526 lines
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526 lines
57 KiB
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<document ID-CLB-Dataset="20962" ID-DOI="http://dx.doi.org/10.3897/zookeys.968.56057" ID-GBIF-Dataset="1a953974-667a-4e16-8086-e725d10113f1" ID-PMC="PMC7511452" ID-Pensoft-Pub="1313-2970-968-143" ID-Pensoft-UUID="EA1B5615859A5AE9BB0E416F06BD2DF5" ID-PubMed="33005085" ID-ZooBank="5A7620751B91472BB5CA6F99C4B6600E" ModsDocID="1313-2970-968-143" checkinTime="1600294162043" checkinUser="pensoft" docAuthor="Kai, Yoshiaki, Murasaki, Kenta, Misawa, Ryo, Fukui, Atsushi, Morikawa, Eisuke & Narimatsu, Yoji" docDate="2020" docId="894DC248300858D6BD307AEF22B020D2" docLanguage="en" docName="ZooKeys 968: 143-159" docOrigin="ZooKeys 968" docSource="http://dx.doi.org/10.3897/zookeys.968.56057" docTitle="Paraliparis mandibularis Kido 1985" docType="treatment" docVersion="6" id="EA1B5615859A5AE9BB0E416F06BD2DF5" lastPageNumber="143" masterDocId="EA1B5615859A5AE9BB0E416F06BD2DF5" masterDocTitle="A new species of snailfish of the genus Paraliparis (Liparidae) from the western North Pacific, with a redescription of the poorly known species Paraliparis mandibularis" masterLastPageNumber="159" masterPageNumber="143" pageNumber="143" updateTime="1732848875551" updateUser="ExternalLinkService">
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<mods:title id="5F5A8F98C97A1599BBADE6E90EC2643D">A new species of snailfish of the genus Paraliparis (Liparidae) from the western North Pacific, with a redescription of the poorly known species Paraliparis mandibularis</mods:title>
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<mods:namePart id="42A9D35A3651FED41DDBAA7B8F8F3C3A">Kai, Yoshiaki</mods:namePart>
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<mods:affiliation id="30F0DFB7B9B5CF75FFC9C2DF5769F10B">Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Nagahama, Maizuru, Kyoto 625 - 0086, Japan</mods:affiliation>
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<mods:nameIdentifier id="0973E3ADA497C0F91DD4DAFEA9CBF539" type="orcid">https://orcid.org/0000-0002-0991-5173</mods:nameIdentifier>
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<mods:nameIdentifier id="E833FB2582A94965BE3C1B406DC4F2F7" type="email">mebaru@kais.kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="9F524FBC2F7D3483369C530BA0EC69C2">Murasaki, Kenta</mods:namePart>
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<mods:affiliation id="52AE38E7C8623077A70835039A9D8C72">Institute of Oceanic Research and Development, Tokai University, 3 - 20 - 1 Orido Shimizu, Shizuoka 424 - 8610, Japan</mods:affiliation>
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<mods:namePart id="92EE291F912C6BAFA75FDFF947547049">Misawa, Ryo</mods:namePart>
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<mods:affiliation id="9388FCBCF1D707B3119EDA917BA15A52">Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, 25 - 259 Same, Hachinohe, Aomori, 031 - 0841, Japan</mods:affiliation>
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<mods:namePart id="4212DE6679DFDB568DABF06FF990D1FA">Fukui, Atsushi</mods:namePart>
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<mods:affiliation id="FD076B03D1AD8093BD2A1899A2971520">School of Marine Science and Technology, Tokai University, 3 - 20 - 1 Orido, Shimizu, Shizuoka 424 - 8610, Japan</mods:affiliation>
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<mods:namePart id="DC9CB44C18CDFE3A9044949F620E1261">Morikawa, Eisuke</mods:namePart>
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<mods:affiliation id="7AB386DCCBE973F3A9429087D22AF2ED">Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, 25 - 259 Same, Hachinohe, Aomori, 031 - 0841, Japan</mods:affiliation>
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<mods:namePart id="F587777B28E34E05445B1974C505B3F2">Narimatsu, Yoji</mods:namePart>
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<mods:affiliation id="DCE7D4CA00959BA8CD55D3E00C6B381A">Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, 25 - 259 Same, Hachinohe, Aomori, 031 - 0841, Japan</mods:affiliation>
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<mods:classification id="597F67010615A0BBF38A0AFE70BE8F0C">journal article</mods:classification>
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<mods:identifier id="33658C5B89D6EAC884DF564CB64EBB95" type="DOI">http://dx.doi.org/10.3897/zookeys.968.56057</mods:identifier>
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<treatment id="894DC248300858D6BD307AEF22B020D2" ID-GBIF-Taxon="167616163" LSID="urn:lsid:plazi:treatment:894DC248300858D6BD307AEF22B020D2" httpUri="http://treatment.plazi.org/id/894DC248300858D6BD307AEF22B020D2" lastPageNumber="143" pageId="0" pageNumber="143" scope_family="Liparidae">
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<subSubSection id="683832141B1AF149DADDA06F4F31B5D9" pageId="0" pageNumber="143" type="nomenclature">
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<paragraph id="9C04758C2666106D394F3E53F45BEAE2" pageId="0" pageNumber="143">
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<taxonomicName id="E1E5BE8044E849070D833142E5C3152C" ID-CoL="4CZDL" LSID="894DC248-3008-58D6-BD30-7AEF22B020D2" authority="Kido, 1985" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">Paraliparis mandibularis Kido, 1985</taxonomicName>
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<figureCitation id="23F7A6F4B5AE9C5EF36EF5BA80D62475" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B teeth on lower jaw C, D pectoral girdle A, C Paraliparis flammeus sp. nov., FAKU 147158, paratype, 61.2 mm SL B, D P. mandibularis, BSKU 44286, 110.0 + mm SL. Abbreviations: S, scapula; PR, proximal radials; C, coracoid; DR, distal radials. Scale bars: 0.5 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.968.56057.figure3" httpUri="https://binary.pensoft.net/fig/453540" pageId="0" pageNumber="143">Figs 3B, D</figureCitation>
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<figureCitation id="C112AC4461A524E79A0E57E74D69F173" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paraliparis mandibularis, BSKU 43451, 128.1 mm SL, fresh specimen. Photo by BSKU." figureDoi="10.3897/zookeys.968.56057.figure6" httpUri="https://binary.pensoft.net/fig/453543" pageId="0" pageNumber="143">, 6 [Japanese name: Ago-inkiuo]</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EAAA9A7B0EF1F00150257DCB27594F64" pageId="0" pageNumber="143" type="reference_group">
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<paragraph id="D97D7F087D4C32644312A0DC2698B9DC" pageId="0" pageNumber="143">
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<taxonomicName id="4FE2BD855FA8BA0F05C9BC9D6DD37D09" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">Paraliparis mandibularis</taxonomicName>
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<bibRefCitation id="62ABD143505D12A63C9E06DEBE440226" author="Kido, K" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="143" pagination="362 - 368" refId="B23" refString="Kido, K, 1985. New and rare species of the genus Paraliparis (family Liparididae) from southern Japan. Japanese Journal of Ichthyology 31: 362 - 368" title="New and rare species of the genus Paraliparis (family Liparididae) from southern Japan." volume="31" year="1985">Kido 1985</bibRefCitation>
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: 362, figs 2-4, Tosa Bay, Kochi, Japan;
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<bibRefCitation id="F543007A0F493FC8B8E20DABBCFEBEA8" author="Kido, K" journalOrPublisher="Memoirs of the Faculty of Fisheries Hokkaido University" pageId="0" pageNumber="143" pagination="125 - 256" refId="B24" refString="Kido, K, 1988. Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memoirs of the Faculty of Fisheries Hokkaido University 35: 125 - 256" title="Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan." volume="35" year="1988">Kido 1988</bibRefCitation>
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: 234, fig. 57 (modified from
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<bibRefCitation id="0F465EC3B605E9A9B343D80A1C04362A" author="Kido, K" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="143" pagination="362 - 368" refId="B23" refString="Kido, K, 1985. New and rare species of the genus Paraliparis (family Liparididae) from southern Japan. Japanese Journal of Ichthyology 31: 362 - 368" title="New and rare species of the genus Paraliparis (family Liparididae) from southern Japan." volume="31" year="1985">Kido 1985</bibRefCitation>
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);
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<bibRefCitation id="6096005988A47F89ECC0ACCEA3A4FDF9" author="Shinohara, G" journalOrPublisher="National Museum of Nature and Science Monographs" pageId="0" pageNumber="143" pagination="283 - 343" refId="B36" refString="Shinohara, G, Endo, H, Matsuura, K, Machida, Y, Honda, H, 2001. Annotated checklist of the deepwater fishes from Tosa Bay, Japan. National Museum of Nature and Science Monographs 20: 283 - 343" title="Annotated checklist of the deepwater fishes from Tosa Bay, Japan." volume="20" year="2001">Shinohara et al. 2001</bibRefCitation>
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: 320, listed, Tosa Bay, Kochi, Japan;
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<bibRefCitation id="3A5196BF8CB7DADC2426856589295677" author="Nakabo, T" editor="Nakabo, T" journalOrPublisher="Third Edition. Tokai University Press, Hadano, 1205 - 1218" pageId="0" pageNumber="143" pagination="2072 - 2076" refId="B32" refString="Nakabo, T, Kai, Y, 2013. Liparidae. In: Nakabo, T, Ed., Fishes of Japan with Pictorial Keys to the Species. Third Edition. Tokai University Press, Hadano, 1205-1218: 2072 - 2076" title="Liparidae." volumeTitle="Fishes of Japan with Pictorial Keys to the Species." year="2013">Nakabo and Kai 2013</bibRefCitation>
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: 1217, key, unnumbered fig., Tosa Bay, Kochi, Japan.
