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<document id="F509539A951DAC628BB39A891D6DAE42" ID-DOI="10.5281/zenodo.202391" ID-GBIF-Dataset="3bb2877d-4319-471f-a6bd-30113a270bcc" ID-ISSN="1175-5326" ID-Zenodo-Dep="202391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460249745227" checkinUser="plazi" docAuthor="Martens, Jochen" docDate="2011" docId="894A41406F435639D4F7FBD823A5F8B5" docLanguage="en" docName="zt02783p051.pdf" docOrigin="Zootaxa 2783" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Centetostoma Kratochvil 1958" docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="757339386F40563AD460FFFD2267FFD3" masterDocTitle="The Centetostoma scabriculum complex — a group of three cryptic species (Arachnida: Opiliones: Nemastomatidae)" masterLastPageNumber="51" masterPageNumber="35" pageNumber="38" updateTime="1698242487167" updateUser="plazi">
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<mods:title id="88D9F82193453336B97863CB4F953A6C">The Centetostoma scabriculum complex — a group of three cryptic species (Arachnida: Opiliones: Nemastomatidae)</mods:title>
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<mods:namePart id="F0F49C3AE864E6D30EED22F02B30B944">Martens, Jochen</mods:namePart>
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<mods:date id="9C5782A6DCC0215FF2C627778ED6998A">2011</mods:date>
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<treatment id="894A41406F435639D4F7FBD823A5F8B5" ID-DOI="http://doi.org/10.5281/zenodo.6194501" ID-GBIF-Taxon="119391324" ID-Zenodo-Dep="6194501" LSID="urn:lsid:plazi:treatment:894A41406F435639D4F7FBD823A5F8B5" httpUri="http://treatment.plazi.org/id/894A41406F435639D4F7FBD823A5F8B5" lastPageNumber="38" pageId="3" pageNumber="38">
<subSubSection id="49F9A3DD6F435639D4F7FBD82074FBED" box="[151,531,1060,1087]" pageId="3" pageNumber="38" type="nomenclature">
<paragraph id="015CF0566F435639D4F7FBD82074FBED" blockId="3.[151,531,1060,1087]" box="[151,531,1060,1087]" pageId="3" pageNumber="38">
<heading id="5A14473A6F435639D4F7FBD82074FBED" bold="true" box="[151,531,1060,1087]" fontSize="11" level="1" pageId="3" pageNumber="38" reason="1">
<taxonomicName id="C6E38BD56F435639D4F7FBD82074FBED" ID-CoL="7NWQH" authority="Kratochvil, 1958" authorityName="Kratochvil" authorityYear="1958" box="[151,531,1060,1087]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="33972C446F435639D4F7FBD82074FBED" bold="true" box="[151,531,1060,1087]" pageId="3" pageNumber="38">
<emphasis id="33972C446F435639D4F7FBD8235CFBEC" bold="true" box="[151,315,1061,1087]" italics="true" pageId="3" pageNumber="38">Centetostoma</emphasis>
<bibRefCitation id="65728DA76F435639D523FBD92074FBED" author="Kratochvil" box="[323,531,1060,1086]" pageId="3" pageNumber="38" refString="Kratochvil, J. (1958) Hohlenweberknechte Bulgariens (Palpatores - Nemastomatidae). Prace, Brn e nske Zakladny C eskoslovenske Akademie V e d, 30, 523 - 576." type="journal article" year="1958">Kratochvíl, 1958</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="49F9A3DD6F435639D4F7FB92237CF8CE" pageId="3" pageNumber="38" type="materials_examined">
<paragraph id="015CF0566F435639D4F7FB922102FB57" blockId="3.[151,869,1134,1157]" box="[151,869,1134,1157]" pageId="3" pageNumber="38">
<typeStatus id="DE584EF46F435639D4F7FB9222AFFB57" box="[151,200,1135,1156]" pageId="3" pageNumber="38">Type</typeStatus>
species:
<taxonomicName id="C6E38BD56F435639D546FB8D201AFB57" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[294,637,1134,1157]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="centetes">
<emphasis id="33972C446F435639D546FB8D2392FB56" box="[294,501,1136,1157]" italics="true" pageId="3" pageNumber="38">Nemastoma centetes</emphasis>
<bibRefCitation id="65728DA76F435639D59CFB93201AFB57" author="Simon" box="[508,637,1134,1156]" pageId="3" pageNumber="38" refString="Simon, E. (1879) Les Ordres des Chernetes, Scorpiones et Opiliones. Les Arachnides de France, 7, 1 - 332." type="journal article" year="1879">Simon, 1879</bibRefCitation>
</taxonomicName>
(original designation).
</paragraph>
<paragraph id="015CF0566F435639D4F7FB452174FB26" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
The genus is here defined comprising four species, one from the western Alps (
<taxonomicName id="C6E38BD56F435639D06DFB4426EFFB03" box="[1037,1160,1209,1232]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="centetes">
<emphasis id="33972C446F435639D06DFB4426EFFB03" box="[1037,1160,1209,1232]" italics="true" pageId="3" pageNumber="38">C. centetes</emphasis>
</taxonomicName>
) and three from the Pyrenees (
<taxonomicName id="C6E38BD56F435639D485FB2323F7FB26" box="[229,400,1245,1269]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="scabriculum">
<emphasis id="33972C446F435639D485FB2323F7FB26" box="[229,400,1245,1269]" italics="true" pageId="3" pageNumber="38">C. scabriculum</emphasis>
</taxonomicName>
,
<taxonomicName id="C6E38BD56F435639D5FEFB232046FB26" box="[414,545,1245,1269]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="ventalloi">
<emphasis id="33972C446F435639D5FEFB232046FB26" box="[414,545,1245,1269]" italics="true" pageId="3" pageNumber="38">C. ventalloi</emphasis>
</taxonomicName>
,
<taxonomicName id="C6E38BD56F435639D64FFB2320DAFB26" box="[559,701,1245,1269]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="juberthiei" status="sp. nov.">
<emphasis id="33972C446F435639D64FFB2320DAFB26" box="[559,701,1245,1269]" italics="true" pageId="3" pageNumber="38">C. juberthiei</emphasis>
</taxonomicName>
<emphasis id="33972C446F435639D6A4FB202174FB26" bold="true" box="[708,787,1244,1269]" pageId="3" pageNumber="38">
<taxonomicNameLabel id="28A4913F6F435639D6A4FB202163FB26" box="[708,772,1244,1269]" pageId="3" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
).
</emphasis>
</paragraph>
<paragraph id="015CF0566F435639D4A5FAFC233DFA7E" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
Genital morphology (
<figureCitation id="99D8ECD36F435639D5D5FAFC2050FACA" box="[437,567,1281,1305]" captionStart="FIGURES 11 13" captionStartId="8.[151,264,1302,1325]" captionTargetBox="[532,1052,183,1253]" captionTargetId="figure@8.[507,1064,165,1270]" captionTargetPageId="8" captionText="FIGURES 11 13. Penes, dorsal view. 11: Centetostoma ventalloi (SW Foix, CJM 1789). 12: C. scabriculum (Col dAubisque, CJM 3918). 13: C. juberthiei sp. n. (Col de Port, CJM 1726)." httpUri="https://zenodo.org/record/202395/files/figure.png" pageId="3" pageNumber="38">Figs 1113</figureCitation>
): Penial truncus straight, at the basis bulb-like enlarged and bifid, thus the two portions of penial muscles confined to one half of the bulb each; truncus slender, more or less parallel-sided, glans symmetrically tapering to the end, opening of the sperm duct at the very tip; below the glans, an alate (wing-like) structure, slender and rounded (in
<emphasis id="33972C446F435639D678FA8C2084FA5B" box="[536,739,1392,1416]" italics="true" pageId="3" pageNumber="38">
<taxonomicName id="C6E38BD56F435639D678FA8C2012FA5B" box="[536,629,1393,1416]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="centetes">centetes</taxonomicName>
,
<taxonomicName id="C6E38BD56F435639D6E1FA8D2084FA5B" box="[641,739,1392,1416]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="ventalloi">ventalloi</taxonomicName>
</emphasis>
) or strongly extended and pointed sidewards (in
<taxonomicName id="C6E38BD56F435639D16BFA8D27F2FA5B" box="[1291,1429,1392,1416]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="scabriculum">
<emphasis id="33972C446F435639D16BFA8D27F2FA5B" box="[1291,1429,1392,1416]" italics="true" pageId="3" pageNumber="38">scabriculum</emphasis>
</taxonomicName>
,
<taxonomicName id="C6E38BD56F435639D4F7FA682363FA7E" box="[151,260,1429,1453]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="juberthiei" status="sp. nov.">
<emphasis id="33972C446F435639D4F7FA682363FA7E" box="[151,260,1429,1453]" italics="true" pageId="3" pageNumber="38">juberthiei</emphasis>
</taxonomicName>
<emphasis id="33972C446F435639D56BFA68232DFA7E" bold="true" box="[267,330,1429,1453]" pageId="3" pageNumber="38">
<taxonomicNameLabel id="28A4913F6F435639D56BFA68232DFA7E" box="[267,330,1429,1453]" pageId="3" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="015CF0566F435639D4A5FA4421A6FA25" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
Apophysis of male chelicerae (
<figureCitation id="99D8ECD36F435639D646FA4420FAFA02" box="[550,669,1465,1489]" captionStart="FIGURES 5 10" captionStartId="7.[151,264,1940,1963]" captionTargetBox="[159,1434,404,1899]" captionTargetId="figure@7.[151,1435,397,1914]" captionTargetPageId="7" captionText="FIGURES 5 10. Right 3 chelicerae. 5, 7, 9: lateral (left column) and medial view (right column); 6, 8, 10: dorsal view. 5 6: Centetostoma ventalloi (Col de la Core, CJM 1792). 7 8: C. scabriculum (Torla, CJM 1799). 9 10: C. juberthiei sp. n. (Lac des Bouillouses, CJM 1792)." httpUri="https://zenodo.org/record/202394/files/figure.png" pageId="3" pageNumber="38">Figs 510</figureCitation>
): Well developed, concave medially and (only in the Pyrenean species) split into two parts, a distal and a proximal one (from dorsal view).
</paragraph>
<paragraph id="015CF0566F435639D4A5F9FE2288F993" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
Pedipalps: Slender, only in two species (
<taxonomicName id="C6E38BD56F435639D6F5F9FE2178F9C8" box="[661,799,1539,1563]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="scabriculum">
<emphasis id="33972C446F435639D6F5F9FE2178F9C8" box="[661,799,1539,1563]" italics="true" pageId="3" pageNumber="38">scabriculum</emphasis>
</taxonomicName>
and
<taxonomicName id="C6E38BD56F435639D739F9FE21A1F9C8" box="[857,966,1539,1563]" class="Arachnida" family="Nemastomatidae" genus="Centetostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="juberthiei" status="sp. nov.">
<emphasis id="33972C446F435639D739F9FE21A1F9C8" box="[857,966,1539,1563]" italics="true" pageId="3" pageNumber="38">juberthiei</emphasis>
</taxonomicName>
<emphasis id="33972C446F435639D7AEF9F92677F9CF" bold="true" box="[974,1040,1540,1564]" pageId="3" pageNumber="38">
<taxonomicNameLabel id="28A4913F6F435639D7AEF9F92677F9CF" box="[974,1040,1540,1564]" pageId="3" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) with a spur on the patella mediodistally.
</paragraph>
<paragraph id="015CF0566F435639D4A5F9B32757F9B6" blockId="3.[151,1436,1208,1894]" box="[197,1328,1613,1638]" pageId="3" pageNumber="38">
Legs: Short in the Pyrenean species, (relatively) longer and considerably more slender in
<taxonomicName id="C6E38BD56F435639D0CEF9B3274EF9B6" box="[1198,1321,1614,1637]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="centetes">
<emphasis id="33972C446F435639D0CEF9B3274EF9B6" box="[1198,1321,1614,1637]" italics="true" pageId="3" pageNumber="38">C. centetes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="015CF0566F435639D4A5F98F2309F900" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
Armature of the dorsal side (
<figureCitation id="99D8ECD36F435639D673F98C201AF95A" box="[531,637,1649,1673]" captionStart="FIGURES 2 4" captionStartId="5.[151,264,798,821]" captionTargetBox="[157,1424,185,770]" captionTargetId="figure@5.[151,1436,177,776]" captionTargetPageId="5" captionText="FIGURES 2 4. Dorsal side of body, ♂. 2: Centetostoma ventalloi (Col de la Core, CJM 752). 3: C. scabriculum (Torla, CJM 1799). 4: C. juberthiei sp. n. (Lac des Bouillouses, CJM 1792)." httpUri="https://zenodo.org/record/202393/files/figure.png" pageId="3" pageNumber="38">Figs 24</figureCitation>
): Besides a differing granulation covering the scutum, a more or less indistinct tubercle pair is on each area I-V of the scutum; these are strongly developed with additional long thorns only in
<taxonomicName id="C6E38BD56F435639D48CF9412300F900" box="[236,359,1724,1747]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="centetes">
<emphasis id="33972C446F435639D48CF9412300F900" box="[236,359,1724,1747]" italics="true" pageId="3" pageNumber="38">C. centetes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="015CF0566F435639D4A5F922237CF8CE" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
<emphasis id="33972C446F435639D4A5F922232AF92B" bold="true" box="[197,333,1759,1784]" pageId="3" pageNumber="38">Coloration.</emphasis>
Depending on age brownish to blackish without silvery marks. This holds true also in
<taxonomicName id="C6E38BD56F435639D140F91C27FCF92B" box="[1312,1435,1761,1784]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="centetes">
<emphasis id="33972C446F435639D140F91C27FCF92B" box="[1312,1435,1761,1784]" italics="true" pageId="3" pageNumber="38">C. centetes</emphasis>
</taxonomicName>
of the Alps.
</paragraph>
</subSubSection>
<subSubSection id="49F9A3DD6F435639D4A5F8D423A5F8B5" pageId="3" pageNumber="38" type="distribution">
<paragraph id="015CF0566F435639D4A5F8D423A5F8B5" blockId="3.[151,1436,1208,1894]" pageId="3" pageNumber="38">
<emphasis id="33972C446F435639D4A5F8D42339F891" bold="true" box="[197,350,1833,1858]" pageId="3" pageNumber="38">Distribution.</emphasis>
One species in the western Alps, three at medium to high altitude in the Pyrenees except for the western-most part (
<figureCitation id="99D8ECD36F435639D50FF8B323D6F8B5" box="[367,433,1870,1894]" captionStart="FIGURE 1" captionStartId="2.[151,250,1911,1934]" captionTargetBox="[177,1412,1106,1889]" captionTargetId="figure@2.[177,1413,1106,1889]" captionTargetPageId="2" captionText="FIGURE 1. Pyrenean distribution of Centetostoma scabriculum (squares, blue), C. ventalloi (dots, green) and C. juberthiei sp. n. (triangles, red). Note two sympatric occurrences of C. ventalloi and C. juberthiei sp. n. with overlaying symbols. The western-most overlaying symbols (scabriculum / juberthiei sp. n.) represent the type locality of C. scabriculum (Saint-Sauveur) and denote a false record for C. juberthiei sp. n." httpUri="https://zenodo.org/record/202392/files/figure.png" pageId="3" pageNumber="38">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>