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<document id="9889C9BEA45E1D3D492F41F8DF59E055" ID-CLB-Dataset="25961" ID-DOI="10.11646/zootaxa.4658.1.1" ID-GBIF-Dataset="31d6e081-95c3-4086-a45a-c11555dda3aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373040" ID-ZooBank="ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566364910789" checkinUser="plazi" docAuthor="Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed &amp; Barbero, Enrico" docDate="2019" docId="8861532B88401836FF760FD6FABEC557" docLanguage="en" docName="zootaxa.4658.1.1.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus (Micronthophagus) ochreatus d'Orbigny 1898" docType="treatment" docVersion="13" lastPageNumber="21" masterDocId="74582B5388531822FFE10957FFA3C74B" masterDocTitle="An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="20" updateTime="1698751869822" updateUser="ExternalLinkService">
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<mods:title id="EC1069E4C90900F12E93A9CB8C5506BF">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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<mods:namePart id="CE26F96AFD8A02630BD98ECF0B93CC36">Ziani, Stefano</mods:namePart>
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<mods:namePart id="E65D56FB8AFECFC6D4818B35663C43CC">Abdel-Dayem, Mahmoud Saleh</mods:namePart>
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<mods:namePart id="AEEACD4560E9B1E116EBB29BEA884855">Aldhafer, Hathal Mohammed</mods:namePart>
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<mods:namePart id="82A9806C8A8453DF2E62C3159F49A8FC">Barbero, Enrico</mods:namePart>
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<mods:date id="94F0A316878CE40986E578E9862DD034">2019</mods:date>
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<treatment id="8861532B88401836FF760FD6FABEC557" ID-GBIF-Taxon="159217249" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557" httpUri="http://treatment.plazi.org/id/8861532B88401836FF760FD6FABEC557" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="48D2B1B688401831FF760FD6FC22C1D0" box="[151,897,1665,1692]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="0077E23D88401831FF760FD6FC22C1D0" blockId="19.[151,897,1665,1727]" box="[151,897,1665,1692]" pageId="19" pageNumber="20">
<heading id="5B3F555188401831FF760FD6FC22C1D0" bold="true" box="[151,897,1665,1692]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="C7C899BE88401831FF760FD6FC22C1D0" ID-CoL="49ML6" authority="d'Orbigny, 1898" authorityName="d'Orbigny" authorityYear="1898" box="[151,897,1665,1692]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ochreatus" subGenus="Micronthophagus">
<emphasis id="32BC3E2F88401831FF760FD6FC22C1D0" bold="true" box="[151,897,1665,1692]" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FF760FD6FE9FC1D7" bold="true" box="[151,316,1665,1692]" italics="true" pageId="19" pageNumber="20">Onthophagus</emphasis>
(
<emphasis id="32BC3E2F88401831FEAC0FD6FD85C1D7" bold="true" box="[333,550,1665,1692]" italics="true" pageId="19" pageNumber="20">Micronthophagus</emphasis>
)
<emphasis id="32BC3E2F88401831FDD60FD6FD0DC1D7" bold="true" box="[567,686,1665,1692]" italics="true" pageId="19" pageNumber="20">ochreatus</emphasis>
dOrbigny, 1898
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688401831FF760FF2FF5AC1F4" box="[151,249,1700,1727]" pageId="19" pageNumber="20" type="description">
<paragraph id="0077E23D88401831FF760FF2FF5AC1F4" blockId="19.[151,897,1665,1727]" box="[151,249,1700,1727]" pageId="19" pageNumber="20">
[
<figureCitation id="98F3FEB888401831FF7F0FF2FF51C1F4" box="[158,242,1700,1727]" captionStart="FIGURES 1418" captionStartId="23.[151,264,1872,1897]" captionTargetBox="[293,1306,179,1808]" captionTargetId="figure@23.[243,1344,166,1852]" captionTargetPageId="23" captionText="FIGURES 1418. Onthophagini of Arabian Peninsula. 14, Onthophagus (Micronthophagus) ochreatus dOrbigny, 1898: YE (Al Hudaydah) 10 km W Al Mansuriyah; 15, Onthophagus (Onthophagiellus) millingeni dOrbigny, 1898: YE (Sanaa) Haddah; 16, Onthophagus (s. str.) falsus Gillet, 1925: SA (Riyadh) Wadi Al Dawaser; 17, Onthophagus (Proagoderus) quadriarmatus Fairmaire, 1892, ♂: YE (Al Mahrah) Wadi N of Damqut vill.; 18, Onthophagus (Proagoderus) quadriarmatus Fairmaire, 1892, ♀: YE (Al Mahrah) Wadi N of Damqut vill. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373051/files/figure.png" pageId="19" pageNumber="20">Fig. 14</figureCitation>
]
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688401836FF760FBBFABEC557" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="0077E23D88401831FF760FBBFA83C038" blockId="19.[151,1437,1772,2015]" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FF760FBBFECBC04C" bold="true" box="[151,360,1772,1799]" pageId="19" pageNumber="20">Literature quotes</emphasis>
.
<collectingCountry id="78DFA2AD88401831FE940FBBFDACC04F" box="[373,527,1772,1799]" name="Saudi Arabia" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FE940FBBFE14C04F" bold="true" box="[373,439,1772,1799]" pageId="19" pageNumber="20">SAUDI</emphasis>
<taxonomicName id="C7C899BE88401831FE5D0FBBFDACC04F" baseAuthorityName="Gestro" baseAuthorityYear="1889" box="[444,527,1772,1799]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="32BC3E2F88401831FE5D0FBBFDACC04F" bold="true" box="[444,527,1772,1799]" pageId="19" pageNumber="20">ARABIA</emphasis>
</taxonomicName>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88401831FDFF0FBAFD45C04C" box="[542,742,1773,1799]" country="Saudi Arabia" name="Makkah" pageId="19" pageNumber="20">Makkah Province</collectingRegion>
, Jeddah (dOrbigny 1913a).
<collectingRegion id="C20C2CDF88401831FBC40FBAFB67C04C" box="[1061,1220,1773,1799]" country="Saudi Arabia" name="'Asir" pageId="19" pageNumber="20">Asir Province</collectingRegion>
: Wadi Shaqa (Scott
<emphasis id="32BC3E2F88401831FF760E45FF0EC060" box="[151,173,1810,1835]" italics="true" pageId="19" pageNumber="20">in</emphasis>
<bibRefCitation id="64599FCC88401831FF540E46FEEEC060" author="Paulian, R." box="[181,333,1809,1835]" pageId="19" pageNumber="20" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
).
<collectingRegion id="C20C2CDF88401831FE830E46FDAAC060" box="[354,521,1809,1835]" country="Saudi Arabia" name="Jizan" pageId="19" pageNumber="20">Jizan Province</collectingRegion>
, Al Hakimah, Gizan (
<bibRefCitation id="64599FCC88401831FCE00E46FC34C060" author="Zunino, M." box="[769,919,1809,1835]" pageId="19" pageNumber="20" pagination="408 - 416" refId="ref40252" refString="Zunino, M. (1981) Insects of Saudi Arabia Coleoptera: Fam. Scarabaeidae Tribe Onthophagini. Fauna of Saudi Arabia, 3, 408 - 416." type="journal article" year="1981">Zunino 1981</bibRefCitation>
).
<collectingCountry id="78DFA2AD88401831FC4D0E46FC5BC063" box="[940,1016,1809,1835]" name="Yemen" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FC4D0E46FC5BC063" bold="true" box="[940,1016,1809,1835]" pageId="19" pageNumber="20">YEMEN</emphasis>
</collectingCountry>
: without locality (dOrbigny 1913a).
<collectingCountry id="78DFA2AD88401831FF760E63FF7FC007" box="[151,220,1844,1871]" name="Oman" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FF760E63FF7FC007" bold="true" box="[151,220,1844,1871]" pageId="19" pageNumber="20">OMAN</emphasis>
</collectingCountry>
: without locality, as
<taxonomicName id="C7C899BE88401831FE200E62FC1EC004" authority=", (Janikova 1998)" authorityName=", (Janikova" authorityYear="1998" box="[449,957,1845,1871]" class="Insecta" family="Scarabaeidae" genus="Micronthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ochreatus">
<emphasis id="32BC3E2F88401831FE200E62FD5AC004" box="[449,761,1845,1871]" italics="true" pageId="19" pageNumber="20">Micronthophagus ochreatus</emphasis>
, (
<bibRefCitation id="64599FCC88401831FCEF0E62FC16C004" author="Janikova, M." box="[782,949,1845,1871]" pageId="19" pageNumber="20" refId="ref38111" refString="Janikova, M. (1998) Coleoptera. Beetles Checklist. Al Nahda Printing Press, Muscat, 39 pp." type="book" year="1998">Janikova 1998</bibRefCitation>
)
</taxonomicName>
.
<collectingCountry id="78DFA2AD88401831FC260E62FB6CC007" box="[967,1231,1844,1871]" name="United Arab Emirates" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FC260E62FB6CC007" bold="true" box="[967,1231,1844,1871]" pageId="19" pageNumber="20">
<subScript id="9C4CE07888401831FC260E62FBB9C007" attach="left" box="[967,1050,1845,1871]" fontSize="7" pageId="19" pageNumber="20">UNITED</subScript>
ARAB EMIRATES
</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88401831FB3A0E62FF1BC038" country="United Arab Emirates" name="Abu Zaby" pageId="19" pageNumber="20">Abu Dhabi Emirate</collectingRegion>
(
<bibRefCitation id="64599FCC88401831FF270E0EFE6AC038" author="Tigar, B. J. &amp; Osborne, P. E." box="[198,457,1881,1907]" pageId="19" pageNumber="20" pagination="159 - 170" refId="ref39693" refString="Tigar, B. J. &amp; Osborne, P. E. (1999) Patterns of biomass and diversity of aerial insects in Abu Dhabi's sandy deserts. Journal of Arid Environments, 43, 159 - 170. https: // doi. org / 10.1006 / jare. 1999.054 2" type="journal article" year="1999">Tigar &amp; Osborne 1999</bibRefCitation>
); Al-Ajban (
<bibRefCitation id="64599FCC88401831FDB50E0EFCE6C038" author="Kral, D. &amp; Batelka, J." box="[596,837,1881,1907]" pageId="19" pageNumber="20" pagination="78 - 168" refId="ref38267" refString="Kral, D. &amp; Batelka, J. (2017) Order Coleoptera, superfamily Scarabaeoidea. Arthropod fauna of the UAE, 6, 78 - 168." type="journal article" year="2017">Král &amp; Batelka 2017</bibRefCitation>
). Without locality (
<bibRefCitation id="64599FCC88401831FBFA0E0EFAB7C038" author="Gillett, M. P. T. &amp; Gillett, C. P. D. T." box="[1051,1300,1880,1907]" pageId="19" pageNumber="20" pagination="352 - 356" refId="ref37609" refString="Gillett, M. P. T. &amp; Gillett, C. P. D. T. (2005) Insects &amp; other arthropods, pp. 168 - 194. Species checklist, In: Aspinall, S. &amp; Hellyer, P. (Eds.), The Emirates. A natural history. P. Vine, Trident Press, London, pp. 352 - 356." type="book chapter" year="2005">Gillett &amp; Gillett 2005</bibRefCitation>
).
</paragraph>
<paragraph id="0077E23D88401836FF260E2BFE54C6C0" blockId="19.[151,1437,1772,2015]" lastBlockId="20.[151,1437,152,540]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FF260E2BFE0BC0DC" bold="true" box="[199,424,1916,1943]" pageId="19" pageNumber="20">Material examined</emphasis>
.
<materialsCitation id="B0A0E86088401831FE540E2BFDECC0F0" ID-GBIF-Occurrence="2350485792" collectingDate="2013-10-11" collectionCode="KSMA" collectorName="I Rasool" country="Saudi Arabia" elevation="402" location="Wadi Yabah" municipality="Al" pageId="19" pageNumber="20" specimenCount="1" stateProvince="'Asir">
<collectingCountry id="78DFA2AD88401831FE540E2BFDEDC0DF" box="[437,590,1916,1943]" name="Saudi Arabia" pageId="19" pageNumber="20">
<emphasis id="32BC3E2F88401831FE540E2BFE55C0DF" bold="true" box="[437,502,1916,1943]" pageId="19" pageNumber="20">SAUDI</emphasis>
<taxonomicName id="C7C899BE88401831FE1B0E2BFDEDC0DF" baseAuthorityName="Gestro" baseAuthorityYear="1889" box="[506,590,1916,1943]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="32BC3E2F88401831FE1B0E2BFDEDC0DF" bold="true" box="[506,590,1916,1943]" pageId="19" pageNumber="20">ARABIA</emphasis>
</taxonomicName>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88401831FDBB0E2AFD55C0DC" box="[602,758,1917,1943]" country="Saudi Arabia" name="'Asir" pageId="19" pageNumber="20">Asir Province</collectingRegion>
,
<collectingMunicipality id="E013784788401831FD1F0E2AFCB9C0DC" box="[766,794,1917,1943]" pageId="19" pageNumber="20">Al</collectingMunicipality>
Magardah,
<location id="0517B4E688401831FC7C0E2AFB86C0DC" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688401831FC7C0E2AFB86C0DC" box="[925,1061,1917,1943]" country="Saudi Arabia" municipality="Al" name="Wadi Yabah" pageId="19" pageNumber="20" stateProvince="'Asir">Wadi Yabah</location>
,
<quantity id="C7304FD888401831FBCE0E2AFBD6C0DC" box="[1071,1141,1917,1943]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="19" pageNumber="20" unit="m" value="402.0">
<elevation id="8BE5050E88401831FBCE0E2AFBD6C0DC" box="[1071,1141,1917,1943]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="19" pageNumber="20" unit="m" value="402.0">402 m</elevation>
</quantity>
(1914.911N 4147.255E),
<date id="7476C4FD88401831FF760EF6FEAAC0F0" box="[151,265,1953,1979]" pageId="19" pageNumber="20" value="2013-10-11">
<collectingDate id="64323D1588401831FF760EF6FEAAC0F0" box="[151,265,1953,1979]" pageId="19" pageNumber="20" value="2013-10-11">11.x.2013</collectingDate>
</date>
,
<collectorName id="AD3D87EB88401831FEF50EF6FED0C0F0" box="[276,371,1953,1979]" pageId="19" pageNumber="20">I Rasool</collectorName>
leg.
<specimenCount id="16CE29B488401831FE4B0EF6FE42C0F0" box="[426,481,1953,1979]" pageId="19" pageNumber="20" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888401831FE110EF6FDE4C0F0" box="[496,583,1953,1979]" pageId="19" pageNumber="20">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088401831FDBB0EF6FE80C094" ID-GBIF-Occurrence="2350485683" collectingDate="2017-04-03" collectionCode="SZCM" collectorName="P. Rapuzzi" country="Saudi Arabia" elevation="200" location="Ainah" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Asir Province">
<quantity id="C7304FD888401831FDBB0EF6FD30C0F0" box="[602,659,1953,1979]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="km" value="6.0">6 km</quantity>
S
<location id="0517B4E688401831FD4E0EF6FD55C0F0" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688401831FD4E0EF6FD55C0F0" box="[687,758,1953,1979]" country="Saudi Arabia" name="Ainah" pageId="19" pageNumber="20" stateProvince="Asir Province">Ainah</location>
,
<quantity id="C7304FD888401831FCE30EF6FCEAC0F0" box="[770,841,1953,1979]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="200.0">
<elevation id="8BE5050E88401831FCE30EF6FCEAC0F0" box="[770,841,1953,1979]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="200.0">200 m</elevation>
</quantity>
(175528N 422212E),
<date id="7476C4FD88401831FB8B0EF6FB76C0F0" box="[1130,1237,1953,1979]" pageId="19" pageNumber="20" value="2017-04-03">
<collectingDate id="64323D1588401831FB8B0EF6FB76C0F0" box="[1130,1237,1953,1979]" pageId="19" pageNumber="20" value="2017-04-03">3.iv.2017</collectingDate>
</date>
,
<collectorName id="AD3D87EB88401831FB010EF6FAF4C0F0" box="[1248,1367,1953,1979]" pageId="19" pageNumber="20">P. Rapuzzi</collectorName>
leg.
<specimenCount id="16CE29B488401831FA6F0EF6FF1AC094" pageId="19" pageNumber="20" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888401831FF290E92FEB9C094" box="[200,282,1989,2015]" pageId="19" pageNumber="20">SZCM</collectionCode>
)
</materialsCitation>
; “
<materialsCitation id="B0A0E86088401831FED80E92FB40C094" ID-GBIF-Occurrence="2350486112" box="[313,1251,1988,2015]" collectingDate="1935-10-17" collectionCode="NHMUK" collectorName="R. C M. Darling" country="Saudi Arabia" location="Um Shaeby" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Asir Province">
<location id="0517B4E688401831FED80E92FE1CC094" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688401831FED80E92FE1CC094" box="[313,447,1989,2015]" country="Saudi Arabia" name="Um Shaeby" pageId="19" pageNumber="20" stateProvince="Asir Province">Um Shaeby</location>
[locality not found],
<date id="7476C4FD88401831FD530E93FC87C094" box="[690,804,1988,2015]" pageId="19" pageNumber="20" value="1935-10-17">
<collectingDate id="64323D1588401831FD530E93FC87C094" box="[690,804,1988,2015]" pageId="19" pageNumber="20" value="1935-10-17">17.x.1935</collectingDate>
</date>
,
<collectorName id="AD3D87EB88401831FCD00E92FC48C094" box="[817,1003,1989,2015]" pageId="19" pageNumber="20">R. C M. Darling</collectorName>
leg.
<specimenCount id="16CE29B488401831FBC50E92FBF8C094" box="[1060,1115,1989,2015]" pageId="19" pageNumber="20" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888401831FB8B0E92FB78C094" box="[1130,1243,1989,2015]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="20" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E86088401836FB0E0E92FD9DC7F8" ID-GBIF-Occurrence="2350485813" collectingDate="1936-05" collectionCode="NHMUK" collectorName="R. C M. Darling" country="Saudi Arabia" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Asir Province">
<emphasis id="32BC3E2F88401831FB0E0E92FA98C094" box="[1263,1339,1989,2015]" italics="true" pageId="19" pageNumber="20">ibidem</emphasis>
,
<date id="7476C4FD88401831FAAB0E92FA34C094" box="[1354,1431,1989,2015]" pageId="19" pageNumber="20" value="1936-05">
<collectingDate id="64323D1588401831FAAB0E92FA34C094" box="[1354,1431,1989,2015]" pageId="19" pageNumber="20" value="1936-05">v.1936</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FF7609CEFEEFC7F8" box="[151,332,153,179]" pageId="20" pageNumber="21">R. C M. Darling</collectorName>
leg.
<specimenCount id="16CE29B488471836FE6309CEFE1BC7F8" box="[386,440,153,179]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FE2709CEFD94C7F8" box="[454,567,153,179]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="20" pageNumber="21" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FDA909CEFD65C79C" ID-GBIF-Occurrence="2350485682" collectingDate="2007-04-08" collectionCode="NMPC" collectorName="S. Kadlec" country="Yemen" elevation="110" location="10 km W Al Mansuriyah" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Al Hudaydah">
<collectingCountry id="78DFA2AD88471836FDA909CEFD37C7FB" box="[584,660,153,179]" name="Yemen" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FDA909CEFD37C7FB" bold="true" box="[584,660,153,179]" pageId="20" pageNumber="21">
<subScript id="9C4CE07888471836FDA909CEFD37C7FB" attach="left" box="[584,660,153,179]" fontSize="7" pageId="20" pageNumber="21">YEMEN</subScript>
</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88471836FD4109CEFC68C7F8" box="[672,971,153,179]" country="Yemen" name="Al Hudaydah" pageId="20" pageNumber="21">Al Hudaydah Governorate</collectingRegion>
,
<location id="0517B4E688471836FC3409CEFB48C7F8" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688471836FC3409CEFB48C7F8" box="[981,1259,153,179]" country="Yemen" name="10 km W Al Mansuriyah" pageId="20" pageNumber="21" stateProvince="Al Hudaydah">
<quantity id="C7304FD888471836FC3409CEFBB8C7F8" box="[981,1051,153,179]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="km" value="10.0">10 km</quantity>
W Al Mansuriyah
</location>
,
<quantity id="C7304FD888471836FB1409CEFA99C7F8" box="[1269,1338,153,179]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="m" value="110.0">
<elevation id="8BE5050E88471836FB1409CEFA99C7F8" box="[1269,1338,153,179]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="m" value="110.0">110 m</elevation>
</quantity>
(1443N 4312E),
<date id="7476C4FD88471836FF1E09EAFEC9C79C" box="[255,362,189,215]" pageId="20" pageNumber="21" value="2007-04-08">
<collectingDate id="64323D1588471836FF1E09EAFEC9C79C" box="[255,362,189,215]" pageId="20" pageNumber="21" value="2007-04-08">8.iv.2007</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FE9609EAFE46C79C" box="[375,485,189,215]" pageId="20" pageNumber="21">S. Kadlec</collectorName>
leg.
<specimenCount id="16CE29B488471836FDC109EAFDFBC79C" box="[544,600,189,215]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FD8909EAFD1DC79C" box="[616,702,189,215]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FD3309EAFDC2C7B0" ID-GBIF-Occurrence="2350485684" collectingDate="2007-04-12" collectionCode="NMPC" collectorName="S. Kadlec" country="Yemen" elevation="1677" location="Hammam Ali" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Dhamar">
<collectingRegion id="C20C2CDF88471836FD3309EAFC67C79C" box="[722,964,189,215]" country="Yemen" name="Dhamar" pageId="20" pageNumber="21">Dhamar Governorate</collectingRegion>
,
<location id="0517B4E688471836FC3109EAFBCFC79C" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688471836FC3109EAFBCFC79C" box="[976,1132,189,215]" country="Yemen" name="Hammam Ali" pageId="20" pageNumber="21" stateProvince="Dhamar">Hammam Ali</location>
,
<quantity id="C7304FD888471836FB9809EAFB73C79C" box="[1145,1232,189,215]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="20" pageNumber="21" unit="m" value="1677.0">
<elevation id="8BE5050E88471836FB9809EAFB73C79C" box="[1145,1232,189,215]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="20" pageNumber="21" unit="m" value="1677.0">1677 m</elevation>
</quantity>
(1440N 4410E),
<date id="7476C4FD88471836FF7609B6FEB3C7B0" box="[151,272,225,251]" pageId="20" pageNumber="21" value="2007-04-12">
<collectingDate id="64323D1588471836FF7609B6FEB3C7B0" box="[151,272,225,251]" pageId="20" pageNumber="21" value="2007-04-12">12.iv.2007</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FEFA09B6FE24C7B0" box="[283,391,225,251]" pageId="20" pageNumber="21">S. Kadlec</collectorName>
leg.
<specimenCount id="16CE29B488471836FE5F09B6FE56C7B0" box="[446,501,225,251]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FDE209B6FDFAC7B0" box="[515,601,225,251]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FD8A09B6FDF2C654" ID-GBIF-Occurrence="2350485812" collectingDate="2005-10-29" collectionCode="NMPC" collectorName="S. Kadlec" country="Yemen" elevation="1794" location="20 km NW Dhawran" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Hadramawt">
<collectingRegion id="C20C2CDF88471836FD8A09B6FCDEC7B0" box="[619,893,225,251]" country="Yemen" name="Hadramawt" pageId="20" pageNumber="21">Hadramaut Governorate</collectingRegion>
,
<location id="0517B4E688471836FC6909B6FBD7C7B0" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688471836FC6909B6FBD7C7B0" box="[904,1140,225,251]" country="Yemen" name="20 km NW Dhawran" pageId="20" pageNumber="21" stateProvince="Hadramawt">
<quantity id="C7304FD888471836FC6909B6FC6CC7B0" box="[904,975,225,251]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="km" value="20.0">20 km</quantity>
NW Dhawran
</location>
,
<quantity id="C7304FD888471836FB9E09B6FB77C7B0" box="[1151,1236,225,251]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="20" pageNumber="21" unit="m" value="1794.0">
<elevation id="8BE5050E88471836FB9E09B6FB77C7B0" box="[1151,1236,225,251]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="20" pageNumber="21" unit="m" value="1794.0">1794 m</elevation>
</quantity>
(1440N 4413E),
<date id="7476C4FD88471836FF760853FEA9C654" box="[151,266,260,287]" pageId="20" pageNumber="21" value="2005-10-29">
<collectingDate id="64323D1588471836FF760853FEA9C654" box="[151,266,260,287]" pageId="20" pageNumber="21" value="2005-10-29">29.x.2005</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FEF20852FEDDC654" box="[275,382,261,287]" pageId="20" pageNumber="21">S. Kadlec</collectorName>
leg.
<specimenCount id="16CE29B488471836FE500852FE45C654" box="[433,486,261,287]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FE120852FDEAC654" box="[499,585,261,287]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FDB80852FA3AC654" ID-GBIF-Occurrence="2350485794" box="[601,1433,260,287]" collectingDate="1935-10-14" collectionCode="NHMUK" collectorName="R. C. M. Darling" country="Yemen" elevation="1794" isTailEnumeration="true" location="20 km NW Dhawran" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Lahij">
<collectingRegion id="C20C2CDF88471836FDB80852FC8BC654" box="[601,808,261,287]" country="Yemen" name="Lahij" pageId="20" pageNumber="21">Lahij Governorate</collectingRegion>
,
<collectingRegion id="C20C2CDF88471836FCD00852FCCCC654" box="[817,879,261,287]" country="Yemen" name="Lahij" pageId="20" pageNumber="21">Lahij</collectingRegion>
,
<date id="7476C4FD88471836FC990853FC49C654" box="[888,1002,260,287]" pageId="20" pageNumber="21" value="1935-10-14">
<collectingDate id="64323D1588471836FC990853FC49C654" box="[888,1002,260,287]" pageId="20" pageNumber="21" value="1935-10-14">14.x.1935</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FC150852FB0EC654" box="[1012,1197,261,287]" pageId="20" pageNumber="21">R. C. M. Darling</collectorName>
leg.
<specimenCount id="16CE29B488471836FB3E0852FAB7C654" box="[1247,1300,261,287]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FAC00852FA31C654" box="[1313,1426,261,287]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="20" pageNumber="21" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FF76087FFE99C62C" ID-GBIF-Occurrence="2350485810" collectingDate="1995-07-07" collectionCode="SOIC" collectorName="B. Skule &amp; M. D. Gallagher" country="Oman" location="Wahiba Sands" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Shamal ash Sharqiyah">
<collectingCountry id="78DFA2AD88471836FF76087FFF7FC60B" box="[151,220,296,323]" name="Oman" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF76087FFF7FC60B" bold="true" box="[151,220,296,323]" pageId="20" pageNumber="21">OMAN</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF88471836FF08087EFD87C608" box="[233,548,297,323]" country="Oman" name="Shamal ash Sharqiyah" pageId="20" pageNumber="21">Ash Sharqiyah Governorate</collectingRegion>
,
<location id="0517B4E688471836FDCF087EFD6EC608" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688471836FDCF087EFD6EC608" box="[558,717,297,323]" country="Oman" name="Wahiba Sands" pageId="20" pageNumber="21" stateProvince="Shamal ash Sharqiyah">Wahiba Sands</location>
(2139N 5918E), 6/
<date id="7476C4FD88471836FC55087FFB85C608" box="[948,1062,296,323]" pageId="20" pageNumber="21" value="1995-07-07">
<collectingDate id="64323D1588471836FC55087FFB85C608" box="[948,1062,296,323]" pageId="20" pageNumber="21" value="1995-07-07">7.vii.1995</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FBD0087EFB31C608" box="[1073,1170,297,323]" pageId="20" pageNumber="21">B. Skule</collectorName>
&amp;
<collectorName id="AD3D87EB88471836FB54087EFACFC608" box="[1205,1388,297,323]" pageId="20" pageNumber="21">M. D. Gallagher</collectorName>
leg. 5 exx. (
<collectionCode id="66D97AF888471836FF0D081AFE91C62C" box="[236,306,333,359]" country="Oman" httpUri="http://grbio.org/cool/txkx-ji9v" name="Natural History Museum, National Insect Collection" pageId="20" pageNumber="21">SOIC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E86088471836FEA4081AFE50C6C0" ID-GBIF-Occurrence="2350486001" collectingDate="1994-09-30" collectionCode="MGCB" collectorName="M. P. T. Gillett" country="United Arab Emirates" county="Sharja Emirate" location="Ain" municipality="Naqa" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Ash Sharqiyah Governorate">
<collectingCountry id="78DFA2AD88471836FEA4081AFDF3C62F" box="[325,592,332,359]" name="United Arab Emirates" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FEA4081AFDF3C62F" bold="true" box="[325,592,332,359]" pageId="20" pageNumber="21">
<subScript id="9C4CE07888471836FEA4081AFE34C62F" attach="left" box="[325,407,333,359]" fontSize="7" pageId="20" pageNumber="21">UNITED</subScript>
ARAB EMIRATES
</emphasis>
</collectingCountry>
:
<collectingCounty id="E9169AB188471836FD81081AFCA8C62C" box="[608,779,333,359]" pageId="20" pageNumber="21">Sharja Emirate</collectingCounty>
,
<collectingMunicipality id="E013784788471836FCF6081AFCF1C62C" box="[791,850,333,359]" pageId="20" pageNumber="21">Naqa</collectingMunicipality>
Suhunah—Al
<location id="0517B4E688471836FC1B081AFB87C62C" LSID="urn:lsid:plazi:treatment:8861532B88401836FF760FD6FABEC557:0517B4E688471836FC1B081AFB87C62C" box="[1018,1060,333,359]" country="United Arab Emirates" county="Sharja Emirate" municipality="Naqa" name="Ain" pageId="20" pageNumber="21" stateProvince="Ash Sharqiyah Governorate">Ain</location>
(2518N 5542E),
<date id="7476C4FD88471836FB17081AFAD3C62C" box="[1270,1392,333,359]" pageId="20" pageNumber="21" value="1994-09-30">
<collectingDate id="64323D1588471836FB17081AFAD3C62C" box="[1270,1392,333,359]" pageId="20" pageNumber="21" value="1994-09-30">30.ix.1994</collectingDate>
</date>
,
<collectorName id="AD3D87EB88471836FA9D081AFEB5C6C0" pageId="20" pageNumber="21">M. P. T. Gillett</collectorName>
leg.
<specimenCount id="16CE29B488471836FEAC0826FE27C6C0" box="[333,388,369,395]" pageId="20" pageNumber="21" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF888471836FE730826FE48C6C0" box="[402,491,369,395]" pageId="20" pageNumber="21">MGCB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0077E23D88471836FF2608C3FABEC557" blockId="20.[151,1437,152,540]" pageId="20" pageNumber="21">
<emphasis id="32BC3E2F88471836FF2608C3FEFAC6E4" bold="true" box="[199,345,404,431]" pageId="20" pageNumber="21">Distribution</emphasis>
.
<emphasis id="32BC3E2F88471836FE8A08C3FDF4C6E4" bold="true" box="[363,599,404,431]" pageId="20" pageNumber="21">Arabian Peninsula:</emphasis>
<collectingCountry id="78DFA2AD88471836FD8308C2FD05C6E4" box="[610,678,405,431]" name="Oman" pageId="20" pageNumber="21">Oman</collectingCountry>
(
<bibRefCitation id="64599FCC88471836FD5908C2FCC0C6E4" author="Janikova, M." box="[696,867,405,431]" pageId="20" pageNumber="21" refId="ref38111" refString="Janikova, M. (1998) Coleoptera. Beetles Checklist. Al Nahda Printing Press, Muscat, 39 pp." type="book" year="1998">Janikova 1998</bibRefCitation>
),
<collectingCountry id="78DFA2AD88471836FC9A08C2FBB0C6E4" box="[891,1043,405,431]" name="Saudi Arabia" pageId="20" pageNumber="21">
Saudi
<taxonomicName id="C7C899BE88471836FC2508C2FBB0C6E4" baseAuthorityName="Gestro" baseAuthorityYear="1889" box="[964,1043,405,431]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="35" phylum="Arthropoda" rank="genus">Arabia</taxonomicName>
</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FBC008C2FBCCC6E4" box="[1057,1135,405,431]" name="Yemen" pageId="20" pageNumber="21">Yemen</collectingCountry>
(dOrbigny 1913a), Unit- ed Arab Emirates (
<bibRefCitation id="64599FCC88471836FE8E08EEFDDAC698" author="Tigar, B. J. &amp; Osborne, P. E." box="[367,633,441,467]" pageId="20" pageNumber="21" pagination="159 - 170" refId="ref39693" refString="Tigar, B. J. &amp; Osborne, P. E. (1999) Patterns of biomass and diversity of aerial insects in Abu Dhabi's sandy deserts. Journal of Arid Environments, 43, 159 - 170. https: // doi. org / 10.1006 / jare. 1999.054 2" type="journal article" year="1999">Tigar &amp; Osborne 1999</bibRefCitation>
).
<emphasis id="32BC3E2F88471836FD6E08EEFC97C698" bold="true" box="[655,820,440,467]" pageId="20" pageNumber="21">North Africa:</emphasis>
<collectingCountry id="78DFA2AD88471836FCDC08EEFC01C698" box="[829,930,441,467]" name="Morocco" pageId="20" pageNumber="21">Morocco</collectingCountry>
(
<bibRefCitation id="64599FCC88471836FC5208EEFBE0C698" author="Arrow, G. J." box="[947,1091,441,467]" pageId="20" pageNumber="21" refId="ref36526" refString="Arrow, G. J. (1931) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor &amp; Francis, London, 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
).
<emphasis id="32BC3E2F88471836FBB808EFFB0DC698" bold="true" box="[1113,1198,440,467]" pageId="20" pageNumber="21">Africa:</emphasis>
<collectingCountry id="78DFA2AD88471836FB5908EEFAB6C698" box="[1208,1301,441,467]" name="Djibouti" pageId="20" pageNumber="21">Djibouti</collectingCountry>
(dOrbigny 1898b),
<collectingCountry id="78DFA2AD88471836FF16088AFEE5C6BC" box="[247,326,477,503]" name="Eritrea" pageId="20" pageNumber="21">Eritrea</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FEB5088AFE15C6BC" box="[340,438,477,503]" name="Ethiopia" pageId="20" pageNumber="21">Ethiopia</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FE25088AFD83C6BC" box="[452,544,477,503]" name="Gambia" pageId="20" pageNumber="21">Gambia</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FDCF088AFD0FC6BC" box="[558,684,477,503]" name="Mauritania" pageId="20" pageNumber="21">Mauritania</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FD58088AFCB7C6BC" box="[697,788,477,503]" name="Senegal" pageId="20" pageNumber="21">Senegal</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FCC2088AFC21C6BC" box="[803,898,477,503]" name="Somalia" pageId="20" pageNumber="21">Somalia</collectingCountry>
,
<collectingCountry id="78DFA2AD88471836FC6E088AFC76C6BC" box="[911,981,477,503]" name="Sudan" pageId="20" pageNumber="21">Sudan</collectingCountry>
(
<bibRefCitation id="64599FCC88471836FC06088AFB3BC6B3" author="Balthasar, V." box="[999,1176,477,504]" pageId="20" pageNumber="21" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar 1963</bibRefCitation>
).
<emphasis id="32BC3E2F88471836FB4F088BFB4FC6BC" bold="true" box="[1198,1260,476,503]" pageId="20" pageNumber="21">Asia:</emphasis>
<collectingCountry id="78DFA2AD88471836FB17088AFA93C6BC" box="[1270,1328,477,503]" name="India" pageId="20" pageNumber="21">India</collectingCountry>
(Scott
<emphasis id="32BC3E2F88471836FA670889FA3FC6BC" box="[1414,1436,478,503]" italics="true" pageId="20" pageNumber="21">in</emphasis>
<bibRefCitation id="64599FCC88471836FF760B56FE8EC550" author="Paulian, R." box="[151,301,513,539]" pageId="20" pageNumber="21" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
),
<collectingCountry id="78DFA2AD88471836FEA30B56FE02C550" box="[322,417,513,539]" name="Pakistan" pageId="20" pageNumber="21">Pakistan</collectingCountry>
(
<bibRefCitation id="64599FCC88471836FE4E0B56FD9FC550" author="Arrow, G. J." box="[431,572,513,539]" pageId="20" pageNumber="21" refId="ref36526" refString="Arrow, G. J. (1931) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor &amp; Francis, London, 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
),
<collectingCountry id="78DFA2AD88471836FDB10B56FD1DC550" box="[592,702,513,539]" name="Sri Lanka" pageId="20" pageNumber="21">Sri Lanka</collectingCountry>
(
<bibRefCitation id="64599FCC88471836FD2D0B56FCFAC550" author="Arrow, G. J." box="[716,857,513,539]" pageId="20" pageNumber="21" refId="ref36526" refString="Arrow, G. J. (1931) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor &amp; Francis, London, 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
; Scott
<emphasis id="32BC3E2F88471836FC440B55FC18C550" box="[933,955,514,539]" italics="true" pageId="20" pageNumber="21">in</emphasis>
<bibRefCitation id="64599FCC88471836FC230B56FBFBC550" author="Paulian, R." box="[962,1112,513,539]" pageId="20" pageNumber="21" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
;
<bibRefCitation id="64599FCC88471836FB850B56FAB2C557" author="Balthasar, V." box="[1124,1297,513,540]" pageId="20" pageNumber="21" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar 1963</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>