293 lines
30 KiB
XML
293 lines
30 KiB
XML
<document id="F49AB50DA27F964BFC4E6E59D6D37C5D" ID-ISSN="1464-5262" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979642237" checkinUser="felipe" docAuthor="Yves" docDate="2003" docId="894C1850FFC1FF8AFE2EB225A175FCF1" docLanguage="en" docName="JNatHist.37.20.2487-2519.pdf" docOrigin="Journal of Natural History 37" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Holothuria (Mertensiothuria) papillifera Heding" docType="treatment" docVersion="3" lastPageNumber="2508" masterDocId="75756028FFD2FF9CFFBBB05AA33CFF94" masterDocTitle="The holothurian subgenus Mertensiothuria (Aspidochirotida: Holothuriidae) revisited" masterLastPageNumber="2519" masterPageNumber="2487" pageNumber="2505" updateTime="1699132880048" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="3963EC052CA98932415026BF16A2B57C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="39795579DC4F1092DE0AE2C9D9BAB6F6">
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<mods:title id="67EAFACC2A1BC2B3A812AFF5108439D7">The holothurian subgenus Mertensiothuria (Aspidochirotida: Holothuriidae) revisited</mods:title>
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</mods:titleInfo>
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<mods:name id="472D5EC23CD6C59C3D790BDD607DB470" type="personal">
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<mods:role id="5712F46425DC3915B80C0EB81CCB0EFD">
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<mods:roleTerm id="88AC51AB36FD36E8285EEAD6FDF1B3CE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DCA628D3526C6702FBBD75E5A8C8FFA8">Yves</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="17D7E94CCE858535EB8F6B36224A3B53">text</mods:typeOfResource>
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<mods:titleInfo id="72FBD1234B3CBDB9B8932FB818F73893">
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<mods:title id="69CB21529CF571814A038F0770DA4BAF">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="E0913F486786221A28AEE1BD63008388">
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<mods:date id="6B352F26667CE66B226C29EAE1CD3E9F">2003</mods:date>
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<mods:detail id="762BC3C77740A430E265640FBB088DA5" type="volume">
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<mods:number id="91DA4E4474EF1952367DACE3676C5CA9">37</mods:number>
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</mods:detail>
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<mods:extent id="F793DE1743F00E83E6778A134FDD45F9" unit="page">
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<mods:start id="3D1D633DF2519B1C1861397964EA25F6">2487</mods:start>
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<mods:end id="032629BF9B4C92974949345602997B45">2519</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="EA20D47B6D8C2C441458667B9FC1DB09">journal article</mods:classification>
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<mods:identifier id="40593318CD15B5F05DC1F384B55B4529" type="ISSN">1464-5262</mods:identifier>
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</mods:mods>
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<treatment id="894C1850FFC1FF8AFE2EB225A175FCF1" ID-DOI="http://doi.org/10.5281/zenodo.5271010" ID-Zenodo-Dep="5271010" LSID="urn:lsid:plazi:treatment:894C1850FFC1FF8AFE2EB225A175FCF1" httpUri="http://treatment.plazi.org/id/894C1850FFC1FF8AFE2EB225A175FCF1" lastPageId="22" lastPageNumber="2508" pageId="19" pageNumber="2505">
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<subSubSection id="49FFFACDFFC1FF8FFE2EB225A0FAFD2C" pageId="19" pageNumber="2505" type="nomenclature">
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||
<paragraph id="015AA946FFC1FF8FFE2EB225A7A4FD0C" blockId="19.[405,1176,639,664]" box="[405,1176,639,664]" pageId="19" pageNumber="2505">
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<heading id="5A121E2AFFC1FF8FFE2EB225A7A4FD0C" box="[405,1176,639,664]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="2505" reason="2">
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||
<taxonomicName id="C6E5D2C5FFC1FF8FFE2EB225A086FD0C" authority="Heding" authorityName="Heding" box="[405,954,639,664]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="19" pageNumber="2505" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
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||
<emphasis id="33917554FFC1FF8FFE2EB225A061FD03" bold="true" box="[405,861,639,663]" italics="true" pageId="19" pageNumber="2505">Holothuria (Mertensiothuria) papillifera</emphasis>
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Heding
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</taxonomicName>
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inMortensen, 1938
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</heading>
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</paragraph>
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<paragraph id="015AA946FFC1FF8FFDD3B2FAA0FAFD2C" blockId="19.[313,1268,672,780]" box="[616,966,672,696]" pageId="19" pageNumber="2505">(figures 7A–M, 8A–J, 12H–K)</paragraph>
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</subSubSection>
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||
<subSubSection id="49FFFACDFFC1FF8FFE82B29BA711FD43" box="[313,1069,705,728]" pageId="19" pageNumber="2505" type="description">
|
||
<paragraph id="015AA946FFC1FF8FFE82B29BA711FD43" blockId="19.[313,1268,672,780]" box="[313,1069,705,728]" pageId="19" pageNumber="2505">
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<taxonomicName id="C6E5D2C5FFC1FF8FFE82B29BA15AFD43" authority="Heding" authorityName="Heding" box="[313,614,705,727]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="19" pageNumber="2505" phylum="Echinodermata" rank="species" species="papillifera">
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<emphasis id="33917554FFC1FF8FFE82B29BA12FFD42" box="[313,531,705,726]" italics="true" pageId="19" pageNumber="2505">Holothuria papillifera</emphasis>
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Heding
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</taxonomicName>
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<emphasis id="33917554FFC1FF8FFDD5B298A1BCFD43" box="[622,640,706,727]" italics="true" pageId="19" pageNumber="2505">in</emphasis>
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Mortensen, 1938: 55, pl. XI, figures 1–3.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="49FFFACDFFC1FF8FFE82B286A762FCD2" pageId="19" pageNumber="2505" type="reference_group">
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||
<paragraph id="015AA946FFC1FF8FFE82B286A1DEFC98" blockId="19.[313,1268,672,780]" pageId="19" pageNumber="2505">
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||
<taxonomicName id="C6E5D2C5FFC1FF8FFE82B286A0B6FD65" authority=": Rowe, 1969: 149" authorityName="Rowe" authorityPageNumber="149" authorityYear="1969" box="[313,906,732,754]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="19" pageNumber="2505" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
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<emphasis id="33917554FFC1FF8FFE82B286A1EDFD65" box="[313,721,732,753]" italics="true" pageId="19" pageNumber="2505">Holothuria (Mertensiothuria) papillifera</emphasis>
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: Rowe, 1969: 149
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</taxonomicName>
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; Clark and Rowe, 1971: 176; Price, 1982: 11 Samyn
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<emphasis id="33917554FFC1FF8FFDBEB2ADA10DFC98" box="[517,561,759,780]" italics="true" pageId="19" pageNumber="2505">et al</emphasis>
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||
., 2001: 104, 107.
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||
</paragraph>
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||
<paragraph id="015AA946FFC1FF8FFEDAB377A762FCD2" blockId="19.[313,1269,813,1730]" box="[353,1118,813,838]" pageId="19" pageNumber="2505">
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<emphasis id="33917554FFC1FF8FFEDAB377A134FCD1" box="[353,520,813,837]" italics="true" pageId="19" pageNumber="2505">Original name.</emphasis>
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<taxonomicName id="C6E5D2C5FFC1FF8FFD99B377A04CFCD1" authority="Heding" authorityName="Heding" box="[546,880,813,837]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="19" pageNumber="2505" phylum="Echinodermata" rank="species" species="papillifera">
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<emphasis id="33917554FFC1FF8FFD99B377A029FCD1" box="[546,789,813,837]" italics="true" pageId="19" pageNumber="2505">Holothuria papillifera</emphasis>
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Heding
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</taxonomicName>
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<emphasis id="33917554FFC1FF8FFCC2B374A0B1FCD1" box="[889,909,814,837]" italics="true" pageId="19" pageNumber="2505">in</emphasis>
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||
Mortensen, 1938.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="49FFFACDFFC1FF8AFEDAB317A175FCF1" lastPageId="22" lastPageNumber="2508" pageId="19" pageNumber="2505" type="description">
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||
<paragraph id="015AA946FFC1FF8FFEDAB317A1DCFC31" blockId="19.[313,1269,813,1730]" pageId="19" pageNumber="2505">
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<emphasis id="33917554FFC1FF8FFEDAB317A102FCF1" box="[353,574,845,869]" italics="true" pageId="19" pageNumber="2505">Name-bearing type.</emphasis>
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<typeStatus id="DE5E17E4FFC1FF8FFDE3B317A18FFCF1" box="[600,691,845,869]" pageId="19" pageNumber="2505" type="syntype">Syntype</typeStatus>
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series ZM Karaf no. 85,
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<date id="755B8F86FFC1FF8FFC64B317A767FCF2" box="[991,1115,845,870]" pageId="19" pageNumber="2505" value="1937-10-25">25.10.1937</date>
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;
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<typeStatus id="DE5E17E4FFC1FF8FFBD3B317A7F2FCF1" box="[1128,1230,845,869]" pageId="19" pageNumber="2505" type="lectotype">lectotype</typeStatus>
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by present designation, ZM Karaf no. 85/1 (largest specimen of the type series),
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<typeStatus id="DE5E17E4FFC1FF8FFE82B3D7A2E1FC31" box="[313,477,909,933]" pageId="19" pageNumber="2505" type="paralectotype">paralectotypes</typeStatus>
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ZM Karaf no. 85/2,3.
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</paragraph>
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||
<paragraph id="015AA946FFC1FF8FFEDAB3F4A1CCFC51" blockId="19.[313,1269,813,1730]" box="[353,752,941,966]" pageId="19" pageNumber="2505">
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<materialsCitation id="B18DA31BFFC1FF8FFEDAB3F4A1D0FC51" box="[353,748,941,966]" country="Egypt" location="Egypt" pageId="19" pageNumber="2505" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="33917554FFC1FF8FFEDAB3F4A2C7FC51" box="[353,507,941,965]" italics="true" pageId="19" pageNumber="2505">
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<typeStatus id="DE5E17E4FFC1FF8FFEDAB3F4A2A5FC51" box="[353,409,942,965]" pageId="19" pageNumber="2505">Type</typeStatus>
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locality.
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</emphasis>
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||
<collectingCountry id="79F2E9D6FFC1FF8FFDAEB3F4A165FC52" box="[533,601,942,966]" name="Egypt" pageId="19" pageNumber="2505">Egypt</collectingCountry>
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(Ghardaqa)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="015AA946FFC1FF8FFEDAB397A24AFB91" blockId="19.[313,1269,813,1730]" pageId="19" pageNumber="2505">
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<emphasis id="33917554FFC1FF8FFEDAB397A136FC71" box="[353,522,973,997]" italics="true" pageId="19" pageNumber="2505">Current status.</emphasis>
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<taxonomicName id="C6E5D2C5FFC1FF8FFD9FB397A776FC71" authority="Heding" authorityName="Heding" box="[548,1098,973,997]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="19" pageNumber="2505" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
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<emphasis id="33917554FFC1FF8FFD9FB397A0D1FC71" box="[548,1005,973,997]" italics="true" pageId="19" pageNumber="2505">Holothuria (Mertensiothuria) papillifera</emphasis>
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Heding
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||
</taxonomicName>
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||
<emphasis id="33917554FFC1FF8FFBEFB394A754FC71" box="[1108,1128,974,997]" italics="true" pageId="19" pageNumber="2505">in</emphasis>
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||
Mortensen, 1938.
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||
</paragraph>
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||
<paragraph id="015AA946FFC1FF8FFEDAB457A731FBB1" blockId="19.[313,1269,813,1730]" box="[353,1037,1037,1061]" pageId="19" pageNumber="2505">
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||
<emphasis id="33917554FFC1FF8FFEDAB457A101FBB1" box="[353,573,1037,1061]" italics="true" pageId="19" pageNumber="2505">Material examined.</emphasis>
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||
ZM
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||
<typeStatus id="DE5E17E4FFC1FF8FFD30B457A1CDFBB1" box="[651,753,1037,1061]" pageId="19" pageNumber="2505" type="lectotype">lectotype</typeStatus>
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||
and
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||
<specimenCount id="17E362CFFFC1FF8FFC95B457A735FBB1" box="[814,1033,1037,1061]" pageId="19" pageNumber="2505" type="generic" typeStatus="paralectotypes">two paralectotypes</specimenCount>
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||
.
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</paragraph>
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||
<paragraph id="015AA946FFC1FF8FFEDAB477A058FAB0" blockId="19.[313,1269,813,1730]" pageId="19" pageNumber="2505">
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||
<emphasis id="33917554FFC1FF8FFEDAB477A2D4FBD0" box="[353,488,1069,1092]" italics="true" pageId="19" pageNumber="2505">Description.</emphasis>
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||
Medium size to large specimens (
|
||
<typeStatus id="DE5E17E4FFC1FF8FFC07B477A71EFBD1" box="[956,1058,1069,1093]" pageId="19" pageNumber="2505" type="lectotype">lectotype</typeStatus>
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||
225×
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||
<quantity id="C61D04A3FFC1FF8FFBC3B477A7CCFBD0" box="[1144,1264,1069,1093]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="19" pageNumber="2505" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 mm</quantity>
|
||
;
|
||
<typeStatus id="DE5E17E4FFC1FF8FFE82B416A2E1FBF0" box="[313,477,1100,1124]" pageId="19" pageNumber="2505" type="paralectotype">paralectotypes</typeStatus>
|
||
165×
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||
<quantity id="C61D04A3FFC1FF8FFD96B416A19EFBF0" box="[557,674,1100,1125]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="19" pageNumber="2505" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 mm</quantity>
|
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and 160×
|
||
<quantity id="C61D04A3FFC1FF8FFC91B417A09FFBF1" box="[810,931,1101,1125]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="19" pageNumber="2505" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 mm</quantity>
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); body club-shaped, narrow anteriorly, much larger posteriorly. Colour of living specimen uniform greyish brown. Colour in alcohol brown dorsally (figure 12H), beige to light brown ventrally (figure 12K). Mouth ventral, anus terminal. Twenty short, yellowish tentacles surrounded by a circle of papillae. Dorsally, numerous, large conical papillae without alignment (figure 12J). Ventrally, tube feet densely crowded in the ambulacral as well as in the interambulacral areas (figure 12J).
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||
</paragraph>
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||
<paragraph id="015AA946FFC1FF8FFEDAB576A080FA37" blockId="19.[313,1269,813,1730]" pageId="19" pageNumber="2505">
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||
Calcareous ring (figure 7A) typical of the subgenus
|
||
<taxonomicName id="C6E5D2C5FFC1FF8FFC7CB576A746FAD0" box="[967,1146,1324,1348]" class="Holothuroidea" family="Holothuriidae" genus="Mertensiothuria" kingdom="Animalia" order="Holothuriida" pageId="19" pageNumber="2505" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="33917554FFC1FF8FFC7CB576A746FAD0" box="[967,1146,1324,1348]" italics="true" pageId="19" pageNumber="2505">Mertensiothuria</emphasis>
|
||
</taxonomicName>
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||
; two long Polian vesicles (one-sixth of body length) and two very short (±2% of body length), contorted stone canals (figure 7B); tentacle ampullae very short (2–2.5% body length). Cuvierian tubules numerous, whitish, very thin.
|
||
</paragraph>
|
||
<paragraph id="015AA946FFC1FF8AFEDAB5F1A022FF7C" blockId="19.[313,1269,813,1730]" lastBlockId="22.[313,1269,176,869]" lastPageId="22" lastPageNumber="2508" pageId="19" pageNumber="2505">
|
||
<emphasis id="33917554FFC1FF8FFEDAB5F1A286FA57" box="[353,442,1451,1475]" italics="true" pageId="19" pageNumber="2505">Ossicles</emphasis>
|
||
: in the body wall tables and buttons. Dorsally, tables few with disc
|
||
<quantity id="C61D04A3FFC1FF8FFE82B590A2ADFA76" box="[313,401,1482,1507]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="7.5" metricValueMin="4.5" pageId="19" pageNumber="2505" unit="m" value="60.0" valueMax="75.0" valueMin="45.0">
|
||
45–75
|
||
<emphasis id="33917554FFC1FF8FFE39B591A2ADFA76" bold="true" box="[386,401,1483,1506]" italics="true" pageId="19" pageNumber="2505">m</emphasis>
|
||
</quantity>
|
||
m across, round to quadrangular (figure 7C), perforated by four central and 9–10 peripheral holes; four pillars united by one cross-beam and ending in a small crown of spines; table height ±
|
||
<quantity id="C61D04A3FFC1FF8FFD0CB650A1D9F9B6" box="[695,741,1546,1571]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="2505" unit="m" value="50.0">
|
||
50
|
||
<emphasis id="33917554FFC1FF8FFD6DB651A1D9F9B6" bold="true" box="[726,741,1547,1570]" italics="true" pageId="19" pageNumber="2505">m</emphasis>
|
||
</quantity>
|
||
m. Buttons regular with three to five pairs of holes (three pairs being the most frequent),
|
||
<quantity id="C61D04A3FFC1FF8FFCDBB670A084F9D6" box="[864,952,1578,1603]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="19" pageNumber="2505" unit="m" value="70.0" valueMax="85.0" valueMin="55.0">
|
||
55–85
|
||
<emphasis id="33917554FFC1FF8FFC12B671A084F9D6" bold="true" box="[937,952,1579,1602]" italics="true" pageId="19" pageNumber="2505">m</emphasis>
|
||
</quantity>
|
||
m long (figure 7D); a few irregular buttons (figure 7E). In the ventral body wall same types of ossicle, tables (figure 7F) being more numerous and buttons (figure 7G) smaller than dorsally. At the base of the dorsal papillae same ossicles as in the dorsal body wall. At the top of the papillae, tables (figure 7H), buttons (figure 7J) generally with three pairs of holes and rods,
|
||
<quantity id="C61D04A3FFC7FF89FE4DB5F1A15BFA57" box="[502,615,1451,1475]" metricMagnitude="2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="21" pageNumber="2507" unit="m" value="215.0" valueMax="230.0" valueMin="200.0">
|
||
200–230
|
||
<emphasis id="33917554FFC7FF89FDE3B5F6A15BFA57" bold="true" box="[600,615,1452,1475]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
</quantity>
|
||
m long, with lateral processes (figure 7K). In the dorsal papillae of the
|
||
<typeStatus id="DE5E17E4FFC7FF89FE52B591A1BAFA77" box="[489,646,1483,1507]" pageId="21" pageNumber="2507" type="paralectotype">paralectotype</typeStatus>
|
||
,
|
||
<quantity id="C61D04A3FFC7FF89FD28B590A1D0FA77" box="[659,748,1482,1507]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" pageId="21" pageNumber="2507" unit="mm" value="165.0">165 mm</quantity>
|
||
long, the buttons have generally four to five pairs of holes and lateral processes of the rods are perforated (figure 7L). In the tube feet buttons (figure 7M), irregular perforated plates (figure 8A), large, regular perforated plates, from 70×
|
||
<quantity id="C61D04A3FFC7FF89FD29B670A183F9D6" box="[658,703,1578,1603]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="21" pageNumber="2507" unit="m" value="85.0">
|
||
85
|
||
<emphasis id="33917554FFC7FF89FD0BB671A183F9D6" bold="true" box="[688,703,1579,1602]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
</quantity>
|
||
m to 95×
|
||
<quantity id="C61D04A3FFC7FF89FC84B671A046F9D6" box="[831,890,1579,1603]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="2507" unit="m" value="120.0">
|
||
120
|
||
<emphasis id="33917554FFC7FF89FCD0B671A046F9D6" bold="true" box="[875,890,1579,1602]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
</quantity>
|
||
m (figure 8B), and small tables (figure 8C),
|
||
<quantity id="C61D04A3FFC7FF89FE71B610A11DF9F6" box="[458,545,1610,1635]" metricMagnitude="1" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="21" pageNumber="2507" unit="m" value="52.5" valueMax="55.0" valueMin="50.0">
|
||
50–55
|
||
<emphasis id="33917554FFC7FF89FDA9B611A11DF9F6" bold="true" box="[530,545,1611,1634]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
</quantity>
|
||
m across. In the paralectoptype
|
||
<quantity id="C61D04A3FFC7FF89FC1EB610A0C1F9F6" box="[933,1021,1610,1635]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" pageId="21" pageNumber="2507" unit="mm" value="165.0">165 mm</quantity>
|
||
long, ossicles of the tube feet somewhat larger, especially the regular and irregular perforated plates (figure 8D and E, respectively). In the tentacles rods, spiny at the extremities (figure 8F, G),
|
||
<quantity id="C61D04A3FFC7FF89FE55B6F3A16FF956" box="[494,595,1705,1730]" metricMagnitude="2" metricUnit="m" metricValue="2.65" metricValueMax="5.0" metricValueMin="0.3" pageId="21" pageNumber="2507" unit="m" value="265.0" valueMax="500.0" valueMin="30.0">
|
||
30–500
|
||
<emphasis id="33917554FFC7FF89FDFFB6F1A16FF956" bold="true" box="[580,595,1707,1730]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
</quantity>
|
||
m long. The smallest rods,
|
||
<quantity id="C61D04A3FFC7FF89FC2DB6F3A0D2F956" box="[918,1006,1705,1730]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="21" pageNumber="2507" unit="m" value="37.5" valueMax="45.0" valueMin="30.0">
|
||
30–45
|
||
<emphasis id="33917554FFC7FF89FC64B6F1A0D2F956" bold="true" box="[991,1006,1707,1730]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
</quantity>
|
||
m long, C- or S-shape (figure 8H). In the longitudinal muscles numerous regular to irregular C-shape and O-ring ossicles (figure 8J),
|
||
<quantity id="C61D04A3FFC4FF8AFDD5B095A1FAFF73" box="[622,710,207,232]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="6.5" metricValueMin="1.0" pageId="22" pageNumber="2508" unit="m" value="37.5" valueMax="65.0" valueMin="10.0">
|
||
10–65
|
||
<emphasis id="33917554FFC4FF8AFD0CB08AA1FAFF73" bold="true" box="[695,710,208,231]" italics="true" pageId="22" pageNumber="2508">m</emphasis>
|
||
</quantity>
|
||
m long.
|
||
</paragraph>
|
||
<caption id="559AF9CEFFC6FF88FE82B597A1F6F922" pageId="20" pageNumber="2506" startId="20.[313,329,1485,1506]" targetBox="[317,1266,176,1457]" targetPageId="20">
|
||
<paragraph id="015AA946FFC6FF88FE82B597A1F6F922" blockId="20.[313,1268,1484,1719]" pageId="20" pageNumber="2506">
|
||
F. 7.
|
||
<taxonomicName id="C6E5D2C5FFC6FF88FE2FB596A041FA76" authority="Heding" authorityName="Heding" box="[404,893,1484,1506]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="20" pageNumber="2506" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC6FF88FE2FB596A017FA75" box="[404,811,1484,1505]" italics="true" pageId="20" pageNumber="2506">Holothuria (Mertensiothuria) papillifera</emphasis>
|
||
Heding
|
||
</taxonomicName>
|
||
<emphasis id="33917554FFC6FF88FC3FB597A0AAFA76" box="[900,918,1485,1506]" italics="true" pageId="20" pageNumber="2506">in</emphasis>
|
||
Mortensen, 1938. (A) Calcareous ring (r, radial plate; ir, interradial plate) (L=225 mm); (B) stone canal (L=225 mm); (C) tables of dorsal body wall (L=225 mm); (D) regular buttons of dorsal body wall (L=225 mm); (E) irregular buttons of dorsal body wall (L=225 mm); (F) tables of ventral body wall (L=225 mm); (G) buttons of ventral body wall (L=225 mm); (H) tables of top of dorsal papillae (L=225 mm); (J) buttons of top of dorsal papillae (L= 225 mm); (K) rods of top of dorsal papillae (L=225 mm); (L) rods of top of dorsal papillae (L=165 mm); (M) buttons of tube feet (L=225 mm). Scale bars: (A, B) 5 mm; (C–J, M) 50
|
||
<emphasis id="33917554FFC6FF88FE45B6FBA137F922" bold="true" box="[510,523,1697,1718]" italics="true" pageId="20" pageNumber="2506">m</emphasis>
|
||
m; (K, L) 100
|
||
<emphasis id="33917554FFC6FF88FD19B6FBA193F922" bold="true" box="[674,687,1697,1718]" italics="true" pageId="20" pageNumber="2506">m</emphasis>
|
||
m.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="559AF9CEFFC7FF89FE82B495A790FAFD" pageId="21" pageNumber="2507" startId="21.[313,329,1231,1252]" targetBox="[317,1266,175,1202]" targetPageId="21">
|
||
<paragraph id="015AA946FFC7FF89FE82B495A790FAFD" blockId="21.[313,1268,1231,1386]" pageId="21" pageNumber="2507">
|
||
F. 8.
|
||
<taxonomicName id="C6E5D2C5FFC7FF89FE2EB495A0BAFB71" authority="Heding" authorityName="Heding" box="[405,902,1231,1253]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="21" pageNumber="2507" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC7FF89FE2EB495A00DFB70" box="[405,817,1231,1252]" italics="true" pageId="21" pageNumber="2507">Holothuria (Mertensiothuria) papillifera</emphasis>
|
||
Heding
|
||
</taxonomicName>
|
||
<emphasis id="33917554FFC7FF89FC34B48AA09DFB71" box="[911,929,1232,1253]" italics="true" pageId="21" pageNumber="2507">in</emphasis>
|
||
Mortensen, 1938. (A) Irregular perforated plates of ventral tube feet (L=225 mm); (B) regular perforated plates of ventral tube feet (L=225 mm); (C) tables of ventral tube feet (L=225 mm); (D) regular perforated plate of ventral tube feet (L=165 mm); (E) irregular perforated plates of ventral tube feet (L=165 mm); (F–H) rods of tentacles (L=225 mm); (J) ossicles of longitudinal muscles (L=225 mm). Scale bars: (F) 200
|
||
<emphasis id="33917554FFC7FF89FC0DB50EA0FFFAFD" bold="true" box="[950,963,1364,1385]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
m; (A–E, G–J) 50
|
||
<emphasis id="33917554FFC7FF89FB3FB50EA7ADFAFD" bold="true" box="[1156,1169,1364,1385]" italics="true" pageId="21" pageNumber="2507">m</emphasis>
|
||
m.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="015AA946FFC4FF8AFEDAB0AAA1B8FE93" blockId="22.[313,1269,176,869]" box="[353,644,240,263]" pageId="22" pageNumber="2508">
|
||
<emphasis id="33917554FFC4FF8AFEDAB0AAA2E9FE93" box="[353,469,240,263]" italics="true" pageId="22" pageNumber="2508">Diagnosis.</emphasis>
|
||
Present note.
|
||
</paragraph>
|
||
<paragraph id="015AA946FFC4FF8AFEDAB155A16AFEB3" blockId="22.[313,1269,176,869]" box="[353,598,271,295]" pageId="22" pageNumber="2508">
|
||
<emphasis id="33917554FFC4FF8AFEDAB155A2FEFEB3" box="[353,450,271,295]" italics="true" pageId="22" pageNumber="2508">Ecology.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="015AA946FFC4FF8AFEDAB175A018FEF3" blockId="22.[313,1269,176,869]" pageId="22" pageNumber="2508">
|
||
<emphasis id="33917554FFC4FF8AFEDAB175A1B0FED3" box="[353,652,303,327]" italics="true" pageId="22" pageNumber="2508">Geographical distribution.</emphasis>
|
||
Up to now only recorded from the
|
||
<collectingRegion id="C32167A4FFC4FF8AFB31B175A7C9FED3" box="[1162,1269,303,327]" country="Egypt" name="Al Bahr al Ahmar" pageId="22" pageNumber="2508">Red Sea</collectingRegion>
|
||
(Ghardaqa, Abu Sadaf and Abu Fanadir).
|
||
</paragraph>
|
||
<paragraph id="015AA946FFC4FF8AFEDAB135A13DFE33" blockId="22.[313,1269,176,869]" pageId="22" pageNumber="2508">
|
||
<emphasis id="33917554FFC4FF8AFEDAB135A10FFE13" box="[353,563,367,391]" italics="true" pageId="22" pageNumber="2508">Proposed status.</emphasis>
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFDF6B135A781FE13" authority="Heding" authorityName="Heding" box="[589,1213,367,391]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFDF6B135A77BFE13" box="[589,1095,367,391]" italics="true" pageId="22" pageNumber="2508">Holothuria (Mertensiothuria) papillifera</emphasis>
|
||
Heding
|
||
</taxonomicName>
|
||
<emphasis id="33917554FFC4FF8AFB5BB12AA7C8FE13" box="[1248,1268,368,391]" italics="true" pageId="22" pageNumber="2508">in</emphasis>
|
||
Mortensen, 1938.
|
||
</paragraph>
|
||
<paragraph id="015AA946FFC4FF8AFEDAB1F5A7CDFDB2" blockId="22.[313,1269,176,869]" pageId="22" pageNumber="2508">
|
||
<emphasis id="33917554FFC4FF8AFEDAB1F5A2E2FE52" box="[353,478,431,454]" italics="true" pageId="22" pageNumber="2508">Comments.</emphasis>
|
||
The original description of
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFC89B1EAA737FE53" box="[818,1035,431,455]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFC89B1EAA737FE53" box="[818,1035,431,455]" italics="true" pageId="22" pageNumber="2508">H. (M.) papillifera</emphasis>
|
||
</taxonomicName>
|
||
given by Heding (
|
||
<emphasis id="33917554FFC4FF8AFB5BB1EAA7C8FE53" box="[1248,1268,432,455]" italics="true" pageId="22" pageNumber="2508">in</emphasis>
|
||
Mortensen, 1938) in a footnote (without any illustration) is very succinct. Since the type series is available in the collection of the ZM, we take the opportunity to describe and illustrate the species, and to designate a
|
||
<typeStatus id="DE5E17E4FFC4FF8AFC19B254A734FDB2" box="[930,1032,526,550]" pageId="22" pageNumber="2508" type="lectotype">lectotype</typeStatus>
|
||
and
|
||
<typeStatus id="DE5E17E4FFC4FF8AFBFEB254A7D1FDB2" box="[1093,1261,526,550]" pageId="22" pageNumber="2508" type="paralectotype">paralectotypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="015AA946FFC4FF8AFEDAB274A175FCF1" blockId="22.[313,1269,176,869]" pageId="22" pageNumber="2508">
|
||
Heding (
|
||
<emphasis id="33917554FFC4FF8AFE7EB275A2E5FDD2" box="[453,473,559,582]" italics="true" pageId="22" pageNumber="2508">in</emphasis>
|
||
Mortensen, 1938) suggested that
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFCDFB275A702FDD2" box="[868,1086,558,582]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFCDFB275A702FDD2" box="[868,1086,558,582]" italics="true" pageId="22" pageNumber="2508">H. (M.) papillifera</emphasis>
|
||
</taxonomicName>
|
||
is very close to
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFE82B215A11EFDF2" baseAuthorityName="Brandt" baseAuthorityYear="1835" box="[313,546,590,614]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="leucospilota" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFE82B215A11EFDF2" box="[313,546,590,614]" italics="true" pageId="22" pageNumber="2508">H. (M.) leucospilota</emphasis>
|
||
</taxonomicName>
|
||
. Rowe (1969) suggested it is possibly conspecific with
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFB2EB215A287FD12" baseAuthorityName="Brandt" baseAuthorityYear="1835" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="leucospilota" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFB2EB215A287FD12" italics="true" pageId="22" pageNumber="2508">H. (M.) leucospilota</emphasis>
|
||
</taxonomicName>
|
||
. Examination of the
|
||
<typeStatus id="DE5E17E4FFC4FF8AFD0DB235A1DAFD13" box="[694,742,623,647]" pageId="22" pageNumber="2508">type</typeStatus>
|
||
series confirms that both species are close to each other. Moreover, presence or absence of Cuvierian tubules is no longer a distinctive character (Heding
|
||
<emphasis id="33917554FFC4FF8AFD2BB2F5A198FD52" box="[656,676,687,710]" italics="true" pageId="22" pageNumber="2508">in</emphasis>
|
||
Mortensen, 1938) since both species possess them. The main characters separating
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFD0BB295A0B0FD72" box="[688,908,718,742]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="papillifera" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFD0BB295A0B0FD72" box="[688,908,718,742]" italics="true" pageId="22" pageNumber="2508">H. (M.) papillifera</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="C6E5D2C5FFC4FF8AFC63B295A7FAFD72" baseAuthorityName="Brandt" baseAuthorityYear="1835" box="[984,1222,718,742]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="22" pageNumber="2508" phylum="Echinodermata" rank="species" species="leucospilota" subGenus="Mertensiothuria">
|
||
<emphasis id="33917554FFC4FF8AFC63B295A7FAFD72" box="[984,1222,718,742]" italics="true" pageId="22" pageNumber="2508">H. (M.) leucospilota</emphasis>
|
||
</taxonomicName>
|
||
are the greyish brown colour of the body wall, the long, soft dorsal papillae, the tables with a smooth disc margin, a very narrow crown of spines at the top of the tables, the rods of the dorsal papillae with lateral perforated processes and the presence of ossicles in the tentacles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |