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128 lines
10 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.352.6052" ID-GBIF-Dataset="99647020-9754-4dd3-8bf9-0db3657c1b5f" ID-PMC="PMC3837398" ID-Pensoft-Pub="1313-2970-352-51" ID-PubMed="24294092" ID-ZBK="FD8FE06F82C341D89C9512943B957BC6" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-352-51" ModsDocOrigin="ZooKeys 352" ModsDocTitle="A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus" checkinTime="1451246663146" checkinUser="pensoft" docAuthor="Sokolov, Igor M." docDate="2013" docId="8803DE8B77A821B2443C4F318824AC1E" docLanguage="en" docName="ZooKeys 352: 51-92" docOrigin="ZooKeys 352" docSource="http://dx.doi.org/10.3897/zookeys.352.6052" docTitle="Zapotecanillus nanus Sokolov, 2013, sp. n." docType="treatment" docUuid="E3677571-83C4-4836-9252-BC1B5C11129E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="61" masterDocId="FFFCE9690B65E958B312FF9EFFA8B926" masterDocTitle="A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus" masterLastPageNumber="92" masterPageNumber="51" pageNumber="59" updateTime="1668156898856" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>352</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>92</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.352.6052</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.352.6052</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-352-51</mods:identifier>
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<mods:identifier type="ZBK">FD8FE06F82C341D89C9512943B957BC6</mods:identifier>
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<mods:identifier type="ZooBank">FD8FE06F82C341D89C9512943B957BC6</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152049654" LSID="urn:lsid:zoobank.org:act:E3677571-83C4-4836-9252-BC1B5C11129E" httpUri="http://treatment.plazi.org/id/8803DE8B77A821B2443C4F318824AC1E" lastPageId="10" lastPageNumber="61" pageId="8" pageNumber="59">
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<subSubSection pageId="8" pageNumber="59" type="nomenclature">
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<paragraph pageId="8" pageNumber="59">
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<taxonomicName LSID="http://zoobank.org/E3677571-83C4-4836-9252-BC1B5C11129E" class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus nanus" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="nanus">Zapotecanillus nanus</taxonomicName>
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<taxonomicNameLabel pageId="8" pageNumber="59">sp. n.</taxonomicNameLabel>
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Figs 19, 20, 25, 39-41, 61, 69, 73, 77, 90-92, 94
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="59" type="type material">
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<paragraph pageId="8" pageNumber="59">Type material.</paragraph>
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<paragraph pageId="8" pageNumber="59">
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HOLOTYPE, male, in NMNH, point-mounted, labeled: \ MEX. Oax., 15mi. S. Valle Nacional, 4000' 21.V.1971 S.Peck Ber.204, leaf litter \ Borrowed Ex. J.M. Campbell \ USNMNH 2051867\. PARATYPES (27 ex., 4♂4♀ were dissected), 4 ex. labeled same as holotype, except one specimen, which has an additional label: \
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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n. sp. det. T.L.Erwin 76\, where italicized font means handwriting.; 12 ex. labeled: MEXICO: Oaxaca, 15mi. S. Valle Nacional, 4000' 21 May 1971, leaf litter, S. Peck \ THOMAS C. BARR COLLECTION 2011 Acc. No. 38014\; 1 ex. labeled: \ MEXICO: Oaxaca 15.1mi S Valle Nacional 4300' VIII-11-18-1973 \ under bark hardwood A.Newton \ Borrowed ex. MCZ \ Loan from USNMNH 2051867\; 10 ex. labeled: \ MEXICO. Oaxaca 18.7mi S Valle Nacional 5200' 17 Aug.1973\ A.Newton Collector \ Loan from USNMNH 2051867\ (deposited in CAS, CMNH, NMNH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="59" type="specific epithet">
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<paragraph pageId="8" pageNumber="59">Specific epithet.</paragraph>
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<paragraph pageId="8" pageNumber="59">The specific epithet is a Latin adjective, nanus, in the masculine form, meaning dwarf, miniature, and refers to the small size of the beetles.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="59" type="type locality">
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<paragraph pageId="8" pageNumber="59">Type locality.</paragraph>
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<paragraph pageId="8" pageNumber="59">Mexico, Oaxaca, 15mi S from Valle Nacional.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="59" type="recognition">
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<paragraph pageId="8" pageNumber="59">Recognition.</paragraph>
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<paragraph pageId="8" pageNumber="59">Adults of this new species are distinguished from those of other species of the genus by the combination of small size and brunneorufous color; and males can be further distinguished by the shape of the median lobe (Fig. 39).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60" type="description">
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<paragraph pageId="9" pageNumber="60">
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<pageBreakToken pageId="9" pageNumber="60" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="60">
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Size. Small for genus (SBL range 1.02-1.16 mm, mean 1.10
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.049 mm, n=12).
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</paragraph>
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<paragraph pageId="9" pageNumber="60">
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Habitus. Body form (Fig. 25) moderately convex, slightly elongate (WE/SBL 0.41
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.09), head of normal proportions for genus (WH/WPm 0.75
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.016), pronotum narrow compared to elytra (WPm/WE 0.75
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.028).
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</paragraph>
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<paragraph pageId="9" pageNumber="60">Color. Body monocolorous, brunneorufous, appendages testaceous.</paragraph>
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<paragraph pageId="9" pageNumber="60">Microsculpture. Microlines partially effaced on disc of pronotum.</paragraph>
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<paragraph pageId="10" pageNumber="61">
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<pageBreakToken pageId="10" pageNumber="61" start="start">Prothorax</pageBreakToken>
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. Pronotum relatively long (LP/LE 0.43
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.020) and moderately transverse (WPm/LP 1.26
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.030), with margins straight and markedly constricted posteriorly (WPm/WPp 1.38
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.044). Basal margin slightly oblique laterally. Posterior angles small, contour of posterior angles obtuse (112-126°) without or with 1 small denticle in front of the angles.
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</paragraph>
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<paragraph pageId="10" pageNumber="61">
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Elytra. Slightly convex, not depressed along suture, comparatively wide (WE/LE 0.72
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.025). Margins rounded, moderately divergent in basal half, evenly rounded to apex in apical third, maximal width of elytra slightly behind the midpoint.
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</paragraph>
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<paragraph pageId="10" pageNumber="61">Legs. 1st male protarsomere markedly dilated apico-laterally (Fig. 20).</paragraph>
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<paragraph pageId="10" pageNumber="61">Male genitalia. Median lobe of aedeagus (Fig. 39), with small, slightly elongated apex, angulately rounded at tip. Dorsal plate 1 long, with apical pointed attenuation of moderate length. Dorsal plate 2 joined to plate 1 at its middle ventrally, where it forms a distinct protuberance. Ventral sclerites with sides divergent ventrally, trianguloid in shape. Right paramere short and moderately narrow (Fig. 41). Left paramere with distinct apical constriction (Fig. 40). Ring sclerite with long handle-like extension, pointed apically (Fig. 61).</paragraph>
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<paragraph pageId="10" pageNumber="61">Female genitalia. Gonocoxite 2 comparatively short, with slightly curved blade and narrowly rounded apex (Fig. 69). Laterotergite with 5-6 setae. Spermatheca standard for genus (Fig. 73).</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="61" type="geographical distribution">
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<paragraph pageId="10" pageNumber="61">Geographical distribution.</paragraph>
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<paragraph pageId="10" pageNumber="61">
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The species is known from a few localities in the Sierra de
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<normalizedToken originalValue="Juárez">Juarez</normalizedToken>
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Range, a part of the Sierra Madre de Oaxaca, along the ~5km stretch of the Rio de Valle Nacional (Figs 77 and 94, white triangles).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="61" type="way of life">
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<paragraph pageId="10" pageNumber="61">Way of life.</paragraph>
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<paragraph pageId="10" pageNumber="61">According to the label data, the elevations of localities range from 4000' to 5200' (1200-1600 m).</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="61" type="relationships">
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<paragraph pageId="10" pageNumber="61">Relationships.</paragraph>
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<paragraph pageId="10" pageNumber="61">
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Aedeagal characters (shape of dorsal plates and left paramere) suggest that
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<taxonomicName class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus nanus" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="nanus">Zapotecanillus nanus</taxonomicName>
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is most closely related to the sympatric
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<taxonomicName class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus oaxacanus" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="oaxacanus">Zapotecanillus oaxacanus</taxonomicName>
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. See also Fig. 90 for cladistic affinities.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |