138 lines
18 KiB
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138 lines
18 KiB
XML
<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="8831BC71A3445ADD8A6E49F4F37F8A27" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupinella sonlaensis Jirapatrasilp & Sutcharit & Panha 2022, comb. nov." docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:mods id="C318FC092061D7F962CD6F416FA51240" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9B6EA0D0972B53874D0CD056AF9BD0CD">
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<mods:title id="47A7ED0BEB6C197AF4BD318768B7E341">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:name id="1329B11464A551A017706CF84DA1E1F3" type="personal">
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<mods:roleTerm id="5760A54A805D818EA7FE63ACE7ACFCB9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="79184FC17EFE1941ED38CBCB88268A8B">Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier id="6268BD6277B525E080036002F2683E4B" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation id="449AFEB8C49E5E056FB79DE71F36E41E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="7A754AD679F84F760E64C2D99A9D203E" type="personal">
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<mods:roleTerm id="D4FF555F32C9C73BC4D91CCFA1FCF1ED">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3CF2721BEBFF6DD431469DB757D756C5">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="175926DC66EE2DB3299A5E114F859FD3">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="3D4C79FFAC6070949C752BCD0E71A949" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:name id="6978CF53906A4942EA77DC90807F51F7" type="personal">
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<mods:roleTerm id="0FF3F218F6EABC39367391982700AA4F">Author</mods:roleTerm>
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<mods:namePart id="271B2904C1E3F8C18B28129AF6BA186A">Panha, Somsak</mods:namePart>
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<mods:affiliation id="C2396D1B01879BACE2C82FD1C313AC28">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="4E70245B864E1498FA6B47F45C75F7DE" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource id="31F0AB57BEE3A0092063F29EA367959E">text</mods:typeOfResource>
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<mods:title id="A8DB1DBA2A080D69F33D0385FEBECDC5">ZooKeys</mods:title>
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<mods:part id="604B433EA6BD89C879A8B52216087C76">
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<mods:date id="5C9E3ED84022265832144DB1D440BA8E">2022</mods:date>
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<mods:number id="E7B121D923996D133B55ECC46ED9EF17">2022-08-25</mods:number>
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<mods:detail id="49D851092BDFED52D072AAD731669D01" type="volume">
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<mods:number id="F35950340197A188002BE7C99FCF618D">1119</mods:number>
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<mods:url id="7E14B738049FCCFB248B46099488E957">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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<mods:classification id="1742198E3B218FD816DBC30FC7D6F168">journal article</mods:classification>
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<mods:identifier id="2751E7BE51C9E59C2521B36682B0B46E" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier id="30EECAA57B742D36169C75CB75260449" type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<treatment id="8831BC71A3445ADD8A6E49F4F37F8A27" ID-GBIF-Taxon="199839068" LSID="urn:lsid:plazi:treatment:8831BC71A3445ADD8A6E49F4F37F8A27" httpUri="http://treatment.plazi.org/id/8831BC71A3445ADD8A6E49F4F37F8A27" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupinellinae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="3D8652E24C0342F7AE6AB5D9037E72A0" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="2DA7B52343C9AFD17BA6826322A32516" pageId="0" pageNumber="1">
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<taxonomicName id="5485C4DB159B6611009C29C44FBAF7BD" ID-CoL="9GG72" LSID="8831BC71-A344-5ADD-8A6E-49F4F37F8A27" authority="(Do, 2017)" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Do" baseAuthorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis" status="comb. nov.">Pupinella sonlaensis (Do, 2017)</taxonomicName>
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<taxonomicNameLabel id="5A1E2A16020BA5A1024ACC20CD087774" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="DE0C5F15EDA25A19CD99082154B7B7DB" pageId="0" pageNumber="1" type="description">
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<paragraph id="CF3BE395E981C8682BC7AE367EDA169D" pageId="0" pageNumber="1">
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<figureCitation id="68F281637DC1811DAE6ECD7B25A30C91" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">Figs 11H</figureCitation>
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<figureCitation id="DE96D73E5BDC3DBB57C02CC68C3963B8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">, 12F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="587CE00C64BF94D0508EAED81D9197F1" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="0226AC1AB5CEB262E9A9861FF4988F7E" pageId="0" pageNumber="1">
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<taxonomicName id="C6A53BB71F7179175E2E2E724A40E29E" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis" status="comb. nov.">Pupina sonlaensis</taxonomicName>
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Do, 2017: 300, 302, figs 2a, 3a. Type locality: limestone karst in Muong Bu Commune, Muong La District, Son La Province, Vietnam.
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</paragraph>
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</subSubSection>
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<subSubSection id="22B2033EF177E4FCE03F576FBEEE3461" pageId="0" pageNumber="1" type="type material">
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<paragraph id="1E8F64F7AE6D52C42E864338C0629435" pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph id="B01D59E80ED4131F7C0BBE3BE237A0D0" pageId="0" pageNumber="1">
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<emphasis id="9552493176EB99E38B8C83F03F821986" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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HNUE-OC 00108 figured in
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<bibRefCitation id="5348349A30A7F717EC96AA3479D45DA3" author="Do, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="299 - 303" refId="B33" refString="Do, DS, 2017. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology 65: 299 - 303" title="Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam." volume="65" year="2017">Do (2017</bibRefCitation>
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: figs 2a, 3a).
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<emphasis id="278970DB17D157EB07A87CCCCC8F2B21" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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ZRC.MOL.9377 (3 shells; Figs
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<figureCitation id="A7E6EAF5B585E58857064BC6012A7883" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11H</figureCitation>
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,
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<figureCitation id="79EA5E40FFFF77D2BCD95DF855F086A1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">12F</figureCitation>
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) from the type locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="916D7CE28CE8BEBFD328F0D35FB034CC" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="0E326E0AC53F81A5E64DEAB642D299D0" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="B4AFFC6B7B0E71220E6EA63636ED4113" pageId="0" pageNumber="1">Shell ovate-fusiform; last whorl ca. 60% of shell height. Apertural lip slightly expanded and reflected, thickened cord-like peristome absent; apertural lip when observed from lateral view almost straight. Parietal callus somewhat distinct and cord-like. Parietal tooth sharp with wide base, thickened and covering posterior canal. Anterior canal funnel-like, appearing as a slit on the inside, widened on outer margin, bordered by a thickened columellar margin. Umbilicus closed.</paragraph>
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</subSubSection>
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<subSubSection id="980F912879D1C94C7EC179D5FFC6B634" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="A234EF9CAB350ED9FBAF2EF4792354E3" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="05E402C417F9F44DCDC5D1C8EFD6F80E" pageId="0" pageNumber="1">
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<taxonomicName id="E6C398A13B7C852C054FAF6C2945DF8E" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Do" baseAuthorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis">
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<emphasis id="349BB12E7BF0525FA881EC6E25B1A5C8" italics="true" pageId="0" pageNumber="1">Pupinella sonlaensis</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName id="1A869D7E6BF23632A4AEED40A36947F3" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
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<emphasis id="A3B7F36D680CB75131A1E11D884446A6" italics="true" pageId="0" pageNumber="1">P. mansuyi</emphasis>
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</taxonomicName>
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in shell size, but differs in having an ovate-fusiform shell shape with a less thickened parietal tooth, as well as a less thickened, expanded, and reflected apertural lip without a thickened cord-like inner peristome.
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</paragraph>
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</subSubSection>
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<subSubSection id="EBC7D69A7982B6D75A5A5DED64A55F93" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="30410884923BACF394CEB64BD35E2596" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="6A4D8C4C9D177C9B3EE71DC7903DD2BD" pageId="0" pageNumber="1">
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Muong La District, Thuan Chau District, and Van Ho District, Son La Province, Vietnam (
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<bibRefCitation id="7FF5CAAD92F36E36944AF9D5D2CCF9A0" author="Do, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="299 - 303" refId="B33" refString="Do, DS, 2017. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology 65: 299 - 303" title="Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam." volume="65" year="2017">Do 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="18F9A696D8F980257DB5E3CD2AEF2DF6" pageId="0" pageNumber="1" type="remarks">
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<paragraph id="3F4585E07902DDA2A8FEAA8B04B15782" pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph id="570BA40F10F8EECC3BA63E57EB85894E" pageId="0" pageNumber="1">
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This taxon is allocated to the genus
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<taxonomicName id="F3A677E7EEEC733EDC96BC4C9142CF60" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis id="0E9F03C574E7B2C749CB35073B5BDC65" italics="true" pageId="0" pageNumber="1">Pupinella</emphasis>
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</taxonomicName>
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due to the presence of a funnel-like anterior canal, which is the diagnostic character of this genus. The paratype figured in this study is similar to
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<taxonomicName id="B325EF1464694D0C60EF79AFA0A5FAD2" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
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<emphasis id="D027B47CBD3545FAA518C717C410D9E3" italics="true" pageId="0" pageNumber="1">P. mansuyi</emphasis>
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</taxonomicName>
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in having a triangular parietal tooth covering the posterior canal and an expanded and reflected apertural lip with somewhat cord-like inner peristome, although with less thickening, and the shell has a less elongate shape. However, the holotype of
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<taxonomicName id="254763E92D06FA78A98042697EEBD58F" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis">
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<emphasis id="AC65AA9E4FE1120A6720AD2729CE4763" italics="true" pageId="0" pageNumber="1">P. sonlaensis</emphasis>
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</taxonomicName>
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figured in
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<bibRefCitation id="50DDC0FED6F7345C77BB83F3DB5E9ACE" author="Do, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="299 - 303" refId="B33" refString="Do, DS, 2017. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology 65: 299 - 303" title="Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam." volume="65" year="2017">Do (2017</bibRefCitation>
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: figs 2a, 3a) has an ovate-fusiform shell with a thickened, wide-based parietal tooth not covering the posterior canal, and a slightly expanded and reflected apertural lip without a thickened cord-like inner peristome. A thorough examination of the specimens would clarify whether the type series contain more than one taxon or whether the validity of this taxon should be reassessed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |