327 lines
36 KiB
XML
327 lines
36 KiB
XML
<document id="AC5BE290A99152408E8EB6608128101B" ID-CLB-Dataset="53332" ID-DOI="10.11646/zootaxa.5288.1.1" ID-GBIF-Dataset="1bb291cc-9442-4973-8dd8-9cc79a0a432b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958875" ID-ZooBank="7ABCE6CB-D92C-4B11-87F2-263B7163EEF2" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1684793450573" checkinUser="plazi" docAuthor="Caldara, Roberto & Košťál, Michael" docDate="2023" docId="890F87E5FFD2FFEAC4ACBBDDFF42FC47" docLanguage="en" docName="zootaxa.5288.1.1.pdf" docOrigin="Zootaxa 5288 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5288.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cionus congoanus Hustache 1932" docType="treatment" docVersion="5" lastPageNumber="12" masterDocId="7536FF9DFFDBFFE1C43BB90BFFBBFFAA" masterDocTitle="A Taxonomic Revision Of The Afrotropical Species Of The Weevil Genus Cionus (Coleoptera: Curculionidae)" masterLastPageNumber="98" masterPageNumber="1" pageNumber="10" updateTime="1699506590563" updateUser="plazi">
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<mods:mods id="66CBD92C2253D209BF6157CA94B5D0FC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5BD9A9FA456015DBDD69467DF6FABBEB">
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<mods:title id="3EC915B1B57A9CC32370DE5BA6C0DFC5">A Taxonomic Revision Of The Afrotropical Species Of The Weevil Genus Cionus (Coleoptera: Curculionidae)</mods:title>
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<mods:name id="AAC711CAC85525F603A0617852EF9BBC" type="personal">
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<mods:namePart id="26B7940B48793DB490CD9D259CAC8FCE">Caldara, Roberto</mods:namePart>
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<mods:name id="495B24609EBFEE531BA61EA0C35F52E2" type="personal">
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<mods:roleTerm id="5AF97D077CA1F113144172FBF9364E8B">Author</mods:roleTerm>
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<mods:namePart id="A310E4D5CC85011CF6E5C21A6EE858E8">Košťál, Michael</mods:namePart>
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<mods:nameIdentifier id="CFFEE20CEB19938BB63C307F0CAD2666" type="ORCID">0000-0002-3271-1400</mods:nameIdentifier>
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<mods:nameIdentifier id="4227EAE9DC69D7D84ACDA0327318D9DA" type="email">michael.kostal@iol.cz</mods:nameIdentifier>
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<mods:title id="2EC73AA7682D527E9CCE94B779A630E3">Zootaxa</mods:title>
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<mods:part id="7667FC2938A59D71D9ACA89A4B98DF39">
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<mods:date id="5F944BFDA7CB6BD8B13DDBE4751308CC">2023</mods:date>
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<mods:number id="9DD6287F41ACFB1D2ABE7171081C59AE">2023-05-18</mods:number>
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<mods:number id="C026A9298858630A22452610C38761DD">5288</mods:number>
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<mods:url id="450D69DA16288C1F7299723DA1D3C86F">http://dx.doi.org/10.11646/zootaxa.5288.1.1</mods:url>
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<mods:classification id="B611ACD45DB658917264B46767E82C91">journal article</mods:classification>
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<treatment id="890F87E5FFD2FFEAC4ACBBDDFF42FC47" ID-DOI="http://doi.org/10.5281/zenodo.7967131" ID-GBIF-Taxon="209629215" ID-Zenodo-Dep="7967131" LSID="urn:lsid:plazi:treatment:890F87E5FFD2FFEAC4ACBBDDFF42FC47" httpUri="http://treatment.plazi.org/id/890F87E5FFD2FFEAC4ACBBDDFF42FC47" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection id="49BC6578FFD2FFE8C4ACBBDDFD24FD5A" box="[151,671,726,753]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="011936F3FFD2FFE8C4ACBBDDFD24FD5A" blockId="9.[151,671,726,753]" box="[151,671,726,753]" pageId="9" pageNumber="10">
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<emphasis id="33D2EAE1FFD2FFE8C4ACBBDDFD24FD5A" bold="true" box="[151,671,726,753]" pageId="9" pageNumber="10">
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1.