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</paragraph>
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</subSubSection>
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<subSubSection id="0B58F8BCAD8CEF3B6F475A83290A2001" pageId="0" pageNumber="143" type="materials_examined">
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<paragraph id="D7C8241C18A4C8F055FB5780AF1BB430" pageId="0" pageNumber="143">Materials examined.</paragraph>
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<paragraph id="351145F240817CECE1B143E3B8D959E9" pageId="0" pageNumber="143">
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BSKU 30513 (holotype of
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<taxonomicName id="D34F04736E6EF6710BD68A7CC7A8E729" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
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<emphasis id="18A16569F1F36E3F9C9A35B1A03C9FF6" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
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</taxonomicName>
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), 103.6 mm SL,
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<geoCoordinate id="6F93DC378BEA3F14C0A0F0ED21EC825F" degrees="32.967" direction="north" orientation="latitude" precision="55" value="32.967">32.967°N</geoCoordinate>
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,
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<geoCoordinate id="1E83FF2CD267500E7A44C8E3D124D1CE" degrees="133.533" direction="east" orientation="longitude" precision="55" value="133.533">133.533°E</geoCoordinate>
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, 605 m depth, Tosa Bay, Kochi, Japan; BSKU 43451, 128.1 mm SL,
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<geoCoordinate id="325F6D92F64D6BC4B68D713E31CC9E93" degrees="32.545" direction="north" orientation="latitude" precision="55" value="32.545">32.545°N</geoCoordinate>
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,
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<geoCoordinate id="1CF0CF616B0C06F77066CD99285CF242" degrees="132.433" direction="east" orientation="longitude" precision="55" value="132.433">132.433°E</geoCoordinate>
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, 1,075-1,092 m depth, off Bungo Channel, Ehime, Japan; BSKU 44260, 44262, 44267, 44269 (C&S), 44397, 44398, 111.4-119.1 mm SL, 600 m depth, Tosa Bay, Kochi, Japan.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5539DA5BEAB812BB049D9512A2334AD" pageId="0" pageNumber="143" type="diagnosis">
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<paragraph id="8118AD9B833013707CF726891ED42F59" pageId="0" pageNumber="143">Diagnosis.</paragraph>
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<paragraph id="3FAD79F03CA3186FBA5D6AEADE83FC24" pageId="0" pageNumber="143">
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<taxonomicName id="77F0A65095610D704455FE81FE36F194" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">
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<emphasis id="2F0E9D5674BD96B512FFA9D6DCAEEE9E" italics="true" pageId="0" pageNumber="143">Paraliparis mandibularis</emphasis>
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</taxonomicName>
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is distinguished from other species of
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<taxonomicName id="C39EA9C22214EBC45B369365784521C1" authorityName="Collett" authorityYear="1879" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="genus">
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<emphasis id="89F2B4E83918620586157C03D06D9D26" italics="true" pageId="0" pageNumber="143">Paraliparis</emphasis>
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</taxonomicName>
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by the following combination of characters: mouth oblique; uppermost pectoral-fin base below a horizontal through posterior margin of maxillary; 63-66 vertebrae, 58-61 dorsal-fin rays, 52-54 anal-fin rays, 6 principal caudal-fin rays, and 27-30 pectoral-fin rays. Proximal pectoral radials 4, enlarged and moved to anterior edge of basal lamina. Parietals present. Among North Pacific species, it is similar to
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<taxonomicName id="7ED512CDE47407D1847762421C2EDB66" lsidName="P. flammeus" pageId="0" pageNumber="143" rank="species" species="flammeus">
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<emphasis id="DB9F951AD2359AFB5247859FCD93E2A1" italics="true" pageId="0" pageNumber="143">P. flammeus</emphasis>
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</taxonomicName>
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sp. nov., which differs from the former in having 17-20 pectoral-fin rays, and to
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<taxonomicName id="A1533D7D154DE6EEF02D9A04A7E2B9DE" lsidName="P. mento" pageId="0" pageNumber="143" rank="species" species="mento">
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<emphasis id="BC17FF4D5F8EB485FDCBA45CF8D182C4" italics="true" pageId="0" pageNumber="143">P. mento</emphasis>
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</taxonomicName>
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, which has 5 principal caudal-fin rays.
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</paragraph>
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</subSubSection>
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<subSubSection id="7CA8E1EC80C5DDDFAE9646DFB55E9FEA" pageId="0" pageNumber="143" type="description">
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<paragraph id="DCA0EAEC08B02525E802187136C640D3" pageId="0" pageNumber="143">Description.</paragraph>
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<paragraph id="A296FEA6EC7C1EF3D8C31350866EC55D" pageId="0" pageNumber="143">
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Measurements are shown in Table
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<tableCitation id="402B01B59374069F8C31E1D68BBC3AE4" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements of Paraliparis flammeus sp. nov. and P. mandibularis (means in parentheses)." httpUri="http://table.plazi.org/id/5CBBD459F25EF9A6CA90D8A8D5AAE207" pageId="0" pageNumber="143" tableUuid="5CBBD459F25EF9A6CA90D8A8D5AAE207">1</tableCitation>
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. Body compressed, elongate, deepest at nape, taping posteriorly (Fig.
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<figureCitation id="5D6E437C47568FFA1829A9C61D35CFFB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Maximum likelihood phylogenetic tree of Paraliparis and related genera based on COI sequences (492 bp). Support values (≥ 50 % ML bootstrap probability) indicated along branches. Each node labeled with a registration number (red, determined in this study) or an accession number deposited in INSDC or BOLD. Nectoliparis pelagicus included as out-group." figureDoi="10.3897/zookeys.968.56057.figure5" httpUri="https://binary.pensoft.net/fig/453542" pageId="0" pageNumber="143">5</figureCitation>
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). Skin thin, fragile. Head compressed, dorsal profile strongly sloping from nape to snout. Snout deep, abruptly angled, its length almost equal to orbit diameter; not projecting anterior to upper jaw. Mouth strongly oblique, lower jaw slightly protruding beyond (or almost same length as) upper jaw; premaxillary tooth plates matching mandibular tooth plates; maxilla extending to posterior margin of orbit; oral cleft extending to middle of orbit. Premaxillary teeth simple, in 3-6 oblique rows; diastema narrow between premaxillae. Mandibular teeth simple, in 4 or 5 oblique rows (Fig.
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<figureCitation id="94063D145CFEE7CE3D0FC58F54445884" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B teeth on lower jaw C, D pectoral girdle A, C Paraliparis flammeus sp. nov., FAKU 147158, paratype, 61.2 mm SL B, D P. mandibularis, BSKU 44286, 110.0 + mm SL. Abbreviations: S, scapula; PR, proximal radials; C, coracoid; DR, distal radials. Scale bars: 0.5 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.968.56057.figure3" httpUri="https://binary.pensoft.net/fig/453540" pageId="0" pageNumber="143">3B</figureCitation>
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); diastema absent at lower jaw symphysis. Orbit of moderate size, rounded. Nostril single, without distinct tube, slightly above level of mid-orbit. Cephalic sensory pores small: nasal pores 2, maxillary pores 6, preoperculomandibular pores 7, suprabranchial pore 1; cephalic pore pattern 2-6-7-1. Chin pores paired, openings well separated on skin surface. Coronal pore absent. Gill slit moderately large, upper margin level with mid-orbit, extending ventrally to just above pectoral fin or to level of 1-3 uppermost pectoral-fin rays. Gill rakers 10-12, blunt and small. Tip of opercular flap sharp, directed posteriorly, level with mid-orbit or posterior margin of maxillary.
|
|
</paragraph>
|
|
<caption id="5CBBD459F25EF9A6CA90D8A8D5AAE207" ID-Table-UUID="5CBBD459F25EF9A6CA90D8A8D5AAE207" httpUri="http://table.plazi.org/id/5CBBD459F25EF9A6CA90D8A8D5AAE207" pageId="0" pageNumber="143" start="Table 1" startId="T1">
|
|
<paragraph id="978AEF4AF199CA391DC89D8FAF3AB15A" pageId="0" pageNumber="143">
|
|
<emphasis id="959269762309229221ABDEF67592704D" bold="true" pageId="0" pageNumber="143">Table 1.</emphasis>
|
|
Measurements of
|
|
<taxonomicName id="7BFA4136548A3142DE4E44BB8D197269" authorityName="Kai, Murasaki & Fukui" authorityYear="2020" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis flammeus" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="flammeus">
|
|
<emphasis id="E44E0B7C5DADE16F194E947359D8566C" italics="true" pageId="0" pageNumber="143">Paraliparis flammeus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="3792910B428D4A14A6002C4E3AA891F1" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
|
|
<emphasis id="8636B9C3580CB44D5731F8600DEDAF72" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
|
|
</taxonomicName>
|
|
(means in parentheses).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="D1E11B2F7904AA110A2F36FD5BC64F2E" pageId="0" pageNumber="143">
|
|
<table id="966758FA4457F0427A5A9C2F88357298" pageId="0" pageNumber="143">
|
|
<tr id="9CAAA0FDC0FDB7F874CE519C9726BB4E" pageId="0" pageNumber="143">
|
|
<td id="634EE330B3D84056B8571775A4591418" colspan="1" isEmpty="true" pageId="0" pageNumber="143" rowspan="3">-</td>
|
|
<td id="49AB28897B10AAC6F47FAB22C6E759C1" colspan="2" pageId="0" pageNumber="143" rowspan="1">
|
|
<taxonomicName id="DFC222EC31EB0A4DC4E26EA72837F0CB" authorityName="Kai, Murasaki & Fukui" authorityYear="2020" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis flammeus" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="flammeus">
|
|
<emphasis id="27120E98A559B8973F7D14327521A0EB" bold="true" italics="true" pageId="0" pageNumber="143">Paraliparis flammeus</emphasis>
|
|
</taxonomicName>
|
|
</td>
|
|
<td id="35A6DA59E42D7FA4A867345DBD86F886" colspan="2" pageId="0" pageNumber="143" rowspan="1">
|
|
<taxonomicName id="34BBB55562D1C82837E23E661349C769" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">
|
|
<emphasis id="5C4153CECB8067528BC7084F0A67323A" bold="true" italics="true" pageId="0" pageNumber="143">Paraliparis mandibularis</emphasis>
|
|
</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="E1561FF8145AAC00E1EC3B88C7B48829" pageId="0" pageNumber="143">
|
|
<td id="066DE31FEB98D1073437A32C62BA97B9" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="DB935CA53AC0B5C38D8EF1BDCAE645FB" bold="true" pageId="0" pageNumber="143">Holotype</emphasis>
|
|
</td>
|
|
<td id="A5C4922030226F6F1093063422998332" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="08E5CCC7A2B70672B72F77F644E15992" bold="true" pageId="0" pageNumber="143">Paratypes</emphasis>
|
|
</td>
|
|
<td id="221B9CA726C9DC6CA09B369133EDA4A5" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="793B68EC0DA796401BA084C0107E45C0" bold="true" pageId="0" pageNumber="143">Holotype</emphasis>
|
|
</td>
|
|
<td id="337EB081D2E5E78508381CAE83016E6A" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="45E67D9987E12DED84495E95312BFF1A" bold="true" pageId="0" pageNumber="143">Non-types</emphasis>
|
|
</td>
|
|
</tr>
|
|
<tr id="020D95E7649C4479F726A62C953CC593" pageId="0" pageNumber="143">
|
|
<td id="3C53A088C32757A6780D468B4E4F9BCD" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="7692D1FA7119CCDA3BD6376CCB510530" bold="true" pageId="0" pageNumber="143">MSM-20-52</emphasis>
|
|
</td>
|
|
<td id="EED1AD9570E480B32E7531FA048AED8D" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="2DA81368FAF5B9131D1DD68FD1D39A26" bold="true" pageId="0" pageNumber="143">
|
|
<emphasis id="B44B2065CCFC28FDDE1878164DDEB298" italics="true" pageId="0" pageNumber="143">n</emphasis>
|
|
= 14
|
|
</emphasis>
|
|
</td>
|
|
<td id="B65E5E4E9BBE4DACB3A054EBA132849B" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="601A5FBECA4E3D66BF51587B81BA4ACA" bold="true" pageId="0" pageNumber="143">BSKU 30513</emphasis>
|
|
</td>
|
|
<td id="05612C495FA1B9F8BC06FBEFAE134BF8" colspan="1" pageId="0" pageNumber="143" rowspan="1">
|
|
<emphasis id="BAC7CF9191C8F9F55C2E7FAFADE81DC6" bold="true" pageId="0" pageNumber="143">
|
|
<emphasis id="45A9F5BD243F25029AF182822B153278" italics="true" pageId="0" pageNumber="143">n</emphasis>
|
|
= 6
|
|
</emphasis>
|
|
</td>
|
|
</tr>
|
|
<tr id="04E1BEBF4532F0DB06AEEE98495F1F39" pageId="0" pageNumber="143">
|
|
<td id="6FACD94D8B9A25DE8BB64FCF7CF4B556" colspan="1" pageId="0" pageNumber="143" rowspan="1">Standard length (mm)</td>
|
|
<td id="0CCF8587368F13418E3BE9E581FF1E24" colspan="1" pageId="0" pageNumber="143" rowspan="1">75.8</td>
|
|
<td id="5014568ED435244432DF50894212A684" colspan="1" pageId="0" pageNumber="143" rowspan="1">42.5-80.4</td>
|
|
<td id="A0A669B40529D8F98D67D669A625CC4A" colspan="1" pageId="0" pageNumber="143" rowspan="1">103.6</td>
|
|
<td id="E6F450E427495E870D68E3A68E676CFC" colspan="1" pageId="0" pageNumber="143" rowspan="1">104.7-128.1</td>
|
|
</tr>
|
|
<tr id="65061D76513EE3F3E8FA276F6B31AAA2" pageId="0" pageNumber="143">
|
|
<td id="488BCF9A2E2A91D775CDA3863CCD38A4" colspan="5" pageId="0" pageNumber="143" rowspan="1">In % of standard length</td>
|
|
</tr>
|
|
<tr id="59FAEDF5F755BC93592EC40011E247EC" pageId="0" pageNumber="143">
|
|
<td id="68853030FFC881E037CC109176B936C8" colspan="1" pageId="0" pageNumber="143" rowspan="1">Head length</td>
|
|
<td id="96D8F691042D83D316498818A9A57E4D" colspan="1" pageId="0" pageNumber="143" rowspan="1">21.4</td>
|
|
<td id="78BCBFD4B75ADCB5EBFB6251D3531422" colspan="1" pageId="0" pageNumber="143" rowspan="1">17.2-24.1 (20.8)</td>
|
|
<td id="062EEDDFD261B83E0161C6DAF3BA9D08" colspan="1" pageId="0" pageNumber="143" rowspan="1">19.8</td>
|
|
<td id="C485F11645253818941803F4AEF3597C" colspan="1" pageId="0" pageNumber="143" rowspan="1">18.0-20.7 (19.8)</td>
|
|
</tr>
|
|
<tr id="2AAA6153029E6ABCE27AF2ABA743AB47" pageId="0" pageNumber="143">
|
|
<td id="3180A2D34A72250A311D81489C026FE3" colspan="1" pageId="0" pageNumber="143" rowspan="1">Snout length</td>
|
|
<td id="360A74A03F87803094D9D049173D30FA" colspan="1" pageId="0" pageNumber="143" rowspan="1">6.7</td>
|
|
<td id="CEE42D4511A096BC4C1DE92B0E3F5971" colspan="1" pageId="0" pageNumber="143" rowspan="1">5.2-7.0 (6.2)</td>
|
|
<td id="FB1279CDFE3FDCF33AA4D9D67B5132C6" colspan="1" pageId="0" pageNumber="143" rowspan="1">6.9</td>
|
|
<td id="1C572A5EF35919CAAAE4EA6FA1676BA6" colspan="1" pageId="0" pageNumber="143" rowspan="1">5.6-6.9 (6.1)</td>
|
|
</tr>
|
|
<tr id="837EF6FDAC8D9716D0C1B6D65C8F4E0D" pageId="0" pageNumber="143">
|
|
<td id="F14A2E3FB9F298CFA95730309C469858" colspan="1" pageId="0" pageNumber="143" rowspan="1">Orbit length</td>
|
|
<td id="5304EEEDFDAF8C84BBF8EE1EE7152209" colspan="1" pageId="0" pageNumber="143" rowspan="1">5.8</td>
|
|
<td id="2DD2E4F5D3EAAB4EF09BC29F91221239" colspan="1" pageId="0" pageNumber="143" rowspan="1">4.5-6.2 (5.4)</td>
|
|
<td id="A050812146CAF9EF286236B7218572F5" colspan="1" pageId="0" pageNumber="143" rowspan="1">5.6</td>
|
|
<td id="9E0F4A63668DABE7850071453C9559EE" colspan="1" pageId="0" pageNumber="143" rowspan="1">4.4-6.0 (5.2)</td>
|
|
</tr>
|
|
<tr id="0D0736CB1BFFEF9A540F0CBF4B7AB453" pageId="0" pageNumber="143">
|
|
<td id="B90BF15C91BD12ED63B08BB103C500B8" colspan="1" pageId="0" pageNumber="143" rowspan="1">Interorbital width</td>
|
|
<td id="A5D191629F27C0E91DFB6F2FE3C9E744" colspan="1" pageId="0" pageNumber="143" rowspan="1">7.2</td>
|
|
<td id="24298EFA75DED24329216DA2B2FFADB8" colspan="1" pageId="0" pageNumber="143" rowspan="1">3.6-8.8 (5.8)</td>
|
|
<td id="3364AFEF3131CBD9723FEB9E17B3F9E9" colspan="1" pageId="0" pageNumber="143" rowspan="1">8.0</td>
|
|
<td id="24123F22CCE5AF96794BA3A35DBA32C0" colspan="1" pageId="0" pageNumber="143" rowspan="1">5.9-7.7 (6.5)</td>
|
|
</tr>
|
|
<tr id="2591200F772335376C8E5CFB9BFA14EB" pageId="0" pageNumber="143">
|
|
<td id="6421373E278ED9E722A4AD1A80FCBDBE" colspan="1" pageId="0" pageNumber="143" rowspan="1">Maxilla length</td>
|
|
<td id="A15C2FA1E3C076BB49EC022F30D78405" colspan="1" pageId="0" pageNumber="143" rowspan="1">10.8</td>
|
|
<td id="BD61DC2D1FB2376B34DDA0C9C92373B2" colspan="1" pageId="0" pageNumber="143" rowspan="1">10.6-12.0 (11.1)</td>
|
|
<td id="7B0E543836D2DBEC82B4D1C051D226B0" colspan="1" pageId="0" pageNumber="143" rowspan="1">10.5</td>
|
|
<td id="E601706510C528B9846738C81C100245" colspan="1" pageId="0" pageNumber="143" rowspan="1">9.7-11.1 (10.6)</td>
|
|
</tr>
|
|
<tr id="25567D2DA31A4AE4A5DA2F5FD28895D6" pageId="0" pageNumber="143">
|
|
<td id="3251CF462774BE5853B60AF0C8BBF4E7" colspan="1" pageId="0" pageNumber="143" rowspan="1">Gill slit length</td>
|
|
<td id="DE6565788BE0F5D7CE71686F69456BA8" colspan="1" pageId="0" pageNumber="143" rowspan="1">10.3</td>
|
|
<td id="D048EAAAD96C9D951B4FF2B9FD28DA88" colspan="1" pageId="0" pageNumber="143" rowspan="1">6.4-10.9 (8.4)</td>
|
|
<td id="A4133A2F0B2AD282CC7E9C7E68945A78" colspan="1" pageId="0" pageNumber="143" rowspan="1">6.9</td>
|
|
<td id="2687FF9F163E11B1A993F18DE3BC88E8" colspan="1" pageId="0" pageNumber="143" rowspan="1">7.5-10.5 (9.1)</td>
|
|
</tr>
|
|
<tr id="B8072AC1B196F45B5A02F6B328A2FFB4" pageId="0" pageNumber="143">
|
|
<td id="3FE0B75AC26F066A13560A9308FE0A72" colspan="1" pageId="0" pageNumber="143" rowspan="1">Body depth</td>
|
|
<td id="BCE1B7DF30D526A4F5525D2CE1969FEC" colspan="1" pageId="0" pageNumber="143" rowspan="1">23.0</td>
|
|
<td id="6C69A95F4DA6E8CA2E32DEE5EAB903B0" colspan="1" pageId="0" pageNumber="143" rowspan="1">13.5-21.7 (18.6)</td>
|
|
<td id="B56AD622E1D5634939B0FB58F7BA38F6" colspan="1" pageId="0" pageNumber="143" rowspan="1">16.1</td>
|
|
<td id="8108F3DA74FFAB578DA10F43D17D9D58" colspan="1" pageId="0" pageNumber="143" rowspan="1">16.0-24.9 (19.8)</td>
|
|
</tr>
|
|
<tr id="6ECA395A577E0395CF274C3EFA999717" pageId="0" pageNumber="143">
|
|
<td id="5E8FDF2E82876A8F3BE1A836558B3FDF" colspan="1" pageId="0" pageNumber="143" rowspan="1">Pectoral-fin length</td>
|
|
<td id="1EBB5D976F20C7BB4AA789BAD7FA5A50" colspan="1" pageId="0" pageNumber="143" rowspan="1">26.2</td>
|
|
<td id="A94DC1B59AD46EBFA7911DA53020ADB3" colspan="1" pageId="0" pageNumber="143" rowspan="1">18.5-26.7 (22.4)</td>
|
|
<td id="3EC922E98F8512349773A5AE03BB18B4" colspan="1" pageId="0" pageNumber="143" rowspan="1">Damaged</td>
|
|
<td id="DA3598BAE9C788237F189AE33A2948F7" colspan="1" pageId="0" pageNumber="143" rowspan="1">21.6-26.0 (23.3)</td>
|
|
</tr>
|
|
<tr id="ED8F10CB079AF3EEC2C1322995960ADA" pageId="0" pageNumber="143">
|
|
<td id="12780BA1BF6F026872D2AC2EB95591DD" colspan="1" pageId="0" pageNumber="143" rowspan="1">Pectoral-fin lower lobe length</td>
|
|
<td id="5EFB964EBDCEE65142AADD7E9DCBEF59" colspan="1" pageId="0" pageNumber="143" rowspan="1">19.6</td>
|
|
<td id="5AAECE47EE13AC7516FF869E5C902A94" colspan="1" pageId="0" pageNumber="143" rowspan="1">16.7-23.4 (20.6)</td>
|
|
<td id="2496DBEBA107501B115B675531A51260" colspan="1" pageId="0" pageNumber="143" rowspan="1">Damaged</td>
|
|
<td id="36819790FA5BB1935E73739588DFEDC5" colspan="1" pageId="0" pageNumber="143" rowspan="1">13.8-15.9 (15.1)</td>
|
|
</tr>
|
|
<tr id="56D0FC108C413206358833A9F0EFA665" pageId="0" pageNumber="143">
|
|
<td id="B5D62133EEA440DD670C2491D7A2E4A9" colspan="1" pageId="0" pageNumber="143" rowspan="1">Pectoral-fin notch-ray length</td>
|
|
<td id="E84DB7678DD098529128E0BC0543DAD9" colspan="1" pageId="0" pageNumber="143" rowspan="1">13.3</td>
|
|
<td id="6C82E5B1E271401490F1F481E73929AE" colspan="1" pageId="0" pageNumber="143" rowspan="1">9.1-15.9 (12.6)</td>
|
|
<td id="AC3DF9C1C348AE100F6B8507DC455E03" colspan="1" pageId="0" pageNumber="143" rowspan="1">Damaged</td>
|
|
<td id="F9819185870386B5198D5B87DF0B04B7" colspan="1" pageId="0" pageNumber="143" rowspan="1">9.6-11.8 (10.7)</td>
|
|
</tr>
|
|
<tr id="D8AFC4C43B048F538B017A0E1DB836E1" pageId="0" pageNumber="143">
|
|
<td id="06A95C834A185E70B3501A2E47F6CDE7" colspan="1" pageId="0" pageNumber="143" rowspan="1">Predorsal length</td>
|
|
<td id="0DF055478C583728059DD93B9E832421" colspan="1" pageId="0" pageNumber="143" rowspan="1">22.4</td>
|
|
<td id="E9F0C074DFC745F1D33CE1171A1E6409" colspan="1" pageId="0" pageNumber="143" rowspan="1">20.3-24.5 (22.5)</td>
|
|
<td id="2BE99861175C266909BA0F9B5193ED34" colspan="1" pageId="0" pageNumber="143" rowspan="1">18.5</td>
|
|
<td id="512841DA448832F6AD277A1921254386" colspan="1" pageId="0" pageNumber="143" rowspan="1">19.7-23.7 (21.2)</td>
|
|
</tr>
|
|
<tr id="7E82452C1D6CA0614FAD2D9C779E5269" pageId="0" pageNumber="143">
|
|
<td id="A1443FF38859D0C017D2B7D117BE8AEF" colspan="1" pageId="0" pageNumber="143" rowspan="1">Preanal length</td>
|
|
<td id="0A0A3042233A9A44FEF0D7BB8ED38928" colspan="1" pageId="0" pageNumber="143" rowspan="1">35.3</td>
|
|
<td id="F22700E9F40C77D5D1B5B0AC1BB17A20" colspan="1" pageId="0" pageNumber="143" rowspan="1">29.9-35.2 (29.9)</td>
|
|
<td id="00A4ADE44D8EBA7E24BDFD1FC365713A" colspan="1" pageId="0" pageNumber="143" rowspan="1">33.2</td>
|
|
<td id="CFED6F3959B6DC1BDF9A0A625B3938F2" colspan="1" pageId="0" pageNumber="143" rowspan="1">31.8-36.1 (33.8)</td>
|
|
</tr>
|
|
<tr id="A9B517DCA9AC04D238DFB938E203F028" pageId="0" pageNumber="143">
|
|
<td id="2ABB3D4DA145AE3CFDF45AE41CC7F285" colspan="1" pageId="0" pageNumber="143" rowspan="1">Snout to anus length</td>
|
|
<td id="4459CC149253104FC9D4053F0D291FA8" colspan="1" pageId="0" pageNumber="143" rowspan="1">16.9</td>
|
|
<td id="1E1C644CE233AD6A5DFFE3C2822F9217" colspan="1" pageId="0" pageNumber="143" rowspan="1">13.0-16.7 (14.5)</td>
|
|
<td id="ADD638DBA26FE41AA92E1EBAB6603C2B" colspan="1" pageId="0" pageNumber="143" rowspan="1">13.2</td>
|
|
<td id="78D233189B69E67027A62D00A832AE79" colspan="1" pageId="0" pageNumber="143" rowspan="1">12.0-18.8 (14.6)</td>
|
|
</tr>
|
|
<tr id="D520CBE4E660D8093134C7EA9C5A806E" pageId="0" pageNumber="143">
|
|
<td id="2E38628BDA3A1D177923446003727111" colspan="1" pageId="0" pageNumber="143" rowspan="1">Caudal-fin length</td>
|
|
<td id="4450D8F23CF016147CCBC730CEF95B31" colspan="1" pageId="0" pageNumber="143" rowspan="1">11.8</td>
|
|
<td id="1A26E3DA61889CE6DD6FA6251BC28570" colspan="1" pageId="0" pageNumber="143" rowspan="1">10.8-15.1 (12.9)</td>
|
|
<td id="492DCD4218EA999EAAA54A69FA30ABEF" colspan="1" pageId="0" pageNumber="143" rowspan="1">Damaged</td>
|
|
<td id="C04F5E382210EE2A726D144406E65E81" colspan="1" pageId="0" pageNumber="143" rowspan="1">15.5-19.6 (17.8)</td>
|
|
</tr>
|
|
<tr id="85B1449E1450C98784B1133978D2F3BB" pageId="0" pageNumber="143">
|
|
<td id="F03290A894CE2C5F37E0E10D43AA5E44" colspan="5" pageId="0" pageNumber="143" rowspan="1">In % of head length</td>
|
|
</tr>
|
|
<tr id="F196D52E063D0A1E24CE914DF044A729" pageId="0" pageNumber="143">
|
|
<td id="76850D03C895FAE3401D7243D668AD2A" colspan="1" pageId="0" pageNumber="143" rowspan="1">Snout length</td>
|
|
<td id="95E4A6F58C63E3E06FC54F8E76070E47" colspan="1" pageId="0" pageNumber="143" rowspan="1">31.3</td>
|
|
<td id="4523012E33C05503BB8B01B0C7E7CDF3" colspan="1" pageId="0" pageNumber="143" rowspan="1">24.5-35.5 (29.9)</td>
|
|
<td id="8B862C0639369C0B7F83595280395FB6" colspan="1" pageId="0" pageNumber="143" rowspan="1">34.6</td>
|
|
<td id="BA8903909B0AB65517DDE02367AC3B32" colspan="1" pageId="0" pageNumber="143" rowspan="1">27.4-34.2 (30.9)</td>
|
|
</tr>
|
|
<tr id="05E28E6ED7B35C0A559F7C6F21EA03C8" pageId="0" pageNumber="143">
|
|
<td id="F1C817B09E875875AC17BFEBF29E9B56" colspan="1" pageId="0" pageNumber="143" rowspan="1">Orbit length</td>
|
|
<td id="7272A8CE75CD44B26F188D52545C2CC2" colspan="1" pageId="0" pageNumber="143" rowspan="1">27.1</td>
|
|
<td id="44076008BDCB82FFAA54CBC37F1F5855" colspan="1" pageId="0" pageNumber="143" rowspan="1">21.7-32.3 (26.1)</td>
|
|
<td id="F6679EAD26830A4E126312074E293122" colspan="1" pageId="0" pageNumber="143" rowspan="1">28.2</td>
|
|
<td id="64853218E4292AF450508F2E4F7DA7DE" colspan="1" pageId="0" pageNumber="143" rowspan="1">23.2-29.0 (26.5)</td>
|
|
</tr>
|
|
<tr id="D348697AAC92F68A70B4F92A29455505" pageId="0" pageNumber="143">
|
|
<td id="C01A819CCE1B5E648FB7F54DEBCC5126" colspan="1" pageId="0" pageNumber="143" rowspan="1">Interorbital width</td>
|
|
<td id="7B9FEF487ED16D28F38F41DD58EFEF9F" colspan="1" pageId="0" pageNumber="143" rowspan="1">33.8</td>
|
|
<td id="0945D803D8D13972A739E324A423BD4A" colspan="1" pageId="0" pageNumber="143" rowspan="1">20.6-36.6 (27.8)</td>
|
|
<td id="60A07839D72F508D4F9FB9A3B07919C0" colspan="1" pageId="0" pageNumber="143" rowspan="1">40.5</td>
|
|
<td id="19BA4D140CDE61A9655D32575C3C1F41" colspan="1" pageId="0" pageNumber="143" rowspan="1">29.8-35.1 (33.0)</td>
|
|
</tr>
|
|
<tr id="C1DC9799BD56E32235CD3539FC045273" pageId="0" pageNumber="143">
|
|
<td id="926F35C6750D44AC0E1F115BC17BC065" colspan="1" pageId="0" pageNumber="143" rowspan="1">Maxilla length</td>
|
|
<td id="07F2DDBD3DEE028DBB9BE95E6C0EC6B1" colspan="1" pageId="0" pageNumber="143" rowspan="1">50.5</td>
|
|
<td id="E3357E54125D972E1C4E0E154C56BE8E" colspan="1" pageId="0" pageNumber="143" rowspan="1">43.8-63.7 (53.7)</td>
|
|
<td id="7348E7803F8AD9B1D3C7C89154F3292A" colspan="1" pageId="0" pageNumber="143" rowspan="1">52.8</td>
|
|
<td id="A2D2E479FE15D14A57AB4FF4AE22C8BF" colspan="1" pageId="0" pageNumber="143" rowspan="1">51.8-56.3 (53.5)</td>
|
|
</tr>
|
|
<tr id="27C5A089A6366693702F0AC0034B4716" pageId="0" pageNumber="143">
|
|
<td id="609B86EB8B0BA525F8D6C67D8C45A771" colspan="1" pageId="0" pageNumber="143" rowspan="1">Gill slit length</td>
|
|
<td id="DE93CD32B6A7710925F943F1A629C72B" colspan="1" pageId="0" pageNumber="143" rowspan="1">48.1</td>
|
|
<td id="589B7F037706ACF0D3F35147A9CA9D65" colspan="1" pageId="0" pageNumber="143" rowspan="1">30.5-18.3 (40.5)</td>
|
|
<td id="C6E13136EA12CF3A6CFEAED79049BF9C" colspan="1" pageId="0" pageNumber="143" rowspan="1">34.7</td>
|
|
<td id="B333799C7D44CD2FC31A2672E059EE0F" colspan="1" pageId="0" pageNumber="143" rowspan="1">37.4-51.9 (46.6)</td>
|
|
</tr>
|
|
<tr id="5EAAE801A3A614FDD38090645116102E" pageId="0" pageNumber="143">
|
|
<td id="39488424DB2A0861C98BFC8E8997169B" colspan="5" pageId="0" pageNumber="143" rowspan="1">In % of caudal-fin length</td>
|
|
</tr>
|
|
<tr id="4C83C3C178AD0B03BF16D9486699AD63" pageId="0" pageNumber="143">
|
|
<td id="A5ED82E59811C0F5A8A5BCF339E5867E" colspan="1" pageId="0" pageNumber="143" rowspan="1">Dorsal-fin connection to caudal fin</td>
|
|
<td id="8367C825FC488AD793EF6A64895149B8" colspan="1" pageId="0" pageNumber="143" rowspan="1">37.9</td>
|
|
<td id="FE96057CA139A694DF9CC393ED86E49B" colspan="1" pageId="0" pageNumber="143" rowspan="1">22.3-37.7 (29.6)</td>
|
|
<td id="398D15D33FDA9CB55B43A137D2E409F4" colspan="1" pageId="0" pageNumber="143" rowspan="1">Damaged</td>
|
|
<td id="9AD16E8B0AE7CBC14EFDF13D09282E04" colspan="1" pageId="0" pageNumber="143" rowspan="1">38.4-50.6 (42.9)</td>
|
|
</tr>
|
|
<tr id="7B40DB8B7911F4304220113B40BD3191" pageId="0" pageNumber="143">
|
|
<td id="C478A3980798A0CEF8EF1554CECE3B48" colspan="1" pageId="0" pageNumber="143" rowspan="1">Anal-fin connection to caudal fin</td>
|
|
<td id="62B9220B7E9328B289628F33538AF437" colspan="1" pageId="0" pageNumber="143" rowspan="1">40.1</td>
|
|
<td id="1975BA26666BD11C973773FC28287954" colspan="1" pageId="0" pageNumber="143" rowspan="1">29.2-44.7 (36.9)</td>
|
|
<td id="3DFF87980C5046CCC005F374CCA2D945" colspan="1" pageId="0" pageNumber="143" rowspan="1">Damaged</td>
|
|
<td id="180356FBA99E3FBE00056DB4E73FE2EE" colspan="1" pageId="0" pageNumber="143" rowspan="1">19.6-36.4 (29.8)</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph id="2BCA4049007B975760AF381DCA495E75" pageId="0" pageNumber="143">
|
|
<emphasis id="7E6EB4A60637A06292838D71DE9887BF" bold="true" italics="true" pageId="0" pageNumber="143">Dorsal-fin</emphasis>
|
|
rays 58-63; anteriormost ray above tip of opercle, posteriormost ray attached membranously to dorsalmost caudal-fin ray. Anteriormost dorsal-fin pterygiophore inserted between neural spines 3 and 4 or 4 and 5, bearing a single ray. Anal-fin rays 52-54; posteriormost ray attached membranously to ventralmost caudal-fin ray. Vertebrae 63-66, comprising precaudal 9 and caudal 54-57. Pleural ribs absent. Hypurals and parhypural fused into single plate. Caudal fin slender, posterior margin slightly rounded. Principal caudal-fin rays 6, dorsal principal rays 3, ventral principal rays 3, no procurrent rays. Pyloric caeca 5 or 6, short and finger-like, on left side of visceral cavity. Anus below posterior margin of orbit.
|
|
</paragraph>
|
|
<paragraph id="67658628DDD369962425821F71A82DF6" pageId="0" pageNumber="143">
|
|
<emphasis id="43C132F842606C50D51F951526DB414C" bold="true" italics="true" pageId="0" pageNumber="143">Pectoral fin</emphasis>
|
|
moderately notched, with 27-30 rays; upper lobe with 17-19 rays, extending beyond (or just reaching) anal-fin origin; lower lobe elongate, with 8-13 rays, uppermost ray of lower lobe longest, extending beyond anus, not reaching anal-fin origin. Uppermost pectoral-fin base below a horizontal through posterior margin of maxillary. Lowermost pectoral-fin base below anterior rim of orbit or below midway between tip of snout and anterior rim of orbit. Rays between upper and lower lobes widely spaced.
|
|
</paragraph>
|
|
<paragraph id="390FEA68495FA5CFE42505EF95EAD8F9" pageId="0" pageNumber="143">
|
|
<emphasis id="52D6E8AB5D36252607700B35C33553A7" bold="true" pageId="0" pageNumber="143">
|
|
<emphasis id="620BA156FFBED2F6D5A898630A9AD3AA" italics="true" pageId="0" pageNumber="143">Selected osteological characters</emphasis>
|
|
.
|
|
</emphasis>
|
|
Roof of cranium without distinct crest comprising well ossified frontals, supraoccipital, and parietals. Opercle well ossified, sharpened posteriorly, supporting upper margin of opercular flap. Subopercle thin, comprising two spines forming a V-shape; lower spine supporting lower margin of opercular flap. Subopercle and interopercle attached. Cleithrum broad and robust, dorsal portion elongated. Proximal pectoral radials 4, enlarged occupying almost entire width of cartilaginous basal laminae and moved to anterior edge of basal lamina (Fig.
|
|
<figureCitation id="9A512120B145BDEFEA1C7B6F1B40D6C4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B teeth on lower jaw C, D pectoral girdle A, C Paraliparis flammeus sp. nov., FAKU 147158, paratype, 61.2 mm SL B, D P. mandibularis, BSKU 44286, 110.0 + mm SL. Abbreviations: S, scapula; PR, proximal radials; C, coracoid; DR, distal radials. Scale bars: 0.5 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.968.56057.figure3" httpUri="https://binary.pensoft.net/fig/453540" pageId="0" pageNumber="143">3D</figureCitation>
|
|
). No interradial fenestrae between proximal radials. Scapula with strong helve. Coracoid narrowly triangular with narrow lamina. Distal radials absent.
|
|
</paragraph>
|
|
<paragraph id="5DC89F7B65C75F3DE224E5395B2A7417" pageId="0" pageNumber="143">
|
|
<emphasis id="5E88B726B37DB1F45DF3AB783DD5AFDA" bold="true" pageId="0" pageNumber="143">
|
|
<emphasis id="84B47EC0A940DFA6FB9C7F5FE52447C9" italics="true" pageId="0" pageNumber="143">Coloration</emphasis>
|
|
.
|
|
</emphasis>
|
|
In fresh specimens, head and body pale pink, posterior half of body reddish; dark peritoneum visible through thin skin; dorsal and anal fins pale pink, distally reddish; caudal and pectoral fins red (Fig.
|
|
<figureCitation id="2C5C199EA044DC89A087ABE3D62E5E40" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paraliparis mandibularis, BSKU 43451, 128.1 mm SL, fresh specimen. Photo by BSKU." figureDoi="10.3897/zookeys.968.56057.figure6" httpUri="https://binary.pensoft.net/fig/453543" pageId="0" pageNumber="143">6</figureCitation>
|
|
). In preserved specimens, head, body, and fins pale; peritoneum black, stomach pale or white; orobranchial cavity pale with scattered melanophores.
|
|
</paragraph>
|
|
<caption id="859AA855F854B5567DB872F01B9A67B7" doi="10.3897/zookeys.968.56057.figure6" httpUri="https://binary.pensoft.net/fig/453543" pageId="0" pageNumber="143" start="Figure 6" startId="F6">
|
|
<paragraph id="C29858698AD97EB1DE06A1CC5DE36F3C" pageId="0" pageNumber="143">
|
|
<emphasis id="2F9BDEFC4443861F4AF4615D09F78D28" bold="true" pageId="0" pageNumber="143">Figure 6.</emphasis>
|
|
<taxonomicName id="314AF9D85FEC238AC98847A1DC7249D5" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">
|
|
<emphasis id="A0E2681D2D825FE7FE64D23948BA200F" italics="true" pageId="0" pageNumber="143">Paraliparis mandibularis</emphasis>
|
|
</taxonomicName>
|
|
, BSKU 43451, 128.1 mm SL, fresh specimen. Photo by BSKU.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="481C126C98B0379C0D0BF8178434001C" pageId="0" pageNumber="143" type="distribution">
|
|
<paragraph id="1226ABA6ED29A96E196FDD6E846E24EE" pageId="0" pageNumber="143">Geographical distribution.</paragraph>
|
|
<paragraph id="FDAECDA3BED953E792903C56D4D40CF1" pageId="0" pageNumber="143">
|
|
Western Pacific Ocean; Tosa Bay and Bungo channel, off Shikoku Island, Japan, in depths of 600-1,092 m (
|
|
<bibRefCitation id="4CE50DC867C2855883F9BA1C44B5AAD4" author="Kido, K" journalOrPublisher="Memoirs of the Faculty of Fisheries Hokkaido University" pageId="0" pageNumber="143" pagination="125 - 256" refId="B24" refString="Kido, K, 1988. Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memoirs of the Faculty of Fisheries Hokkaido University 35: 125 - 256" title="Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan." volume="35" year="1988">Kido 1988</bibRefCitation>
|
|
; this study).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="18EEBDF0A3FB2DACC65EB4B62BE12950" pageId="0" pageNumber="143" type="remarks">
|
|
<paragraph id="DDA12DB0319153226054FC0D26C401A7" pageId="0" pageNumber="143">Remarks.</paragraph>
|
|
<paragraph id="1013012DAD10F2C39C4CB8FE618E83B1" pageId="0" pageNumber="143">
|
|
<taxonomicName id="47582A2FBCCF8D25605C860A27E343CD" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">
|
|
<emphasis id="AAE012877BFFB19C3BCFFDABD5C52C40" italics="true" pageId="0" pageNumber="143">Paraliparis mandibularis</emphasis>
|
|
</taxonomicName>
|
|
was originally described by
|
|
<bibRefCitation id="EDAEAA0B3745FE62B0FE9EACA1A9781F" author="Kido, K" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="143" pagination="362 - 368" refId="B23" refString="Kido, K, 1985. New and rare species of the genus Paraliparis (family Liparididae) from southern Japan. Japanese Journal of Ichthyology 31: 362 - 368" title="New and rare species of the genus Paraliparis (family Liparididae) from southern Japan." volume="31" year="1985">Kido (1985)</bibRefCitation>
|
|
on the basis of a single specimen collected from Tosa Bay, Japan. Subsequently,
|
|
<bibRefCitation id="FBE8B85AAD9C1DBEC9B406EF7114502B" author="Kido, K" journalOrPublisher="Memoirs of the Faculty of Fisheries Hokkaido University" pageId="0" pageNumber="143" pagination="125 - 256" refId="B24" refString="Kido, K, 1988. Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memoirs of the Faculty of Fisheries Hokkaido University 35: 125 - 256" title="Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan." volume="35" year="1988">Kido (1988)</bibRefCitation>
|
|
redescribed the species on the basis of the holotype and an additional non-type specimen. However, a number of details, including osteology and fresh coloration have remained unknown to date. Whereas the pectoral girdle in
|
|
<taxonomicName id="A2F19416DE6797DC90FD610165D76B68" authorityName="Collett" authorityYear="1879" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="genus">
|
|
<emphasis id="D86151B9D12454D58AA0095916B97EC3" italics="true" pageId="0" pageNumber="143">Paraliparis</emphasis>
|
|
</taxonomicName>
|
|
species generally has a reduced number and size of radials (
|
|
<bibRefCitation id="AC2D7FD85893F46CD753824F75F0A7AC" author="Andriashev, AP" journalOrPublisher="Cybium" pageId="0" pageNumber="143" pagination="255 - 266" publicationUrl="http://sfi-cybium.fr/fr/review-recent-studies-southern-ocean-liparidae-teleostei-scorpaeniformes" refId="B1" refString="Andriashev, AP, 1998. A review of recent studies of Southern Ocean Liparidae (Teleostei: Scorpaeniformes). Cybium 22: 255 - 266, http://sfi-cybium.fr/fr/review-recent-studies-southern-ocean-liparidae-teleostei-scorpaeniformes" title="A review of recent studies of Southern Ocean Liparidae (Teleostei: Scorpaeniformes)." url="http://sfi-cybium.fr/fr/review-recent-studies-southern-ocean-liparidae-teleostei-scorpaeniformes" volume="22" year="1998">Andriashev 1998</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="D6BFDC9680931DAD07D12D2B2A217A1F" DOI="https://doi.org/10.11646/zootaxa.4642.1.1" author="Orr, JW" journalOrPublisher="Zootaxa" pageId="0" pageNumber="143" pagination="1 - 79" refId="B34" refString="Orr, JW, Spies, I, Stevenson, DE, Longo, GC, Kai, Y, Ghodes, S, Hollowed, M, 2019. Molecular phylogenetics of snailfishes (Liparidae: Cottoidei) based on mtDNA and RADseq genomic analyses, with comments on selected morphological characters. Zootaxa 4642: 1 - 79, DOI: https://doi.org/10.11646/zootaxa.4642.1.1" title="Molecular phylogenetics of snailfishes (Liparidae: Cottoidei) based on mtDNA and RADseq genomic analyses, with comments on selected morphological characters." url="https://doi.org/10.11646/zootaxa.4642.1.1" volume="4642" year="2019">Orr et al. 2019</bibRefCitation>
|
|
), that of
|
|
<taxonomicName id="1758D12093717376E9696D53165DDB47" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
|
|
<emphasis id="DAE340A554DC176A0B72B43D57D5DE36" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
|
|
</taxonomicName>
|
|
has four broad robust radials (Fig.
|
|
<figureCitation id="D76E0BAE5A86BC57591297BDA5729C5E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B teeth on lower jaw C, D pectoral girdle A, C Paraliparis flammeus sp. nov., FAKU 147158, paratype, 61.2 mm SL B, D P. mandibularis, BSKU 44286, 110.0 + mm SL. Abbreviations: S, scapula; PR, proximal radials; C, coracoid; DR, distal radials. Scale bars: 0.5 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.968.56057.figure3" httpUri="https://binary.pensoft.net/fig/453540" pageId="0" pageNumber="143">3D</figureCitation>
|
|
). The species is also characterized by a pair of parietals in the cranium, such being absent in some other species of
|
|
<taxonomicName id="D0B9FDA77C2D00BE9ACE027B44F55CE6" authorityName="Collett" authorityYear="1879" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="genus">
|
|
<emphasis id="762A6BB3FFF4FE533AEF923448BE4AE7" italics="true" pageId="0" pageNumber="143">Paraliparis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="6D264A48AC218A281A5BA3BD9BA197A3" author="Kido, K" journalOrPublisher="Memoirs of the Faculty of Fisheries Hokkaido University" pageId="0" pageNumber="143" pagination="125 - 256" refId="B24" refString="Kido, K, 1988. Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memoirs of the Faculty of Fisheries Hokkaido University 35: 125 - 256" title="Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan." volume="35" year="1988">Kido 1988</bibRefCitation>
|
|
). Although
|
|
<bibRefCitation id="3721BAA690BFF686E603B3D28B590499" author="Kido, K" journalOrPublisher="Memoirs of the Faculty of Fisheries Hokkaido University" pageId="0" pageNumber="143" pagination="125 - 256" refId="B24" refString="Kido, K, 1988. Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memoirs of the Faculty of Fisheries Hokkaido University 35: 125 - 256" title="Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan." volume="35" year="1988">Kido (1988)</bibRefCitation>
|
|
described
|
|
<taxonomicName id="6B9AC52AC043EF7CA2CCECFAC72DE00A" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
|
|
<emphasis id="9B21E0A83EDBDDF7C1EC4FE9221D3093" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
|
|
</taxonomicName>
|
|
as having a diastema at the symphysis of both the upper and lower jaws, we could find no obvious diastema at the lower jaw symphysis in the specimens examined here, including the holotype. In addition to the differences described above between
|
|
<taxonomicName id="AFB29964F4F1051AAC50402156EA34EC" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
|
|
<emphasis id="25625A6D044DC4A7A1F7DA69D9AF3EBE" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="FE1D5DB3B9E7BD7F704F8BC04AD452B3" lsidName="P. flammeus" pageId="0" pageNumber="143" rank="species" species="flammeus">
|
|
<emphasis id="97B88C7B51E8A4096DADB3AEFA7AAE74" italics="true" pageId="0" pageNumber="143">P. flammeus</emphasis>
|
|
</taxonomicName>
|
|
, the former is further distinguished from the latter by the enlarged pectoral radials occupying almost entire width of the cartilaginous basal laminae and moved to the anterior edge of basal lamina in the pectoral girdle (vs moderately large and medial) and the parietals in the cranium (vs. absent).
|
|
<taxonomicName id="C23F12868709ECE979DC3A9D98F00FB6" authorityName="Kido" authorityYear="1985" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis mandibularis" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="mandibularis">
|
|
<emphasis id="AF0F5AC9E2601535D47BB176733A09CA" italics="true" pageId="0" pageNumber="143">Paraliparis mandibularis</emphasis>
|
|
</taxonomicName>
|
|
differs from other species with an oblique mouth and a low positioned pectoral fin, viz.
|
|
<taxonomicName id="DFA0CB2FF6279E3AE5170EC7A2F06625" lsidName="P. angustifrons" pageId="0" pageNumber="143" rank="species" species="angustifrons">
|
|
<emphasis id="7C6E5B469F7BFEBB070A8BB52671411A" italics="true" pageId="0" pageNumber="143">P. angustifrons</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="EE9C71B09BFB44D474F918005D55A986" lsidName="P. membranaceus" pageId="0" pageNumber="143" rank="species" species="membranaceus">
|
|
<emphasis id="C56F5268A4D30D386CD557C11756D339" italics="true" pageId="0" pageNumber="143">P. membranaceus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="BA1737868A2FE36EFA2D2725AF8F012A" lsidName="P. molinai" pageId="0" pageNumber="143" rank="species" species="molinai">
|
|
<emphasis id="85ABBCB5CB9880F68AB7E078AFAB9AB5" italics="true" pageId="0" pageNumber="143">P. molinai</emphasis>
|
|
</taxonomicName>
|
|
(see Remarks under
|
|
<taxonomicName id="7E2C3371FA5DA7F58AE03E9C8D23A7AC" lsidName="P. flammeus" pageId="0" pageNumber="143" rank="species" species="flammeus">
|
|
<emphasis id="1131E54A3F50D0809A835B6D9EA6C00D" italics="true" pageId="0" pageNumber="143">P. flammeus</emphasis>
|
|
</taxonomicName>
|
|
), in having 27-30 pectoral-fin rays (vs 37 in
|
|
<taxonomicName id="67E2AC3CA2117C600B8D7FB947C96F67" lsidName="P. angustifrons" pageId="0" pageNumber="143" rank="species" species="angustifrons">
|
|
<emphasis id="57F809679B9EA0435391FB12D0D250EA" italics="true" pageId="0" pageNumber="143">P. angustifrons</emphasis>
|
|
</taxonomicName>
|
|
, ca 25 in
|
|
<taxonomicName id="D5DD908D2FCC0C9088DA152D9A910BF4" lsidName="P. membranaceus" pageId="0" pageNumber="143" rank="species" species="membranaceus">
|
|
<emphasis id="CA4C08FA12FF8F206450DDE100EFFC5C" italics="true" pageId="0" pageNumber="143">P. membranaceus</emphasis>
|
|
</taxonomicName>
|
|
, 24 in
|
|
<taxonomicName id="DA0E9BA97105C123E2619A6EEE01A0A0" lsidName="P. molinai" pageId="0" pageNumber="143" rank="species" species="molinai">
|
|
<emphasis id="5E4524281E8F580C1CDCDA8661D57412" italics="true" pageId="0" pageNumber="143">P. molinai</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
<taxonomicName id="7C69A742A12B2F544684801E553D13E9" family="Liparidae" genus="Paraliparis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraliparis membranaceus" order="Scorpaeniformes" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="membranaceus">
|
|
<emphasis id="363845D536C8C8A9CA8BA908E157F294" italics="true" pageId="0" pageNumber="143">Paraliparis membranaceus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="3E93AA3E61A97DD1FF43ED7AC423A7F5" lsidName="P. molinai" pageId="0" pageNumber="143" rank="species" species="molinai">
|
|
<emphasis id="68275B3315470CDD4485E8F58423667E" italics="true" pageId="0" pageNumber="143">P. molinai</emphasis>
|
|
</taxonomicName>
|
|
have similar pectoral-fin ray numbers to
|
|
<taxonomicName id="6098E483DAE33E9810089EC1FE715811" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
|
|
<emphasis id="66353847FF8ECF71294F848D5AA7C2B7" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
|
|
</taxonomicName>
|
|
but have a reduced caudal fin (4 rays in
|
|
<taxonomicName id="6697B7C186413DCE247E409EBFE67B25" lsidName="P. molinai" pageId="0" pageNumber="143" rank="species" species="molinai">
|
|
<emphasis id="881689749AC4AFA44D63F0E78EEFF3B1" italics="true" pageId="0" pageNumber="143">P. molinai</emphasis>
|
|
</taxonomicName>
|
|
, 2 or 3 in
|
|
<taxonomicName id="5F7CAD3C2125C59B5060C8959D622B31" lsidName="P. membranaceus" pageId="0" pageNumber="143" rank="species" species="membranaceus">
|
|
<emphasis id="35835209C8768BCCDABDCB210B8AF99C" italics="true" pageId="0" pageNumber="143">P. membranaceus</emphasis>
|
|
</taxonomicName>
|
|
vs 6 in
|
|
<taxonomicName id="6CAB289348F8D705204D4A691F0BF1C9" lsidName="P. mandibularis" pageId="0" pageNumber="143" rank="species" species="mandibularis">
|
|
<emphasis id="31AC77C60C5E20EDA17252853B6DA7A6" italics="true" pageId="0" pageNumber="143">P. mandibularis</emphasis>
|
|
</taxonomicName>
|
|
) (
|
|
<bibRefCitation id="87A9DB67FB217F52193E3FB862CF8830" DOI="https://doi.org/10.11646/zootaxa.1019.1.1" author="Stein, DL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="143" pagination="1 - 25" refId="B39" refString="Stein, DL, 2005. Descriptions of four new species, redescription of Paraliparis membranaceus, and additional data on species of the fish family Liparidae (Pisces, Scorpaeniformes) from the west coast of South America and the Indian Ocean. Zootaxa 1019: 1 - 25, DOI: https://doi.org/10.11646/zootaxa.1019.1.1" title="Descriptions of four new species, redescription of Paraliparis membranaceus, and additional data on species of the fish family Liparidae (Pisces, Scorpaeniformes) from the west coast of South America and the Indian Ocean." url="https://doi.org/10.11646/zootaxa.1019.1.1" volume="1019" year="2005">Stein 2005</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |