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<document id="81289B00E376984D5B4ABA390999E9A9" ID-CLB-Dataset="25961" ID-DOI="10.11646/zootaxa.4658.1.1" ID-GBIF-Dataset="31d6e081-95c3-4086-a45a-c11555dda3aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373040" ID-ZooBank="ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566364910789" checkinUser="plazi" docAuthor="Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed &amp; Barbero, Enrico" docDate="2019" docId="8861532B88471836FF760AAEFC3CC038" docLanguage="en" docName="zootaxa.4658.1.1.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus (Onthophagiellus) millingeni d'Orbigny 1898" docType="treatment" docVersion="12" lastPageNumber="21" masterDocId="74582B5388531822FFE10957FFA3C74B" masterDocTitle="An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="21" updateTime="1698751869822" updateUser="ExternalLinkService">
<mods:mods id="7C03564896F8EBD2319A36DBEFC610C2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A35B076ACA0C07E8E341A14DE6070637">
<mods:title id="9656E9733367E4039A32F295E7AB67E2">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
</mods:titleInfo>
<mods:name id="CD07C30F364376C4B76BE764B2E10547" type="personal">
<mods:role id="1AF50E50573C303A63A24FD05B93FFFA">
<mods:roleTerm id="C4E1B5176766E660D35C214465783F09">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B372E0FA7E85E0FD4EE937176057EC53">Ziani, Stefano</mods:namePart>
</mods:name>
<mods:name id="5C2506032E6859859345CC36E520B891" type="personal">
<mods:role id="0B0C27D6EC81F5E00619D496597DB905">
<mods:roleTerm id="314EF90D6A28FF3A8453C1610F135021">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D67DCDE7736583C4810E7102A91B89C">Abdel-Dayem, Mahmoud Saleh</mods:namePart>
</mods:name>
<mods:name id="4B99036915DE80527FD5B0E33E3C78BC" type="personal">
<mods:role id="AE1CD12399F243D8B6490966AAAB39DA">
<mods:roleTerm id="E5E3825508F9136316F3E3FBA7C03E59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2623839E73C75E6DF1B79F9A4F34F69E">Aldhafer, Hathal Mohammed</mods:namePart>
</mods:name>
<mods:name id="E5D948E4AD75B8209BFA0F30213203BF" type="personal">
<mods:role id="5408843C96F5FF11F3C3505D42D34CF4">
<mods:roleTerm id="BC747E93319C3B15563B3D782582AB04">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0BF9FC39CE51668AD05234145D95133F">Barbero, Enrico</mods:namePart>
</mods:name>
<mods:typeOfResource id="ACDA879AFA4CE2C6AD801D6C2D9D19AD">text</mods:typeOfResource>
<mods:relatedItem id="E4A634B23D93E9794DE02273B3461892" type="host">
<mods:titleInfo id="B681F0115A99E7385A133979B1C04974">
<mods:title id="0DF7348781A64D7C00B7C78CAD04A229">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9FAF92C2D5FD14D7E65AE419688256B6">
<mods:date id="7E93CE76B5E1DA1FD632696CE2FA76FD">2019</mods:date>
<mods:detail id="DD5074D771E63BC60F07E10F988F3D7A" type="pubDate">
<mods:number id="D540B8868339DEC03D1787362B5C4A8F">2019-08-21</mods:number>
</mods:detail>
<mods:detail id="551B0F37DBC64C8CCEDD14A49219362B" type="volume">
<mods:number id="8B0F85A99E69AE525806BAC7939097C3">4658</mods:number>
</mods:detail>
<mods:detail id="370BFEB96893432D2303C1EEA4FCE2E8" type="issue">
<mods:number id="53669AAF5A095205E6F885DB388B3CEF">1</mods:number>
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<mods:start id="C6D57771319A73CBD4D9F3B433646A00">1</mods:start>
<mods:end id="50D4FC6DDB737AAFD70A569D6A597E13">36</mods:end>
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<mods:classification id="B5C5ED48728A41CF962E384D0F1BB1D1">journal article</mods:classification>
<mods:identifier id="CBEBFC1BB4CCCE35007114BD97FC564E" type="CLB-Dataset">25961</mods:identifier>
<mods:identifier id="BFF91BC668294BC4C5D6B6057C924F9A" type="DOI">10.11646/zootaxa.4658.1.1</mods:identifier>
<mods:identifier id="DDE4739758C637C635F26FAAFA815557" type="GBIF-Dataset">31d6e081-95c3-4086-a45a-c11555dda3aa</mods:identifier>
<mods:identifier id="0DDE4CB3AF919B1CACD712D939BDF18B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="46C7A89AF33370B54C1DADF116D44253" type="Zenodo-Dep">3373040</mods:identifier>
<mods:identifier id="48BD084A98838723A24B461C61039988" type="ZooBank">ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21</mods:identifier>
</mods:mods>
<treatment id="8861532B88471836FF760AAEFC3CC038" ID-DOI="http://doi.org/10.5281/zenodo.5667084" ID-GBIF-Taxon="159217240" ID-Zenodo-Dep="5667084" LSID="urn:lsid:plazi:treatment:8861532B88471836FF760AAEFC3CC038" httpUri="http://treatment.plazi.org/id/8861532B88471836FF760AAEFC3CC038" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="48D2B1B688471836FF760AAEFCD4C358" box="[151,887,1017,1044]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="0077E23D88471836FF760AAEFCD4C358" blockId="20.[151,887,1017,1079]" box="[151,887,1017,1044]" pageId="20" pageNumber="21">
<heading id="5B3F555188471836FF760AAEFCD4C358" bold="true" box="[151,887,1017,1044]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="C7C899BE88471836FF760AAEFCD4C358" ID-CoL="49MM2" authority="d'Orbigny, 1898" authorityName="d'Orbigny" authorityYear="1898" box="[151,887,1017,1044]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="millingeni" subGenus="Onthophagiellus">
<emphasis id="32BC3E2F88471836FF760AAEFCD4C358" bold="true" box="[151,887,1017,1044]" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF760AAEFE9FC35F" bold="true" box="[151,316,1017,1044]" italics="true" pageId="20" pageNumber="21">Onthophagus</emphasis>
(
<emphasis id="32BC3E2F88471836FEAC0AAEFDB4C35F" bold="true" box="[333,535,1017,1044]" italics="true" pageId="20" pageNumber="21">Onthophagiellus</emphasis>
)
<emphasis id="32BC3E2F88471836FDC90AAEFD07C35F" bold="true" box="[552,676,1017,1044]" italics="true" pageId="20" pageNumber="21">millingeni</emphasis>
d’Orbigny, 1898
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688471836FF760D4AFF59C37C" box="[151,250,1052,1079]" pageId="20" pageNumber="21" type="description">
<paragraph id="0077E23D88471836FF760D4AFF59C37C" blockId="20.[151,887,1017,1079]" box="[151,250,1052,1079]" pageId="20" pageNumber="21">
[
<figureCitation id="98F3FEB888471836FF7F0D4AFF51C37C" box="[158,242,1052,1079]" captionStart="FIGURES 14–18" captionStartId="23.[151,264,1872,1897]" captionTargetBox="[293,1306,179,1808]" captionTargetId="figure@23.[243,1344,166,1852]" captionTargetPageId="23" captionText="FIGURES 14–18. Onthophagini of Arabian Peninsula. 14, Onthophagus (Micronthophagus) ochreatus d’Orbigny, 1898: YE (Al Hudaydah) 10 km W Al Mansuriyah; 15, Onthophagus (Onthophagiellus) millingeni d’Orbigny, 1898: YE (Sana’a) Haddah; 16, Onthophagus (s. str.) falsus Gillet, 1925: SA (Riyadh) Wadi Al Dawaser; 17, Onthophagus (Proagoderus) quadriarmatus Fairmaire, 1892, ♂: YE (Al Mahrah) Wadi N of Damqut vill.; 18, Onthophagus (Proagoderus) quadriarmatus Fairmaire, 1892, ♀: YE (Al Mahrah) Wadi N of Damqut vill. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373051/files/figure.png" pageId="20" pageNumber="21">Fig. 15</figureCitation>
]
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688471836FF760D33FC3CC038" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="0077E23D88471836FF760D33FF4FC3E8" blockId="20.[151,1437,1124,1907]" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF760D33FECCC334" bold="true" box="[151,367,1124,1151]" pageId="20" pageNumber="21">Literature quotes</emphasis>
.
<collectingCountry id="78DFA2AD88471836FE650D33FD86C337" box="[388,549,1124,1151]" name="Saudi Arabia" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FE650D33FE65C337" bold="true" box="[388,454,1124,1151]" pageId="20" pageNumber="21">SAUDI</emphasis>
<taxonomicName id="C7C899BE88471836FE330D33FD86C337" authority=": Hejaz" authorityName="Hejaz" box="[466,549,1124,1151]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="32BC3E2F88471836FE330D33FD86C337" bold="true" box="[466,549,1124,1151]" pageId="20" pageNumber="21">ARABIA</emphasis>
</taxonomicName>
</collectingCountry>
: Hejaz, “Hedjaz” (d’Orbigny 1898a).
<collectingCountry id="78DFA2AD88471836FC160D32FBE0C337" box="[1015,1091,1125,1151]" name="Yemen" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FC160D32FBE0C337" bold="true" box="[1015,1091,1125,1151]" pageId="20" pageNumber="21">
<subScript id="9C4CE07888471836FC160D32FBE0C337" attach="left" box="[1015,1091,1125,1151]" fontSize="7" pageId="20" pageNumber="21">YEMEN</subScript>
</emphasis>
</collectingCountry>
: without locality (d’Orbigny 1898a).
</paragraph>
<paragraph id="0077E23D88471836FF260DFBFDEAC164" blockId="20.[151,1437,1124,1907]" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF260DFBFE13C38C" bold="true" box="[199,432,1196,1223]" pageId="20" pageNumber="21">Material examined</emphasis>
.
<materialsCitation id="B0A0E86088471836FE250DFBFF5DC244" ID-GBIF-Occurrence="2350485907" collectingDate="2015-07-31" collectionCode="KSMA" collectorName="H. Al Dhafer &amp; H. Fadl &amp; M. Abdel-Dayem &amp; A. El Gharbawy &amp; A. El Torkey" country="Saudi Arabia" elevation="1614" location="Raydah Nature Reserve" municipality="Abha" pageId="20" pageNumber="21" specimenCount="1" stateProvince="'Asir">
<collectingCountry id="78DFA2AD88471836FE250DFBFDC6C38F" box="[452,613,1196,1223]" name="Saudi Arabia" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FE250DFBFDA5C38F" bold="true" box="[452,518,1196,1223]" pageId="20" pageNumber="21">SAUDI</emphasis>
<taxonomicName id="C7C899BE88471836FDF30DFBFDC6C38F" box="[530,613,1196,1223]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="32BC3E2F88471836FDF30DFBFDC6C38F" bold="true" box="[530,613,1196,1223]" pageId="20" pageNumber="21">ARABIA</emphasis>
</taxonomicName>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88471836FD990DFAFCBFC38C" box="[632,796,1197,1223]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="21">Asir Province</collectingRegion>
,
<collectingMunicipality id="E013784788471836FCCD0DFAFCCFC38C" box="[812,876,1197,1223]" pageId="20" pageNumber="21">Abha</collectingMunicipality>
,
<location id="0517B4E688471836FC9C0DFAFB34C38C" LSID="urn:lsid:plazi:treatment:8861532B88471836FF760AAEFC3CC038:0517B4E688471836FC9C0DFAFB34C38C" box="[893,1175,1197,1223]" country="Saudi Arabia" municipality="Abha" name="Raydah Nature Reserve" pageId="20" pageNumber="21" stateProvince="'Asir">Raydah Nature Reserve</location>
,
<quantity id="C7304FD888471836FB490DFAFAA0C38C" box="[1192,1283,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="1.614" pageId="20" pageNumber="21" unit="m" value="1614.0">
<elevation id="8BE5050E88471836FB490DFAFAA0C38C" box="[1192,1283,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="1.614" pageId="20" pageNumber="21" unit="m" value="1614.0">1614 m</elevation>
</quantity>
(1811.749′N 4223.345′E),
<date id="7476C4FD88471836FED00D87FE10C3A0" box="[305,435,1232,1259]" pageId="20" pageNumber="21" value="2015-07-31">
<collectingDate id="64323D1588471836FED00D87FE10C3A0" box="[305,435,1232,1259]" pageId="20" pageNumber="21" value="2015-07-31">31.vii.2015</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FE200D86FDFAC3A0" box="[449,601,1233,1259]" pageId="20" pageNumber="21">H. Al Dhafer</collectorName>
,
<collectorName id="AD3D87EB88471836FD870D86FD63C3A0" box="[614,704,1233,1259]" pageId="20" pageNumber="21">H. Fadl</collectorName>
,
<collectorName id="AD3D87EB88471836FD2C0D86FC34C3A0" box="[717,919,1233,1259]" pageId="20" pageNumber="21">M. Abdel-Dayem</collectorName>
,
<collectorName id="AD3D87EB88471836FC430D86FBFDC3A0" box="[930,1118,1233,1259]" pageId="20" pageNumber="21">A. El Gharbawy</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FB650D86FAB9C3A0" box="[1156,1306,1233,1259]" pageId="20" pageNumber="21">A. El Torkey</collectorName>
leg. 2 exx. (
<collectionCode id="66D97AF888471836FF7E0DA2FF55C244" box="[159,246,1269,1295]" pageId="20" pageNumber="21">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FEE80DA2FE84C278" ID-GBIF-Occurrence="2350485975" collectingDate="2015-11-17" collectionCode="KSMA" collectorName="H. Al Dhafer &amp; H. Fadl &amp; M. Abdel-Dayem &amp; A. El Gharbawy &amp; A. El Torkey" country="Saudi Arabia" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Asir Province">
<emphasis id="32BC3E2F88471836FEE80DA2FEF6C244" box="[265,341,1269,1295]" italics="true" pageId="20" pageNumber="21">ibidem</emphasis>
,
<date id="7476C4FD88471836FE850DA3FE7DC244" box="[356,478,1268,1295]" pageId="20" pageNumber="21" value="2015-11-17">
<collectingDate id="64323D1588471836FE850DA3FE7DC244" box="[356,478,1268,1295]" pageId="20" pageNumber="21" value="2015-11-17">17.xi.2015</collectingDate>
</date>
, VC,
<collectorName id="AD3D87EB88471836FDFF0DA2FD12C244" box="[542,689,1269,1295]" pageId="20" pageNumber="21">H. Al Dhafer</collectorName>
,
<collectorName id="AD3D87EB88471836FD5D0DA2FCB0C244" box="[700,787,1269,1295]" pageId="20" pageNumber="21">H. Fadl</collectorName>
,
<collectorName id="AD3D87EB88471836FCFF0DA2FC46C244" box="[798,997,1269,1295]" pageId="20" pageNumber="21">M. Abdel-Dayem</collectorName>
,
<collectorName id="AD3D87EB88471836FC0F0DA2FB06C244" box="[1006,1189,1269,1295]" pageId="20" pageNumber="21">A. El Gharbawy</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FB260DA2FAF4C244" box="[1223,1367,1269,1295]" pageId="20" pageNumber="21">A. El Torkey</collectorName>
leg.
<specimenCount id="16CE29B488471836FA6F0DA2FF1AC278" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FF290C4EFEBCC278" box="[200,287,1305,1331]" pageId="20" pageNumber="21">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FED20C4EFD8EC21C" ID-GBIF-Occurrence="2350485910" collectionCode="KSMA" collectorName="H. Al Dhafer &amp; H. Fadl &amp; M. Abdel-Dayem &amp; A. El Torkey &amp; A. El Gharbawy" country="Saudi Arabia" elevation="1851" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Asir Province">
<emphasis id="32BC3E2F88471836FED20C4EFEDCC278" box="[307,383,1305,1331]" italics="true" pageId="20" pageNumber="21">ibidem</emphasis>
,
<quantity id="C7304FD888471836FE6F0C4FFE47C278" box="[398,484,1304,1331]" metricMagnitude="3" metricUnit="m" metricValue="1.8509999999999998" pageId="20" pageNumber="21" unit="m" value="1851.0">
<elevation id="8BE5050E88471836FE6F0C4FFE47C278" box="[398,484,1304,1331]" metricMagnitude="3" metricUnit="m" metricValue="1.8509999999999998" pageId="20" pageNumber="21" unit="m" value="1851.0">1851 m</elevation>
</quantity>
(1811.679′N 4223.691′E),
<collectorName id="AD3D87EB88471836FCF60C4EFC0FC278" box="[791,940,1305,1331]" pageId="20" pageNumber="21">H. Al Dhafer</collectorName>
,
<collectorName id="AD3D87EB88471836FC560C4EFBACC278" box="[951,1039,1305,1331]" pageId="20" pageNumber="21">H. Fadl</collectorName>
,
<collectorName id="AD3D87EB88471836FBFA0C4EFB40C278" box="[1051,1251,1305,1331]" pageId="20" pageNumber="21">M. Abdel-Dayem</collectorName>
,
<collectorName id="AD3D87EB88471836FB0C0C4EFADCC278" box="[1261,1407,1305,1331]" pageId="20" pageNumber="21">A. El Torkey</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FF760C6AFEECC21C" box="[151,335,1341,1367]" pageId="20" pageNumber="21">A. El Gharbawy</collectorName>
leg.
<specimenCount id="16CE29B488471836FE660C6AFE1CC21C" box="[391,447,1341,1367]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FE2F0C6AFD86C21C" box="[462,549,1341,1367]" pageId="20" pageNumber="21">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FDD80C6AFC1FC230" ID-GBIF-Occurrence="2350485880" collectingDate="2014-04-26" collectionCode="KSMA" collectorName="H. Al Dhafer &amp; H. Fadl &amp; M. Abdel-Dayem &amp; A. El Torkey &amp; A. El Gharbawy" country="Saudi Arabia" elevation="1897" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Asir Province">
<emphasis id="32BC3E2F88471836FDD80C6AFD26C21C" box="[569,645,1341,1367]" italics="true" pageId="20" pageNumber="21">ibidem</emphasis>
,
<quantity id="C7304FD888471836FD750C6AFD4AC21C" box="[660,745,1341,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.8969999999999998" pageId="20" pageNumber="21" unit="m" value="1897.0">
<elevation id="8BE5050E88471836FD750C6AFD4AC21C" box="[660,745,1341,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.8969999999999998" pageId="20" pageNumber="21" unit="m" value="1897.0">1897 m</elevation>
</quantity>
(1811.695′N 4223.818′E),
<date id="7476C4FD88471836FBFD0C6AFB37C21C" box="[1052,1172,1341,1367]" pageId="20" pageNumber="21" value="2014-04-26">
<collectingDate id="64323D1588471836FBFD0C6AFB37C21C" box="[1052,1172,1341,1367]" pageId="20" pageNumber="21" value="2014-04-26">26.iv.2014</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FB400C6AFA96C21C" box="[1185,1333,1341,1367]" pageId="20" pageNumber="21">H. Al Dhafer</collectorName>
,
<collectorName id="AD3D87EB88471836FAA00C6AFA3BC21C" box="[1345,1432,1341,1367]" pageId="20" pageNumber="21">H. Fadl</collectorName>
,
<collectorName id="AD3D87EB88471836FF760C36FEC3C230" box="[151,352,1377,1403]" pageId="20" pageNumber="21">M. Abdel-Dayem</collectorName>
,
<collectorName id="AD3D87EB88471836FE8B0C36FE5EC230" box="[362,509,1377,1403]" pageId="20" pageNumber="21">A. El Torkey</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FDC30C36FD78C230" box="[546,731,1377,1403]" pageId="20" pageNumber="21">A. El Gharbawy</collectorName>
leg.
<specimenCount id="16CE29B488471836FCF40C36FCEEC230" box="[789,845,1377,1403]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FCBC0C36FC17C230" box="[861,948,1377,1403]" pageId="20" pageNumber="21">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FC290C36FEC3C2D4" ID-GBIF-Occurrence="2350485853" collectingDate="2015-05-09" collectionCode="KSMA" collectorName="Al Dhafer" country="Saudi Arabia" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Asir Province">
<emphasis id="32BC3E2F88471836FC290C36FBB7C230" box="[968,1044,1377,1403]" italics="true" pageId="20" pageNumber="21">ibidem</emphasis>
,
<date id="7476C4FD88471836FBC50C37FB25C230" box="[1060,1158,1376,1403]" pageId="20" pageNumber="21" value="2015-05-09">
<collectingDate id="64323D1588471836FBC50C37FB25C230" box="[1060,1158,1376,1403]" pageId="20" pageNumber="21" value="2015-05-09">9.v.2015</collectingDate>
</date>
, PT. H.
<collectorName id="AD3D87EB88471836FB020C36FAF4C230" box="[1251,1367,1377,1403]" pageId="20" pageNumber="21">Al Dhafer</collectorName>
,
<emphasis id="32BC3E2F88471836FA820C35FA3EC230" box="[1379,1437,1377,1403]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
leg.
<specimenCount id="16CE29B488471836FF260CD2FF51C2D4" box="[199,242,1413,1439]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
(
<collectionCode id="66D97AF888471836FEE00CD2FEFBC2D4" box="[257,344,1413,1439]" pageId="20" pageNumber="21">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FE8A0CD2FE9FC288" ID-GBIF-Occurrence="2350485909" collectingDate="2015-07-31" collectionCode="KSMA" collectorName="Al Dhafer, H &amp; Fadl &amp; Abdel-Dayem &amp; El Gharbawy &amp; A. El Torkey" country="Saudi Arabia" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Asir Province">
<emphasis id="32BC3E2F88471836FE8A0CD2FE14C2D4" box="[363,439,1413,1439]" italics="true" pageId="20" pageNumber="21">ibidem</emphasis>
,
<date id="7476C4FD88471836FE240CD3FDE4C2D4" box="[453,583,1412,1439]" pageId="20" pageNumber="21" value="2015-07-31">
<collectingDate id="64323D1588471836FE240CD3FDE4C2D4" box="[453,583,1412,1439]" pageId="20" pageNumber="21" value="2015-07-31">31.vii.2015</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FDB00CD2FD45C2D4" box="[593,742,1413,1439]" pageId="20" pageNumber="21">Al Dhafer, H</collectorName>
.
<collectorName id="AD3D87EB88471836FD130CD2FC84C2D4" box="[754,807,1413,1439]" pageId="20" pageNumber="21">Fadl</collectorName>
, M.
<collectorName id="AD3D87EB88471836FCB60CD2FC5AC2D4" box="[855,1017,1413,1439]" pageId="20" pageNumber="21">Abdel-Dayem</collectorName>
, A.
<collectorName id="AD3D87EB88471836FBC50CD2FB1AC2D4" box="[1060,1209,1413,1439]" pageId="20" pageNumber="21">El Gharbawy</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FB3A0CD2FACFC2D4" box="[1243,1388,1413,1439]" pageId="20" pageNumber="21">A. El Torkey</collectorName>
leg.
<specimenCount id="16CE29B488471836FF760CFEFF6DC288" box="[151,206,1449,1475]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FF3C0CFEFE97C288" box="[221,308,1449,1475]" pageId="20" pageNumber="21">KSMA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FEA60CFEFE86C2AC" ID-GBIF-Occurrence="2350486023" collectionCode="NMPC" collectorName="V. Hula &amp; Niedobov" country="Yemen" elevation="2500" location="Bait Bows" pageId="20" pageNumber="21" specimenCount="1" stateProvince="San'a'">
<collectingCountry id="78DFA2AD88471836FEA60CFEFE30C28B" box="[327,403,1449,1475]" name="Yemen" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FEA60CFEFE30C28B" bold="true" box="[327,403,1449,1475]" pageId="20" pageNumber="21">
<subScript id="9C4CE07888471836FEA60CFEFE30C28B" attach="left" box="[327,403,1449,1475]" fontSize="7" pageId="20" pageNumber="21">YEMEN</subScript>
</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88471836FE400CFEFD27C288" box="[417,644,1449,1475]" country="Yemen" name="San'a'" pageId="20" pageNumber="21">Sana’a Governorate</collectingRegion>
,
<location id="0517B4E688471836FD6E0CFEFCA6C288" LSID="urn:lsid:plazi:treatment:8861532B88471836FF760AAEFC3CC038:0517B4E688471836FD6E0CFEFCA6C288" box="[655,773,1449,1475]" country="Yemen" name="Bait Bows" pageId="20" pageNumber="21" stateProvince="San'a'">Bait Bows</location>
dam,
<quantity id="C7304FD888471836FCAB0CFFFC3CC288" box="[842,927,1448,1475]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="m" value="2500.0">
<elevation id="8BE5050E88471836FCAB0CFFFC3CC288" box="[842,927,1448,1475]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="m" value="2500.0">2500 m</elevation>
</quantity>
(1516′N 4412′E),
<collectorName id="AD3D87EB88471836FB8E0CFEFB67C288" box="[1135,1220,1449,1475]" pageId="20" pageNumber="21">V. Hula</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FB090CFEFAF5C288" box="[1256,1366,1449,1475]" pageId="20" pageNumber="21">Niedobov</collectorName>
leg.
<specimenCount id="16CE29B488471836FA6F0CFEFF1AC2AC" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FF260C9AFEBEC2AC" box="[199,285,1485,1511]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FED10C9AFB7BC2AC" ID-GBIF-Occurrence="2350485917" box="[304,1240,1484,1511]" collectingDate="2003-11-20" collectionCode="NMPC" collectorName="D. Kral" country="Yemen" elevation="2413" location="Haddah" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Sana'a Governorate">
<location id="0517B4E688471836FED10C9AFE29C2AC" LSID="urn:lsid:plazi:treatment:8861532B88471836FF760AAEFC3CC038:0517B4E688471836FED10C9AFE29C2AC" box="[304,394,1485,1511]" country="Yemen" name="Haddah" pageId="20" pageNumber="21" stateProvince="Sana'a Governorate">Haddah</location>
,
<quantity id="C7304FD888471836FE750C9AFE4AC2AC" box="[404,489,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.413" pageId="20" pageNumber="21" unit="m" value="2413.0">
<elevation id="8BE5050E88471836FE750C9AFE4AC2AC" box="[404,489,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.413" pageId="20" pageNumber="21" unit="m" value="2413.0">2413 m</elevation>
</quantity>
(1517′24″N 4409′59″), 11/
<date id="7476C4FD88471836FCF80C9AFC30C2AC" box="[793,915,1485,1511]" pageId="20" pageNumber="21" value="2003-11-20">
<collectingDate id="64323D1588471836FCF80C9AFC30C2AC" box="[793,915,1485,1511]" pageId="20" pageNumber="21" value="2003-11-20">20.xi.2003</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FC7F0C9AFC51C2AC" box="[926,1010,1485,1511]" pageId="20" pageNumber="21">D. Král</collectorName>
leg. 2 exx. (
<collectionCode id="66D97AF888471836FB9A0C9AFB72C2AC" box="[1147,1233,1485,1511]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FB020C9AFAA1C140" ID-GBIF-Occurrence="2350485852" collectingDate="2010-11-03" collectionCode="NMPC" collectorName="J. Hajek &amp; Bezdek" country="Yemen" elevation="2510" location="Jabal Haraz mts." pageId="20" pageNumber="21" specimenCount="1" stateProvince="Sana'a Governorate">
<location id="0517B4E688471836FB020C9AFA3EC2AC" LSID="urn:lsid:plazi:treatment:8861532B88471836FF760AAEFC3CC038:0517B4E688471836FB020C9AFA3EC2AC" box="[1251,1437,1485,1511]" country="Yemen" name="Jabal Haraz mts." pageId="20" pageNumber="21" stateProvince="Sana'a Governorate">Jabal Haraz mts.</location>
<locationDeviation id="11C5FB1F88471836FF760CA6FEE6C140" box="[151,325,1521,1547]" location="Manakhah" pageId="20" pageNumber="21">S of Manakhah</locationDeviation>
vill.,
<quantity id="C7304FD888471836FE680CA7FE43C140" box="[393,480,1520,1547]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="20" pageNumber="21" unit="m" value="2510.0">
<elevation id="8BE5050E88471836FE680CA7FE43C140" box="[393,480,1520,1547]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="20" pageNumber="21" unit="m" value="2510.0">2510 m</elevation>
</quantity>
(1504′N 4345′E),
<date id="7476C4FD88471836FD550CA6FC83C140" box="[692,800,1521,1547]" pageId="20" pageNumber="21" value="2010-11-03">
<collectingDate id="64323D1588471836FD550CA6FC83C140" box="[692,800,1521,1547]" pageId="20" pageNumber="21" value="2010-11-03">3.xi.2010</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FCCC0CA6FC28C140" box="[813,907,1521,1547]" pageId="20" pageNumber="21">J. Hájek</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FC500CA6FBA6C140" box="[945,1029,1521,1547]" pageId="20" pageNumber="21">Bezdĕk</collectorName>
leg. 15 exx. (
<collectionCode id="66D97AF888471836FB450CA6FB59C140" box="[1188,1274,1521,1547]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088471836FAEF0CA6FDE6C164" ID-GBIF-Occurrence="2350485862" collectingDate="2009-06-10" collectionCode="NMPC" collectorName="V. Hula" country="Yemen" location="Wadi Dhahr" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Sana'a Governorate">
<location id="0517B4E688471836FAEF0CA6FA3BC140" LSID="urn:lsid:plazi:treatment:8861532B88471836FF760AAEFC3CC038:0517B4E688471836FAEF0CA6FA3BC140" box="[1294,1432,1521,1547]" country="Yemen" name="Wadi Dhahr" pageId="20" pageNumber="21" stateProvince="Sana'a Governorate">Wadi Dhahr</location>
,
<date id="7476C4FD88471836FF760F42FEB2C164" box="[151,273,1557,1583]" pageId="20" pageNumber="21" value="2009-06-10">
<collectingDate id="64323D1588471836FF760F42FEB2C164" box="[151,273,1557,1583]" pageId="20" pageNumber="21" value="2009-06-10">10.vi.2009</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FEFD0F42FED2C164" box="[284,369,1557,1583]" pageId="20" pageNumber="21">V. Hula</collectorName>
leg
<specimenCount id="16CE29B488471836FE430F42FE7BC164" box="[418,472,1557,1583]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FE060F42FD9EC164" box="[487,573,1557,1583]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0077E23D88471836FF260F6FFF43C133" blockId="20.[151,1437,1124,1907]" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF260F6FFEFAC118" bold="true" box="[199,345,1592,1619]" pageId="20" pageNumber="21">Distribution</emphasis>
.
<emphasis id="32BC3E2F88471836FE8D0F6FFDF9C118" bold="true" box="[364,602,1592,1619]" pageId="20" pageNumber="21">Arabian Peninsula:</emphasis>
<collectingCountry id="78DFA2AD88471836FD870F6EFD5CC118" box="[614,767,1593,1619]" name="Saudi Arabia" pageId="20" pageNumber="21">
Saudi
<taxonomicName id="C7C899BE88471836FD510F6EFD5CC118" box="[688,767,1593,1619]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="35" phylum="Arthropoda" rank="genus">Arabia</taxonomicName>
</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FCEF0F6EFCFFC118" box="[782,860,1593,1619]" name="Yemen" pageId="20" pageNumber="21">Yemen</collectingCountry>
(d’Orbigny 1898a).
<emphasis id="32BC3E2F88471836FBB20F6FFB32C118" bold="true" box="[1107,1169,1592,1619]" pageId="20" pageNumber="21">Asia:</emphasis>
<collectingCountry id="78DFA2AD88471836FB7C0F6EFB69C118" box="[1181,1226,1593,1619]" name="Iraq" pageId="20" pageNumber="21">Iraq</collectingCountry>
(
<bibRefCitation id="64599FCC88471836FB3F0F6EFF77C133" author="Ziani, S. &amp; Bezdek, A." pageId="20" pageNumber="21" pagination="180 - 204" refId="ref39999" refString="Ziani, S. &amp; Bezdek, A. (2016) Scarabaeidae, subfamily Scarabaeinae, tribe Onthophagini. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill, Leiden-Boston, pp. 180 - 204." type="book chapter" year="2016">Ziani &amp; Bezděk 2016</bibRefCitation>
).
</paragraph>
<paragraph id="0077E23D88471836FF260FD6FC3CC038" blockId="20.[151,1437,1124,1907]" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF260FD6FE97C1D0" bold="true" box="[199,308,1665,1691]" pageId="20" pageNumber="21">Remarks</emphasis>
. According to
<bibRefCitation id="64599FCC88471836FE030FD6FD03C1D0" author="Palestrini, C." box="[482,672,1665,1691]" pageId="20" pageNumber="21" pagination="173 - 175" refId="ref39143" refString="Palestrini, C. (1982) Description du male de l' Onthophagus missor Balthasar et remarques sur la position systematique de l'espece (Coleoptera, Scarabaeidae). Revue Francaise d'Entomologie (New Series), 4 (4), 173 - 175." type="journal article" year="1982">Palestrini (1982)</bibRefCitation>
<taxonomicName id="C7C899BE88471836FD4B0FD6FCE2C1D0" authorityName="d'Orbigny" authorityYear="1898" box="[682,833,1665,1691]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="millingeni">
<emphasis id="32BC3E2F88471836FD4B0FD6FCE2C1D0" box="[682,833,1665,1691]" italics="true" pageId="20" pageNumber="21">O. millingeni</emphasis>
</taxonomicName>
belongs to the subgenus
<taxonomicName id="C7C899BE88471836FB890FD6FF71C1F4" authority="Balthasar, 1935" authorityName="Balthasar" authorityYear="1935" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subGenus" subGenus="Onthophagiellus">
<emphasis id="32BC3E2F88471836FB890FD6FA80C1D0" box="[1128,1315,1665,1691]" italics="true" pageId="20" pageNumber="21">Onthophagiellus</emphasis>
Balthasar, 1935
</taxonomicName>
. As noted by
<bibRefCitation id="64599FCC88471836FE8A0FF2FDB3C18B" author="Zunino, M." box="[363,528,1701,1728]" pageId="20" pageNumber="21" pagination="11 - 16" refId="ref40183" refString="Zunino, M. (1976) Onthophagus tessulatus Har. e O. millingeni d'Orb. (Coleoptera, Scarabaeoidea). Annali del Museo Civico di Storia Naturale di Genova, 81, 11 - 16." type="journal article" year="1976">Zunino (1976)</bibRefCitation>
,
<bibRefCitation id="64599FCC88471836FDFD0FF2FCF6C1F4" author="Boucomont, A. &amp; Gillet, J. J. E." box="[540,853,1701,1727]" pageId="20" pageNumber="21" pagination="103 - 263" refId="ref36942" refString="Boucomont, A. &amp; Gillet, J. J. E. (1927) Fam. Scarabaeidae. Subfam. Coprinae II, pp. 103 - 263. In: Gillet, J. J. E. &amp; Boucomont, A. (Eds.), Coleopterorum Catalogus, Vol. 19 (3). Scarabaeidae: Coprinae, Termitotroginae. Partes 38 et 90, W. Junk, Berlin, pp. 103 - 263." type="book chapter" year="1927">Boucomont &amp; Gillet (1927)</bibRefCitation>
misunderstood d’Orbigny (1913) and wrongly considered
<taxonomicName id="C7C899BE88471836FF100F9EFDA1C1A8" authorityName="d'Orbigny" authorityYear="1898" box="[241,514,1737,1763]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="millingeni">
<emphasis id="32BC3E2F88471836FF100F9EFDA1C1A8" box="[241,514,1737,1763]" italics="true" pageId="20" pageNumber="21">Onthophagus millingeni</emphasis>
</taxonomicName>
a junior synonym of
<taxonomicName id="C7C899BE88471836FD1A0F9EFB8DC1A8" authority="Harold, 1871" authorityName="Harold" authorityYear="1871" box="[763,1070,1737,1763]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tessulatus">
<emphasis id="32BC3E2F88471836FD1A0F9EFC2EC1A8" box="[763,909,1737,1763]" italics="true" pageId="20" pageNumber="21">O. tessulatus</emphasis>
<bibRefCitation id="64599FCC88471836FC770F9EFB8DC1A8" author="Harold, E." box="[918,1070,1737,1763]" pageId="20" pageNumber="21" pagination="1 - 28" refId="ref37937" refString="Harold, E. (1871) Herausgegeben unter Mitwirkung mehrerer Fachgenossen. I. Verzeichniss der von Dr. Beccari in Bogos gesammelten coprophagen Lamellicornien Vom Herausgeber. Coleopterologische Hefte, 8, 1 - 28. https: // doi. org / 10.1002 / mmnd. 4791871022 9" type="journal article" year="1871">Harold, 1871</bibRefCitation>
</taxonomicName>
, a species distributed in
<collectingCountry id="78DFA2AD88471836FAA80F9EFA3BC1A8" box="[1353,1432,1737,1763]" name="Eritrea" pageId="20" pageNumber="21">Eritrea</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FF760FBAFF55C04C" box="[151,246,1773,1799]" name="Ethiopia" pageId="20" pageNumber="21">Ethiopia</collectingCountry>
and
<collectingCountry id="78DFA2AD88471836FECE0FBAFE2DC04C" box="[303,398,1773,1799]" name="Somalia" pageId="20" pageNumber="21">Somalia</collectingCountry>
. Subsequent authors such as
<bibRefCitation id="64599FCC88471836FD3B0FBAFC29C04C" author="Winkler, A." box="[730,906,1773,1799]" pageId="20" pageNumber="21" pagination="1009 - 1136" refId="ref39902" refString="Winkler, A. (1929) Catalogus Coleopterorum regionis palaearcticae 2 (pars 9). Albert Winkler, Wien, pp. 1009 - 1136." type="book chapter" year="1929">Winkler (1929)</bibRefCitation>
,
<bibRefCitation id="64599FCC88471836FC760FBAFBF5C043" author="Balthasar, V." box="[919,1110,1773,1800]" pageId="20" pageNumber="21" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar (1963)</bibRefCitation>
,
<bibRefCitation id="64599FCC88471836FB820FBAFAACC04C" author="Ferreira, M. C." box="[1123,1295,1773,1799]" pageId="20" pageNumber="21" pagination="1968 - 1969" refId="ref37353" refString="Ferreira, M. C. (1972) Os Escarabideos de Africa (sul do Saara). I. Revista de Entomologia de Mocambique 11 [1968 - 1969], 1 - 1088, 35 anns, 288 pls." type="book chapter" year="1972">Ferreira (1972)</bibRefCitation>
and
<bibRefCitation id="64599FCC88471836FAA90FBAFF41C060" author="Paulian, R." pageId="20" pageNumber="21" pagination="141 - 154" refId="ref39235" refString="Paulian, R. (1980) Insects of Saudi Arabia. Coleoptera: Scarabaeoidea (1 re contribution). Fauna of Saudi Arabia, 2, 141 - 154." type="journal article" year="1980">Paulian (1980)</bibRefCitation>
uncritically followed this synonymy. Actually the two species, even if very close, can be mainly distinguished by the morphology of genital armatures (
<bibRefCitation id="64599FCC88471836FD220E62FCFAC01B" author="Zunino, M." box="[707,857,1845,1872]" pageId="20" pageNumber="21" pagination="11 - 16" refId="ref40183" refString="Zunino, M. (1976) Onthophagus tessulatus Har. e O. millingeni d'Orb. (Coleoptera, Scarabaeoidea). Annali del Museo Civico di Storia Naturale di Genova, 81, 11 - 16." type="journal article" year="1976">Zunino 1976</bibRefCitation>
). All the Arabian records for
<taxonomicName id="C7C899BE88471836FB430E62FA90C004" authorityName="Harold" authorityYear="1871" box="[1186,1331,1845,1871]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tessulatus">
<emphasis id="32BC3E2F88471836FB430E62FA90C004" box="[1186,1331,1845,1871]" italics="true" pageId="20" pageNumber="21">O. tessulatus</emphasis>
</taxonomicName>
are to be attributed to
<taxonomicName id="C7C899BE88471836FEC70E0EFE19C038" authorityName="d'Orbigny" authorityYear="1898" box="[294,442,1881,1907]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="millingeni">
<emphasis id="32BC3E2F88471836FEC70E0EFE19C038" box="[294,442,1881,1907]" italics="true" pageId="20" pageNumber="21">O. millingeni</emphasis>
</taxonomicName>
that is therefore an Iraqi-Arabian endemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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