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<taxonomicName id="C6A64D70FFD2FFE8C48EBBDDFDA8FD5A" ID-CoL="5Z6VT" authority="Hustache" authorityName="Hustache" authorityYear="1932" box="[181,531,726,753]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="congoanus">
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<emphasis id="33D2EAE1FFD2FFE8C48EBBDDFE2DFD5B" bold="true" box="[181,406,726,753]" italics="true" pageId="9" pageNumber="10">Cionus congoanus</emphasis>
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Hustache
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</taxonomicName>
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(
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<figureCitation id="999D2A76FFD2FFE8C619BBDDFD2CFD5A" box="[546,663,726,752]" captionStart="FIGURES 1–2" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[182,1406,184,1879]" captionTargetId="figure-31@10.[178,1410,179,1883]" captionTargetPageId="10" captionText="FIGURES 1–2. FIGURE 1. Cionus congoanus. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. FIGURE 2. Cionus coniungens sp. n. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7958877" httpUri="https://zenodo.org/record/7958877/files/figure.png" pageId="9" pageNumber="10">Figs 1a–f</figureCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="49BC6578FFD2FFE8C4ACBA10FD8BFC98" box="[151,560,794,819]" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="011936F3FFD2FFE8C4ACBA10FD8BFC98" blockId="9.[151,560,794,819]" box="[151,560,794,819]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="21B611DDFFD2FFE8C4ACBA10FD8BFC98" box="[151,560,794,819]" pageId="9" pageNumber="10">
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<treatmentCitation id="800710E2FFD2FFE8C4ACBA10FD97FC98" author="Hustache, A." box="[151,556,794,819]" page="59" pageId="9" pageNumber="10" year="1932">
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<taxonomicName id="C6A64D70FFD2FFE8C4ACBA10FD97FC98" ID-CoL="5Z6VT" authority="Hustache, 1932 a: 59" authorityName="Hustache" authorityPageNumber="59" authorityYear="1932" box="[151,556,794,819]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="congoanus">
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<emphasis id="33D2EAE1FFD2FFE8C4ACBA10FEE8FC98" box="[151,339,795,818]" italics="true" pageId="9" pageNumber="10">Cionus congoanus</emphasis>
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<bibRefCitation id="65374B02FFD2FFE8C562BA10FD97FC98" author="Hustache, A." box="[345,556,794,819]" pageId="9" pageNumber="10" pagination="28 - 109" refId="ref85753" refString="Hustache, A. (1932 a) Curculionides nouveaux de I'Afrique Equatoriale (IIe partie). Sbornik entomologickeho oddeleni narodniho Musea v Praze, 10, 28 - 109." type="journal article" year="1932">Hustache, 1932a: 59</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="49BC6578FFD2FFE8C4ACBA50FEC2FBAE" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="011936F3FFD2FFE8C4ACBA50FCA7FCDE" blockId="9.[151,1437,858,2037]" box="[151,796,858,884]" pageId="9" pageNumber="10">
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<emphasis id="33D2EAE1FFD2FFE8C4ACBA50FE96FCDE" box="[151,301,858,884]" italics="true" pageId="9" pageNumber="10">
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<typeStatus id="DE1D8851FFD2FFE8C4ACBA50FF70FCDE" box="[151,203,859,884]" pageId="9" pageNumber="10">Type</typeStatus>
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locality.
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</emphasis>
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Dindi (
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<collectingCountry id="79B17663FFD2FFE8C5BABA51FCABFCDE" box="[385,784,858,884]" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">Democratic Republic of the Congo</collectingCountry>
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).
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</paragraph>
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<paragraph id="011936F3FFD2FFE8C4FCBA74FEC2FBAE" blockId="9.[151,1437,858,2037]" pageId="9" pageNumber="10">
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<emphasis id="33D2EAE1FFD2FFE8C4FCBA74FEF7FC32" box="[199,332,895,920]" italics="true" pageId="9" pageNumber="10">
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<typeStatus id="DE1D8851FFD2FFE8C4FCBA74FF40FC32" box="[199,251,895,920]" pageId="9" pageNumber="10">Type</typeStatus>
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series.
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</emphasis>
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This species was described from
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<specimenCount id="17A0FD7AFFD2FFE8C6F3BA75FCCBFC32" box="[712,880,894,920]" pageId="9" pageNumber="10" type="generic">five specimens</specimenCount>
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collected at Dindi in
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<collectingCountry id="79B17663FFD2FFE8C05FBA75FEABFC16" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">the Democratic Republic of the Congo</collectingCountry>
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. We did not find these specimens but only
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<specimenCount id="17A0FD7AFFD2FFE8C73BBAA8FC20FC16" box="[768,923,930,957]" pageId="9" pageNumber="10" type="generic">one specimen</specimenCount>
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labelled “Muséum Paris Ogooué Lambarèné [
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<collectingCountry id="79B17663FFD2FFE8C49ABACDFF4AFC4A" box="[161,241,966,992]" name="Gabon" pageId="9" pageNumber="10">Gabon</collectingCountry>
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], R. Ellenberger 1912 /
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<taxonomicName id="C6A64D70FFD2FFE8C5C3BACDFC8EFC4A" authority="Hustache" authorityName="Hustache" authorityYear="1932" box="[504,821,966,993]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="congoanus">Cionus congoanus Hustache</taxonomicName>
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det. ” (
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<collectionCode id="67B7AE36FFD2FFE8C7BDBACDFC64FC4A" box="[902,991,966,992]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
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) perfectly corresponding to the detailed original description.
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</paragraph>
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</subSubSection>
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<subSubSection id="49BC6578FFD2FFE8C4FCBD04FBEEFBDB" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="011936F3FFD2FFE8C4FCBD04FBEEFBDB" blockId="9.[151,1437,858,2037]" pageId="9" pageNumber="10">
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<emphasis id="33D2EAE1FFD2FFE8C4FCBD04FEFBFB82" box="[199,320,1039,1064]" italics="true" pageId="9" pageNumber="10">Diagnosis.</emphasis>
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Pronotum with four protuberances emphasized by tufts of yellowish scales. Prosternum with distinct emargination. Elytra with interstria 3 moderately carinate at base, others with protuberances and tufts of scales, striae distinctly sinuate. Tibiae with mucro in male. Claws symmetrical in both sexes.
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</paragraph>
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</subSubSection>
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<subSubSection id="49BC6578FFD2FFEAC4EABD71FB90FF53" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
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<paragraph id="011936F3FFD2FFE8C4EABD71FBF2F85E" blockId="9.[151,1437,858,2037]" pageId="9" pageNumber="10">
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<emphasis id="33D2EAE1FFD2FFE8C4EABD71FEC8FB3E" box="[209,371,1146,1172]" italics="true" pageId="9" pageNumber="10">Redescription.</emphasis>
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Male (
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<figureCitation id="999D2A76FFD2FFE8C5FDBD71FD81FB3E" box="[454,570,1146,1172]" captionStart="FIGURES 1–2" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[182,1406,184,1879]" captionTargetId="figure-31@10.[178,1410,179,1883]" captionTargetPageId="10" captionText="FIGURES 1–2. FIGURE 1. Cionus congoanus. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. FIGURE 2. Cionus coniungens sp. n. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7958877" httpUri="https://zenodo.org/record/7958877/files/figure.png" pageId="9" pageNumber="10">Figs 1a–b</figureCitation>
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).
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<emphasis id="33D2EAE1FFD2FFE8C66ABD71FD2CFB3E" bold="true" box="[593,663,1146,1172]" pageId="9" pageNumber="10">Body:</emphasis>
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stout, globose.
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<emphasis id="33D2EAE1FFD2FFE8C775BD71FC2CFB3E" bold="true" box="[846,919,1146,1172]" pageId="9" pageNumber="10">Head:</emphasis>
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rostrum stout, short (Rl/Rw 3.45–3.60; Rl/Pl 1.21–1.26), black, in lateral view regularly moderately curved; in dorsal view slightly gradually widening from base to apex, strongly punctate-carinate, from base to near apex with recumbent to suberect, moderately dense, moderately long (l/w 5–6), intermixed grayish and brown scales. Head between eyes narrow, 0.4× as wide as rostrum at base. Eyes flat. Antennae blackish, inserted between middle and apical third; scape long (l/w 8); funicle half as long as scape, S1 slightly more robust than and as long as S2, 2.5× as long as wide,
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<geoCoordinate id="64925034FFD2FFE8C7CABC25FB82FAE3" box="[1009,1081,1326,1353]" degrees="2" direction="south" minutes="2.5" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="-2.0416667">S2 2.5</geoCoordinate>
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× as long as wide, S3–5 as long as wide; club distinctly elongated, oval, moderately longer than funicle, densely pubescent.
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<emphasis id="33D2EAE1FFD2FFE8C0A3BC59FAA2FAC6" bold="true" box="[1176,1305,1362,1388]" pageId="9" pageNumber="10">Pronotum:</emphasis>
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black, with four protuberances, median ones stouter than lateral ones, with dense irregular moderately deep punctures, intervals among punctures shining, smooth, moderately visible between recumbent, moderately dense, intermixed whitish and black scales, darker scales forming tufts on protuberances and two short vittae between median protuberances and base (l/w 4–7); subconical, distinctly transverse (Pw/Pl 1.48–1.55), widest at basal third, sides moderately rounded in basal half and slightly concave in apical half, weakly convex on disc.
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<emphasis id="33D2EAE1FFD2FFE8C7FABF0DFBECF98A" bold="true" box="[961,1111,1542,1568]" pageId="9" pageNumber="10">Prosternum:</emphasis>
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anterior margin with distinct emargination, almost reaching procoxae.
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<emphasis id="33D2EAE1FFD2FFE8C64FBF21FC8CF9EE" bold="true" box="[628,823,1578,1604]" pageId="9" pageNumber="10">Scutellar shield:</emphasis>
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triangular, with dense whitish scales.
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<emphasis id="33D2EAE1FFD2FFE8C0C2BF21FAF5F9EE" bold="true" box="[1273,1358,1578,1604]" pageId="9" pageNumber="10">Elytra:</emphasis>
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black, globose, slightly wider than long (El/Ew 0.90–0.95) widest at midlength, moderately wider than pronotum (Ew/Pw 1.30–1.36), humeri strongly prominent, rounded, sides moderately rounded, distinctly convex on disc; interstriae strongly sinuate, flat to distinctly convex, partly strongly callous (interstriae 3, 5, 7 and 9), moderately visible behind sparse, irregularly arranged, shallow, small punctures; scales whitish (sparse) and black, darker scales covering convex parts of interstriae, long (l/w 7–10), recumbent but erect on calli, moderately dense; striae moderately visible, with irregularly arranged deep punctures, half as wide as interstriae.
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<emphasis id="33D2EAE1FFD2FFE8C77EBE09FC25F8B6" bold="true" box="[837,926,1794,1820]" pageId="9" pageNumber="10">Venter:</emphasis>
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Mesosternal process flat, weakly emarginated. Metathoracic episterna with distinct spot of long, whitish scales. Abdomen with dense punctures, irregular in size and arrangement, and clearly visible between sparse, hair-like, whitish scales, of same shape; V1 and V2 moderately convex, V1 1.6× longer than V2; V1+2 4.9× longer than V3+4, the latter ones 0.5× as long as V5.
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<emphasis id="33D2EAE1FFD2FFE8C13FBE65FAFFF822" bold="true" box="[1284,1348,1902,1928]" pageId="9" pageNumber="10">Legs:</emphasis>
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femora black, with stout sharp teeth, with indistinct striae of moderately dense whitish and black scales; tibiae black, all with mucro; tarsi black, onychium as long as T1–3 combined; claws symmetrical.
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<emphasis id="33D2EAE1FFD2FFE8C03EBEBDFBF6F87A" bold="true" box="[1029,1101,1974,2000]" pageId="9" pageNumber="10">Penis:</emphasis>
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<figureCitation id="999D2A76FFD2FFE8C06DBEBDFB78F87A" box="[1110,1219,1974,2000]" captionStart="FIGURES 1–2" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[182,1406,184,1879]" captionTargetId="figure-31@10.[178,1410,179,1883]" captionTargetPageId="10" captionText="FIGURES 1–2. FIGURE 1. Cionus congoanus. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. FIGURE 2. Cionus coniungens sp. n. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7958877" httpUri="https://zenodo.org/record/7958877/files/figure.png" pageId="9" pageNumber="10">Figs 1d–f</figureCitation>
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, its body gradually narrowed to apex, with blunted tip. Flagellum moderately long, bifurcated at its base.
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</paragraph>
|
||
<caption id="55D9667BFFD1FFEBC4ACBE7EFC75F853" ID-DOI="http://doi.org/10.5281/zenodo.7958877" ID-Zenodo-Dep="7958877" httpUri="https://zenodo.org/record/7958877/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1909,1933]" targetBox="[182,1406,184,1879]" targetPageId="10" targetType="figure">
|
||
<paragraph id="011936F3FFD1FFEBC4ACBE7EFC75F853" blockId="10.[151,1437,1909,2041]" pageId="10" pageNumber="11">
|
||
<emphasis id="33D2EAE1FFD1FFEBC4ACBE7EFE79F827" bold="true" box="[151,450,1909,1933]" pageId="10" pageNumber="11">FIGURES 1–2. FIGURE 1.</emphasis>
|
||
<taxonomicName id="C6A64D70FFD1FFEBC5F2BE7EFD3CF827" authorityName="Hustache" authorityYear="1932" box="[457,647,1909,1933]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="congoanus">
|
||
<emphasis id="33D2EAE1FFD1FFEBC5F2BE7EFD3CF827" box="[457,647,1909,1933]" italics="true" pageId="10" pageNumber="11">Cionus congoanus</emphasis>
|
||
</taxonomicName>
|
||
. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view.
|
||
<emphasis id="33D2EAE1FFD1FFEBC0F6BE92FAF7F81B" bold="true" box="[1229,1356,1945,1969]" pageId="10" pageNumber="11">FIGURE 2.</emphasis>
|
||
<taxonomicName id="C6A64D70FFD1FFEBC16FBE92FEB1F87F" authority="Caldara & Košťál, 2023" authorityName="Caldara & Košťál" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="coniungens" status="sp. nov.">
|
||
<emphasis id="33D2EAE1FFD1FFEBC16FBE92FEB1F87F" italics="true" pageId="10" pageNumber="11">Cionus coniungens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="33D2EAE1FFD1FFEBC52ABEB6FEF0F87F" bold="true" box="[273,331,1981,2005]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="28E1579AFFD1FFEBC52ABEB6FEF0F87F" box="[273,331,1981,2005]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="011936F3FFD0FFEAC4FCB99CFB81FF1B" blockId="11.[151,1437,150,1005]" box="[199,1082,150,177]" pageId="11" pageNumber="12">
|
||
Female. Rostrum slightly longer (Rl/Rw 3.65–3.75; Rl/Pl 1.27–1.30) (
|
||
<figureCitation id="999D2A76FFD0FFEAC7E1B99CFB91FF1B" box="[986,1066,151,177]" captionStart="FIGURES 1–2" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[182,1406,184,1879]" captionTargetId="figure-31@10.[178,1410,179,1883]" captionTargetPageId="10" captionText="FIGURES 1–2. FIGURE 1. Cionus congoanus. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. FIGURE 2. Cionus coniungens sp. n. a: body in dorsal view (male), b: body in lateral view (male), c: rostrum in lateral view (female), d: penis in ventral view, e: penis in lateral view, f: apex of penis in dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7958877" httpUri="https://zenodo.org/record/7958877/files/figure.png" pageId="11" pageNumber="12">Fig. 1c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="011936F3FFD0FFEAC4FCB9B0FB90FF53" blockId="11.[151,1437,150,1005]" pageId="11" pageNumber="12">
|
||
Variability. Length
|
||
<quantity id="C65E9B16FFD0FFEAC59EB9B0FD89FF7F" box="[421,562,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="11" pageNumber="12" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.7–4.2 mm</quantity>
|
||
. The specimens examined showed low variability. The light scales covering scutellum, a part of metathoracic episterna and of elytra are sometimes yellowish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49BC6578FFD0FFEAC4FCB808FB79FDCB" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="011936F3FFD0FFEAC4FCB808FB79FDCB" blockId="11.[151,1437,150,1005]" pageId="11" pageNumber="12">
|
||
<emphasis id="33D2EAE1FFD0FFEAC4FCB808FE8AFEB7" box="[199,305,259,285]" italics="true" pageId="11" pageNumber="12">Remarks.</emphasis>
|
||
The pronotal and elytral protuberances appear more distinctly pronounced due to the tufts of scales covering them. They actually are small in specimens without scales. This is the unique Afrotropical species with tibiae bearing mucro at least in male and one of the few with symmetrical claws also in the male. As regards to the separation of the genera, the inclusion of this species in
|
||
<taxonomicName id="C6A64D70FFD0FFEAC73BB864FCF4FE23" box="[768,847,367,393]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC73BB864FCF4FE23" box="[768,847,367,393]" italics="true" pageId="11" pageNumber="12">Cionus</emphasis>
|
||
</taxonomicName>
|
||
is questionable, because these characters are unusual in this genus (
|
||
<bibRefCitation id="65374B02FFD0FFEAC501B898FDC2FE07" author="Caldara, R. & Korotyaev, B. A." box="[314,633,403,429]" pageId="11" pageNumber="12" pagination="183 - 195" refId="ref85179" refString="Caldara, R. & Korotyaev, B. A. (2002) Taxonomic revision and reconstructed phylogeny of the weevil genus Nanomicrophyes Pic, 1908 (Coleoptera: Curculionidae, Curculioninae). Koleopterologische Rundschau, 72, 183 - 195." type="journal article" year="2002">Caldara & Korotyaev 2002</bibRefCitation>
|
||
): a mucro was observed only in
|
||
<taxonomicName id="C6A64D70FFD0FFEAC7CAB898FBE6FE07" box="[1009,1117,403,429]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="helleri">
|
||
<emphasis id="33D2EAE1FFD0FFEAC7CAB898FBE6FE07" box="[1009,1117,403,429]" italics="true" pageId="11" pageNumber="12">C. helleri</emphasis>
|
||
</taxonomicName>
|
||
from Far East and
|
||
<collectingCountry id="79B17663FFD0FFEAC17BB898FA38FE07" box="[1344,1411,403,429]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
|
||
, a character never emphasized previously (
|
||
<bibRefCitation id="65374B02FFD0FFEAC66BB8BCFCECFE7B" author="Kostal, M. & Caldara, R." box="[592,855,439,465]" pageId="11" pageNumber="12" pagination="1 - 144" refId="ref86052" refString="Kostal, M. & Caldara, R. (2019) Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini). Zootaxa, 4631 (1), 1 - 144. https: // doi. org / 10.11646 / zootaxa. 4631.1.1" type="journal article" year="2019">Košťál & Caldara 2019</bibRefCitation>
|
||
).
|
||
<taxonomicName id="C6A64D70FFD0FFEAC75CB8BCFB8FFE7A" authorityName="Hustache" authorityYear="1932" box="[871,1076,439,465]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="congoanus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC75CB8BCFB8FFE7A" box="[871,1076,439,465]" italics="true" pageId="11" pageNumber="12">Cionus congoanus</emphasis>
|
||
</taxonomicName>
|
||
might be related to species of the genus
|
||
<taxonomicName id="C6A64D70FFD0FFEAC4E4B8D0FE80FE5F" box="[223,315,475,501]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="83" phylum="Arthropoda" rank="subGenus" subGenus="Cleopus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC4E4B8D0FE80FE5F" box="[223,315,475,501]" italics="true" pageId="11" pageNumber="12">Cleopus</emphasis>
|
||
</taxonomicName>
|
||
, which possess both mentioned characters but lack the emargination of the prosternum, or to species of the genera
|
||
<taxonomicName id="C6A64D70FFD0FFEAC515B8F4FE79FDB3" authorityName="Suffrian" authorityYear="1854" box="[302,450,511,537]" class="Insecta" family="Curculionidae" genus="Stereonychus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC515B8F4FE79FDB3" box="[302,450,511,537]" italics="true" pageId="11" pageNumber="12">Stereonychus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C6A64D70FFD0FFEAC5CDB8F4FD13FDB3" authorityName="Morimoto" authorityYear="1962" box="[502,680,511,537]" class="Insecta" family="Curculionidae" genus="Stereonychidius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC5CDB8F4FD13FDB3" box="[502,680,511,537]" italics="true" pageId="11" pageNumber="12">Stereonychidius</emphasis>
|
||
</taxonomicName>
|
||
which are characterized by a single tarsal claw. It is noteworthy that the presence of protuberances on pronotum and elytra and large elytra seem to approach
|
||
<taxonomicName id="C6A64D70FFD0FFEAC053BB28FABBFD96" authorityName="Hustache" authorityYear="1932" box="[1128,1280,547,573]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="congoanus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC053BB28FABBFD96" box="[1128,1280,547,573]" italics="true" pageId="11" pageNumber="12">C. congoanus</emphasis>
|
||
</taxonomicName>
|
||
more to some species from
|
||
<collectingCountry id="79B17663FFD0FFEAC517BB4CFE0EFDCB" box="[300,437,583,609]" name="Madagascar" pageId="11" pageNumber="12">Madagascar</collectingCountry>
|
||
, such as
|
||
<taxonomicName id="C6A64D70FFD0FFEAC622BB4CFD2DFDCB" authorityName="Caldara & Košťál" authorityYear="2023" box="[537,662,583,609]" class="Insecta" family="Curculionidae" genus="Cionus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="callosus">
|
||
<emphasis id="33D2EAE1FFD0FFEAC622BB4CFD2DFDCB" box="[537,662,583,609]" italics="true" pageId="11" pageNumber="12">C. callosus</emphasis>
|
||
</taxonomicName>
|
||
, than to other species from the African mainland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49BC6578FFD0FFEAC4FCBB60FDECFD2F" box="[199,599,619,645]" pageId="11" pageNumber="12" type="biology_ecology">
|
||
<paragraph id="011936F3FFD0FFEAC4FCBB60FDECFD2F" blockId="11.[151,1437,150,1005]" box="[199,599,619,645]" pageId="11" pageNumber="12">
|
||
<emphasis id="33D2EAE1FFD0FFEAC4FCBB60FE3EFD2E" box="[199,389,619,645]" italics="true" pageId="11" pageNumber="12">Biological notes.</emphasis>
|
||
No available data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49BC6578FFD0FFEAC4FCBB84FDB7FD67" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="011936F3FFD0FFEAC4FCBB84FDB7FD67" blockId="11.[151,1437,150,1005]" pageId="11" pageNumber="12">
|
||
<emphasis id="33D2EAE1FFD0FFEAC4FCBB84FEE3FD03" box="[199,344,655,681]" italics="true" pageId="11" pageNumber="12">Distribution.</emphasis>
|
||
West, Central and East Africa (
|
||
<collectingCountry id="79B17663FFD0FFEAC6D8BB84FC8CFD03" box="[739,823,655,681]" name="Guinea-Bissau" pageId="11" pageNumber="12">Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="79B17663FFD0FFEAC773BB84FC7BFD03" box="[840,960,655,681]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
|
||
,
|
||
<collectingCountry id="79B17663FFD0FFEAC7E9BB84FB13FD03" box="[978,1192,655,681]" name="Equatorial Guinea" pageId="11" pageNumber="12">Equatorial Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="79B17663FFD0FFEAC081BB84FAB3FD03" box="[1210,1288,655,681]" name="Gabon" pageId="11" pageNumber="12">Gabon</collectingCountry>
|
||
.
|
||
<collectingCountry id="79B17663FFD0FFEAC121BB84FE23FD67" name="Democratic Republic of the Congo" pageId="11" pageNumber="12">Democratic Republic of the Congo</collectingCountry>
|
||
,
|
||
<collectingCountry id="79B17663FFD0FFEAC59FBBB8FDBBFD67" box="[420,512,691,717]" name="Uganda" pageId="11" pageNumber="12">Uganda</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49BC6578FFD0FFEAC4FCBBDCFF42FC47" pageId="11" pageNumber="12" type="materials_examined">
|
||
<paragraph id="011936F3FFD0FFEAC4FCBBDCFF42FC47" blockId="11.[151,1437,150,1005]" pageId="11" pageNumber="12">
|
||
<emphasis id="33D2EAE1FFD0FFEAC4FCBBDCFDAFFD5B" box="[199,532,727,753]" italics="true" pageId="11" pageNumber="12">Non-type material examined.</emphasis>
|
||
<emphasis id="33D2EAE1FFD0FFEAC626BBDDFD2DFD5A" bold="true" box="[541,662,726,752]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="79B17663FFD0FFEAC626BBDDFD2AFD5A" box="[541,657,726,752]" name="Guinea-Bissau" pageId="11" pageNumber="12">GUINEA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Guinea (3,
|
||
<collectionCode id="67B7AE36FFD0FFEAC71FBBDCFCC6FD5B" box="[804,893,727,753]" pageId="11" pageNumber="12">BMNH</collectionCode>
|
||
).
|
||
<emphasis id="33D2EAE1FFD0FFEAC7A9BBDDFBFEFD5A" bold="true" box="[914,1093,726,752]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="79B17663FFD0FFEAC7A9BBDDFBFBFD5A" box="[914,1088,726,752]" name="Cameroon" pageId="11" pageNumber="12">CAMEROON</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Buea,
|
||
<quantity id="C65E9B16FFD0FFEAC0A2BBDCFB48FD5B" box="[1177,1267,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="m" value="1000.0">1000 m</quantity>
|
||
,
|
||
<date id="75181033FFD0FFEAC139BBDCFA23FD5B" box="[1282,1432,727,753]" pageId="11" pageNumber="12" value="1912-03-02" valueMax="1912-03-07" valueMin="1912-03-02">2.–7.III.1912</date>
|
||
, leg. V. Rothkirch (2,
|
||
<collectionCode id="67B7AE36FFD0FFEAC5A8BBF0FE52FCBF" box="[403,489,763,789]" pageId="11" pageNumber="12">ZMHB</collectionCode>
|
||
).
|
||
<emphasis id="33D2EAE1FFD0FFEAC5C4BBF1FC84FCBE" bold="true" box="[511,831,762,788]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="79B17663FFD0FFEAC5C4BBF1FC82FCBE" box="[511,825,762,788]" name="Equatorial Guinea" pageId="11" pageNumber="12">EQUATORIAL GUINEA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Fernando Po [= Bioko], leg. Báguera (1,
|
||
<collectionCode id="67B7AE36FFD0FFEAC109BBF0FA2BFCBF" box="[1330,1424,763,789]" pageId="11" pageNumber="12">MCNM</collectionCode>
|
||
); Nkolentungan,
|
||
<date id="75181033FFD0FFEAC57FBA14FE1BFC93" box="[324,416,799,825]" pageId="11" pageNumber="12" value="1907-11">XI.1907</date>
|
||
–
|
||
<date id="75181033FFD0FFEAC594BA14FDBAFC93" box="[431,513,799,825]" pageId="11" pageNumber="12" value="1908-05">V.1908</date>
|
||
, leg. Tessmann (1,
|
||
<collectionCode id="67B7AE36FFD0FFEAC6E3BA14FC95FC93" box="[728,814,799,825]" pageId="11" pageNumber="12">ZMHB</collectionCode>
|
||
).
|
||
<emphasis id="33D2EAE1FFD0FFEAC77ABA15FC0AFC92" bold="true" box="[833,945,798,824]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="79B17663FFD0FFEAC77ABA15FC17FC92" box="[833,940,798,824]" name="Gabon" pageId="11" pageNumber="12">GABON</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Ogooué Lambarèné, 1912, leg. Ellenberger (1,
|
||
<collectionCode id="67B7AE36FFD0FFEAC486BA48FEA3FCF7" box="[189,280,835,861]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="12">MNHN</collectionCode>
|
||
).
|
||
<emphasis id="33D2EAE1FFD0FFEAC512BA49FCD6FCF6" bold="true" box="[297,877,834,861]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="79B17663FFD0FFEAC512BA49FCD3FCF6" box="[297,872,834,861]" name="Democratic Republic of the Congo" pageId="11" pageNumber="12">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
P.N.A., secteur nord riv. Musinini affl. Byangolo,
|
||
<quantity id="C65E9B16FFD0FFEAC4ACBA6DFF49FC2B" box="[151,242,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="11" pageNumber="12" unit="m" value="1225.0">1225 m</quantity>
|
||
,
|
||
<date id="75181033FFD0FFEAC53ABA6CFE3BFC2B" box="[257,384,870,897]" pageId="11" pageNumber="12" value="1958-04-14">14.IV.1958</date>
|
||
, leg. Vanschuytbroeck (1,
|
||
<collectionCode id="67B7AE36FFD0FFEAC68DBA6CFCB5FC2B" box="[694,782,871,897]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="11" pageNumber="12" type="Museum">MRAC</collectionCode>
|
||
).
|
||
<emphasis id="33D2EAE1FFD0FFEAC718BA6DFC1CFC2A" bold="true" box="[803,935,870,896]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="79B17663FFD0FFEAC718BA6DFC19FC2A" box="[803,930,870,896]" name="Uganda" pageId="11" pageNumber="12">UGANDA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
NW Beni, W of Ruwenzori, forest,
|
||
<date id="75181033FFD0FFEAC16FBA6CFA27FC2B" box="[1364,1436,871,897]" pageId="11" pageNumber="12" value="1908-01">I.1908</date>
|
||
(3,
|
||
<collectionCode id="67B7AE36FFD0FFEAC4FBBA80FEADFC0F" box="[192,278,907,933]" pageId="11" pageNumber="12">ZMHB</collectionCode>
|
||
); Bugoma forest, Unyoro,
|
||
<quantity id="C65E9B16FFD0FFEAC66BBA80FD16FC0F" box="[592,685,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.1277599999999999" pageId="11" pageNumber="12" unit="ft" value="3700.0">3700 ft.</quantity>
|
||
,
|
||
<date id="75181033FFD0FFEAC686BA81FCE8FC0F" box="[701,851,906,933]" pageId="11" pageNumber="12" value="1911-12-01" valueMax="1911-12-05" valueMin="1911-12-01">1–5.XII.1911</date>
|
||
(1,
|
||
<collectionCode id="67B7AE36FFD0FFEAC7BCBA80FC5BFC0F" box="[903,992,907,933]" pageId="11" pageNumber="12">BMNH</collectionCode>
|
||
);
|
||
<collectingRegion id="C362F811FFD0FFEAC7CCBA80FBE4FC0F" box="[1015,1119,907,933]" country="Uganda" name="Kampala" pageId="11" pageNumber="12">Kampala</collectingRegion>
|
||
, Nambadzidza,
|
||
<date id="75181033FFD0FFEAC122BA80FA2CFC0F" box="[1305,1431,907,933]" pageId="11" pageNumber="12" value="1931-04-19">19.IV.1931</date>
|
||
, leg. Hopkins (2,
|
||
<collectionCode id="67B7AE36FFD0FFEAC56CBAA4FE0BFC63" box="[343,432,943,969]" pageId="11" pageNumber="12">BMNH</collectionCode>
|
||
); Kawanda,
|
||
<date id="75181033FFD0FFEAC605BAA4FD05FC63" box="[574,702,943,969]" pageId="11" pageNumber="12" value="1942-03-10">10.III.1942</date>
|
||
, leg. Taylor (2,
|
||
<collectionCode id="67B7AE36FFD0FFEAC74EBAA4FC75FC63" box="[885,974,943,969]" pageId="11" pageNumber="12">BMNH</collectionCode>
|
||
); Mboga, SW of Albert-See,
|
||
<date id="75181033FFD0FFEAC121BAA4FACDFC63" box="[1306,1398,943,969]" pageId="11" pageNumber="12" value="1908-03">III.1908</date>
|
||
(1,
|
||
<collectionCode id="67B7AE36FFD0FFEAC4ACBAD8FF56FC47" box="[151,237,979,1005]" pageId="11" pageNumber="12">ZMHB</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |