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<document id="C5F69A7888A21A6D58D104BA0103C7CB" ID="10.11646/zootaxa.4169.3.7" ID-DOI="10.11646/zootaxa.4169.3.7" ID-GBIF-Dataset="faeafa7a-a105-4443-bbc3-2bea598d8cd5" ID-ISSN="1175-5326" ID-Zenodo-Dep="258027" ID-ZooBank="8A1FB454-AC73-4D54-A5CB-D12D6CD82AC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474440730591" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2016" docId="8706BE21FFB9FF9B1DC8FE7CFD1EA048" docLanguage="en" docName="zootaxa.4169.3.7.pdf" docOrigin="Zootaxa 4169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexachaetus lateralis subsp. lateralis Guerin" docType="treatment" docVersion="7" lastPageNumber="543" masterDocId="7B3FC659FFBAFF981D5FFF88FFF0A546" masterDocTitle="Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)" masterLastPageNumber="554" masterPageNumber="540" pageNumber="543" updateTime="1698386596725" updateUser="plazi">
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<mods:title id="712C77A13CF5683DA4FB540B59347AD9">Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)</mods:title>
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<mods:namePart id="D9DD086E447808A51D10A26371617EB1">Tian, Mingyi</mods:namePart>
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<mods:namePart id="7E66BFA446F24E534F30325E6B2D95BA">Deuve, Thierry</mods:namePart>
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<mods:date id="9A96F699D143D4D9EB8ED938B6B14C11">2016</mods:date>
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<treatment id="8706BE21FFB9FF9B1DC8FE7CFD1EA048" ID-DOI="http://doi.org/10.5281/zenodo.6075817" ID-GBIF-Taxon="124552514" ID-Zenodo-Dep="6075817" LSID="urn:lsid:plazi:treatment:8706BE21FFB9FF9B1DC8FE7CFD1EA048" httpUri="http://treatment.plazi.org/id/8706BE21FFB9FF9B1DC8FE7CFD1EA048" lastPageNumber="543" pageId="3" pageNumber="543">
<subSubSection id="47B55CBCFFB9FF9B1DC8FE7CFEF7A769" pageId="3" pageNumber="543" type="nomenclature">
<paragraph id="0F100F37FFB9FF9B1DC8FE7CFD35A748" blockId="3.[151,709,500,559]" box="[151,709,500,526]" pageId="3" pageNumber="543">
<heading id="5458B85BFFB9FF9B1DC8FE7CFD35A748" bold="true" box="[151,709,500,526]" fontSize="11" level="1" pageId="3" pageNumber="543" reason="1">
<taxonomicName id="C8AF74B4FFB9FF9B1DC8FE7CFD35A748" ID-CoL="7H949" authority="(Guerin), 1843" authorityName="Guerin" baseAuthorityName="Guerin" baseAuthorityYear="1843" box="[151,709,500,526]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="lateralis" subSpecies="lateralis">
<emphasis id="3DDBD325FFB9FF9B1DC8FE7CFD35A748" bold="true" box="[151,709,500,526]" pageId="3" pageNumber="543">
<emphasis id="3DDBD325FFB9FF9B1DC8FE7CFDF6A748" bold="true" box="[151,518,500,526]" italics="true" pageId="3" pageNumber="543">Hexachaetus lateralis lateralis</emphasis>
(Guérin), 1843
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="0F100F37FFB9FF9B1DC8FD9EFEF7A769" blockId="3.[151,709,500,559]" box="[151,263,534,559]" pageId="3" pageNumber="543">
(
<figureCitation id="979413B2FFB9FF9B1DC1FD9EFF0CA769" box="[158,252,534,559]" captionStart="FIGURES 1 4" captionStartId="4.[151,264,1491,1514]" captionTargetBox="[395,1186,207,1455]" captionTargetId="figure@4.[386,1201,195,1468]" captionTargetPageId="4" captionText="FIGURES 1 4. Hexachaetus lateralis Guérin (1. H. lateralis lateralis Guérin, habitus, male (photo: E. Kirschhofer); 2. H. lateralis maindroni Tian et Deuve, habitus, male, holotype; 3. Median lobe of aedeagus of H. lateralis maindroni, lateral view; 4. ibid. apical lamella, dorsal view; Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258028/files/figure.png" pageId="3" pageNumber="543">Figure 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1DC8FDD1FCCAA788" pageId="3" pageNumber="543" type="reference_group">
<paragraph id="0F100F37FFB9FF9B1DC8FDD1FD5FA729" blockId="3.[151,881,601,654]" box="[151,687,601,623]" pageId="3" pageNumber="543">
<treatmentCitationGroup id="2FBF2819FFB9FF9B1DC8FDD1FD5FA729" box="[151,687,601,623]" pageId="3" pageNumber="543">
<taxonomicName id="C8AF74B4FFB9FF9B1DC8FDD1FD5FA729" ID-CoL="6T99X" authority="Guerin, in Delessert, 1843: 35" authorityName="Guerin, in Delessert" authorityPageNumber="35" authorityYear="1843" box="[151,687,601,623]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB9FF9B1DC8FDD1FE81A729" box="[151,369,601,623]" italics="true" pageId="3" pageNumber="543">Orthogonius lateralis</emphasis>
: Guérin, in
<treatmentCitation id="8E0E2926FFB9FF9B1CB4FDD1FD5FA729" author="Delessert" box="[491,687,601,623]" page="35" pageId="3" pageNumber="543" year="1843">
<bibRefCitation id="6B3E72C6FFB9FF9B1CB4FDD1FD7CA729" author="Delessert" box="[491,652,601,623]" pageId="3" pageNumber="543" refString="Delessert, A. (1843) Souvenirs d'un voyage dans l'Inde execute de 1834 a 1839. II. Fortin-Masson, Paris, 107 pp." type="book" year="1843">Delessert, 1843</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="0F100F37FFB9FF9B1DC8FDF0FC81A7C8" blockId="3.[151,881,601,654]" box="[151,881,632,654]" pageId="3" pageNumber="543">
<treatmentCitationGroup id="2FBF2819FFB9FF9B1DC8FDF0FC81A7C8" box="[151,881,632,654]" pageId="3" pageNumber="543">
<taxonomicName id="C8AF74B4FFB9FF9B1DC8FDF0FE25A7C8" ID-CoL="3LG2F" authority="Guerin" authorityName="Guerin" box="[151,469,632,654]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB9FF9B1DC8FDF0FE84A7C8" box="[151,372,632,654]" italics="true" pageId="3" pageNumber="543">Hexachaetus lateralis</emphasis>
(Guérin)
</taxonomicName>
:
<treatmentCitation id="8E0E2926FFB9FF9B1C80FDF0FD42A7C8" author="Chaudoir" box="[479,690,632,654]" page="125" pageId="3" pageNumber="543" year="1871">
<bibRefCitation id="6B3E72C6FFB9FF9B1C80FDF0FD8FA7C8" author="Chaudoir" box="[479,639,632,654]" pageId="3" pageNumber="543" refString="Chaudoir, M. de (1871) Essai Monographique sur les Orthogoniens. Bulletin de la Societe entomologique de Belgique, 14, 95 - 130." type="journal article" year="1871">Chaudoir, 1871</bibRefCitation>
: 125
</treatmentCitation>
;
<treatmentCitation id="8E0E2926FFB9FF9B1FE3FDF0FC81A7C8" author="Csiki" box="[700,881,632,654]" page="1592" pageId="3" pageNumber="543" year="1932">
<bibRefCitation id="6B3E72C6FFB9FF9B1FE3FDF0FCC4A7C8" author="Csiki" box="[700,820,632,654]" pageId="3" pageNumber="543" refString="Csiki, E. (1932) Coleopterorum Catalogus: Carabidae III: Harpalinae VII, pars 124. Junk, W., Berlin, pp. 1279 - 1598." type="journal article" year="1932">Csiki, 1932</bibRefCitation>
: 1592
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="0F100F37FFB9FF9B1DC8FD3DFCCAA788" blockId="3.[151,1437,693,1294]" box="[151,826,693,718]" pageId="3" pageNumber="543">
Length: 17.018.0 mm; width: 7.0 mm. Habitus as in
<figureCitation id="979413B2FFB9FF9B1FADFD3DFCC4A788" box="[754,820,693,718]" captionStart="FIGURES 1 4" captionStartId="4.[151,264,1491,1514]" captionTargetBox="[395,1186,207,1455]" captionTargetId="figure@4.[386,1201,195,1468]" captionTargetPageId="4" captionText="FIGURES 1 4. Hexachaetus lateralis Guérin (1. H. lateralis lateralis Guérin, habitus, male (photo: E. Kirschhofer); 2. H. lateralis maindroni Tian et Deuve, habitus, male, holotype; 3. Median lobe of aedeagus of H. lateralis maindroni, lateral view; 4. ibid. apical lamella, dorsal view; Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258028/files/figure.png" pageId="3" pageNumber="543">Fig. 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98FD51FC0EA11C" pageId="3" pageNumber="543" type="diagnosis">
<paragraph id="0F100F37FFB9FF9B1D98FD51FC0EA11C" blockId="3.[151,1437,693,1294]" pageId="3" pageNumber="543">
<emphasis id="3DDBD325FFB9FF9B1D98FD51FEB1A7B4" bold="true" box="[199,321,729,754]" pageId="3" pageNumber="543">Diagnosis.</emphasis>
Large sized; head, pronotum, underside of head, metepimeron, tibiae and tarsi of legs, antennae and margins of elytra dark brown, majority part of elytra, underside, femora and coxae of legs, palps yellowish brown; elytral marking from base to apex along suture, anchoreaform, widest at base, then gradually contracted backwards, wider at middle, then trapped towards apex; strongly shiny, glabrous on disc, without wrinkle or puncture, palps normally subcylindrical, ligula wide, 6-setose; antennae short, extending only at shoulders of elytra; pronotum with lateral expanded margin glabrous and smooth, border unclear, slightly reflexed; elytra elongate-ovate, apex concavely truncated, outer angle distinct; elytral intervals 3 with two setiferous pores, located near base (close to stria 3) and subapex (closed to stria 2) respectively; fore tibia stout, outer apical angle distinct, somewhat rectangular; hind tibial spurs long and sharp, claws pectinate, hind tarsomere 4 bilobed; prosternal process unbordered at apex, abdominal ventrite VII slightly emarginated in median portion of apical margin in male; male genitalia very similar to that of
<taxonomicName id="C8AF74B4FFB9FF9B1F25FBCAFC85A11F" box="[634,885,1089,1113]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="lateralis" subSpecies="maindroni">
<emphasis id="3DDBD325FFB9FF9B1F25FBCAFC85A11F" box="[634,885,1089,1113]" italics="true" pageId="3" pageNumber="543">H. lateralis maindroni</emphasis>
</taxonomicName>
(
<figureCitation id="979413B2FFB9FF9B1EDBFBC9FC01A11C" box="[900,1009,1089,1114]" captionStart="FIGURES 1 4" captionStartId="4.[151,264,1491,1514]" captionTargetBox="[395,1186,207,1455]" captionTargetId="figure@4.[386,1201,195,1468]" captionTargetPageId="4" captionText="FIGURES 1 4. Hexachaetus lateralis Guérin (1. H. lateralis lateralis Guérin, habitus, male (photo: E. Kirschhofer); 2. H. lateralis maindroni Tian et Deuve, habitus, male, holotype; 3. Median lobe of aedeagus of H. lateralis maindroni, lateral view; 4. ibid. apical lamella, dorsal view; Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258028/files/figure.png" pageId="3" pageNumber="543">Figs. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98FBEDFE90A1AC" pageId="3" pageNumber="543" type="materials_examined">
<paragraph id="0F100F37FFB9FF9B1D98FBEDFE90A1AC" blockId="3.[151,1437,693,1294]" pageId="3" pageNumber="543">
<materialsCitation id="BFC7056AFFB9FF9B1D98FBEDFEABA1E7" ID-GBIF-Occurrence="1313880543" collectingDate="26. II-" collectionCode="NHMB" collectorName="Bocak &amp; Bocakova" elevation="450" location="Ketambe" pageId="3" pageNumber="543" specimenCount="1" specimenCount-male="1" stateProvince="Sumatra">
<emphasis id="3DDBD325FFB9FF9B1D98FBEDFEC6A138" bold="true" box="[199,310,1125,1150]" pageId="3" pageNumber="543">Material.</emphasis>
<specimenCount id="19A9C4BEFFB9FF9B1C63FBEEFE7AA138" box="[316,394,1125,1150]" pageId="3" pageNumber="543" type="male">1 male</specimenCount>
, “
<collectingRegion id="CD6BC1D5FFB9FF9B1CC0FBEEFE0FA138" box="[415,511,1126,1150]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="543">Sumatra</collectingRegion>
(
<collectingRegion id="CD6BC1D5FFB9FF9B1F52FBEDFDBEA138" box="[525,590,1125,1150]" country="Indonesia" name="Aceh" pageId="3" pageNumber="543">Aceh</collectingRegion>
),
<location id="0A7059ECFFB9FF9B1F3EFBEDFD39A138" LSID="urn:lsid:plazi:treatment:8706BE21FFB9FF9B1DC8FE7CFD1EA048:0A7059ECFFB9FF9B1F3EFBEDFD39A138" box="[609,713,1125,1150]" name="Ketambe" pageId="3" pageNumber="543" stateProvince="Sumatra">Ketambe</location>
, Leuser Nat. Park,
<elevation id="8482E804FFB9FF9B1EC2FBEDFC16A13B" box="[925,998,1125,1150]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="543" unit="m" value="450.0">
<quantity id="C857A2D2FFB9FF9B1EC2FBEDFC16A13B" box="[925,998,1125,1150]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="543" unit="m" value="450.0">450 m</quantity>
</elevation>
,
<collectingDate id="6B55D01FFFB9FF9B1EAEFBEDFBCDA138" box="[1009,1085,1125,1150]" pageId="3" pageNumber="543">
<date id="7B1129F7FFB9FF9B1EAEFBEDFBCDA138" box="[1009,1085,1125,1150]" pageId="3" pageNumber="543">26. II-</date>
</collectingDate>
1, III, 1991,
<collectorName id="A25A6AE1FFB9FF9B199EFBEDFAF9A138" box="[1217,1289,1125,1150]" pageId="3" pageNumber="543">Bocák</collectorName>
&amp;
<collectorName id="A25A6AE1FFB9FF9B1873FBEDFA6CA138" box="[1324,1436,1125,1150]" pageId="3" pageNumber="543">Bocáková</collectorName>
leg.”, in
<collectionCode id="69BE97F2FFB9FF9B1C5BFB02FEABA1E7" box="[260,347,1162,1185]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="3" pageNumber="543">NHMB</collectionCode>
</materialsCitation>
;
<materialsCitation id="BFC7056AFFB9FF9B1C32FB02FDA8A180" ID-GBIF-Occurrence="1313880537" collectionCode="MCG" collectorName="Museo Civico di Genova" country="Malaysia" location="Museo Civico di Genova" pageId="3" pageNumber="543" specimenCount="1" specimenCount-female="1">
<specimenCount id="19A9C4BEFFB9FF9B1C32FB02FE27A1E4" box="[365,471,1161,1186]" pageId="3" pageNumber="543" type="female">1 female</specimenCount>
, “
<collectingCountry id="77B84FA7FFB9FF9B1CACFB01FDA7A1E4" box="[499,599,1161,1186]" name="Malaysia" pageId="3" pageNumber="543">Malacca</collectingCountry>
, Coll. Castelnau”, “
<collectorName id="A25A6AE1FFB9FF9B1E19FB02FB89A1E4" box="[838,1145,1161,1186]" pageId="3" pageNumber="543">
<location id="0A7059ECFFB9FF9B1E19FB02FB89A1E4" LSID="urn:lsid:plazi:treatment:8706BE21FFB9FF9B1DC8FE7CFD1EA048:0A7059ECFFB9FF9B1E19FB02FB89A1E4" box="[838,1145,1161,1186]" country="Malaysia" name="Museo Civico di Genova" pageId="3" pageNumber="543">Museo Civico di Genova</location>
</collectorName>
”, “
<taxonomicName id="C8AF74B4FFB9FF9B19FEFB01FE58A180" authority="Guerin, H. E. Andrewes" authorityName="Guerin, H. E. Andrewes" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB9FF9B19FEFB01FA6CA1E7" box="[1185,1436,1161,1185]" italics="true" pageId="3" pageNumber="543">Hexachaetus lateralis</emphasis>
Guérin, H. E. Andrewes
</taxonomicName>
det.”, in
<collectionCode id="69BE97F2FFB9FF9B1F4BFB26FDA8A180" box="[532,600,1198,1222]" country="Italy" httpUri="http://grbio.org/cool/3pg5-vsyp" name="Museo Civico DI Storia Naturale 'Giacomo Doria'" pageId="3" pageNumber="543">MCG</collectionCode>
</materialsCitation>
; 1 female, “Collection P. Dupuis”, “
<taxonomicName id="C8AF74B4FFB9FF9B1EACFB25FF23A1AC" authority="Det. P. Dupuis, 1914" authorityName="Det. P. Dupuis" authorityYear="1914" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB9FF9B1EACFB25FB19A183" box="[1011,1257,1197,1221]" italics="true" pageId="3" pageNumber="543">Hexachaetus lateralis</emphasis>
Det. P. Dupuis, 1914
</taxonomicName>
”, in IRSNB.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98FB7DFD1EA048" box="[199,750,1269,1294]" pageId="3" pageNumber="543" type="distribution">
<paragraph id="0F100F37FFB9FF9B1D98FB7DFD1EA048" blockId="3.[151,1437,693,1294]" box="[199,750,1269,1294]" pageId="3" pageNumber="543">
<emphasis id="3DDBD325FFB9FF9B1D98FB7DFE90A048" bold="true" box="[199,352,1269,1294]" pageId="3" pageNumber="543">Distribution.</emphasis>
<collectingCountry id="77B84FA7FFB9FF9B1C38FB7DFE3EA048" box="[359,462,1269,1294]" name="Malaysia" pageId="3" pageNumber="543">Malaysia</collectingCountry>
and
<collectingCountry id="77B84FA7FFB9FF9B1F5BFB7DFD81A048" box="[516,625,1269,1294]" name="Indonesia" pageId="3" pageNumber="543">Indonesia</collectingCountry>
(
<figureCitation id="979413B2FFB9FF9B1FDFFB7DFD24A048" box="[640,724,1269,1294]" captionStart="FIGURE 20" captionStartId="12.[151,250,1789,1811]" captionTargetBox="[226,1360,198,1765]" captionTargetId="figure@12.[223,1363,195,1768]" captionTargetPageId="12" captionText="FIGURE 20. Distribution of Hexachaetus species (square: angulatus species group; round: lateralis species group; star: laevissimus species group; heart: vietnamensis species group; triangle: mulan species group; a. H. lateralis lateralis; b. H. lateralis maindroni; c. H. maculatus; d. H. kirschenhoferi n. sp.; e. H. brunki n. sp.; f. H. laevissimus; g. H. vietnamensis n. sp.; h. H. prodigus; i. H. mulan n. sp.)" httpUri="https://zenodo.org/record/258032/files/figure.png" pageId="3" pageNumber="543">Fig. 20</figureCitation>
a).
</paragraph>
</subSubSection>
</treatment>
</document>