456 lines
38 KiB
XML
456 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.70.97117" ID-Pensoft-Pub="1860-1324-1-69" ID-Pensoft-UUID="CA0CBE818BFA5DFA80DBD42521D5DBE3" ID-ZooBank="99BBA4C8ED2048879952B61CC25309D4" ModsDocID="1860-1324-70-1-69" checkinTime="1679031804748" checkinUser="pensoft" docAuthor="Tran, Anh Duc, Zettel, Herbert & Sites, Robert W." docDate="2023" docId="894CBD714AF7598192F85A2FA0A1A60F" docLanguage="en" docName="DeutEntomolZeit 70(1): 69-111" docOrigin="Deutsche Entomologische Zeitschrift 70 (1)" docPubDate="2023-03-16" docSource="http://dx.doi.org/10.3897/dez.70.97117" docTitle="Eotrechus steineri Tran & Zettel & Sites 2023, sp. nov." docType="treatment" docUuid="33671505-47A3-4991-B071-BF9A8042AB5D" docUuidSource="ZooBank" docVersion="2" id="CA0CBE818BFA5DFA80DBD42521D5DBE3" lastPageNumber="69" masterDocId="CA0CBE818BFA5DFA80DBD42521D5DBE3" masterDocTitle="Revision of the genus Eotrechus Kirkaldy (Hemiptera, Heteroptera, Gerridae), with descriptions of six new species" masterLastPageNumber="111" masterPageNumber="69" pageNumber="69" updateTime="1679032168055" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Eotrechus Kirkaldy (Hemiptera, Heteroptera, Gerridae), with descriptions of six new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tran, Anh Duc</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9605-0739</mods:nameIdentifier>
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<mods:affiliation>Faculty of Biology, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier type="email">tran.anhduc@hus.edu.vn</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zettel, Herbert</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7760-0472</mods:nameIdentifier>
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<mods:affiliation>Herbert Zettel, 2 nd Zoological Department (Entomology), Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sites, Robert W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3895-813X</mods:nameIdentifier>
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<mods:affiliation>Enns Entomology Museum, University of Missouri, Columbia, MO 65211, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>70</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>111</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/dez.70.97117</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.70.97117</mods:identifier>
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||
<mods:identifier type="Pensoft-Pub">1860-1324-1-69</mods:identifier>
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<mods:identifier type="ZooBank">99BBA4C8ED2048879952B61CC25309D4</mods:identifier>
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<mods:identifier type="Pensoft-UUID">CA0CBE818BFA5DFA80DBD42521D5DBE3</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:33671505-47A3-4991-B071-BF9A8042AB5D" httpUri="http://treatment.plazi.org/id/894CBD714AF7598192F85A2FA0A1A60F" lastPageNumber="69" pageId="0" pageNumber="69">
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<subSubSection pageId="0" pageNumber="69" type="nomenclature">
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||
<paragraph pageId="0" pageNumber="69">
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<taxonomicName LSID="https://zoobank.org/33671505-47A3-4991-B071-BF9A8042AB5D" authority="Tran & Zettel & Sites, 2023" authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus steineri" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="steineri" status="sp. nov.">Eotrechus steineri</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="69">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="69" type="description">
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||
<paragraph pageId="0" pageNumber="69">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eotrechus steineri sp. nov., habitus. A. Apterous male; B. Apterous female." figureDoi="10.3897/dez.70.97117.figure3" httpUri="https://binary.pensoft.net/fig/826092" pageId="0" pageNumber="69">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eotrechus kalidasa, specimen from India (A-C) and E. steineri sp. nov. (D-F). A, D. Pygophore, lateral view; B, E. Proctiger, dorsal view; C, F. Paramere, lateral view (A, B, D, E. Same scale; C, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure4" httpUri="https://binary.pensoft.net/fig/826093" pageId="0" pageNumber="69">, 4D-F</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eotrechus steineri sp. nov. (A-E) and E. kalidasa (F). A. Fore leg of male; B. Abdomen of male, ventral view; C, D. Posterior part of abdomen of female, dorsal and lateral views; E, F. Paramere, lateral view (B-D. Same scale; E, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure5" httpUri="https://binary.pensoft.net/fig/826094" pageId="0" pageNumber="69">, 5A-E</figureCitation>
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Distribution records of species of Eotrechus. Squares represent E. kalidasa species group; circles represent E. pingae species group; triangles represent species unassigned to a group." figureDoi="10.3897/dez.70.97117.figure25" httpUri="https://binary.pensoft.net/fig/826114" pageId="0" pageNumber="69">, 25</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="69" type="reference_group">
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName authority="Tran & Zettel & Sites, 2023" authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus kalidasa" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="kalidasa" status="sp. nov.">Eotrechus kalidasa</taxonomicName>
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(non Kirkaldy, 1902):
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<bibRefCitation author="Vitheepradit, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="1 - 19" publicationUrl="https://www.mapress.com/zootaxa/2007f/zt01478p019.pdf" refId="B19" refString="Vitheepradit, A, Sites, RW, 2007. A review of Eotrechus Kirkaldy (Hemiptera: Heteroptera: Gerridae) of Thailand with descriptions of three new species. Zootaxa 1478: 1 - 19, https://www.mapress.com/zootaxa/2007f/zt01478p019.pdf" title="A review of Eotrechus Kirkaldy (Hemiptera: Heteroptera: Gerridae) of Thailand with descriptions of three new species." url="https://www.mapress.com/zootaxa/2007f/zt01478p019.pdf" volume="1478" year="2007">Vitheepradit and Sites (2007</bibRefCitation>
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: 18-19) (diagnosis, notes);
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3860.1.2" author="Nakthong, LA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="47 - 63" refId="B9" refString="Nakthong, LA, Vitheepradit, A, Sites, RW, 2014. Key to the species of Eotrechinae (Hemiptera: Heteroptera: Gerridae) of Thailand and review of the fauna of the Phetchabun mountain range. Zootaxa 3860 (1): 47 - 63, DOI: https://doi.org/10.11646/zootaxa.3860.1.2" title="Key to the species of Eotrechinae (Hemiptera: Heteroptera: Gerridae) of Thailand and review of the fauna of the Phetchabun mountain range." url="https://doi.org/10.11646/zootaxa.3860.1.2" volume="3860" year="2014">Nakthong et al. (2014</bibRefCitation>
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: 59-60, fig. 42) (key, diagnosis, notes). Misidentifications.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="materials_examined">
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<paragraph pageId="0" pageNumber="69">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<materialsCitation collectingDate="2006-02-10" collectorName="H. Steiner" country="Laos" county="Cave Project" latitude="20.765833" location="Vieng Phoukha" longLatPrecision="1" longitude="101.0138" municipality="Tham Kuat" specimenCount="♂" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="69">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Laos">Laos</collectingCountry>
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•
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<specimenCount>♂</specimenCount>
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(apterous);
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<location LSID="urn:lsid:plazi:treatment:894CBD714AF7598192F85A2FA0A1A60F:82B0016DD621D8FEDFEB21AF46ABD2A8" country="Laos" county="Cave Project" latitude="20.765833" longLatPrecision="1" longitude="101.0138" municipality="Tham Kuat" name="Vieng Phoukha">Vieng Phoukha</location>
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,
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<collectingMunicipality>Tham Kuat</collectingMunicipality>
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(F47-119/010);
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<geoCoordinate degrees="20" direction="north" minutes="45.950" orientation="latitude" precision="1" value="20.765833">20°45.950'N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="0.828" orientation="longitude" precision="1" value="101.0138">101°0.828'E</geoCoordinate>
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;
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<collectingDate value="2006-02-10">10 Feb. 2006</collectingDate>
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;
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<collectorName>H. Steiner</collectorName>
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leg.; 132/06; N-Lao-Europ.
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<collectingCounty>Cave Project</collectingCounty>
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2006; NHMW.
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Laos" location="Laos" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="Paratypes">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="69">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Laos">Laos</collectingCountry>
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•
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<specimenCount type="male">3 ♂♂</specimenCount>
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,
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<specimenCount type="female">2 ♀♀</specimenCount>
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(apterous); same locality data as
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<typeStatus>holotype</typeStatus>
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; NHMW
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</materialsCitation>
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•
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<materialsCitation collectingDate="2005-02-03" collectorName="H. Steiner" country="Laos" county="Tham Suea" latitude="19.4488" location="Cave Project" longLatPrecision="1" longitude="102.43595" municipality="Tham Nam Lot" specimenCount="1" specimenCount-female="1" stateProvince="Luang Prabang" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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(apterous);
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<collectingRegion country="Laos" name="Louangphabang">Luang Prabang</collectingRegion>
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,
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<collectingCounty>Tham Suea</collectingCounty>
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,
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<collectingMunicipality>Tham Nam Lot</collectingMunicipality>
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;
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<geoCoordinate degrees="19" direction="north" minutes="26.928" orientation="latitude" precision="1" value="19.4488">19°26.928'N</geoCoordinate>
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,
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<geoCoordinate degrees="102" direction="east" minutes="26.157" orientation="longitude" precision="1" value="102.43595">102°26.157'E</geoCoordinate>
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;
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<collectingDate value="2005-02-03">3 Feb. 2005</collectingDate>
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;
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<collectorName>H. Steiner</collectorName>
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leg.; 069/05; N-Lao-Europ.
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<location LSID="urn:lsid:plazi:treatment:894CBD714AF7598192F85A2FA0A1A60F:348743A8CC0B951766C129D827755826" country="Laos" county="Tham Suea" latitude="19.4488" longLatPrecision="1" longitude="102.43595" municipality="Tham Nam Lot" name="Cave Project" stateProvince="Luang Prabang">Cave Project</location>
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2005; NHMW
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</materialsCitation>
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||
•
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<materialsCitation collectingDate="2012-01-18" collectorName="Tham A Pa, H. Steiner" country="Laos" latitude="20.830717" location="Northern Lao-European Cave Project" longLatPrecision="1" longitude="101.06222" municipality="Vieng Phoukha" specimenCount="1" specimenCount-female="1" stateProvince="Luang Nam Tha Province" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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(apterous);
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<collectingRegion country="Laos" name="Louangnamtha">Luang Nam Tha Province</collectingRegion>
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,
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<collectingMunicipality>Vieng Phoukha</collectingMunicipality>
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;
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<collectorName>Tham A Pa</collectorName>
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(F47-119-026);
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<geoCoordinate degrees="20" direction="north" minutes="49.843" orientation="latitude" precision="1" value="20.830717">20°49.843'N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="3.733" orientation="longitude" precision="1" value="101.06222">101°3.733'E</geoCoordinate>
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;
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<collectingDate value="2012-01-18">18 Jan. 2012</collectingDate>
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;
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<collectorName>H. Steiner</collectorName>
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leg.; 027/12;
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<location LSID="urn:lsid:plazi:treatment:894CBD714AF7598192F85A2FA0A1A60F:6BC9AC477F0B05C0C2916986B154B8F3" country="Laos" latitude="20.830717" longLatPrecision="1" longitude="101.06222" municipality="Vieng Phoukha" name="Northern Lao-European Cave Project" stateProvince="Luang Nam Tha Province">Northern Lao-European Cave Project</location>
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2012; NHMW
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</materialsCitation>
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||
•
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<materialsCitation collectingDate="2012-01-19" collectorName="H. Steiner" country="Laos" county="Vieng Phouka" latitude="20.722784" location="Northern Lao-European Cave Project" longLatPrecision="1" longitude="101.01605" municipality="Um Chuck Njet" specimenCount="1" specimenCount-female="1" stateProvince="Luang Nam Tha Province" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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(apterous);
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<collectingRegion country="Laos" name="Louangnamtha">Luang Nam Tha Province</collectingRegion>
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,
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<collectingCounty>Vieng Phouka</collectingCounty>
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,
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<collectingMunicipality>Um Chuck Njet</collectingMunicipality>
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(F47-119-029);
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||
<geoCoordinate degrees="20" direction="north" minutes="43.367" orientation="latitude" precision="1" value="20.722784">20°43.367'N</geoCoordinate>
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||
,
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||
<geoCoordinate degrees="101" direction="east" minutes="0.963" orientation="longitude" precision="1" value="101.01605">101°0.963'E</geoCoordinate>
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;
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<collectingDate value="2012-01-19">19 Jan. 2012</collectingDate>
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;
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<collectorName>H. Steiner</collectorName>
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leg.; 138/12;
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<location LSID="urn:lsid:plazi:treatment:894CBD714AF7598192F85A2FA0A1A60F:74E7E172C2EECC4CA8C0B9A4D88ACB3F" country="Laos" county="Vieng Phouka" latitude="20.722784" longLatPrecision="1" longitude="101.01605" municipality="Um Chuck Njet" name="Northern Lao-European Cave Project" stateProvince="Luang Nam Tha Province">Northern Lao-European Cave Project</location>
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2012; NHMW
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph pageId="0" pageNumber="69">
|
||
<materialsCitation collectingDate="2002-09-21" collectingDateMax="2002-10-21" collectingDateMin="2002-09-21" collectorName="Vitheepradit" country="Thailand" determinerName="Kirkaldy" elevation="1600" latitude="16.95" location="Man Daeng Noi" longLatPrecision="1280" longitude="101.05" municipality="Phu Hin Rongkla National Park" specimenCount="1" specimenCount-male="1" stateProvince="Loei Province">
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||
<collectingCountry name="Thailand">Thailand</collectingCountry>
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||
•
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<specimenCount type="male">1 ♂</specimenCount>
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(macropterous);
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<collectingRegion country="Thailand" name="Loei">Loei Province</collectingRegion>
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||
,
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||
<collectingMunicipality>Phu Hin Rongkla National Park</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:894CBD714AF7598192F85A2FA0A1A60F:CC201CE30AEC8EC4C1245DE0DF23F694" country="Thailand" latitude="16.95" longLatPrecision="1280" longitude="101.05" municipality="Phu Hin Rongkla National Park" name="Man Daeng Noi" stateProvince="Loei Province">Man Daeng Noi</location>
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||
at trail;
|
||
<geoCoordinate degrees="16" direction="north" minutes="57" orientation="latitude" precision="925" value="16.95">16°57'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="101" direction="east" minutes="03" orientation="longitude" precision="925" value="101.05">101°03'E</geoCoordinate>
|
||
;
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</quantity>
|
||
a.s.l.
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||
</elevation>
|
||
;
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||
<collectingDate value="2002-09-21" valueMax="2002-10-21" valueMin="2002-09-21">21 Sep.-21 Oct. 2002</collectingDate>
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||
; coll: CMU team, leg.;
|
||
<taxonomicName authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus kalidasa" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus kalidasa</emphasis>
|
||
</taxonomicName>
|
||
<determinerName>Kirkaldy</determinerName>
|
||
, det
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||
<collectorName>Vitheepradit</collectorName>
|
||
2005; UMC
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation collectingDate="2009-04-17" collectorName="Sites, Vitheepradit, Prommi" country="Thailand" elevation="532" latitude="19.19005" location="Namtok Sa Pan" longLatPrecision="1" longitude="101.1975" specimenCount="1" specimenCount-male="1" stateProvince="Nan Province">
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
(macropterous);
|
||
<collectingRegion country="Thailand" name="Nan">Nan Province</collectingRegion>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:894CBD714AF7598192F85A2FA0A1A60F:F73C0A7B7849F660A2BEC2B80E1704EE" country="Thailand" latitude="19.19005" longLatPrecision="1" longitude="101.1975" name="Namtok Sa Pan" stateProvince="Nan Province">Namtok Sa Pan</location>
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||
, stream with waterfall;
|
||
<geoCoordinate degrees="19" direction="north" minutes="11.403" orientation="latitude" precision="1" value="19.19005">19°11.403'N</geoCoordinate>
|
||
,
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||
<geoCoordinate degrees="101" direction="east" minutes="11.850" orientation="longitude" precision="1" value="101.1975">101°11.850'E</geoCoordinate>
|
||
;
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||
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.32" unit="m" value="532.0">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.32" unit="m" value="532.0">532 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectingDate value="2009-04-17">17 Apr. 2009</collectingDate>
|
||
;
|
||
<collectorName>Sites</collectorName>
|
||
,
|
||
<collectorName>Vitheepradit</collectorName>
|
||
,
|
||
<collectorName>Prommi</collectorName>
|
||
leg.; L-1043;
|
||
<taxonomicName authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus kalidasa" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus kalidasa</emphasis>
|
||
</taxonomicName>
|
||
, det
|
||
<collectorName>Vitheepradit</collectorName>
|
||
2017; UMC
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="description">
|
||
<paragraph pageId="0" pageNumber="69">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">Size: apterous males, length 9.25-9.75 (holotype 9.25), width 2.07-2.28 (holotype 2.12); macropterous male, length 10.32, width 2.12; apterous females, length 9.92-10.50, width 2.25-2.36.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Colour (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eotrechus steineri sp. nov., habitus. A. Apterous male; B. Apterous female." figureDoi="10.3897/dez.70.97117.figure3" httpUri="https://binary.pensoft.net/fig/826092" pageId="0" pageNumber="69">3</figureCitation>
|
||
). Apterous form, dorsum of body chiefly brown to dark-brown or black, with yellow markings on head and thorax, covered with golden pubescence; venter chiefly pale-brown. Head with two lateral yellow marks at anterior section and a transverse yellow mark on posterior margin. Antennae yellowish-brown to brown. Pronotum with a yellow midline stripe. Mesonotum with yellow midline stripe thinner posteriorly and usually not reaching posterior margin. Metanotum and abdominal tergum mainly without yellow marking. Propleuron with a longitudinal yellow stripe. Meso- and metapleura black, without yellow marking. All pleura covered with golden pubescence. All coxae chiefly light-coloured (usually light-yellowish); all other leg segments yellowish-brown to brown; apical parts of all femora dark-brown to black.
|
||
</paragraph>
|
||
<caption doi="10.3897/dez.70.97117.figure3" httpUri="https://binary.pensoft.net/fig/826092" pageId="0" pageNumber="69" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus steineri" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="steineri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus steineri</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., habitus.
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A.</emphasis>
|
||
Apterous male;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B.</emphasis>
|
||
Apterous female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Structural characteristics: Apterous male (holotype): Head width across eyes 1.76; interocular width 0.69; eye kidney-shaped in dorsal view, length of eye 0.76. Antennae ca. 1.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
body length (9.94: 9.25), lengths of segments I-IV: 2.64: 2.46: 2.25: 2.59; segment I without black spines. Pronotum broader than long, shorter than head length (1.17: 1.45). Posterior two-thirds of mesonotum strongly swollen, forming a hump. Lengths of mesosternum and metasternum: 1.81 and 1.17. Lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II) as follows: fore leg: 3.86: 2.91: 0.52: 0.48; middle leg: 6.50: 6.67: 0.41: 0.49; hind leg: 7.25: 8.50: 0.40: 0.48. Fore leg (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eotrechus steineri sp. nov. (A-E) and E. kalidasa (F). A. Fore leg of male; B. Abdomen of male, ventral view; C, D. Posterior part of abdomen of female, dorsal and lateral views; E, F. Paramere, lateral view (B-D. Same scale; E, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure5" httpUri="https://binary.pensoft.net/fig/826094" pageId="0" pageNumber="69">5A</figureCitation>
|
||
): trochanter with one or two long, fine setae; femur slender, length ca. 11.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum width (3.86: 0.34), proximal part slightly thicker distal part, flexor side with few long, fine setae about evenly spaced along its length; tibia straight, without short, stout setae on distal third; tarsus covered with dense, short, soft, yellowish setae (denser on ventral surface). Middle and hind femora slender and shorter than body; middle and hind femora with scattered short, stout setae on ventral surface; middle and hind tibiae with very long, stout setae scattered along their lengths. Claws stout, lengths of fore, mid- and hind claws: 0.18: 0.22: 0.21. Abdomen relatively long, total length of abdominal sterna II-VII: 3.06, sterna II-VII weakly depressed medially, not forming distinct groove, depression on sternum VII widened posteriorly. Sternum VII ca. 0.66
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
the length of two preceding sterna combined (0.63: 0.96), posterior margin with shallow median notch (ca. 0.15
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
the length of sternum VII) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eotrechus steineri sp. nov. (A-E) and E. kalidasa (F). A. Fore leg of male; B. Abdomen of male, ventral view; C, D. Posterior part of abdomen of female, dorsal and lateral views; E, F. Paramere, lateral view (B-D. Same scale; E, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure5" httpUri="https://binary.pensoft.net/fig/826094" pageId="0" pageNumber="69">5B</figureCitation>
|
||
). Genitalia (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eotrechus kalidasa, specimen from India (A-C) and E. steineri sp. nov. (D-F). A, D. Pygophore, lateral view; B, E. Proctiger, dorsal view; C, F. Paramere, lateral view (A, B, D, E. Same scale; C, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure4" httpUri="https://binary.pensoft.net/fig/826093" pageId="0" pageNumber="69">4D-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eotrechus steineri sp. nov. (A-E) and E. kalidasa (F). A. Fore leg of male; B. Abdomen of male, ventral view; C, D. Posterior part of abdomen of female, dorsal and lateral views; E, F. Paramere, lateral view (B-D. Same scale; E, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure5" httpUri="https://binary.pensoft.net/fig/826094" pageId="0" pageNumber="69">5B, E</figureCitation>
|
||
): ventral length of abdominal segment VIII: 0.41. Pygophore with two long, relatively slender posterolateral projections which curve dorsad. Paramere long, with proximal third wider, ca. 2.0
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as distal part; distal part slender, directed slightly dorsad, apex rounded. Proctiger simple, longer than wide, with broadly rounded apex, posterolaterally slightly produced as broad processes.
|
||
</paragraph>
|
||
<caption doi="10.3897/dez.70.97117.figure4" httpUri="https://binary.pensoft.net/fig/826093" pageId="0" pageNumber="69" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus kalidasa" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus kalidasa</emphasis>
|
||
</taxonomicName>
|
||
, specimen from India (
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A-C</emphasis>
|
||
) and
|
||
<taxonomicName lsidName="E. steineri" pageId="0" pageNumber="69" rank="species" species="steineri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. steineri</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<emphasis bold="true" pageId="0" pageNumber="69">D-F</emphasis>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A, D.</emphasis>
|
||
Pygophore, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B, E.</emphasis>
|
||
Proctiger, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C, F.</emphasis>
|
||
Paramere, lateral view (
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A, B, D, E.</emphasis>
|
||
Same scale;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C, F.</emphasis>
|
||
Same scale).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/dez.70.97117.figure5" httpUri="https://binary.pensoft.net/fig/826094" pageId="0" pageNumber="69" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus steineri" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="steineri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus steineri</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A-E</emphasis>
|
||
) and
|
||
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A.</emphasis>
|
||
Fore leg of male;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B.</emphasis>
|
||
Abdomen of male, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C, D.</emphasis>
|
||
Posterior part of abdomen of female, dorsal and lateral views;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E, F.</emphasis>
|
||
Paramere, lateral view (
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B-D.</emphasis>
|
||
Same scale;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E, F.</emphasis>
|
||
Same scale).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Apterous female: Head width across eyes 1.89; interocular width 0.75; eye kidney-shaped in dorsal view, length 0.82. Antennae subequal to body length (10.33: 10.50), lengths of segments I-IV: 2.69: 2.51: 2.36: 2.77; segment I without black spines. Pronotum shorter than head length (1.17: 1.55). Posterior two-thirds of mesonotum strongly swollen, forming a hump. Lengths of mesosternum and metasternum: 1.99 and 1.37. Lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 3.97: 3.06: 0.65: 0.52; middle leg: 6.83: 6.96: 0.44: 0.53; hind leg: 7.42: 8.83: 0.45: 0.54. Fore trochanter with two or three long, fine setae on ventral side. Fore femur similar to males, length ca. 11.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum width (3.97: 0.35). Fore tibia similar to males. Middle and hind legs similar to males. Claws stout, lengths of fore, mid- and hind claws: 0.21: 0.22: 0.23. Total length of abdominal sterna II-VII ca. 0.46
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
body length (4.82: 10.5). Sternum VII ca. 0.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
length of two preceding sterna combined (1.17: 1.55), posterior margin straight. Connexival corners of sternum VII broadly rounded. Genitalia almost completely concealed by abdomen in lateral view, only tip of proctiger visible, proctiger broader than long, directed ventrad (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eotrechus steineri sp. nov. (A-E) and E. kalidasa (F). A. Fore leg of male; B. Abdomen of male, ventral view; C, D. Posterior part of abdomen of female, dorsal and lateral views; E, F. Paramere, lateral view (B-D. Same scale; E, F. Same scale)." figureDoi="10.3897/dez.70.97117.figure5" httpUri="https://binary.pensoft.net/fig/826094" pageId="0" pageNumber="69">5C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Macropterous male: Similar to apterous male but with following differences: Head ground colour dark-brown to black, with orange markings at middle of posterior margin and at anterodorsal margin of eye extending halfway to base of antenna; pronotum dark-brown to black, with orange longitudinal stripe on midline; propleuron with orange longitudinal stripe posterior to eye; wings chiefly dark-brown with medium-brown patches, reddish-brown veins, and linear series of golden setae. Pronotum length (including pronotal lobe) 3.08, humeral width 1.92. Fore wing length 6.52, wing veins similar to
|
||
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.15602" author="Matsuda, R" journalOrPublisher="The University of Kansas Science Bulletin" pageId="0" pageNumber="69" pagination="25 - 632" refId="B8" refString="Matsuda, R, 1960. Morphology, evolution and a classification of the Gerridae (Hemiptera - Heteroptera). The University of Kansas Science Bulletin 41 (2): 25 - 632, DOI: https://doi.org/10.5962/bhl.part.15602" title="Morphology, evolution and a classification of the Gerridae (Hemiptera - Heteroptera)." url="https://doi.org/10.5962/bhl.part.15602" volume="41" year="1960">Matsuda 1960</bibRefCitation>
|
||
: fig. 553;
|
||
<bibRefCitation author="Andersen, NM" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="69" pagination="1 - 25" refId="B2" refString="Andersen, NM, 1982. Semiterrestrial water striders of the genera Eotrechus Kirkaldy and Chimarrhometra Bianchi (Insecta, Hemiptera, Gerridae). Steenstrupia 9: 1 - 25" title="Semiterrestrial water striders of the genera Eotrechus Kirkaldy and Chimarrhometra Bianchi (Insecta, Hemiptera, Gerridae)." volume="9" year="1982">Andersen 1982</bibRefCitation>
|
||
: fig. 8).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">Macropterous female: Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="etymology">
|
||
<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">This species is dedicated to the German speleologist Dr Helmut Steiner who discovered it during one of his expeditions to northern Laos.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="remarks">
|
||
<paragraph pageId="0" pageNumber="69">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<taxonomicName authorityName="Tran & Zettel & Sites" authorityYear="2023" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus steineri" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="steineri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus steineri</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated from
|
||
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. fuscus" pageId="0" pageNumber="69" rank="species" species="fuscus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. fuscus</emphasis>
|
||
</taxonomicName>
|
||
by the structure of the paramere, the relative lengths of the antennal segments, and the relative lengths of the fore tarsal segments. This new species can also be recognised by the large hump on the mesonotum. A mesonotal hump is also present in
|
||
<taxonomicName lsidName="E. fuscus" pageId="0" pageNumber="69" rank="species" species="fuscus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. fuscus</emphasis>
|
||
</taxonomicName>
|
||
, but is much less conspicuous than that of this new species. Detailed differences between the three species are summarised in Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparative morphology of E. kalidasa, E. fuscus, and E. steineri sp. nov." httpUri="http://table.plazi.org/id/EF594DCD37796B0AEC53DFC488CF38A9" pageId="0" pageNumber="69" tableUuid="EF594DCD37796B0AEC53DFC488CF38A9">1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="distribution">
|
||
<paragraph pageId="0" pageNumber="69">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Laos: Luang Nam Tha, Luang Prabang; Thailand: Loei (the specimen from Thailand reported as
|
||
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Vitheepradit, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="1 - 19" publicationUrl="https://www.mapress.com/zootaxa/2007f/zt01478p019.pdf" refId="B19" refString="Vitheepradit, A, Sites, RW, 2007. A review of Eotrechus Kirkaldy (Hemiptera: Heteroptera: Gerridae) of Thailand with descriptions of three new species. Zootaxa 1478: 1 - 19, https://www.mapress.com/zootaxa/2007f/zt01478p019.pdf" title="A review of Eotrechus Kirkaldy (Hemiptera: Heteroptera: Gerridae) of Thailand with descriptions of three new species." url="https://www.mapress.com/zootaxa/2007f/zt01478p019.pdf" volume="1478" year="2007">Vitheepradit and Sites (2007)</bibRefCitation>
|
||
belongs to this new species) (Fig.
|
||
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Distribution records of species of Eotrechus. Squares represent E. kalidasa species group; circles represent E. pingae species group; triangles represent species unassigned to a group." figureDoi="10.3897/dez.70.97117.figure25" httpUri="https://binary.pensoft.net/fig/826114" pageId="0" pageNumber="69">25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">
|
||
<taxonomicName authorityName="Andersen" authorityYear="1982" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus petraeus" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="petraeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus petraeus</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Diagnosis.</emphasis>
|
||
Male: fore femur incrassate, usually with a large basal tubercle bearing a spot of black, minute setae, or only with a spot of black, minute setae without distinct basal tubercle; posterior margin of sternum VII medially notched, about two-fifths of sternum VII length; pygophore with lateral tubercles with long setae; caudal projection of pygophore subtriangular or elongate with pointed apex; proctiger laterally bearing long setae, posterolaterally with round lobes or pointed processes, apex of proctiger angular. Female: tergum VIII usually produced posterolaterally; sternum VII about as long as two preceding sterna combined.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Species included.</emphasis>
|
||
<taxonomicName lsidName="E. petraeus" pageId="0" pageNumber="69" rank="species" species="petraeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. petraeus</emphasis>
|
||
</taxonomicName>
|
||
Andersen, 1982,
|
||
<taxonomicName lsidName="E. kerberos" pageId="0" pageNumber="69" rank="species" species="kerberos">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. kerberos</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName lsidName="E. romglao" pageId="0" pageNumber="69" rank="species" species="romglao">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. romglao</emphasis>
|
||
</taxonomicName>
|
||
Vitheepradit & Sites, 2007, and
|
||
<taxonomicName lsidName="E. thai" pageId="0" pageNumber="69" rank="species" species="thai">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. thai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |