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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1171.104142" ID-Pensoft-Pub="1313-2970-1171-1" ID-Pensoft-UUID="7559C2D42DE85144AEFFC98A16172F97" ID-ZooBank="267D0D4559CA4A18A08034768E652607" ModsDocID="1313-2970-1171-1" checkinTime="1689969829564" checkinUser="pensoft" docAuthor="Short, Andrew Edward Z. &amp; Giron, Jennifer C." docDate="2023" docId="8911FCCD32625A149B31EFFFE605F6A0" docLanguage="en" docName="ZooKeys 1171: 1-112" docOrigin="ZooKeys 1171" docPubDate="2023-07-20" docSource="http://dx.doi.org/10.3897/zookeys.1171.104142" docTitle="Novochares aperito Short &amp; Girón 2023, sp. nov." docType="treatment" docUuid="0DE84FFA-2FB4-44C6-AC75-2B80606B148E" docUuidSource="ZooBank" docVersion="1" id="7559C2D42DE85144AEFFC98A16172F97" lastPageNumber="1" masterDocId="7559C2D42DE85144AEFFC98A16172F97" masterDocTitle="Revision of the Neotropical water scavenger beetle genus Novochares Giron &amp; Short (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="112" masterPageNumber="1" pageNumber="1" updateTime="1689969829564" updateUser="pensoft">
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<mods:title id="A57BF2EADD0D477C209FD4B7267D7FA2">Revision of the Neotropical water scavenger beetle genus Novochares Giron &amp; Short (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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<mods:namePart id="E553FC9629ED74F62D40042400620E73">Short, Andrew Edward Z.</mods:namePart>
<mods:nameIdentifier id="42F188270DD7B60D2A5A92B8CD96811C" type="ORCID">https://orcid.org/0000-0002-7467-7116</mods:nameIdentifier>
<mods:affiliation id="A2AF9E3DA6E5AE2C48BF99B3A99BD6F5">Department of Entomology &amp; Nematology, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
<mods:nameIdentifier id="E9549E72BF1E7EB1F44216DA4912469B" type="email">aezshort@ku.edu</mods:nameIdentifier>
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<mods:namePart id="389D7BA42E373212A843E4B56DB413A1">Giron, Jennifer C.</mods:namePart>
<mods:nameIdentifier id="804F6DA77C69285459C02D1103B0EC9D" type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
<mods:affiliation id="3501E87D35FE3025B9B3BDA8C23CE97D">Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
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<mods:date id="3EA384D1573C27A0B498BD6D9A4C79CF">2023</mods:date>
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<mods:number id="2572E13E444334D02E7C2B1EBD38C4AE">1171</mods:number>
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<subSubSection id="DE769779A8797F20295A2ACB8CD9C2A9" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="C881C64DA0B250CCDAE492ED8001434B" pageId="0" pageNumber="1">
<taxonomicName id="3B8E0EB15F95DE125FAF30D1E37FB5E7" LSID="https://zoobank.org/0DE84FFA-2FB4-44C6-AC75-2B80606B148E" authority="Short &amp; Girón, 2023" authorityName="Short &amp; Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares aperito" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aperito" status="sp. nov.">Novochares aperito</taxonomicName>
<taxonomicNameLabel id="C968F6A1DF38B1859F81024F9BBE0703" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D5BE4598FFEF856A4E10A6139CBC8CA3" pageId="0" pageNumber="1" type="description">
<paragraph id="108BA73EE20661810AC9D9D3E08A207D" pageId="0" pageNumber="1">
<figureCitation id="F37BB32A1DE14105E465D64703B69C11" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Novochares spp. A-C N. aperito D-F N. orchis G-K N. minor G-J Peru K Venezuela A, D, G, K dorsal view B, E, H ventral view C, F, J oblique view I lateral view." figureDoi="10.3897/zookeys.1171.104142.figure11" httpUri="https://binary.pensoft.net/fig/881178" pageId="0" pageNumber="1">Figs 11A-C</figureCitation>
<figureCitation id="F98E6DF29713BDB1243EB856414F5201" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Novochares spp. A N. aperito (red), N. minor (yellow), N. orchis (blue) B N. bidens (red), N. furcatus (yellow), N. garfo (blue), N. tenedor (green)." figureDoi="10.3897/zookeys.1171.104142.figure13" httpUri="https://binary.pensoft.net/fig/881180" pageId="0" pageNumber="1">, 13A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C0F3304A9AE39B8463E70E58ADE8220" pageId="0" pageNumber="1" type="type material">
<paragraph id="DF502B7249A3076AF8CE16A290AD27B0" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="C17856F6E856A7DAB3DB43A0B2812C8F" pageId="0" pageNumber="1">
<emphasis id="86A1BB969EAB6A65A0EDE8267F81CFF1" bold="true" pageId="0" pageNumber="1">
<emphasis id="5476FA520D33EC600C50AE3325BC1C4F" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
(male)
</emphasis>
: &quot;BOLIVIA: Santa Cruz Dept./ Potrerillos del Guenda,/ Preserva Natural,
<geoCoordinate id="033740CD6FF99EC4568275270D2F1AD2" degrees="17" direction="south" minutes="40" orientation="latitude" precision="925" value="-17.666666">17°40'S</geoCoordinate>
,
<geoCoordinate id="E0B3236AEF21519FF49EC31D3E46D557" degrees="63" direction="west" minutes="27" orientation="longitude" precision="925" value="-63.45">63°27'W</geoCoordinate>
, 370m, 12-13-X-2007/ ex. BL/MV, A.R Cline &amp; J.E./ Wappes BOL1Cline07 004.5&quot; (SEMC).
</paragraph>
</subSubSection>
<subSubSection id="FF894B3FA18AC85FB88288E13ECD246A" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="624E3F407E6FDAB92D6B2D93B0703E64" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="94130646E8BE475003C9023CB38B17AD" pageId="0" pageNumber="1">See species group diagnosis.</paragraph>
</subSubSection>
<subSubSection id="B3F9F368B8AC7A8D04F36C8A8A9590DD" pageId="0" pageNumber="1" type="description">
<paragraph id="EA52EE220584094020A7B9699E207071" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="3901915787FE9D302B0CB3AAE2EA924E" pageId="0" pageNumber="1">
Body length 4.9 mm.
<emphasis id="012D18D40500ADDA06923DF356E6C38E" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
: Dorsal surfaces orange, with paler clypeus and margins of pronotum.
<emphasis id="EAD5EAFA34DE615C95CE5E1CD48A0F6A" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: Maxillary palps slightly longer than width of head, uniformly orange in color.
<emphasis id="6338DA716955CC6FAE65ECAE8755383A" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
: Ground punctation on pronotum and elytra relatively dense and very shallowly impressed. Elytra without rows of serial punctures, each with very faint rows of weakly marked systematic punctures on lateral 1/2. Prosternum weakly medially convex. Posterior elevation of mesoventrite transversely elevated.
<emphasis id="2D4D512F182CE7E02727E75CE4AF4252" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
: Apical emargination of fifth ventrite relatively deep, U-shaped.
<emphasis id="942D496C6DA77542D1C7269717D81BB1" bold="true" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
: (Fig.
<figureCitation id="AD077C0BC06C1D6DF909FFF88CD0059D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Novochares spp. A-C N. aperito D-F N. orchis G-K N. minor G-J Peru K Venezuela A, D, G, K dorsal view B, E, H ventral view C, F, J oblique view I lateral view." figureDoi="10.3897/zookeys.1171.104142.figure11" httpUri="https://binary.pensoft.net/fig/881178" pageId="0" pageNumber="1">11A-C</figureCitation>
) 2.3
<normalizedToken id="3ACE494C6776C1B6AD98E436C7496034" originalValue="×">x</normalizedToken>
longer than wide, with outer lateral margins of parameres slightly convex along basal 1/2, then weakly sinuate up to apical region; apical region of each paramere rounded, with outer margin somewhat hook-like; at closest point, dorsal inner margins of parameres separated by distance 0.33
<normalizedToken id="ECB94DA7A9D77C45052E9CCB2AC7E355" originalValue="×">x</normalizedToken>
greatest width of a paramere; dorsal plate of median lobe with neck 0.7
<normalizedToken id="F4E5459DFF54FE0E96C32D1862057F0C" originalValue="×">x</normalizedToken>
as broad as base; arms of dorsal plate of median lobe not clearly indicated; apical region of dorsal plate of median lobe dorsally concave, apically pointed; ventral plate of median lobe membranous, neck of dorsal plate; basal piece 0.3
<normalizedToken id="19211507FDAA9953739F04DC15622E23" originalValue="×">x</normalizedToken>
length of a paramere. In lateral view, aedeagus straight at base, with ventral outline of parameres 3
<normalizedToken id="AD7B3C588A9710F21C99681EEE1CD74B" originalValue="×">x</normalizedToken>
longer than greatest width near base; dorsal outline of aedeagus in lateral view slightly convex along basal 1/4, then oblique to apical region; ventral outline of aedeagus in lateral view straight.
</paragraph>
</subSubSection>
<subSubSection id="48D7F4D8C7434602AACAF1A0C34DB528" pageId="0" pageNumber="1" type="etymology">
<paragraph id="36E43A41095F75EA0602841E4A75EF6E" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="E3761EBF9C33E66E033C3F324750770B" pageId="0" pageNumber="1">
<emphasis id="353E4B8B446C6E39396E411D50D7537B" italics="true" pageId="0" pageNumber="1">Aperito</emphasis>
(L.) meaning to open, referring to the distinctive shape of the apical region of the parameres which resemble a bottle opener.
</paragraph>
</subSubSection>
<subSubSection id="79B3CBB18856116A534B427759E6B3B2" pageId="0" pageNumber="1" type="distribution">
<paragraph id="A27068E3DF4AE603A61DB26A70BD6727" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="D7E290524B0A48519A1A74BB45A1C0F0" pageId="0" pageNumber="1">
Only known from the type locality in Bolivia (Fig.
<figureCitation id="D6F22E329B326D2ED96CC74C515FAF18" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Novochares spp. A N. aperito (red), N. minor (yellow), N. orchis (blue) B N. bidens (red), N. furcatus (yellow), N. garfo (blue), N. tenedor (green)." figureDoi="10.3897/zookeys.1171.104142.figure13" httpUri="https://binary.pensoft.net/fig/881180" pageId="0" pageNumber="1">13A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E8746EA0B6367A4BE871191803C74E50" pageId="0" pageNumber="1" type="habitat">
<paragraph id="82769B1686856ED2DEDB28D099574890" pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph id="CCDB5BAF95CA511276BC958B16B9DB62" pageId="0" pageNumber="1">Nothing is known about the habitat of this species as it was collected at lights.</paragraph>
</subSubSection>
<subSubSection id="3E73592B0B9206A25D0E3C51DDEE4B85" pageId="0" pageNumber="1" type="remarks">
<paragraph id="2BB9F231BEB88223719681120F0E94A6" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="AC8953BD90670BE33FAE7F3DCA8E541C" pageId="0" pageNumber="1">Among the thousands of specimens studied here, we only found one specimen of this species, which might be an indicator of their rarity in nature, or a reflection of the lack of sampling in the lowlands of Bolivia. The specimen is in modest condition, with one maxillary palp and a few tarsi missing.</paragraph>
</subSubSection>
<subSubSection id="B16B0AB40A5D054A49F0AFC9D0668820" pageId="0" pageNumber="1" type="novochares garfo species group">
<paragraph id="FDD3A18007D5F4A487F04A033F188BC4" pageId="0" pageNumber="1">
<taxonomicName id="AA85972DCB6AC4B050B9E5AE49D5B6C2" authorityName="Short &amp; Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares garfo" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="garfo">
<emphasis id="FFDF32B74362519B95C9C60FEAEA6290" italics="true" pageId="0" pageNumber="1">Novochares garfo</emphasis>
</taxonomicName>
species group
</paragraph>
<paragraph id="1D12595DEE1480EA798CF8025CE48CA8" pageId="0" pageNumber="1">
<emphasis id="79E640814317DF40ADC33CA1705684BC" bold="true" pageId="0" pageNumber="1">Species group diagnosis.</emphasis>
Body length 4.9-6.5 mm.
<emphasis id="04D88DAA57BFA08789417033DEE7034B" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
: Dorsal surfaces pale brown (orange to yellowish), with slightly paler margins of pronotum and elytra, sometimes also clypeus.
<emphasis id="226D5238E2B439340189705E232875C3" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: Maxillary palps slightly to 1.3
<normalizedToken id="70A1C0A09A15C38E0A10B46E7611B315" originalValue="×">x</normalizedToken>
longer than width of head, uniformly orange or yellow in color.
<emphasis id="57F0C187D206366F2EA3B8D1E9E85504" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
: Ground punctation on pronotum and elytra relatively dense and shallowly impressed. Elytra without rows of serial punctures, each with very faint rows (sometimes one dorsal and two or three lateral, more usually only lateral) of scarce and weakly marked systematic punctures. Prosternum medially broadly and weakly convex. Posterior elevation of mesoventrite broadly elevated, somewhat transverse, often with glabrous longitudinal ridge extending anteriorly.
<emphasis id="1613B3574DD8AEF1FA50D84583D38D76" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
: Apical emargination of fifth ventrite relatively deep, U- or V-shaped.
<emphasis id="034B870B7660EA592C946C22086D5CA6" bold="true" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
: (Fig.
<figureCitation id="FA4594D7906D7549A50E2FBC49393799" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagi of the Novochares garfo species group A-C N. bidens D-F N. furcatus G-J N. garfo K-M N. tenedor A, D, G, K dorsal view B, E, H, L ventral view C, I, M lateral view F, J oblique view." figureDoi="10.3897/zookeys.1171.104142.figure12" httpUri="https://binary.pensoft.net/fig/881179" pageId="0" pageNumber="1">12</figureCitation>
) Overall shape sub-rectangular or oval to pear-like, joint basal margins of parameres truncate to medially pointed or medially emarginate in dorsal view; outer margin and apical region of each paramere weakly to strongly pointed; parameres longer than median lobe, with apex rounded; parameres with apical region variable in degree of sclerotization; dorsal inner margin of each paramere straight to sinuate; dorsal plate of median lobe (in dorsal view) forming a narrow neck; notch between arms variable; arms usually parallel, variable in shape and length; gonopore placed at or near base of dorsal plate of median lobe; ventral plate of median lobe moderately sclerotized, triangular, slender, and apically rounded, not reaching base of fork of dorsal plate; basal piece nearly 0.3
<normalizedToken id="0C3608ECC162B460DFDD838F48590F2D" originalValue="×">x</normalizedToken>
length of a paramere. In lateral view, aedeagus somewhat triangular, straight to weakly oblique at base, with ventral outline of parameres 3-4
<normalizedToken id="A483423F2A9D4063D435C012B327458C" originalValue="×">x</normalizedToken>
longer than greatest width.
</paragraph>
<caption id="DCB77EAE9058A4DBD9E5A39F169132CA" doi="10.3897/zookeys.1171.104142.figure12" httpUri="https://binary.pensoft.net/fig/881179" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph id="FC1BA002C768435C8BEE9E4B4DB8D7C2" pageId="0" pageNumber="1">
<emphasis id="D920B9BC5E73DD149C28FAAF304DE8B0" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
Aedeagi of the
<taxonomicName id="C1D99965BF1DE8727FF7D5C7FDBAEF1E" authorityName="Short &amp; Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares garfo" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="garfo">
<emphasis id="F782D9F588672A7D294955DAAB319CC5" italics="true" pageId="0" pageNumber="1">Novochares garfo</emphasis>
</taxonomicName>
species group
<emphasis id="9B9D6B1F7663FD416B9991FCA55EE3DB" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
<taxonomicName id="D25992FF9C8EE729C89C486CDE1848F0" lsidName="N. bidens" pageId="0" pageNumber="1" rank="species" species="bidens">
<emphasis id="BA3025DD6BF1F9E1FFDA1553D73CA26F" italics="true" pageId="0" pageNumber="1">N. bidens</emphasis>
</taxonomicName>
<emphasis id="E2C8E0D49DEED44A6BC999C6F0F67B6E" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<taxonomicName id="33F2148240748585BD83D4D6965CCB83" lsidName="N. furcatus" pageId="0" pageNumber="1" rank="species" species="furcatus">
<emphasis id="7D595614B8A4D5EDF2F99A325F015921" italics="true" pageId="0" pageNumber="1">N. furcatus</emphasis>
</taxonomicName>
<emphasis id="E634F03E7D5927FA801A6F765F290F34" bold="true" pageId="0" pageNumber="1">G-J</emphasis>
<taxonomicName id="647D689755B21E1352500D537290A306" lsidName="N. garfo" pageId="0" pageNumber="1" rank="species" species="garfo">
<emphasis id="8C799113AEE47C17BEC2213E3C757289" italics="true" pageId="0" pageNumber="1">N. garfo</emphasis>
</taxonomicName>
<emphasis id="77FE1703661750DF0190888037C9494F" bold="true" pageId="0" pageNumber="1">K-M</emphasis>
<taxonomicName id="BE20F3F3BEA58349FC098AF9C31D0DF0" lsidName="N. tenedor" pageId="0" pageNumber="1" rank="species" species="tenedor">
<emphasis id="51814E9515B337EBB3C48A769207A8AC" italics="true" pageId="0" pageNumber="1">N. tenedor</emphasis>
</taxonomicName>
<emphasis id="F90B2F232B34624357C8A65E46FDCB6F" bold="true" pageId="0" pageNumber="1">A, D, G, K</emphasis>
dorsal view
<emphasis id="FA67F911D389691E9E8B089150A0E1C0" bold="true" pageId="0" pageNumber="1">B, E, H, L</emphasis>
ventral view
<emphasis id="F44C8CB2E65CE1BD0D72AEB7E53B8C3A" bold="true" pageId="0" pageNumber="1">C, I, M</emphasis>
lateral view
<emphasis id="8B078FA4E88B51C6AC131BB6272F7B9D" bold="true" pageId="0" pageNumber="1">F, J</emphasis>
oblique view.
</paragraph>
</caption>
<paragraph id="0E108D5355D697A546A393230FF0D77A" pageId="0" pageNumber="1">
<emphasis id="3FA0F74DCDE50582FFF1F57BB5F4749A" bold="true" pageId="0" pageNumber="1">Composition.</emphasis>
The
<taxonomicName id="08A67DFD77AD17504C55218F146A5F26" authorityName="Short &amp; Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares garfo" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="garfo">
<emphasis id="F635B25EFD93EBAAEAFCD66079871486" italics="true" pageId="0" pageNumber="1">Novochares garfo</emphasis>
</taxonomicName>
species group is composed of four species:
<taxonomicName id="DC2BCFA62F26CB29ED315EF1CA496FF6" authorityName="Short &amp; Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares bidens" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="CAA1906B3DDAACAB6F3E1C968B71E943" italics="true" pageId="0" pageNumber="1">Novochares bidens</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="8511E17B983DAAAF7B81D661CB325577" lsidName="N. furcatus" pageId="0" pageNumber="1" rank="species" species="furcatus">
<emphasis id="8EFFE751B04F24917B104A9CDCE01D4E" italics="true" pageId="0" pageNumber="1">N. furcatus</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="38EDEA9D76E9D55E48CDD1BC43F1CDB3" lsidName="N. garfo" pageId="0" pageNumber="1" rank="species" species="garfo">
<emphasis id="4CEC104974EFD5A569A7F3335BAAF100" italics="true" pageId="0" pageNumber="1">N. garfo</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName id="084A89496B3942E963C62A2619C4A398" lsidName="N. tenedor" pageId="0" pageNumber="1" rank="species" species="tenedor">
<emphasis id="05CA49D688BAE08DFBC9B64FE0588A27" italics="true" pageId="0" pageNumber="1">N. tenedor</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph id="935AB55DE8E1B92D680945CB19F3D512" pageId="0" pageNumber="1">
<emphasis id="787F20AA8FBF7267B5C470FAC561EE39" bold="true" pageId="0" pageNumber="1">Remarks.</emphasis>
All four species in this group are relatively light brown/yellow in dorsal coloration, making them resemble the much more common members of the
<taxonomicName id="37980E81A6038300751A193EB2DE3131" baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares abbreviatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="abbreviatus">
<emphasis id="FEBF3DBD36F606B31AA0CAE753706DBF" italics="true" pageId="0" pageNumber="1">Novochares abbreviatus</emphasis>
</taxonomicName>
species group. Species in this group can be recognized by the relatively simple shape of the dorsal plate of the median lobe which resembles a bifid fork in all four species, as well as the shape of the parameres, which are typically straighter and with only a very small tooth near the apex (except
<taxonomicName id="0AD18CAF5A44FDF94019E7769FF6276A" lsidName="N. tenedor" pageId="0" pageNumber="1" rank="species" species="tenedor">
<emphasis id="D92DC9B1BDF848FA400C62F2ADCBD0A8" italics="true" pageId="0" pageNumber="1">N. tenedor</emphasis>
</taxonomicName>
, in which the apex is more curved and well developed).
</paragraph>
</subSubSection>
</treatment>
</document>