the newly described fossil species as a species-specific character.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |