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<document ID-DOI="http://doi.org/10.5281/zenodo.3748525" ID-GBIF-Dataset="6384b520-7e9f-4874-a414-76c2e9b01d74" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3748525" approvalRequired="1" checkinTime="1586358796837" checkinUser="plazi" docAuthor="Burgin, Connor" docDate="2019" docId="885887A2FFFF8A18F894F3EDFACDD152" docLanguage="en" docName="hbmw_9_Rhinolophidae.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Rhinolophus francisi Soisook 2015" docType="treatment" docVersion="24" lastPageNumber="328" masterDocId="7461FFDAFFCF8A29FFCCFFA1FF85D963" masterDocTitle="Rhinolophidae" masterLastPageNumber="332" masterPageNumber="280" pageNumber="327" updateTime="1658949228820" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhinolophidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burgin, Connor</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Mittermeier, Russell A.</mods:namePart>
</mods:name>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>332</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3748525</mods:identifier>
<mods:identifier type="GBIF-Dataset">6384b520-7e9f-4874-a414-76c2e9b01d74</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3748525</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3809062" ID-GBIF-Taxon="163549924" ID-Zenodo-Dep="3809062" LSID="urn:lsid:plazi:treatment:885887A2FFFF8A18F894F3EDFACDD152" httpUri="http://treatment.plazi.org/id/885887A2FFFF8A18F894F3EDFACDD152" lastPageId="49" lastPageNumber="328" pageId="48" pageNumber="327">
<subSubSection box="[1880,2863,3148,3209]" pageId="48" pageNumber="327" type="vernacular_names">
<paragraph blockId="48.[1880,3459,3148,3209]" box="[1880,2863,3148,3209]" pageId="48" pageNumber="327">
<figureCitation box="[1880,1952,3148,3209]" captionStart="On" captionStartId="46.[166,208,4390,4417]" captionTargetBox="[152,3486,163,4684]" captionTargetPageId="45" captionText="On following pages: 94. Thomas's Horseshoe Bat (Rhinolophus thomasi); 95. Lesser Horseshoe Bat (Rhinolophus hipposideros); 96. rancis's Woolly Horseshoe Bat (Rhinolophus trancisi); 97. Lesser Woolly Horseshoe Bat (Rhinolophus sedulus); 98. Trefoil Horseshoe Bat (Rhinolophus trifoliatus); 99. Northern Woolly Horseshoe Bat (Rhinolophus perniger); 100. Selangor Woolly Horseshoe Bat (Rhinolophus luctoides); 101. Malaysian Woolly Horseshoe Bat (Rhinolophus mono); 102. Great Woolly Horseshoe Bat (Rhinolophus luctus); 1.03. Beddome's Woolly Horseshoe Bat (Rhinolophus beddomei); 104. Formosan Woolly Horseshoe Bat (Rhinolophus formosaë); 105. Thailand Horseshoe Bat (Rhinolophus thailandensis); 106. Dobson's Horseshoe Bat (Rhinolophus yunanensis); 107. Chiew Kwee's Horseshoe Bat (Rhinolophus chiewkweeae); 108. Pearson's Horseshoe Bat (Rhinolophus pearsonii); 109. Mitred Horseshoe Bat (Rhinolophus mitratus)." figureDoi="http://doi.org/10.5281/zenodo.3748537" httpUri="https://zenodo.org/record/3748537/files/figure.png" pageId="48" pageNumber="327">96</figureCitation>
. Franciss
<emphasis bold="true" box="[2240,2436,3148,3209]" pageId="48" pageNumber="327">Woolly</emphasis>
Horseshoe
<emphasis bold="true" box="[2767,2863,3148,3209]" pageId="48" pageNumber="327">Bat</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[2927,3459,3148,3209]" pageId="48" pageNumber="327" type="nomenclature">
<paragraph blockId="48.[1880,3459,3148,3209]" box="[2927,3459,3148,3209]" pageId="48" pageNumber="327">
<taxonomicName authority="Soisook" authorityName="Soisook" authorityYear="2015" box="[2927,3459,3148,3209]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="francisi">
<emphasis box="[2927,3459,3148,3209]" italics="true" pageId="48" pageNumber="327">Rhinolophus francisi</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[1880,3302,3224,3251]" pageId="48" pageNumber="327" type="vernacular_names">
<paragraph blockId="48.[1880,3302,3224,3251]" box="[1880,3302,3224,3251]" pageId="48" pageNumber="327">
French: Rhinolophe de rancis /
<emphasis bold="true" box="[2317,2440,3224,3251]" pageId="48" pageNumber="327">German:</emphasis>
Francis-Wollhaar-Hufeisennase /
<emphasis bold="true" box="[2886,3012,3224,3251]" pageId="48" pageNumber="327">Spanish:</emphasis>
Herradura de Francis
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="327" type="reference_group">
<paragraph blockId="48.[2697,3502,3325,3890]" pageId="48" pageNumber="327">
Taxonomy.
<taxonomicName authority="Soisook" authorityName="Soisook" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="frantisi">
<emphasis box="[2927,3291,3325,3364]" italics="true" pageId="48" pageNumber="327">Rhinolophus frantisi</emphasis>
Soisook
<emphasis box="[3473,3502,3325,3364]" italics="true" pageId="48" pageNumber="327">et</emphasis>
<emphasis box="[2701,2752,3377,3416]" italics="true" pageId="48" pageNumber="327">al,</emphasis>
2015
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="327" type="materials_examined">
<paragraph blockId="48.[2697,3502,3325,3890]" pageId="48" pageNumber="327">
<materialsCitation ID-GBIF-Occurrence="2597534024" country="Malaysia" elevation="1600" latitude="5.5666666" location="Gunung Trus Madi" longLatPrecision="1305" longitude="116.48333" municipality="Gunung Trus Madi" pageId="48" pageNumber="327" specimenCount="1" stateProvince="Sabah" typeStatus="type">
<collectingMunicipality box="[2898,3264,3377,3416]" pageId="48" pageNumber="327">Gunung Trus Madi</collectingMunicipality>
,
<collectingRegion box="[3286,3406,3377,3416]" country="Malaysia" name="Sabah" pageId="48" pageNumber="327">Sabah</collectingRegion>
,
<collectingCountry name="Malaysia" pageId="48" pageNumber="327">Malaysia</collectingCountry>
(approx.
<geoCoordinate box="[3005,3140,3429,3468]" degrees="5" direction="north" minutes="34" orientation="latitude" pageId="48" pageNumber="327" precision="925" value="5.5666666">5°34N</geoCoordinate>
,
<geoCoordinate box="[3172,3348,3429,3468]" degrees="116" direction="east" minutes="29" orientation="longitude" pageId="48" pageNumber="327" precision="925" value="116.48333">
116°29
<emphasis box="[3314,3348,3429,3468]" italics="true" pageId="48" pageNumber="327">E</emphasis>
</geoCoordinate>
), at an elevation of about
<quantity box="[3053,3201,3482,3521]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="48" pageNumber="327" unit="m" value="1600.0">
<elevation box="[3053,3201,3482,3521]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="48" pageNumber="7" unit="m" value="1600.0">1,600 m</elevation>
</quantity>
a.s.l.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="327" type="discussion">
<paragraph blockId="48.[2697,3502,3325,3890]" lastBlockId="48.[1877,3500,3901,4624]" pageId="48" pageNumber="327">
<taxonomicName box="[2698,3053,3535,3574]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="frantisi">
<emphasis box="[2698,3053,3535,3574]" italics="true" pageId="48" pageNumber="327">Rhinolophus frantisi</emphasis>
</taxonomicName>
is included in the
<emphasis box="[3422,3498,3535,3574]" italics="true" pageId="48" pageNumber="327">
<emphasis box="[2701,2796,3588,3627]" italics="true" pageId="48" pageNumber="327">trifoliatus</emphasis>
</emphasis>
species group, along with.
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[3361,3487,3588,3627]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="sedulus">
<emphasis box="[3361,3487,3588,3627]" italics="true" pageId="48" pageNumber="327">sedulus</emphasis>
</taxonomicName>
,.
<emphasis box="[2750,3494,3640,3679]" italics="true" pageId="48" pageNumber="327">
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[2750,2911,3640,3679]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="trifoliatus">trifoliatus</taxonomicName>
,
<taxonomicName authorityName="Hodgson" authorityYear="1843" box="[2928,3112,3640,3679]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="perniger">R.perniger</taxonomicName>
,
<taxonomicName box="[3130,3324,3640,3679]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="luctoides">R.luctoides</taxonomicName>
,
<taxonomicName box="[3342,3488,3640,3679]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="mono">R.mono</taxonomicName>
,
</emphasis>
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[2698,2852,3693,3732]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="330" phylum="Chordata" rank="species" species="luctus">
<emphasis box="[2698,2747,3693,3732]" italics="true" pageId="48" pageNumber="327">R.</emphasis>
luctus
</taxonomicName>
,
<emphasis box="[2872,3081,3693,3732]" italics="true" pageId="48" pageNumber="327">
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[2872,3075,3693,3732]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="beddomei">R. beddomei</taxonomicName>
,
</emphasis>
and
<emphasis box="[3179,3385,3693,3732]" italics="true" pageId="48" pageNumber="327">
<taxonomicName authorityName="Sanborn" authorityYear="1939" box="[3179,3381,3693,3732]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="formosae">R formosae</taxonomicName>
.
</emphasis>
It appears to be
<emphasis box="[2910,3006,3746,3785]" italics="true" pageId="48" pageNumber="327">sister</emphasis>
to the rest of the
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[3336,3499,3746,3785]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="trifoliatus">
<emphasis box="[3336,3499,3746,3785]" italics="true" pageId="48" pageNumber="327">trifoliatus</emphasis>
</taxonomicName>
species group. The
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[3058,3219,3799,3838]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="trifoliatus">
<emphasis box="[3058,3219,3799,3838]" italics="true" pageId="48" pageNumber="327">trifoliatus</emphasis>
</taxonomicName>
group appears to be close to the
<taxonomicName authorityName="Borkhausen" authorityYear="1797" box="[3045,3246,3851,3890]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="hipposideros">
<emphasis box="[3045,3246,3851,3890]" italics="true" pageId="48" pageNumber="327">hipposideros</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Horsfield" authorityYear="1851" box="[3342,3499,3851,3890]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="pearsonii">
<emphasis box="[3342,3499,3851,3890]" italics="true" pageId="48" pageNumber="327">pearsonii</emphasis>
</taxonomicName>
species groups, based on mitochondrial genes; they form a clade that is basal to the rest of the Asiatic clade of
<taxonomicName authorityName="Horsfield" authorityYear="1823" box="[2284,2497,3953,3992]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis box="[2284,2497,3953,3992]" italics="true" pageId="48" pageNumber="327">Rhinolophus</emphasis>
</taxonomicName>
or is sister to the rest
<emphasis box="[2894,3135,3953,3992]" italics="true" pageId="48" pageNumber="327">of hinolophus,</emphasis>
based on nuclear genes, although further genetic studies are needed to confirm the
<emphasis box="[3087,3271,4006,4045]" italics="true" pageId="48" pageNumber="327">respective</emphasis>
positions of the species groups within the genus. Specimens ofthis
<emphasis box="[2874,3004,4059,4098]" italics="true" pageId="48" pageNumber="327">species</emphasis>
from
<collectingCountry box="[3112,3284,4059,4098]" name="Thailand" pageId="48" pageNumber="327">Thailand</collectingCountry>
were previously attributed to
<emphasis box="[2228,2438,4112,4151]" italics="true" pageId="48" pageNumber="327">
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[2228,2432,4112,4151]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="beddomei">R.beddomei</taxonomicName>
,
</emphasis>
in 2010, but proved to represent
<emphasis box="[3066,3250,4112,4151]" italics="true" pageId="48" pageNumber="327">
<emphasis bold="true" box="[3066,3104,4112,4151]" italics="true" pageId="48" pageNumber="327">A</emphasis>
<taxonomicName box="[3111,3250,4112,4151]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="frantisi">frantisi</taxonomicName>
</emphasis>
The current distribution of this
<emphasis box="[2226,2355,4164,4203]" italics="true" pageId="48" pageNumber="327">species</emphasis>
is limited to two
<emphasis bold="true" box="[2664,2783,4164,4203]" italics="true" pageId="48" pageNumber="327">widely</emphasis>
separate areas; the species may also be present in
<collectingCountry box="[2071,2234,4217,4256]" name="Vietnam" pageId="48" pageNumber="327">Vietnam</collectingCountry>
, based on genetic evidence that placed three Vietnamese specimens as sister to the nominate race. Race
<emphasis box="[2573,2782,4269,4308]" italics="true" pageId="48" pageNumber="327">thailandicus</emphasis>
may ultimately prove to be a distinct
<emphasis box="[1877,2008,4322,4361]" italics="true" pageId="48" pageNumber="327">species</emphasis>
when more specimens are available for comparison. Two subspecies recognized.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3750104" ID-Zenodo-Dep="3750104" httpUri="https://zenodo.org/record/3750104/files/figure.png" inLine="true" pageId="48" pageNumber="327" targetBox="[1876,2667,3326,3882]" targetPageId="48">
<paragraph blockId="48.[1877,3500,3901,4624]" box="[1878,2414,4375,4414]" pageId="48" pageNumber="327">
<emphasis bold="true" box="[1878,2414,4375,4414]" pageId="48" pageNumber="327">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="48.[1877,3500,3901,4624]" pageId="48" pageNumber="327">
<taxonomicName authority="Soisook" authorityName="Soisook" box="[1878,2236,4427,4466]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata" rank="species" species="frantisi">
<emphasis box="[1878,1947,4427,4466]" italics="true" pageId="48" pageNumber="327">R.f.</emphasis>
<emphasis box="[1953,2087,4427,4466]" italics="true" pageId="48" pageNumber="327">frantisi</emphasis>
Soisook
</taxonomicName>
<emphasis box="[2246,2331,4427,4466]" italics="true" pageId="48" pageNumber="327">et al,</emphasis>
2015 - known only from a few localities in N,
<emphasis bold="true" box="[3166,3229,4427,4466]" pageId="48" pageNumber="327">
<collectionCode box="[3166,3229,4427,4466]" pageId="48" pageNumber="327">SW</collectionCode>
</emphasis>
&amp; S
<location LSID="urn:lsid:plazi:treatment:885887A2FFFF8A18F894F3EDFACDD152:052E606FFFFF8A19F2C0EEEAF218C811" box="[3340,3485,4427,4466]" name="Borneo" pageId="48" pageNumber="327">Borneo</location>
.
<taxonomicName box="[1879,2587,4480,4519]" class="Mammalia" family="Rhinolophidae" kingdom="Animalia" order="Chiroptera" pageId="48" pageNumber="327" phylum="Chordata">
<emphasis box="[1879,2176,4480,4519]" italics="true" pageId="48" pageNumber="327">
<collectionCode box="[1879,1915,4480,4519]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="48" pageNumber="327">R</collectionCode>
.f. thailandicus
</emphasis>
<collectorName box="[2181,2328,4480,4519]" pageId="48" pageNumber="327">Soisook</collectorName>
&amp;
<emphasis bold="true" box="[2378,2481,4480,4519]" pageId="48" pageNumber="327">
<collectorName box="[2378,2481,4480,4519]" pageId="48" pageNumber="327">Bates</collectorName>
</emphasis>
, 2015
</taxonomicName>
— known only from
<typeStatus box="[2958,3035,4480,4519]" pageId="48" pageNumber="327">type</typeStatus>
locality
<emphasis bold="true" box="[3187,3225,4480,4519]" pageId="48" pageNumber="327">in</emphasis>
<collectionCode box="[3235,3299,4480,4519]" pageId="48" pageNumber="327">SW</collectionCode>
<collectorName pageId="48" pageNumber="327">Thailand. Genetic</collectorName>
data suggest that the species may also occur in
<emphasis bold="true" box="[2977,3143,4532,4571]" pageId="48" pageNumber="327">
<collectingCountry box="[2977,3143,4532,4571]" name="Vietnam" pageId="48" pageNumber="327">Vietnam</collectingCountry>
</emphasis>
, but this requires more research.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="49" pageNumber="328" type="description">
<paragraph blockId="49.[190,1817,359,2104]" pageId="49" pageNumber="328">
Descriptive
<emphasis bold="true" box="[419,531,359,398]" pageId="49" pageNumber="328">notes.</emphasis>
<emphasis box="[546,767,359,398]" italics="true" pageId="49" pageNumber="328">Head-body</emphasis>
63-
<quantity box="[838,953,359,398]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="49" pageNumber="328" unit="mm" value="4.0">4 mm</quantity>
, tail 29-
<quantity box="[1111,1289,359,398]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.8" metricValueMin="0.9" pageId="49" pageNumber="328" unit="mm" value="23.5" valueMax="38.0" valueMin="9.0">9-38 mm</quantity>
, ear 23-
<quantity box="[1448,1626,359,398]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="49" pageNumber="328" unit="mm" value="17.5" valueMax="27.0" valueMin="8.0">8-27 mm</quantity>
, hindfoot 13-5-15-
<quantity box="[353,468,411,450]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="49" pageNumber="328" unit="mm" value="5.0">5 mm</quantity>
, forearm 52-9-54-
<quantity box="[818,930,411,450]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="49" pageNumber="328" unit="mm" value="7.0">7 mm</quantity>
; weight
<quantity box="[1090,1237,411,450]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="49" pageNumber="328" unit="g" value="17.0" valueMax="18.0" valueMin="16.0">16-18 g</quantity>
. Fur is long and woolly; dorsal and ventral pelage has dark grayish brown appearance, each hair being whitish brown basally and grayish brown at the dp. Ears are medium-sized. Noseleaf is dark brown and the lancet is high and slightly concave in the middle, with a pointed tip; connecting process is long and projects anteriorly; sella is broader basally,
<emphasis box="[1489,1648,627,666]" italics="true" pageId="49" pageNumber="328">abruptly</emphasis>
concave in the middle, parallel-sided at top with a rounded
<emphasis box="[1177,1234,680,719]" italics="true" pageId="49" pageNumber="328">off</emphasis>
tip, and has two large circular lappets on either side of base; horseshoe is
<emphasis box="[1034,1206,733,772]" italics="true" pageId="49" pageNumber="328">relatively</emphasis>
wide (12-2-12-
<quantity box="[1501,1617,733,772]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="49" pageNumber="328" unit="mm" value="4.0">4 mm</quantity>
) but does not
<emphasis box="[274,483,786,825]" italics="true" pageId="49" pageNumber="328">completely</emphasis>
cover muzzle, and has a deep median emargination. Lower lip has one mental groove. Skull is robust (zygomatic width is greater than mastoid width); anterior median swellings
<emphasis box="[695,843,893,932]" italics="true" pageId="49" pageNumber="328">are well</emphasis>
inflated and bulbous; posterior swellings are small and uninflated; frontal depression is deep; supraorbital ridges are very conspicuous; sagittal crest is well defined; zygomatics are strong with a distinct triangular dorsal arch at midpoint. C1 is tall; P2 is very small with distinct cusp, and is within the tooth row; P3 is minute and almost
<emphasis box="[657,868,1107,1146]" italics="true" pageId="49" pageNumber="328">completely</emphasis>
extruded from tooth row; P2 and P4 are touching or nearly so.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="328" type="biology_ecology">
<paragraph blockId="49.[190,1817,359,2104]" pageId="49" pageNumber="328">
Habitat In Borneo, Franciss
<emphasis box="[760,893,1213,1252]" italics="true" pageId="49" pageNumber="328">Woolly</emphasis>
Horseshoe Bat was collected in montane forest and undisturbed evergreen forest in Sabah, and undisturbed and logged evergreen forest in Kalimantan; also, over a seasonal stream surrounded by dense primary evergreen forest in
<collectingCountry box="[479,658,1373,1412]" name="Thailand" pageId="49" pageNumber="328">Thailand</collectingCountry>
. It was found at
<quantity box="[967,1085,1373,1412]" metricMagnitude="2" metricUnit="m" metricValue="4.31" pageId="49" pageNumber="328" unit="m" value="431.0">431 m</quantity>
in
<collectingCountry box="[1145,1320,1373,1412]" name="Thailand" pageId="49" pageNumber="328">Thailand</collectingCountry>
and at
<quantity box="[1464,1600,1373,1412]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="328" unit="m" value="1600.0">1600 m</quantity>
in Sabah.
</paragraph>
<paragraph blockId="49.[190,1817,359,2104]" box="[192,867,1426,1465]" pageId="49" pageNumber="328">
<emphasis bold="true" box="[192,373,1426,1465]" pageId="49" pageNumber="328">Food and</emphasis>
Feeding. No information.
</paragraph>
<paragraph blockId="49.[190,1817,359,2104]" box="[192,695,1479,1518]" pageId="49" pageNumber="328">Breeding. No information.</paragraph>
<paragraph blockId="49.[190,1817,359,2104]" pageId="49" pageNumber="328">
<emphasis bold="true" box="[190,510,1532,1571]" pageId="49" pageNumber="328">Activity patterns.</emphasis>
Call shape is FM/CF/FM with a peak F of 49-2—50 kHz, durations of 20-2—39-9 milliseconds, and interpulse intervals of 30-7—126 milliseconds, in Kalimantan and
<collectingCountry box="[432,609,1639,1678]" name="Thailand" pageId="49" pageNumber="328">Thailand</collectingCountry>
.
</paragraph>
<paragraph blockId="49.[190,1817,359,2104]" box="[193,1459,1692,1731]" pageId="49" pageNumber="328">
<emphasis bold="true" box="[193,881,1692,1731]" pageId="49" pageNumber="328">Movements, Home range and Social</emphasis>
organization. No information.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="328" type="conservation">
<paragraph blockId="49.[190,1817,359,2104]" pageId="49" pageNumber="328">
<emphasis bold="true" box="[193,670,1746,1785]" pageId="49" pageNumber="328">Status and Conservation.</emphasis>
<emphasis box="[686,760,1746,1785]" italics="true" pageId="49" pageNumber="328">Not</emphasis>
assessed on
<emphasis box="[1009,1354,1746,1785]" italics="true" pageId="49" pageNumber="328">The IUCN ed List,</emphasis>
due to its recent
<emphasis box="[1698,1814,1746,1785]" italics="true" pageId="49" pageNumber="328">recognition</emphasis>
as a species. Franciss
<emphasis box="[744,877,1798,1837]" italics="true" pageId="49" pageNumber="328">Woolly</emphasis>
Horseshoe Bat is
<emphasis box="[1230,1404,1798,1837]" italics="true" pageId="49" pageNumber="328">currently</emphasis>
known from
<emphasis box="[1669,1752,1798,1837]" italics="true" pageId="49" pageNumber="328">only</emphasis>
six records, two in Sabah and three in Kalimantan, in Borneo, and one in
<collectingCountry box="[1585,1764,1851,1890]" name="Thailand" pageId="49" pageNumber="328">Thailand</collectingCountry>
. It is probably more
<emphasis box="[539,662,1904,1943]" italics="true" pageId="49" pageNumber="328">widely</emphasis>
distributed, and should be considered data deficient until more information is available. The species is probably threatened by habitat destruction throughout its range, and perhaps by hunting in some areas.
</paragraph>
</subSubSection>
<subSubSection box="[195,1352,2070,2097]" pageId="49" pageNumber="328" type="reference_group">
<paragraph blockId="49.[190,1817,359,2104]" box="[195,1352,2070,2097]" pageId="49" pageNumber="328">
<emphasis bold="true" box="[195,401,2070,2097]" pageId="49" pageNumber="328">Bibliography.</emphasis>
<emphasis box="[413,531,2070,2097]" italics="true" pageId="49" pageNumber="328">Soisook</emphasis>
, Niyomwan et al. (2010), Soisook, Struebig et al. (2015).
</paragraph>
</subSubSection>
</treatment>
</document>