201 lines
18 KiB
XML
201 lines
18 KiB
XML
<document id="C498401D2BE0AB7542597ADE082C4866" ID-CLB-Dataset="29914" ID-DOI="http://dx.doi.org/10.3897/zookeys.766.24523" ID-GBIF-Dataset="3232638c-2666-4380-b236-0af5bb1f76b4" ID-PMC="PMC6013545" ID-Pensoft-Pub="1313-2970-766-79" ID-PubMed="29942172" ID-ZooBank="87FD3EE451E046A1BA5E03FA451140E7" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-766-79" ModsDocOrigin="ZooKeys 766" ModsDocTitle="Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru" checkinTime="1529036602696" checkinUser="pensoft" docAuthor="Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M. & Torres-Carvajal, Omar" docDate="2018" docId="88FB43CB53DB280639129EFCDF2D17D3" docLanguage="en" docName="ZooKeys 766: 79-147" docOrigin="ZooKeys 766" docSource="http://dx.doi.org/10.3897/zookeys.766.24523" docTitle="Dipsas peruana Boettger 1898" docType="treatment" docVersion="6" lastPageNumber="101" masterDocId="4170FF9E0C19FF8DBD79FFD3FFBCFFCE" masterDocTitle="Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru" masterLastPageNumber="147" masterPageNumber="79" pageNumber="100" updateTime="1701376524545" updateUser="ExternalLinkService">
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<mods:title id="F545D3B96C31608BEE97F2063343AC3F">Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru</mods:title>
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<mods:namePart id="CC5C6FB585E8139C15A7E903741889C3">Arteaga, Alejandro</mods:namePart>
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<mods:namePart id="BEE79148619C4B7A711CE9E254C6BE20">Salazar-Valenzuela, David</mods:namePart>
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<mods:namePart id="0A592AAAF327977DB3FCFE6E5B7D5685">Mebert, Konrad</mods:namePart>
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<mods:namePart id="95769F1D3DC0F6771953FF75A9DBD8DA">Penafiel, Nicolas</mods:namePart>
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<mods:namePart id="4784CF3F0B82C506BC87DBB26A937DBF">Aguiar, Gabriela</mods:namePart>
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<mods:namePart id="E07A124F8DEAE0F482AB9BB3B80FEF81">Sa ́ nchez-Nivicela, Juan C.</mods:namePart>
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<mods:namePart id="D2399C77675A818D14D0A2C77C0D0E15">Pyron, R. Alexander</mods:namePart>
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<mods:namePart id="8867312336D872710E8893F96B0DFAC7">Yanez-Munoz, Mario H.</mods:namePart>
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<mods:namePart id="1FC2F7F8DAFA794BAC67E768689AA0A2">Venegas, Pablo J.</mods:namePart>
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<mods:namePart id="AD8AC6B8A2496A52BEB151AE3CB6A131">Guayasamin, Juan M.</mods:namePart>
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<mods:namePart id="D81B3B2B9432ED4BC72701CB7D7749E2">Torres-Carvajal, Omar</mods:namePart>
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<mods:typeOfResource id="7ED20C6266059C027D2BF959BA388119">text</mods:typeOfResource>
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<mods:title id="0414880CE797B9F4BFC4F3E3E9BB059E">ZooKeys</mods:title>
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<mods:date id="44E785F2724F75C4C6881C1332508F84">2018</mods:date>
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<mods:detail id="E0FE9171821342369A4FFF9F4A3BCF4F" type="volume">
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<mods:number id="358B4351544D185D9D0F1A2C10C948CD">766</mods:number>
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<mods:start id="942B4D44EBEAFA3260DC36BF549A3129">79</mods:start>
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<mods:end id="08435BDE206777405B98DB1F287632A1">147</mods:end>
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<mods:url id="FB182FF94E32320221526747A25D89F9">http://dx.doi.org/10.3897/zookeys.766.24523</mods:url>
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</mods:location>
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<mods:classification id="6B9CC1FF1B964DCA3FE07666CB4FA999">journal article</mods:classification>
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<mods:identifier id="1CAC360B20879FB320DD930748A5C315" type="DOI">http://dx.doi.org/10.3897/zookeys.766.24523</mods:identifier>
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<mods:identifier id="24BC1DD1A5D9717541883AEA9B27114A" type="Pensoft-Pub">1313-2970-766-79</mods:identifier>
