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<document id="CFF88A006F7396C115E66EFC8EA5F290" ID-CLB-Dataset="21035" ID-DOI="10.5281/zenodo.4004389" ID-GBIF-Dataset="03168ca4-19e8-4ef5-a000-0e23e0c1a0d2" ID-ISSN="0253-116X" ID-Zenodo-Dep="4004389" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598542413075" checkinUser="valdenar" docAuthor="Smetana, Tephens &amp; Shavrin, A." docDate="2018" docId="8948DC625D6EFFACFF36FF26FF3FA01E" docLanguage="en" docName="Linzer.biol.Beitr.50.1.0825-0836.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Quedius (Microsaurus) amurensis Smetana &amp; Shavrin, 2018, nov.sp." docType="treatment" docVersion="7" lastPageNumber="831" masterDocId="7571A41A5D6BFFAAFF84FFFEFFCBA633" masterDocTitle="Contribution to the knowledge of the genus Quedius S, 1829 of Siberia and Russian Far East (Coleoptera: Staphylinidae: Staphylinini: Quediina)" masterLastPageNumber="836" masterPageNumber="825" pageNumber="830" updateTime="1698845623269" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5095EFCBB769991B75AB5ADC15B977B7">Contribution to the knowledge of the genus Quedius S, 1829 of Siberia and Russian Far East (Coleoptera: Staphylinidae: Staphylinini: Quediina)</mods:title>
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<mods:namePart id="FD92A4B3C6472AF514894390171F2BB4">Smetana, Tephens</mods:namePart>
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<mods:title id="F7C4C761584C2499F71CFD46A566AFB2">Linzer biologische Beiträge</mods:title>
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<mods:date id="F6BDC6CC00E7AFA140D59A3EAF6006D2">2018</mods:date>
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<paragraph id="015E6D745D6EFFAFFF36FF26FCBBA6DD" blockId="5.[178,880,216,238]" box="[178,880,216,238]" pageId="5" pageNumber="830">
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<emphasis id="3395B1665D6EFFAFFF36FF26FCBBA6DD" bold="true" box="[178,880,216,238]" pageId="5" pageNumber="830">
<taxonomicName id="C6E116F75D6EFFAFFF36FF26FDD3A6DD" ID-CoL="4R2K2" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[178,536,216,238]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="amurensis" subGenus="Microsaurus">
<emphasis id="3395B1665D6EFFAFFF36FF26FEC2A6DD" bold="true" box="[178,265,216,238]" italics="true" pageId="5" pageNumber="830">Quedius</emphasis>
(
<emphasis id="3395B1665D6EFFAFFE9CFF27FE56A6DD" bold="true" box="[280,413,217,238]" italics="true" pageId="5" pageNumber="830">Microsaurus</emphasis>
)
<emphasis id="3395B1665D6EFFAFFE28FF27FDD3A6DD" bold="true" box="[428,536,217,238]" italics="true" pageId="5" pageNumber="830">amurensis</emphasis>
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SMETANA
<taxonomicNameLabel id="28A60C1D5D6EFFAFFD09FF27FD12A6DD" box="[653,729,217,238]" pageId="5" pageNumber="830" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="99DA71F15D6EFFAFFD62FF26FC8EA6DD" box="[742,837,216,238]" captionStart="Figs 1-11" captionStartId="3.[178,218,1403,1425]" captionTargetBox="[198,1071,221,1371]" captionTargetId="figure@3.[178,1085,212,1385]" captionTargetPageId="3" captionText="Figs 1-11: 1-6. Quedius conviva nov.sp. (1)- apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of underside of paramere with sensory peg setae; (6) tergite 10 of female genital segment. 7-11. Quedius amurensis nov.sp. (7) apical portion of male sternite 8; (8) tergite 10 of male genital segment (9) sternite 9 of male genital segment; (10) aedoeagus, ventral view; (11) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004391" httpUri="https://zenodo.org/record/4004391/files/figure.png" pageId="5" pageNumber="830">Figs 7-11</figureCitation>
,
<figureCitation id="99DA71F15D6EFFAFFCD4FF26FCA2A6DD" box="[848,873,216,238]" captionStart="Figs 22-23" captionStartId="6.[178,218,1433,1455]" captionTargetBox="[178,1085,212,1416]" captionTargetId="figure@6.[178,1085,212,1416]" captionTargetPageId="6" captionText="Figs 22-23: (22) Type locality of Quedius conviva nov.sp.; (23) Type locality of Quedius amurensis nov.sp." figureDoi="http://doi.org/10.5281/zenodo.4004395" httpUri="https://zenodo.org/record/4004395/files/figure.png" pageId="5" pageNumber="830">23</figureCitation>
)
</emphasis>
</heading>
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<paragraph id="015E6D745D6EFFAFFF36FF01FCE7A71F" blockId="5.[178,1086,254,428]" pageId="5" pageNumber="830">
T y p e l o c a l i t y.
<collectingCountry id="79F62DE45D6EFFAFFE1AFF01FE3BA727" box="[414,496,255,276]" name="Russia" pageId="5" pageNumber="830">RUSSIA</collectingCountry>
: Amurskaya obl., Skovorodinskiy r-n., lev. ber. r. Urka, ok
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S p. Erofei Pavlovich, N 54˚ 0224.84 E 121˚ 58 30.69, h=
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.
</specimenCount>
</paragraph>
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T y p e m a t e r i a l.
<typeStatus id="DE5AD3D65D6EFFAFFE22FEC8FE3CA77F" box="[422,503,310,332]" pageId="5" pageNumber="830" type="holotype">Holotype</typeStatus>
(
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),
<collectingCountry id="79F62DE45D6EFFAFFDADFEC9FDB1A77F" box="[553,634,311,332]" name="Russia" pageId="5" pageNumber="830">RUSSIA</collectingCountry>
: &quot;Amurskaya obl., Skovorodinskiy r-n., lev. ber. r. Urka, ok
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S p. Erofei Pavlovich, 54˚ 0224.84 N
<specimenCount id="17E7A6FD5D6EFFAFFD74FEAAFCCAA75A" box="[752,769,340,361]" pageId="5" pageNumber="830" type="female"></specimenCount>
121˚ 58 30.69E, h=
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, 28-
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<collectingDate id="651BB25C5D6EFFAFFF5BFE99FE88A74F" box="[223,323,359,380]" pageId="5" pageNumber="830" value="2014-08-29">29.08.2014</collectingDate>
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,
<collectorName id="AC1408A25D6EFFAFFEC9FE99FE0FA74F" box="[333,452,358,380]" pageId="5" pageNumber="830">A.V. Shavrin</collectorName>
,
<collectorName id="AC1408A25D6EFFAFFE4AFE99FDBCA74F" box="[462,631,358,380]" pageId="5" pageNumber="830">I. V. Enushchenko</collectorName>
&quot;. [
<collectingRegion id="C325A3965D6EFFAFFD09FE99FD08A74F" box="[653,707,359,380]" country="Russia" name="Amur" pageId="5" pageNumber="830">Amur</collectingRegion>
<collectingCounty id="E83F15F85D6EFFAFFD4DFE99FD3EA74F" box="[713,757,359,380]" pageId="5" pageNumber="830">Area</collectingCounty>
,
<collectingMunicipality id="E13AF70E5D6EFFAFFC84FE98FC1FA74F" box="[768,980,358,380]" pageId="5" pageNumber="830">Skorovodinskiy District</collectingMunicipality>
, left side of
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, about
<locationDeviation id="10EC74565D6EFFAFFEF9FE81FDBEA7A7" box="[381,629,382,404]" location="Yerofey Pavlovitch" pageId="5" pageNumber="830">
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N Yerofey Pavlovitch
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, 54˚ 0224.84 N / 121˚ 58 30.69E, h=
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<elevation id="8ACC8A475D6EFFAFFC5FFE80FBDDA7A7" box="[987,1046,382,404]" metricMagnitude="2" metricUnit="m" metricValue="5.52" pageId="5" pageNumber="830" unit="m" value="552.0">552 m</elevation>
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, 28-
<date id="755F4BB45D6EFFAFFF5BFE69FE88A79F" box="[223,323,407,428]" pageId="5" pageNumber="830" value="2014-08-29">
<collectingDate id="651BB25C5D6EFFAFFF5BFE69FE88A79F" box="[223,323,407,428]" pageId="5" pageNumber="830" value="2014-08-29">29.08.2014</collectingDate>
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,
<collectorName id="AC1408A25D6EFFAFFEC8FE69FE09A79F" box="[332,450,406,428]" pageId="5" pageNumber="830">A.V. Shavrin</collectorName>
,
<collectorName id="AC1408A25D6EFFAFFE4EFE69FDA7A79F" box="[458,620,406,428]" pageId="5" pageNumber="830">I. V. Enushchenko</collectorName>
]. Deposited in ZMM.
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</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FE45FDF3A43A" blockId="5.[178,1085,442,521]" pageId="5" pageNumber="830">
D i a g n o s i s. Species similar in external characters to both
<taxonomicName id="C6E116F75D6EFFAFFCBEFE44FC78A7FC" authorityName="Sharp" authorityYear="1874" box="[826,947,442,464]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="3395B1665D6EFFAFFCBEFE44FC87A7FC" box="[826,844,442,463]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFCDDFE44FC78A7FC" box="[857,947,442,463]" italics="true" pageId="5" pageNumber="830">simulans</emphasis>
</taxonomicName>
<bibRefCitation id="657010855D6EFFAFFC3EFE44FBF6A7FC" author="SHARP D" box="[954,1085,442,464]" pageId="5" pageNumber="830" pagination="1 - 103" refId="ref6211" refString="SHARP D. (1874): The Staphylinidae of Japan. - The Transactions of the Entomolog ical Society of London 1874: 1 - 103." type="journal article" year="1874">SHARP, 1874</bibRefCitation>
and
<taxonomicName id="C6E116F75D6EFFAFFF59FE29FEA4A7DF" box="[221,367,471,492]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFF59FE29FF24A7DF" box="[221,239,471,492]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFF78FE29FEA4A7DF" box="[252,367,471,492]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
<bibRefCitation id="657010855D6EFFAFFEF2FE29FDD0A7DE" author="SMETANA A" box="[374,539,471,493]" pageId="5" pageNumber="830" pagination="113 - 132" refId="ref6349" refString="SMETANA A. (1996): Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 5. Genus Quedius STEPHENS, 1829. Subgenus Microsaurus DEJEAN, 1833. Section 4. - Bulletion of t he National Science Museum Tokyo, Serie A (Zoology) 22: 113 - 132." type="journal article" year="1996">SMETANA, 1996</bibRefCitation>
, but different mainly by the aedoeagus with paramere wide, covering the entire median lobe.
</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FDE6FBE6A578" blockId="5.[178,1085,536,843]" pageId="5" pageNumber="830">
D e s c r i p t i o n. In all external characters similar to
<taxonomicName id="C6E116F75D6EFFAFFD7AFDE6FC59A41E" box="[766,914,536,557]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFD7AFDE6FCDBA41E" box="[766,784,536,557]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFC9BFDE6FC59A41E" box="[799,914,536,557]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
, but different as follows: size smaller than average specimens of
<taxonomicName id="C6E116F75D6EFFAFFD2CFDCBFCF7A47A" box="[680,828,564,586]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFD2CFDCBFD71A479" box="[680,698,565,586]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFD4DFDCAFCF7A47A" box="[713,828,564,585]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
, head shorter and wider, distinctly wider than long (ratio 1.20, corresponding ratio for
<taxonomicName id="C6E116F75D6EFFAFFCD8FDAFFC3FA455" box="[860,1012,593,614]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFCD8FDAFFCA5A455" box="[860,878,593,614]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFC05FDAFFC3FA455" box="[897,1012,593,614]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
1.12); microsculpture distinctly finer and more superficial; antenna similar to that of
<taxonomicName id="C6E116F75D6EFFAFFBA0FD90FEEEA4AC" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFBA0FD90FBFDA4B0" box="[1060,1078,622,643]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFF36FD74FEEEA4AC" box="[178,293,650,671]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
, but distinctly more slender and shorter, with outer segments 7-10 distinctly transverse. Pronotum shorter and wider, more markedly wider than long (ratio 1.19, corresponding ratio for
<taxonomicName id="C6E116F75D6EFFAFFE29FD3DFD88A4EB" box="[429,579,707,728]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFE29FD3DFE74A4EB" box="[429,447,707,728]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFE54FD3DFD88A4EB" box="[464,579,707,728]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
1.10); left dorsal row with only two, right one with three punctures; scutellum with only a few punctures; elytra markedly shorter than those of
<taxonomicName id="C6E116F75D6EFFAFFE8FFD02FE55A522" box="[267,414,764,785]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFE8FFD02FED6A522" box="[267,285,764,785]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFEAFFD02FE55A522" box="[299,414,764,785]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
, at suture about as long as, at sides longer than pronotum at midline (ratio 1.15); punctation and setation less dense than in
<taxonomicName id="C6E116F75D6EFFAFFC97FCE7FC6DA51D" box="[787,934,793,814]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFC97FCE7FCEEA51D" box="[787,805,793,814]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFCB7FCE7FC6DA51D" box="[819,934,793,814]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
; punctation of abdominal tergites similar to that of
<taxonomicName id="C6E116F75D6EFFAFFDA7FCCBFD7EA579" box="[547,693,821,843]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFDA7FCCBFDFEA579" box="[547,565,821,842]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFDC6FCCBFD7EA579" box="[578,693,821,842]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
, but in general somewhat less dense.
</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FCA4FC6DA2CC" blockId="5.[178,1085,858,1279]" pageId="5" pageNumber="830">
Male. First four segments of front tarsus moderately dilated, sub-bilobed, each densely covered with tenent setae ventrally; segment 2 about as wide as apex of tibia; segment 4 narrower than preceding segments. Sternite 7 with slight medioapical sinuation; sternite 8 with four macrosetae at right side and three on left side, apical margin with wide, moderately deep, obtusely triangular medioapical emargination, a long, narrow area before emargination, reaching basal third of sternite, devoid of pubescence (
<figureCitation id="99DA71F15D6EFFAFFC68FC16FBFBA5CD" box="[1004,1072,1000,1022]" captionStart="Figs 1-11" captionStartId="3.[178,218,1403,1425]" captionTargetBox="[198,1071,221,1371]" captionTargetId="figure@3.[178,1085,212,1385]" captionTargetPageId="3" captionText="Figs 1-11: 1-6. Quedius conviva nov.sp. (1)- apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of underside of paramere with sensory peg setae; (6) tergite 10 of female genital segment. 7-11. Quedius amurensis nov.sp. (7) apical portion of male sternite 8; (8) tergite 10 of male genital segment (9) sternite 9 of male genital segment; (10) aedoeagus, ventral view; (11) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004391" httpUri="https://zenodo.org/record/4004391/files/figure.png" pageId="5" pageNumber="830">Fig. 7</figureCitation>
); tergite 10 evenly narrowed toward arcuate apex, with a few long setae at apex and some shorter setae in front of them (
<figureCitation id="99DA71F15D6EFFAFFE6DFBDFFDECA20B" box="[489,551,1057,1080]" captionStart="Figs 1-11" captionStartId="3.[178,218,1403,1425]" captionTargetBox="[198,1071,221,1371]" captionTargetId="figure@3.[178,1085,212,1385]" captionTargetPageId="3" captionText="Figs 1-11: 1-6. Quedius conviva nov.sp. (1)- apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of underside of paramere with sensory peg setae; (6) tergite 10 of female genital segment. 7-11. Quedius amurensis nov.sp. (7) apical portion of male sternite 8; (8) tergite 10 of male genital segment (9) sternite 9 of male genital segment; (10) aedoeagus, ventral view; (11) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004391" httpUri="https://zenodo.org/record/4004391/files/figure.png" pageId="5" pageNumber="830">Fig. 8</figureCitation>
); sternite 9 not appreciably different from that of
<taxonomicName id="C6E116F75D6EFFAFFBA0FBDCFEEEA260" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFBA0FBDCFBFDA204" box="[1060,1078,1058,1079]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFF36FBC0FEEEA260" box="[178,293,1086,1107]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
(
<figureCitation id="99DA71F15D6EFFAFFEB1FBC0FEBDA267" box="[309,374,1086,1108]" captionStart="Figs 1-11" captionStartId="3.[178,218,1403,1425]" captionTargetBox="[198,1071,221,1371]" captionTargetId="figure@3.[178,1085,212,1385]" captionTargetPageId="3" captionText="Figs 1-11: 1-6. Quedius conviva nov.sp. (1)- apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of underside of paramere with sensory peg setae; (6) tergite 10 of female genital segment. 7-11. Quedius amurensis nov.sp. (7) apical portion of male sternite 8; (8) tergite 10 of male genital segment (9) sternite 9 of male genital segment; (10) aedoeagus, ventral view; (11) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004391" httpUri="https://zenodo.org/record/4004391/files/figure.png" pageId="5" pageNumber="830">Fig. 9</figureCitation>
). Aedoeagus (
<figureCitation id="99DA71F15D6EFFAFFD8EFBC0FD49A267" box="[522,642,1086,1108]" captionStart="Figs 1-11" captionStartId="3.[178,218,1403,1425]" captionTargetBox="[198,1071,221,1371]" captionTargetId="figure@3.[178,1085,212,1385]" captionTargetPageId="3" captionText="Figs 1-11: 1-6. Quedius conviva nov.sp. (1)- apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of underside of paramere with sensory peg setae; (6) tergite 10 of female genital segment. 7-11. Quedius amurensis nov.sp. (7) apical portion of male sternite 8; (8) tergite 10 of male genital segment (9) sternite 9 of male genital segment; (10) aedoeagus, ventral view; (11) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004391" httpUri="https://zenodo.org/record/4004391/files/figure.png" pageId="5" pageNumber="830">Figs 10, 11</figureCitation>
) markedly different from those of both
<taxonomicName id="C6E116F75D6EFFAFFBA0FBC0FEC7A25C" authorityName="Sharp" authorityYear="1874" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="3395B1665D6EFFAFFBA0FBC0FBFDA260" box="[1060,1078,1086,1107]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFF36FBA4FEC7A25C" box="[178,268,1114,1135]" italics="true" pageId="5" pageNumber="830">simulans</emphasis>
</taxonomicName>
and
<taxonomicName id="C6E116F75D6EFFAFFEC4FBA5FE1FA25C" box="[320,468,1114,1136]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3395B1665D6EFFAFFEC4FBA5FE99A243" box="[320,338,1115,1136]" italics="true" pageId="5" pageNumber="830">Q</emphasis>
.
<emphasis id="3395B1665D6EFFAFFEE5FBA4FE1FA25C" box="[353,468,1114,1135]" italics="true" pageId="5" pageNumber="830">klapperichi</emphasis>
</taxonomicName>
; median lobe narrow, subparallelsided, anteriorly narrowed into narrow apical part with acute apex; paramere large, wide, entirely covering median lobe, anteriorly narrowed into subacute apex surpasing distinctly apex of median lobe; four apical setae, medial pair considerably longer than lateral pair and two short setae at each lateral margin close to apex; underside with numerous sensory peg setae, forming irregular long row along each lateral margin of apical portion of paramere.
</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FAF3FEA2A310" blockId="5.[178,361,1293,1315]" box="[178,361,1293,1315]" pageId="5" pageNumber="830">Female unknown.</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FACFFE9EA374" blockId="5.[178,341,1329,1351]" box="[178,341,1329,1351]" pageId="5" pageNumber="830">
Length
<quantity id="C619C0915D6EFFAFFE84FACFFE9BA374" box="[256,336,1329,1351]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="830" unit="mm" value="6.8">6.8 mm</quantity>
.
</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FAA8FC05A3BB" blockId="5.[178,1085,1366,1416]" pageId="5" pageNumber="830">
E t y m o l o g y. The specific epithet is the Latinized adjective derived from the name of the river
<collectingRegion id="C325A3965D6EFFAFFEADFA8CFEAFA3B4" box="[297,356,1394,1415]" country="Russia" name="Amur" pageId="5" pageNumber="830">Amur</collectingRegion>
and by adding the suffix -
<emphasis id="3395B1665D6EFFAFFDEBFA8DFD6AA3BB" box="[623,673,1395,1416]" italics="true" pageId="5" pageNumber="830">ensis</emphasis>
, denoting the place of origin.
</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FA69FD98A3FA" blockId="5.[178,1085,1431,1481]" pageId="5" pageNumber="830">
G e o g r a p h i c a l d i s t r i b u t i o n.
<taxonomicName id="C6E116F75D6EFFAFFD11FA69FC96A39F" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[661,861,1431,1452]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="830" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="3395B1665D6EFFAFFD11FA69FC96A39F" box="[661,861,1431,1452]" italics="true" pageId="5" pageNumber="830">Quedius amurensis</emphasis>
</taxonomicName>
is at present known only from the
<typeStatus id="DE5AD3D65D6EFFAFFEC6FA4AFEA5A3FA" box="[322,366,1460,1481]" pageId="5" pageNumber="830">type</typeStatus>
locality in
<collectingRegion id="C325A3965D6EFFAFFE64FA4DFDD0A3FB" box="[480,539,1459,1480]" country="Russia" name="Amur" pageId="5" pageNumber="830">Amur</collectingRegion>
area.
</paragraph>
<paragraph id="015E6D745D6EFFAFFF36FA26FD6BA077" blockId="5.[178,1085,1496,1604]" pageId="5" pageNumber="830">
B i o n o m i c s. The specimen was sifted from litter and mosses at swampy site along a forest road with
<emphasis id="3395B1665D6EFFAFFEF1FA0AFDDEA03A" box="[373,533,1524,1545]" italics="true" pageId="5" pageNumber="830">Ledum palustre</emphasis>
,
<emphasis id="3395B1665D6EFFAFFDA1FA0BFCC1A03A" box="[549,778,1524,1546]" italics="true" pageId="5" pageNumber="830">Vaccinium uliginosum</emphasis>
,
<emphasis id="3395B1665D6EFFAFFC9EFA0AFBDDA039" box="[794,1046,1524,1546]" italics="true" pageId="5" pageNumber="830">Calamagrostis purpurea</emphasis>
,
<emphasis id="3395B1665D6EFFAFFBA2FA0BFEC2A015" italics="true" pageId="5" pageNumber="830">C. obtusata</emphasis>
,
<emphasis id="3395B1665D6EFFAFFE9CF9EFFE9DA015" box="[280,342,1553,1574]" italics="true" pageId="5" pageNumber="830">Carex</emphasis>
spp. and
<emphasis id="3395B1665D6EFFAFFE3BF9EFFDE1A015" box="[447,554,1553,1574]" italics="true" pageId="5" pageNumber="830">Sphagnum</emphasis>
spp. in a mixed forest with
<emphasis id="3395B1665D6EFFAFFCDCF9ECFC20A015" box="[856,1003,1553,1575]" italics="true" pageId="5" pageNumber="830">Larix gmelinii</emphasis>
,
<emphasis id="3395B1665D6EFFAFFC7FF9EFFEC7A070" italics="true" pageId="5" pageNumber="830">Betula fruticosa</emphasis>
,
<emphasis id="3395B1665D6EFFAFFE9DF9D0FE6CA071" box="[281,423,1581,1603]" italics="true" pageId="5" pageNumber="830">B. platyphylla</emphasis>
and
<emphasis id="3395B1665D6EFFAFFE5CF9D3FDC0A071" box="[472,523,1581,1602]" italics="true" pageId="5" pageNumber="830">Salix</emphasis>
spp. (
<figureCitation id="99DA71F15D6EFFAFFDCCF9D3FD5EA077" box="[584,661,1581,1604]" captionStart="Figs 22-23" captionStartId="6.[178,218,1433,1455]" captionTargetBox="[178,1085,212,1416]" captionTargetId="figure@6.[178,1085,212,1416]" captionTargetPageId="6" captionText="Figs 22-23: (22) Type locality of Quedius conviva nov.sp.; (23) Type locality of Quedius amurensis nov.sp." figureDoi="http://doi.org/10.5281/zenodo.4004395" httpUri="https://zenodo.org/record/4004395/files/figure.png" pageId="5" pageNumber="830">Fig. 23</figureCitation>
).
</paragraph>
<caption id="559E3DFC5D6DFFACFF36FA67FE9CA3F5" ID-DOI="http://doi.org/10.5281/zenodo.4004395" ID-Zenodo-Dep="4004395" httpUri="https://zenodo.org/record/4004395/files/figure.png" pageId="6" pageNumber="831" startId="6.[178,218,1433,1455]" targetBox="[178,1085,212,1416]" targetPageId="6">
<paragraph id="015E6D745D6DFFACFF36FA67FE9CA3F5" blockId="6.[178,1085,1433,1478]" pageId="6" pageNumber="831">
<emphasis id="3395B1665D6DFFACFF36FA67FED7A39C" bold="true" box="[178,284,1433,1455]" pageId="6" pageNumber="831">Figs 22-23</emphasis>
: (
<emphasis id="3395B1665D6DFFACFEBCFA64FE84A39C" bold="true" box="[312,335,1434,1455]" pageId="6" pageNumber="831">22</emphasis>
) Type locality of
<taxonomicName id="C6E116F75D6DFFACFD94FA64FD64A39C" authority="Smetana &amp; Shavrin, 2018" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[528,687,1434,1455]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="831" phylum="Arthropoda" rank="species" species="conviva" status="sp. nov.">
<emphasis id="3395B1665D6DFFACFD94FA64FD64A39C" box="[528,687,1434,1455]" italics="true" pageId="6" pageNumber="831">Quedius conviva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28A60C1D5D6DFFACFD39FA64FCCAA39C" box="[701,769,1434,1455]" pageId="6" pageNumber="831" rank="species">nov.sp.</taxonomicNameLabel>
; (
<emphasis id="3395B1665D6DFFACFC9DFA64FCFBA39C" bold="true" box="[793,816,1434,1455]" pageId="6" pageNumber="831">23</emphasis>
) Type locality of
<taxonomicName id="C6E116F75D6DFFACFC76FA64FEC4A3F5" authority="Smetana &amp; Shavrin, 2018" authorityName="Smetana &amp; Shavrin" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="831" phylum="Arthropoda" rank="species" species="amurensis" status="sp. nov.">
<emphasis id="3395B1665D6DFFACFC76FA64FEC4A3F5" italics="true" pageId="6" pageNumber="831">Quedius amurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28A60C1D5D6DFFACFE91FA4FFE9CA3F5" box="[277,343,1457,1478]" pageId="6" pageNumber="831" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="015E6D745D6DFFACFF36FA23FF3FA01E" blockId="6.[178,1085,1500,1581]" pageId="6" pageNumber="831">
C o m m e n t s. The
<typeStatus id="DE5AD3D65D6DFFACFE26FA22FE31A3C1" box="[418,506,1500,1522]" pageId="6" pageNumber="831" type="holotype">holotype</typeStatus>
bears an additional, small setiferous puncture anteriomediad of right posterior frontal puncture and it is missing the last segment of right front tarsus.
</paragraph>
</subSubSection>
</treatment>
</document>