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<document ID-DOI="http://doi.org/10.5281/zenodo.3748525" ID-GBIF-Dataset="6384b520-7e9f-4874-a414-76c2e9b01d74" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3748525" approvalRequired="1" checkinTime="1586358796837" checkinUser="plazi" docAuthor="Burgin, Connor" docDate="2019" docId="885887A2FFFC8A1AF8B1FD56F286D6A0" docLanguage="en" docName="hbmw_9_Rhinolophidae.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Rhinolophus beddomei K. Andersen 1905" docType="treatment" docVersion="21" lastPageNumber="330" masterDocId="7461FFDAFFCF8A29FFCCFFA1FF85D963" masterDocTitle="Rhinolophidae" masterLastPageNumber="332" masterPageNumber="280" pageNumber="330" updateTime="1658949228820" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhinolophidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burgin, Connor</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Mittermeier, Russell A.</mods:namePart>
</mods:name>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>332</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3748525</mods:identifier>
<mods:identifier type="GBIF-Dataset">6384b520-7e9f-4874-a414-76c2e9b01d74</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3748525</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3809044" ID-GBIF-Taxon="163549925" ID-Zenodo-Dep="3809044" LSID="urn:lsid:plazi:treatment:885887A2FFFC8A1AF8B1FD56F286D6A0" httpUri="http://treatment.plazi.org/id/885887A2FFFC8A1AF8B1FD56F286D6A0" lastPageNumber="330" pageId="51" pageNumber="330">
<subSubSection box="[1917,2999,759,820]" pageId="51" pageNumber="330" type="vernacular_names">
<paragraph blockId="51.[1912,3454,759,969]" box="[1917,2999,759,820]" pageId="51" pageNumber="330">
<figureCitation box="[1917,2018,759,820]" captionStart="On" captionStartId="46.[166,208,4390,4417]" captionTargetBox="[152,3486,163,4684]" captionTargetPageId="45" captionText="On following pages: 94. Thomas's Horseshoe Bat (Rhinolophus thomasi); 95. Lesser Horseshoe Bat (Rhinolophus hipposideros); 96. rancis's Woolly Horseshoe Bat (Rhinolophus trancisi); 97. Lesser Woolly Horseshoe Bat (Rhinolophus sedulus); 98. Trefoil Horseshoe Bat (Rhinolophus trifoliatus); 99. Northern Woolly Horseshoe Bat (Rhinolophus perniger); 100. Selangor Woolly Horseshoe Bat (Rhinolophus luctoides); 101. Malaysian Woolly Horseshoe Bat (Rhinolophus mono); 102. Great Woolly Horseshoe Bat (Rhinolophus luctus); 1.03. Beddome's Woolly Horseshoe Bat (Rhinolophus beddomei); 104. Formosan Woolly Horseshoe Bat (Rhinolophus formosaë); 105. Thailand Horseshoe Bat (Rhinolophus thailandensis); 106. Dobson's Horseshoe Bat (Rhinolophus yunanensis); 107. Chiew Kwee's Horseshoe Bat (Rhinolophus chiewkweeae); 108. Pearson's Horseshoe Bat (Rhinolophus pearsonii); 109. Mitred Horseshoe Bat (Rhinolophus mitratus)." figureDoi="http://doi.org/10.5281/zenodo.3748537" httpUri="https://zenodo.org/record/3748537/files/figure.png" pageId="51" pageNumber="330">103</figureCitation>
. Beddomes Woolly Horseshoe Bat
</paragraph>
</subSubSection>
<subSubSection box="[1913,2481,814,875]" pageId="51" pageNumber="330" type="nomenclature">
<paragraph blockId="51.[1912,3454,759,969]" box="[1913,2481,814,875]" pageId="51" pageNumber="330">
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[1913,2481,814,875]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata" rank="species" species="beddomei">
<emphasis box="[1913,2481,814,875]" italics="true" pageId="51" pageNumber="330">Rhinolophus beddomei</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="vernacular_names">
<paragraph blockId="51.[1912,3454,759,969]" box="[1913,3453,886,913]" pageId="51" pageNumber="330">
French:
<emphasis box="[2027,2178,886,913]" italics="true" pageId="51" pageNumber="330">Rhinolophe</emphasis>
de Beddome /
<emphasis bold="true" box="[2388,2511,886,913]" pageId="51" pageNumber="330">German:</emphasis>
Beddome-Wollhaar-Hufeisennase /
<emphasis bold="true" box="[2999,3123,886,913]" pageId="51" pageNumber="330">Spanish:</emphasis>
Herradura
<emphasis box="[3278,3312,886,913]" italics="true" pageId="51" pageNumber="330">de</emphasis>
Beddome
</paragraph>
<paragraph blockId="51.[1912,3454,759,969]" box="[1912,2628,942,969]" pageId="51" pageNumber="330">
<emphasis bold="true" box="[1912,2246,942,969]" pageId="51" pageNumber="330">Other common names:</emphasis>
Bedomme's Horseshoe Bat
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="reference_group">
<paragraph blockId="51.[2729,3540,1037,1609]" pageId="51" pageNumber="330">
Taxonomy.
<taxonomicName authority="K. Andersen, 1905" authorityName="K. Andersen" authorityYear="1905" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata" rank="species" species="beddomei">
<emphasis box="[2977,3380,1037,1076]" italics="true" pageId="51" pageNumber="330">Rhinolophus beddomei</emphasis>
K. Andersen, 1905
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="materials_examined">
<paragraph blockId="51.[2729,3540,1037,1609]" pageId="51" pageNumber="330">
<materialsCitation ID-GBIF-Occurrence="2597534027" country="India" location="Wynaad [= Wayanad], Mysore [= Mysuru], S [outh]" pageId="51" pageNumber="330" specimenCount="1" typeStatus="type">
<location LSID="urn:lsid:plazi:treatment:885887A2FFFC8A1AF8B1FD56F286D6A0:052E606FFFFC8A1AF435FBE0F34FDDFD" country="India" name="Wynaad [= Wayanad], Mysore [= Mysuru], S [outh]" pageId="51" pageNumber="330">
<emphasis box="[3065,3219,1089,1128]" italics="true" pageId="51" pageNumber="330">Wynaad</emphasis>
[= Wayanad], Mysore [=
<emphasis box="[2937,3090,1143,1182]" italics="true" pageId="51" pageNumber="330">Mysuru</emphasis>
], S[outh]
</location>
.
<collectingCountry box="[3295,3403,1143,1182]" name="India" pageId="51" pageNumber="330">India</collectingCountry>
.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="discussion">
<paragraph blockId="51.[2729,3540,1037,1609]" pageId="51" pageNumber="330">
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[2737,3123,1197,1236]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata" rank="species" species="beddomei">
<emphasis box="[2737,3123,1197,1236]" italics="true" pageId="51" pageNumber="330">Rhinolophus beddomei</emphasis>
</taxonomicName>
is included in the
<emphasis box="[3491,3540,1197,1236]" italics="true" pageId="51" pageNumber="330">
<emphasis box="[2729,2866,1249,1288]" italics="true" pageId="51" pageNumber="330">trifoliatus</emphasis>
</emphasis>
species group and appears to be close to
<emphasis box="[2893,3322,1303,1343]" italics="true" pageId="51" pageNumber="330">
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[2893,3114,1303,1342]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata" rank="species" species="trifoliatus">R. trifoliatus</taxonomicName>
,
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[3135,3316,1303,1343]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata" rank="species" species="sedulus">R.sedulus</taxonomicName>
,
</emphasis>
and
<emphasis box="[3423,3539,1303,1342]" italics="true" pageId="51" pageNumber="330">
R.
<emphasis box="[2739,2796,1356,1395]" italics="true" pageId="51" pageNumber="330">luctus.</emphasis>
</emphasis>
Two subspecies are recognized.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.3750118" ID-Zenodo-Dep="3750118" httpUri="https://zenodo.org/record/3750118/files/figure.png" inLine="true" pageId="51" pageNumber="330" targetBox="[1910,2704,1044,1602]" targetPageId="51">
<paragraph blockId="51.[2729,3540,1037,1609]" box="[2736,3276,1411,1450]" pageId="51" pageNumber="330">
<emphasis bold="true" box="[2736,3276,1411,1450]" pageId="51" pageNumber="330">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="51.[2729,3540,1037,1609]" pageId="51" pageNumber="330">
<taxonomicName authority="K. Andersen, 1905" authorityName="K. Andersen" authorityYear="1905" box="[2737,3340,1462,1501]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata" rank="species" species="beddomei">
<emphasis box="[2737,2815,1462,1501]" italics="true" pageId="51" pageNumber="330">R. b.</emphasis>
<emphasis box="[2830,2985,1462,1501]" italics="true" pageId="51" pageNumber="330">beddomei</emphasis>
K. Andersen, 1905
</taxonomicName>
— W &amp; E
<collectingCountry box="[2736,2837,1516,1555]" name="India" pageId="51" pageNumber="330">India</collectingCountry>
(
<collectingRegion box="[2884,3128,1516,1555]" country="India" name="Maharashtra" pageId="51" pageNumber="330">Maharashtra</collectingRegion>
,
<collectingRegion box="[3166,3364,1516,1555]" country="India" name="Karnataka" pageId="51" pageNumber="330">Karnataka</collectingRegion>
,
<collectingRegion box="[3402,3529,1516,1555]" country="India" name="Kerala" pageId="51" pageNumber="330">Kerala</collectingRegion>
, and
<collectingRegion box="[2818,3139,1570,1609]" country="India" name="Andhra Pradesh" pageId="51" pageNumber="330">Andhra Pradesh</collectingRegion>
).
</paragraph>
<paragraph blockId="51.[1911,3538,1623,4035]" box="[1913,2740,1623,1663]" pageId="51" pageNumber="330">
<taxonomicName box="[1913,2506,1623,1662]" class="Mammalia" family="Rhinolophidae" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="330" phylum="Chordata">
<emphasis box="[1913,2151,1623,1662]" italics="true" pageId="51" pageNumber="330">R.b. sobrinus</emphasis>
K Andersen, 1918
</taxonomicName>
-
<collectingCountry box="[2551,2735,1623,1662]" name="Sri Lanka" pageId="51" pageNumber="330">Sri Lanka</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="description">
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
Descriptive
<emphasis bold="true" box="[2146,2259,1674,1713]" pageId="51" pageNumber="330">notes.</emphasis>
Head-body
<quantity box="[2515,2723,1674,1713]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="51" pageNumber="330" unit="mm" value="70.0" valueMax="75.0" valueMin="65.0">65-75 mm</quantity>
, tail
<quantity box="[2823,3031,1674,1713]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.8" metricValueMin="3.7" pageId="51" pageNumber="330" unit="mm" value="42.5" valueMax="48.0" valueMin="37.0">37-48 mm</quantity>
, ear
<quantity box="[3132,3341,1674,1713]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.4" metricValueMin="2.1" pageId="51" pageNumber="330" unit="mm" value="27.5" valueMax="34.0" valueMin="21.0">21-34 mm</quantity>
, hindfoot 12-
<quantity box="[1982,2156,1728,1767]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="51" pageNumber="330" unit="mm" value="9.5" valueMax="18.0" valueMin="1.0">1-18 mm</quantity>
, forearm 54-9—64-
<quantity box="[2515,2627,1728,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="51" pageNumber="330" unit="mm" value="3.0">3 mm</quantity>
. Fur is long and
<emphasis box="[2952,3079,1728,1767]" italics="true" pageId="51" pageNumber="330">woolly</emphasis>
; dorsal and ventral pelage is dark brown or black (hairs with paler tips). Ears are blackish and of medium length. Noseleaf is dark brown or blackish, and has long and pointed or rounded lancet; connecting process is relatively low and broadly rounded; sella is broad and
<emphasis box="[1911,2034,1941,1980]" italics="true" pageId="51" pageNumber="330">widely</emphasis>
rounded off with large circular lateral lappets on either side of base; horseshoe is very wide (
<quantity box="[2283,2432,1994,2033]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="51" pageNumber="330" unit="mm" value="11.0">11 mm</quantity>
), covering muzzle, and has wide median emargination. Lower lip has one mental groove. Skull is robust (zygomatic breadth is larger than mastoid breadth); anterior median swellings are elongated but do not expand anteriorly to upper edge of nasal orifice, formed by a protruding bony rim; posterior swellings are more laterally situated; frontal depression is deep and long; supraorbital crests are sharp; sagittal crest is very well developed; canines are strong but relatively short. P2 is medium-sized and within tooth row or slightly displaced; P4 is small to medium-sized and extruded from the tooth row to a variable degree; P2 and P4 are almost in contact. Chromosomal complement has 2n = 32 and FN = 60.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="biology_ecology">
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
Habitat. Mainly found in dense dry and tropical moist forests. Elevational range is from sea level up to elevations of
<quantity box="[2547,2686,2522,2561]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="51" pageNumber="330" unit="m" value="1070.0">1070 m</quantity>
.
</paragraph>
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
<emphasis bold="true" box="[1914,2094,2578,2617]" pageId="51" pageNumber="330">Food and</emphasis>
Feeding. Beddomes
<emphasis box="[2496,2628,2578,2617]" italics="true" pageId="51" pageNumber="330">Woolly</emphasis>
Horseshoe Bat feeds on a variety of insects
<emphasis box="[3441,3535,2578,2617]" italics="true" pageId="51" pageNumber="330">especially</emphasis>
beetles and termites. It flies low over the ground while foraging, and will forage close to vegetation.
</paragraph>
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
Breeding. A pregnant female with one embryo was collected in mid-January in
<collectingCountry name="Sri Lanka" pageId="51" pageNumber="330">Sri Lanka</collectingCountry>
.
<emphasis box="[2055,2184,2789,2828]" italics="true" pageId="51" pageNumber="330">Young</emphasis>
have been reported with females in
<collectingRegion box="[2892,3023,2789,2828]" country="India" name="Kerala" pageId="51" pageNumber="330">Kerala</collectingRegion>
,
<collectingCountry box="[3042,3149,2789,2828]" name="India" pageId="51" pageNumber="330">India</collectingCountry>
, in
<emphasis bold="true" box="[3220,3390,2789,2828]" pageId="51" pageNumber="330">mid-May</emphasis>
, and in
<collectingCountry box="[1915,2106,2842,2881]" name="Sri Lanka" pageId="51" pageNumber="330">Sri Lanka</collectingCountry>
in mid-March and late April. Females may not breed until two or three years old.
</paragraph>
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
Activity
<emphasis bold="true" box="[2070,2241,2948,2987]" pageId="51" pageNumber="330">patterns.</emphasis>
Beddomes
<emphasis box="[2483,2616,2948,2987]" italics="true" pageId="51" pageNumber="330">Woolly</emphasis>
Horseshoe
<emphasis bold="true" box="[2851,2917,2948,2987]" pageId="51" pageNumber="330">Bat</emphasis>
roosts by day in small caves, dilapidated buildings, tree hollows, wells, culverts, and old and unused tunnels, or clings from overhanging rocky outcrops. It emerges late in the evening from its roosts to forage through the night. Call shape is FM/CF/FM with a peak F recorded at 38-5—38-7 kHz and durations of 48-2—58 milliseconds in the
<emphasis box="[3076,3234,3155,3194]" italics="true" pageId="51" pageNumber="330">Western</emphasis>
Ghats of
<collectingCountry box="[3420,3526,3155,3194]" name="India" pageId="51" pageNumber="330">India</collectingCountry>
; another study in the
<emphasis box="[2341,2499,3210,3249]" italics="true" pageId="51" pageNumber="330">Western</emphasis>
Ghats reported peak frequencies of 41-7-43-3 kHz (mean 42-8 kHz) and durations of 1-4-51-5 milliseconds (mean 25-2 milliseconds).
<emphasis box="[1913,1946,3317,3356]" italics="true" pageId="51" pageNumber="330">A</emphasis>
recording inside a cave in
<collectingCountry box="[2481,2670,3317,3356]" name="Sri Lanka" pageId="51" pageNumber="330">Sri Lanka</collectingCountry>
produced peak frequencies of 41—62-5 kHz but this huge range must be an error, perhaps including more than one species.
</paragraph>
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
<emphasis bold="true" box="[1915,2484,3423,3462]" pageId="51" pageNumber="330">Movements, Home range and</emphasis>
Social
<emphasis bold="true" box="[2625,2868,3423,3462]" pageId="51" pageNumber="330">organization.</emphasis>
Beddomes
<emphasis box="[3108,3241,3423,3462]" italics="true" pageId="51" pageNumber="330">Woolly</emphasis>
Horseshoe Bat roosts alone or in pairs, or sometimes in groups of three or four individuals.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="conservation">
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
<emphasis bold="true" box="[1915,2110,3528,3567]" pageId="51" pageNumber="330">Status and</emphasis>
Conservation. Classified as Least Concern on
<emphasis box="[2979,3231,3528,3567]" italics="true" pageId="51" pageNumber="330">TheIUCNRed</emphasis>
List. Beddomes
<emphasis box="[1913,2045,3580,3619]" italics="true" pageId="51" pageNumber="330">Woolly</emphasis>
Horseshoe Bat is relatively widespread and common throughout much of its distribution. However, in
<collectingCountry box="[2411,2598,3633,3672]" name="Sri Lanka" pageId="51" pageNumber="330">Sri Lanka</collectingCountry>
the species is regionally listed as Vulnerable according to the National Red List for the
<emphasis box="[2672,2817,3686,3725]" italics="true" pageId="51" pageNumber="330">country</emphasis>
, and is mostly restricted to old-growth forests with small populations. The species appears to be threatened most
<emphasis bold="true" box="[3295,3340,3739,3778]" pageId="51" pageNumber="330">by</emphasis>
deforestation for logging and agricultural expansion.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="330" type="reference_group">
<paragraph blockId="51.[1911,3538,1623,4035]" pageId="51" pageNumber="330">
<emphasis bold="true" box="[1917,2123,3850,3877]" pageId="51" pageNumber="330">Bibliography.</emphasis>
Bates &amp; Harrison (1997), Csorba
<emphasis box="[2626,2677,3850,3877]" italics="true" pageId="51" pageNumber="330">eta</emphasis>
/. (2003), Edirisinghe et
<emphasis box="[3037,3068,3850,3877]" italics="true" pageId="51" pageNumber="330">al.</emphasis>
(2016), Koubfnovà et
<emphasis box="[3393,3424,3850,3877]" italics="true" pageId="51" pageNumber="330">al.</emphasis>
(2010), Kusuminda et al. (2013), Molur et
<emphasis box="[2417,2448,3903,3930]" italics="true" pageId="51" pageNumber="330">al.</emphasis>
(2002), Naidu &amp; Gururaj (1984), Raghuram et
<emphasis box="[3130,3161,3903,3930]" italics="true" pageId="51" pageNumber="330">al.</emphasis>
(2014), Rao et al. (2004), Sinha (1973), Soisook, Niyomwan et al. (2010), Soisook, Struebig et al. (2015), Srinivasulu &amp; Molur (2008b), Srinivasulu &amp; Srinivasulu (2012),Topàl &amp; Csorba (1992), Volleth et al. (2017), Wordley
<emphasis box="[3155,3186,4008,4035]" italics="true" pageId="51" pageNumber="330">et</emphasis>
al. (2014).
</paragraph>
</subSubSection>
</treatment>
</document>