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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.894.37862" ID-GBIF-Dataset="b9f6ab2b-1860-4796-b756-167ae5d2c916" ID-PMC="PMC6906170" ID-Pensoft-Pub="1313-2970-894-53" ID-Pensoft-UUID="BAF8B2CC491254A3AC7E08368A2697B5" ID-PubMed="31844409" ID-ZooBank="D11503CA5A574067817904E0C8C162C8" ModsDocID="1313-2970-894-53" checkinTime="1575565036169" checkinUser="pensoft" docAuthor="Pentinsaari, Mikko, Anderson, Robert, Borowiec, Lech, Bouchard, Patrice, Brunke, Adam, Douglas, Hume, Smith, Andrew B. T. &amp; Hebert, Paul D. N." docDate="2019" docId="8885187350DB52769C3883AE92383DC8" docLanguage="en" docName="ZooKeys 894: 53-150" docOrigin="ZooKeys 894" docSource="http://dx.doi.org/10.3897/zookeys.894.37862" docTitle="Tachyporus atriceps Stephens 1832" docType="treatment" docVersion="4" id="BAF8B2CC491254A3AC7E08368A2697B5" lastPageNumber="53" masterDocId="BAF8B2CC491254A3AC7E08368A2697B5" masterDocTitle="DNA barcodes reveal 63 overlooked species of Canadian beetles (Insecta, Coleoptera)" masterLastPageNumber="150" masterPageNumber="53" pageNumber="53" updateTime="1668168181706" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>DNA barcodes reveal 63 overlooked species of Canadian beetles (Insecta, Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pentinsaari, Mikko</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Anderson, Robert</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Borowiec, Lech</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bouchard, Patrice</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Brunke, Adam</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Douglas, Hume</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Andrew B. T.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hebert, Paul D. N.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>894</mods:number>
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<mods:start>53</mods:start>
<mods:end>150</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.894.37862</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.894.37862</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-894-53</mods:identifier>
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<treatment ID-GBIF-Taxon="160631249" LSID="urn:lsid:plazi:treatment:8885187350DB52769C3883AE92383DC8" httpUri="http://treatment.plazi.org/id/8885187350DB52769C3883AE92383DC8" lastPageNumber="53" pageId="0" pageNumber="53">
<subSubSection pageId="0" pageNumber="53" type="nomenclature">
<paragraph pageId="0" pageNumber="53">
<taxonomicName LSID="88851873-50DB-5276-9C38-83AE92383DC8" authority="Stephens, 1832" authorityName="Stephens" authorityYear="1832" class="Insecta" family="Carabidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus atriceps" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="atriceps">Tachyporus atriceps Stephens, 1832</taxonomicName>
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 Tachyporus atriceps Stephens 5 A habitus 5 B elytral chaetotaxy (re-drawn from Assing and Schuelke 2012) and pattern variability 5 C aedeagus, ventral view, M. Schuelke, modified. Abbreviations: OD outer discal row, MD middle discal row, ID inner discal row, S sutural row 6 Amischa decipiens (Sharp) 6 A habitus 6 B spermatheca, re-drawn from Muona (1990). Scale bars: 1.0 mm (5 A; 6 A), 0.5 mm (5 B, C), 0.1 mm (6 B)." figureDoi="10.3897/zookeys.894.37862.figures5-6" httpUri="https://binary.pensoft.net/fig/362140" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures5-6">Figure 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="distribution">
<paragraph pageId="0" pageNumber="53">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="53">
Native to the Palaearctic region, where it is widespread (
<bibRefCitation author="Schuelke, M" editor="Assing, V" journalOrPublisher="PhD Thesis, Michigan State University, Department of Entomology" pageId="0" pageNumber="53" refId="B186" refString="Schuelke, M, Assing, V, Schuelke, M, 2012b. Tachyporinae. In: AssingVSchuelkeM (Eds) Die Kaefer Mitteleuropas, Band 4 (2.Auflage). Spektrum Akademischer Verlag, Heidelberg, 130-199." title="Tachyporinae. In: AssingVSchuelkeM (Eds) Die Kaefer Mitteleuropas, Band 4 (2. Auflage). Spektrum Akademischer Verlag, Heidelberg, 130 - 199." year="2012 b">
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
2012b
</bibRefCitation>
). Adventive in the Nearctic region (British Columbia, Ontario, Quebec, Nova Scotia, New Brunswick, and Prince Edward Island, Canada).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="canadian records">
<paragraph pageId="0" pageNumber="53">Canadian records</paragraph>
<paragraph pageId="0" pageNumber="53">(DNA barcoded specimens). British Columbia: Burnaby, 21-Sep-2015 to 02-Oct-2015 (1 ex, CBG). Ontario: Ancaster, 21-Sep-2015 to 02-Oct-2015 (1 ex, CBG); Ausable-Bayfield Conservation Authority, 30-Jun-2015 (1 ex, CBG); Cambridge, 22-Sep-2014 to 03-Oct-2014 (2 exx, CBG); Cambridge, 24-Apr-2015 to 01-May-2015 (49 exx, CBG); Carillon Park, 06-May-1982 (2 ex, CNC); Courtice, 19-Sep-2016 to 30-Sep-2016 (1 ex, CBG); Guelph, 22-Sep-2014 to 03-Oct-2014 (1 ex, CBG); Guelph, 23-Sep-2013 to 04-Oct-2013 (2 exx, CBG); Guelph, 26-Sep-2014 to 29-Sep-2014 (1 ex, CBG); Hartington, 04-Oct-2017 (1 ex, CBG); Mississauga, 24-May-2016 to 26-May-2016 (1 ex, CBG); Point Pelee National Park, 06-Jul-2015 (1 ex, CBG); Puslinch Township, 19-Sep-2010 to 27-Sep-2010 (2 exx, CBG); Puslinch Township, 24-Oct-2010 to 31-Oct-2010 (1 ex, CBG); Red Rock, 21-Sep-2015 to 02-Oct-2015 (1 ex, CBG); Rondeau Provincial Park, 09-Jul-2015 (1 ex, CBG); Rouge National Urban Park, 03-Jun-2013 to 09-Jun-2013 (13 exx, CBG); Rouge National Urban Park, 15-Sep-2013 (1 ex, CBG); Stayner, 21-Sep-2015 to 02-Oct-2015 (1 ex, CBG). Quebec: Forillon National Park, 16-Sep-2013 to 23-Sep-2013 (2 exx, CBG). New Brunswick: Florenceville-Bristol, 22-Sep-2014 to 03-Oct-2014 (3 exx, CBG); Fredericton, 19-Sep-2016 to 30-Sep-2016 (1 ex, CBG). Nova Scotia: Cape Breton Highlands National Park, 13-Sep-2013 to 20-Sep-2013 (1 ex, CBG); Hubbards, 19-Sep-2016 to 30-Sep-2016 (1 ex, CBG). Prince Edward Island: Prince Edward Island National Park, 11-Sep-2013 to 18-Sep-2013 (1 ex, CBG).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="additional canadian records">
<paragraph pageId="0" pageNumber="53">Additional Canadian records.</paragraph>
<paragraph pageId="0" pageNumber="53">Ontario: Carillon Park, 06-May-1982 (2 exx, CNC); Renfrew County, 4 km SE Cobden, 15-Sep-1980 (1 ex, CNC); Wentworth County, Stoney Creek, 03-Mar-1973 (1 ex, CNC). Quebec: Montreal, 07-Sep-1984 (1 ex, CNC). Nova Scotia: Halifax, 1988 (1 ex, CNC).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="diagnostic information">
<paragraph pageId="0" pageNumber="53">Diagnostic information.</paragraph>
<paragraph pageId="0" pageNumber="53">
Body length 2.4-3.6 mm. Habitus as in
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 Tachyporus atriceps Stephens 5 A habitus 5 B elytral chaetotaxy (re-drawn from Assing and Schuelke 2012) and pattern variability 5 C aedeagus, ventral view, M. Schuelke, modified. Abbreviations: OD outer discal row, MD middle discal row, ID inner discal row, S sutural row 6 Amischa decipiens (Sharp) 6 A habitus 6 B spermatheca, re-drawn from Muona (1990). Scale bars: 1.0 mm (5 A; 6 A), 0.5 mm (5 B, C), 0.1 mm (6 B)." figureDoi="10.3897/zookeys.894.37862.figures5-6" httpUri="https://binary.pensoft.net/fig/362140" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures5-6">Fig. 5A</figureCitation>
. Head black, strongly contrasting with pronotum, elytra with either medial and ovoid lateral markings across disc or disc nearly entirely darkened (
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 Tachyporus atriceps Stephens 5 A habitus 5 B elytral chaetotaxy (re-drawn from Assing and Schuelke 2012) and pattern variability 5 C aedeagus, ventral view, M. Schuelke, modified. Abbreviations: OD outer discal row, MD middle discal row, ID inner discal row, S sutural row 6 Amischa decipiens (Sharp) 6 A habitus 6 B spermatheca, re-drawn from Muona (1990). Scale bars: 1.0 mm (5 A; 6 A), 0.5 mm (5 B, C), 0.1 mm (6 B)." figureDoi="10.3897/zookeys.894.37862.figures5-6" httpUri="https://binary.pensoft.net/fig/362140" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures5-6">Fig. 5B</figureCitation>
). Elytral chaetotaxy as in
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 Tachyporus atriceps Stephens 5 A habitus 5 B elytral chaetotaxy (re-drawn from Assing and Schuelke 2012) and pattern variability 5 C aedeagus, ventral view, M. Schuelke, modified. Abbreviations: OD outer discal row, MD middle discal row, ID inner discal row, S sutural row 6 Amischa decipiens (Sharp) 6 A habitus 6 B spermatheca, re-drawn from Muona (1990). Scale bars: 1.0 mm (5 A; 6 A), 0.5 mm (5 B, C), 0.1 mm (6 B)." figureDoi="10.3897/zookeys.894.37862.figures5-6" httpUri="https://binary.pensoft.net/fig/362140" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures5-6">Fig. 5B</figureCitation>
, with sutural row of punctures and three rows of discal punctures, with two punctures in inner discal row. Aedeagus as in
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 Tachyporus atriceps Stephens 5 A habitus 5 B elytral chaetotaxy (re-drawn from Assing and Schuelke 2012) and pattern variability 5 C aedeagus, ventral view, M. Schuelke, modified. Abbreviations: OD outer discal row, MD middle discal row, ID inner discal row, S sutural row 6 Amischa decipiens (Sharp) 6 A habitus 6 B spermatheca, re-drawn from Muona (1990). Scale bars: 1.0 mm (5 A; 6 A), 0.5 mm (5 B, C), 0.1 mm (6 B)." figureDoi="10.3897/zookeys.894.37862.figures5-6" httpUri="https://binary.pensoft.net/fig/362140" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures5-6">Fig. 5C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="bionomic notes">
<paragraph pageId="0" pageNumber="53">Bionomic notes.</paragraph>
<paragraph pageId="0" pageNumber="53">
This species occurs in a variety of moist to very dry microhabitats (
<bibRefCitation author="Schuelke, M" editor="Assing, V" journalOrPublisher="PhD Thesis, Michigan State University, Department of Entomology" pageId="0" pageNumber="53" refId="B186" refString="Schuelke, M, Assing, V, Schuelke, M, 2012b. Tachyporinae. In: AssingVSchuelkeM (Eds) Die Kaefer Mitteleuropas, Band 4 (2.Auflage). Spektrum Akademischer Verlag, Heidelberg, 130-199." title="Tachyporinae. In: AssingVSchuelkeM (Eds) Die Kaefer Mitteleuropas, Band 4 (2. Auflage). Spektrum Akademischer Verlag, Heidelberg, 130 - 199." year="2012 b">
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
2012b
</bibRefCitation>
). In the Nearctic, this species occurs in disturbed habitats. It was collected by
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.19" author="Brunke, AJ" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="53" pagination="652 - 670" refId="B32" refString="Brunke, AJ, Bahlai, CA, Klimaszewski, J, Hallett, RH, 2014. Rove beetles (Coleoptera: Staphylinidae) in Ontario, Canada soybean agroecosystems: assemblage diversity, composition, seasonality, and habitat use. The Canadian Entomologist 146: 652 - 670" title="Rove beetles (Coleoptera: Staphylinidae) in Ontario, Canada soybean agroecosystems: assemblage diversity, composition, seasonality, and habitat use." url="https://doi.org/10.4039/tce.2014.19" volume="146" year="2014">Brunke et al. (2014)</bibRefCitation>
in soybean fields and adjacent hedgerows in Ontario (misidentified as
<taxonomicName lsidName="T. canadensis" pageId="0" pageNumber="53" rank="species" species="canadensis">
<emphasis italics="true" pageId="0" pageNumber="53">T. canadensis</emphasis>
</taxonomicName>
Campbell, 1979, to which it is similar). The barcoded specimens were collected in suburban residential areas and protected land adjacent to cities, mainly with Malaise or pitfall traps. Some of the barcoded specimens are larvae extracted from soil or leaf litter.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="comments">
<paragraph pageId="0" pageNumber="53">Comments.</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus atriceps" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis italics="true" pageId="0" pageNumber="53">Tachyporus atriceps</emphasis>
</taxonomicName>
has the same elytral chaetotaxy as
<taxonomicName lsidName="T. borealis" pageId="0" pageNumber="53" rank="species" species="borealis">
<emphasis italics="true" pageId="0" pageNumber="53">T. borealis</emphasis>
</taxonomicName>
Campbell, 1979,
<taxonomicName lsidName="T. nimbicola" pageId="0" pageNumber="53" rank="species" species="nimbicola">
<emphasis italics="true" pageId="0" pageNumber="53">T. nimbicola</emphasis>
</taxonomicName>
Campbell, 1979, and
<taxonomicName lsidName="T. canadensis" pageId="0" pageNumber="53" rank="species" species="canadensis">
<emphasis italics="true" pageId="0" pageNumber="53">T. canadensis</emphasis>
</taxonomicName>
Campbell, 1979 but can be separated from the first two by the elytra with discal markings.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus canadensis" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis italics="true" pageId="0" pageNumber="53">Tachyporus canadensis</emphasis>
</taxonomicName>
has a dark red-brown head, bright yellow pronotum, and either a pair of narrow linear lateral dark markings (and medial darkening) or entirely immaculate elytra, while
<taxonomicName lsidName="T. atriceps" pageId="0" pageNumber="53" rank="species" species="atriceps">
<emphasis italics="true" pageId="0" pageNumber="53">T. atriceps</emphasis>
</taxonomicName>
has a deep black head, slightly darkened (dingy yellow-orange) pronotum and lateral elytral markings that are ovoid or entirely fused with the medial marking to form a broad darkened area over much of the elytra. The internal sac sclerite of
<taxonomicName lsidName="T. atriceps" pageId="0" pageNumber="53" rank="species" species="atriceps">
<emphasis italics="true" pageId="0" pageNumber="53">T. atriceps</emphasis>
</taxonomicName>
is similarly shaped to
<taxonomicName lsidName="T. nimbicola" pageId="0" pageNumber="53" rank="species" species="nimbicola">
<emphasis italics="true" pageId="0" pageNumber="53">T. nimbicola</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. borealis" pageId="0" pageNumber="53" rank="species" species="borealis">
<emphasis italics="true" pageId="0" pageNumber="53">T. borealis</emphasis>
</taxonomicName>
(cane-shaped,
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 Tachyporus atriceps Stephens 5 A habitus 5 B elytral chaetotaxy (re-drawn from Assing and Schuelke 2012) and pattern variability 5 C aedeagus, ventral view, M. Schuelke, modified. Abbreviations: OD outer discal row, MD middle discal row, ID inner discal row, S sutural row 6 Amischa decipiens (Sharp) 6 A habitus 6 B spermatheca, re-drawn from Muona (1990). Scale bars: 1.0 mm (5 A; 6 A), 0.5 mm (5 B, C), 0.1 mm (6 B)." figureDoi="10.3897/zookeys.894.37862.figures5-6" httpUri="https://binary.pensoft.net/fig/362140" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures5-6">Fig. 5C</figureCitation>
) while
<taxonomicName lsidName="T. canadensis" pageId="0" pageNumber="53" rank="species" species="canadensis">
<emphasis italics="true" pageId="0" pageNumber="53">T. canadensis</emphasis>
</taxonomicName>
possesses a characteristic arc-shaped sclerite that is not hooked.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus atriceps" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis italics="true" pageId="0" pageNumber="53">Tachyporus atriceps</emphasis>
</taxonomicName>
appears to be common in at least southern Canada and has been present in the Nearctic since at least 1973. This species was probably recently introduced, or has only recently become widespread and common, as it was not detected by
<bibRefCitation DOI="https://doi.org/10.4039/entm111109fv" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="53" pagination="1 - 95" refId="B34" refString="Campbell, JM, 1979. A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America. Memoirs of the Entomological Society of Canada 111: 1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." url="https://doi.org/10.4039/entm111109fv" volume="111" year="1979">Campbell (1979)</bibRefCitation>
.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus atriceps" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis italics="true" pageId="0" pageNumber="53">Tachyporus atriceps</emphasis>
</taxonomicName>
is split into two closely clustered BINs which show no differences in morphology, including male genitalia.
</paragraph>
<caption doi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" start="Figures 7, 8" startId="F5">
<paragraph pageId="0" pageNumber="53">
<emphasis bold="true" pageId="0" pageNumber="53">Figures 7, 8.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="53">7A</emphasis>
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta vaga" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="vaga">
<emphasis italics="true" pageId="0" pageNumber="53">Atheta vaga</emphasis>
</taxonomicName>
(Heer), habitus
<emphasis bold="true" pageId="0" pageNumber="53">7B</emphasis>
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta vaga" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="vaga">
<emphasis italics="true" pageId="0" pageNumber="53">Atheta vaga</emphasis>
</taxonomicName>
, aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="53">7C</emphasis>
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta fanatica" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="fanatica">
<emphasis italics="true" pageId="0" pageNumber="53">Atheta fanatica</emphasis>
</taxonomicName>
Casey, aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="53">7D</emphasis>
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta fanatica" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="fanatica">
<emphasis italics="true" pageId="0" pageNumber="53">Atheta fanatica</emphasis>
</taxonomicName>
, spermatheca
<emphasis bold="true" pageId="0" pageNumber="53">8</emphasis>
<taxonomicName class="Insecta" family="Staphylinidae" genus="Myllaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myllaena infuscata" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis italics="true" pageId="0" pageNumber="53">Myllaena infuscata</emphasis>
</taxonomicName>
Kraatz
<emphasis bold="true" pageId="0" pageNumber="53">8A</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="53">8B</emphasis>
aedeagus, ventral view
<emphasis bold="true" pageId="0" pageNumber="53">8C</emphasis>
aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="53">8D</emphasis>
spermatheca
<emphasis bold="true" pageId="0" pageNumber="53">
8
<normalizedToken originalValue="BD">B-D</normalizedToken>
</emphasis>
V. Assing. Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="53">7A</emphasis>
), 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="53">
7
<normalizedToken originalValue="BD">B-D</normalizedToken>
</emphasis>
), 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="53">8A</emphasis>
), 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="53">
8
<normalizedToken originalValue="BD">B-D</normalizedToken>
</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>