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<mods:identifier id="FA69BB285A52DB576D703612CF2A688A" type="ZooBank">87FD3EE451E046A1BA5E03FA451140E7</mods:identifier>
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</mods:mods>
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<treatment id="88FB43CB53DB280639129EFCDF2D17D3" ID-GBIF-Taxon="145939715" LSID="urn:lsid:plazi:treatment:88FB43CB53DB280639129EFCDF2D17D3" httpUri="http://treatment.plazi.org/id/88FB43CB53DB280639129EFCDF2D17D3" lastPageId="22" lastPageNumber="101" pageId="21" pageNumber="100">
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<subSubSection id="E8ACB5B5C31DFFC22E7AD22E1DD93CE9" pageId="21" pageNumber="100" type="nomenclature">
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<paragraph id="14B9A648517D31FD37FD0A3334CAB216" pageId="21" pageNumber="100">
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<taxonomicName id="5D60524A5AE338F50C21831DFB4DC678" authority="Boettger, 1898" authorityName="Boettger" authorityYear="1898" class="Squamata" family="Dipsadidae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas peruana" order="Squamata" pageId="21" pageNumber="100" phylum="Chordata" rank="species" species="peruana">Dipsas peruana (Boettger, 1898)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="EA1F9ABB61FE121282EECD0D91569B1A" pageId="21" pageNumber="100" type="reference_group">
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<paragraph id="5A8B0628A2E6F3E7606C65742FA97832" pageId="21" pageNumber="100">
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<taxonomicName id="B624FDDB5D97D35EB6A572AF0037FB1B" class="Squamata" family="Dipsadidae" genus="Leptognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptognathus peruana" order="Squamata" pageId="21" pageNumber="100" phylum="Chordata" rank="species" species="peruana">Leptognathus peruana</taxonomicName>
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Boettger, 1898: 128. Holotype SMF 20801, a female from Santa Ana, department of Cuzco, Peru.
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</paragraph>
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<paragraph id="C02F63C4FB461BBDB7B864695CECDFF4" pageId="21" pageNumber="100">
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<taxonomicName id="85F7D7BC500DFDD52582F2B46672783E" class="Squamata" family="Dipsadidae" genus="Leptognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptognathus boettgeri" order="Squamata" pageId="21" pageNumber="100" phylum="Chordata" rank="species" species="boettgeri">Leptognathus boettgeri</taxonomicName>
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Werner, 1901: 11. Holotype MTKD D 1671 M, a female from Chanchamayo, department of
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<normalizedToken id="1E3D8E8B4F47BA872F13434D79DEA0CA" originalValue="Junín">Junin</normalizedToken>
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, Peru.
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</paragraph>
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<paragraph id="56BD8A5E56BCEBA991D200FC15348F90" pageId="21" pageNumber="100">
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<taxonomicName id="BA37BB837629C8BB39FD31734ADFED4E" class="Squamata" family="Dipsadidae" genus="Leptognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptognathus boliviana" order="Squamata" pageId="21" pageNumber="100" phylum="Chordata" rank="species" species="boliviana">Leptognathus boliviana</taxonomicName>
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Werner, 1909: 240. Holotype ZMH, a female from department of Beni, Bolivia.
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</paragraph>
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<paragraph id="BB50098838BE995D3A2990EEE6C0FBAD" pageId="21" pageNumber="100">
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<taxonomicName id="0856AAB78EB4EDD6345D7BF459B39B78" class="Squamata" family="Dipsadidae" genus="Leptognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptognathus polylepis" order="Squamata" pageId="21" pageNumber="100" phylum="Chordata" rank="species" species="polylepis">Leptognathus polylepis</taxonomicName>
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Boulenger, 1912: 422. Holotype BMNH 1946.1.2078, a female from Huancabamba, department of Pasco, Peru.
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</paragraph>
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</subSubSection>
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<subSubSection id="BBFB4DFC43247DB4FA2CAF6B13B960E2" pageId="21" pageNumber="100" type="proposed standard english name">
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<paragraph id="89D11593CAE664A2D7FE9839B9176869" pageId="21" pageNumber="100">Proposed standard English name.</paragraph>
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<paragraph id="D8C10B8446755CFBDB4AAEF2F3AA71B7" pageId="21" pageNumber="100">Peruvian Snail-Eater</paragraph>
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</subSubSection>
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<subSubSection id="B5AFF38A64925311E86E5B44B5066953" pageId="21" pageNumber="100" type="proposed standard spanish name">
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<paragraph id="7C0EFFD55E6216EF858F234F826ACDF7" pageId="21" pageNumber="100">Proposed standard Spanish name.</paragraph>
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<paragraph id="7071494786A67497678DBB80896DF88F" pageId="21" pageNumber="100">Caracolera Peruana</paragraph>
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</subSubSection>
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<subSubSection id="78110930601D220E544E04757C52AE83" lastPageId="22" lastPageNumber="101" pageId="21" pageNumber="100" type="diagnosis">
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<paragraph id="EE2DFD8E4C077C217B787FDC66FB1746" pageId="21" pageNumber="100">Diagnosis.</paragraph>
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<paragraph id="D852A0D521CB4AF7A686055DED866E24" lastPageId="22" lastPageNumber="101" pageId="21" pageNumber="100">
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<taxonomicName id="FA871E3840C80763E149C084A58B0957" class="Squamata" family="Colubridae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas peruana" pageId="21" pageNumber="100" phylum="Chordata" rank="species" species="peruana">Dipsas peruana</taxonomicName>
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differs from all described species of
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<taxonomicName id="0B4B6AA07BD0E9A12A0926895468134F" class="Insecta" family="Lycaenidae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas" order="Lepidoptera" pageId="21" pageNumber="100" phylum="Arthropoda" rank="genus">Dipsas</taxonomicName>
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based on the following combination of characters: (1) 15/15/15 smooth dorsals with moderately enlarged vertebral row; (2) one loreal and one preocular in contact with orbit; (3) 8-9 supralabials with 4-6 or 3-5 contacting orbit; (4) one pair of infralabials in contact behind symphysial; (5) 177-200 ventrals in males, 180-203 in females; (6) 75-127 divided subcaudals in males, 79-105 in females; (7) dorsal and ventral ground color brown to dark brown (light brown in juveniles) with 33-43 blackish brown to complete black, white to cream edged circular to vertically elliptical blotches that are longer than interspaces; head dark brown with dingy cream reticulations and different degrees of whitish edging on the labial scales, and a thin (1-3 scales long) white to light grayish brown irregular nuchal collar; dorsal blotches extending marginally onto ventrals and
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<pageBreakToken id="70908F40FAFBEE56ACA1AB0B4F64A167" pageId="22" pageNumber="101" start="start">rarely</pageBreakToken>
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fusing midventrally; (8) 199 mm SVL in males, 610-725 mm in females; (9) 85 mm TL in males, 155-241 mm in females.
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</paragraph>
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</subSubSection>
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<subSubSection id="C1B4AA400D8F08A0BE07AC90DC005699" pageId="22" pageNumber="101" type="comparisons">
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<paragraph id="91756D009A416EE50C11B630AB4F0052" pageId="22" pageNumber="101">Comparisons.</paragraph>
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<paragraph id="AA8E9AE0EBAB34CE5FA41B9C1248FD05" pageId="22" pageNumber="101">
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<taxonomicName id="8BCA4EC621DEEE105EF3D46AFE5E5FBF" class="Squamata" family="Colubridae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas peruana" pageId="22" pageNumber="101" phylum="Chordata" rank="species" species="peruana">Dipsas peruana</taxonomicName>
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sensu stricto is compared to species previously subsumed under
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<taxonomicName id="A2164BA4C2FC3224A8DCCAD3FF248409" lsidName="D. peruana" pageId="22" pageNumber="101" rank="species" species="peruana">D. peruana</taxonomicName>
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sensu lato:
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<taxonomicName id="ECB9FED849FAC77290205C07A4286F70" lsidName="D. latifrontalis" pageId="22" pageNumber="101" rank="species" species="latifrontalis">D. latifrontalis</taxonomicName>
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,
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<taxonomicName id="A4D1F99C8BA9C0E4601E60B87F1BCD46" lsidName="D. palmeri" pageId="22" pageNumber="101" rank="species" species="palmeri">D. palmeri</taxonomicName>
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, and
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<taxonomicName id="3D0EABFCFD0EC4E125AB06681B319E78" lsidName="D. klebbai" pageId="22" pageNumber="101" rank="species" species="klebbai">D. klebbai</taxonomicName>
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. From
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<taxonomicName id="5C9E364B4A5AA086B04C9626E9D11075" lsidName="D. latifrontalis" pageId="22" pageNumber="101" rank="species" species="latifrontalis">D. latifrontalis</taxonomicName>
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and
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<taxonomicName id="F751E3811B62AE1495389CAD0942E7BF" lsidName="D. palmeri" pageId="22" pageNumber="101" rank="species" species="palmeri">D. palmeri</taxonomicName>
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, it differs in having dorsal blotches along the entire body similar in length or longer than interspaces (shorter than interspaces in
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<taxonomicName id="CE080F7900A30176DF1596F04064D115" lsidName="D. latifrontalis" pageId="22" pageNumber="101" rank="species" species="latifrontalis">D. latifrontalis</taxonomicName>
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and
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<taxonomicName id="E2647C14C3C08DF3558085FDCA672C51" lsidName="D. palmeri" pageId="22" pageNumber="101" rank="species" species="palmeri">D. palmeri</taxonomicName>
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), and in having melanized interspaces in some adult individuals. With the exception of BMNH 1946.1.2078, specimens of
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<taxonomicName id="A8BA5844968EB7FF875A0D50CD8923D7" lsidName="D. peruana" pageId="22" pageNumber="101" rank="species" species="peruana">D. peruana</taxonomicName>
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can be separated from specimens of
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<taxonomicName id="03497BF4DB9C063002D0FE8297D723B8" lsidName="D. klebbai" pageId="22" pageNumber="101" rank="species" species="klebbai">D. klebbai</taxonomicName>
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by possessing at least one of the following characteristics: posterior body blotches similar in length or marginally longer than interspaces (twice to four times as long in
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<taxonomicName id="51BC9BDA8343E45C6636755CFCF20372" lsidName="D. klebbai" pageId="22" pageNumber="101" rank="species" species="klebbai">D. klebbai</taxonomicName>
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); short circular to vertically elliptical body blotches usually only up to 4-7 vertebral scales long; melanized interspaces; dorsal surface of the head not completely black; and dorsal body blotches rarely fused ventrally.
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</paragraph>
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</subSubSection>
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<subSubSection id="1FE4782A3839ABD63495E7586FD97551" pageId="22" pageNumber="101" type="distribution">
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<paragraph id="1C8D7763A24226AA09ACF3FFA83A3BCC" pageId="22" pageNumber="101">Distribution.</paragraph>
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<paragraph id="2D40255BDCBBE727510DDD191BA57AF9" pageId="22" pageNumber="101">Eastern slopes of the Peruvian and Bolivian Andes south of the Huancabamba depression at elevations between 1279 and 2671 m (Fig. 4).</paragraph>
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<caption id="3EC1492DD6A963D29908C978B3B38E7C" pageId="22" pageNumber="101">
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<paragraph id="4B8F716C8E5C838C41B29310A41ABB37" pageId="22" pageNumber="101">
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Figure 13. Adult female holotype of
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<taxonomicName id="A5962D34293EAC2F83F0DAA782F3E0AD" class="Squamata" family="Colubridae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas oswaldobaezi" pageId="22" pageNumber="101" phylum="Chordata" rank="species" species="oswaldobaezi">Dipsas oswaldobaezi</taxonomicName>
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QCAZ 10369 in a dorsal and b ventral view. Scale bar: 1 cm.
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</paragraph>
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</caption>
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<caption id="3A4FFDB6FA2D0AA8EB66E9A05ABD4BDA" pageId="22" pageNumber="101">
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<paragraph id="226B97A924128AC6CC7107D2A9A8AE3D" pageId="22" pageNumber="101">
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Figure 14. Adult female paratype of
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<taxonomicName id="86A32C30A8BD9049629090E03DD02071" class="Squamata" family="Colubridae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas oswaldobaezi" pageId="22" pageNumber="101" phylum="Chordata" rank="species" species="oswaldobaezi">Dipsas oswaldobaezi</taxonomicName>
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. QCAZ 15108.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |