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<document id="0C91C96AE3BE38A446B733B3A0E80DF9" ID-DOI="10.3897/zoologia.38.e65001" ID-ZooBank="A417A7BD-9459-43AD-8FAE-42151BE7D733" ID-publisher-id="65001" URI-arpha="062AA804-1299-5853-8E64-4A9AF7EA1BFC" URI-zenodo_dep_id="5219823" URI-zoobank="http://zoobank.org/A417A7BD-9459-43AD-8FAE-42151BE7D733" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1723475328472" checkinUser="pensoft" docAuthor="de Freitas, Álvaro J. B., Bezerra, Carine A. M., Meneses, Yuri C., Justo, Marcia Cristina N., Viana, Diego C. &amp; Cohen, Simone C." docDate="2021" docId="8676C93C2DF653D391B02643D6BCF6EA" docLanguage="en" docName="Zoologia 38: 1-11" docOrigin="Zoologia 38" docSource="https://zoologia.pensoft.net/article/65001/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Urocleidoides paratriangulus Freitas, Bezerra, Meneses, Justo, Viana &amp; Cohen, 2021, sp. nov." docType="treatment" docUuid="3C18E487-2368-45B7-BB00-2A7F03AAE9AA" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="11" masterDocId="062AA804129958538E644A9AF7EA1BFC" masterDocTitle="Three new species of Urocleidoides (Monogenoidea: Dactylogyridae) parasitizing characiforms (Actinopterygii: Characiformes) in Tocantins River, states of Tocantins and Maranhão, and new record for U. triangulus in Guandu River, state of Rio de Janeiro, Brazil" masterLastPageNumber="11" masterPageNumber="1" pageNumber="1" updateTime="1723478778410" updateUser="admin">
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<mods:title id="B43069B375ED79B9743A80227E75B2AA">Three new species of Urocleidoides (Monogenoidea: Dactylogyridae) parasitizing characiforms (Actinopterygii: Characiformes) in Tocantins River, states of Tocantins and Maranhão, and new record for U. triangulus in Guandu River, state of Rio de Janeiro, Brazil</mods:title>
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<mods:namePart id="6981DBD0F6B44FFCBE48AF97220F51B7">de Freitas, Álvaro J. B.</mods:namePart>
<mods:affiliation id="1E25019D5B509755449624FD7BA6306C">Laboratório de Helmintos Parasitos de Peixes, Instituto Oswaldo Cruz, FIOCRUZ. Avenida Brasil 4365, 21040 - 900 Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:namePart id="8F934975252C9AB52B10186B44A11990">Bezerra, Carine A. M.</mods:namePart>
<mods:affiliation id="DE8ADB09DE25FFBAC42745F2CDA5F8D1">Programa de Pós-Graduação em Ciência Animal, Universidade Estadual do Maranhão. Cidade Universitária Paulo VI, Avenida Lourenço Vieira da Silva 1000, 65055 - 310 São Luís, MA, Brazil.</mods:affiliation>
</mods:name>
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<mods:namePart id="627492A7D046B0AF3AEB07C3B81EAF6E">Meneses, Yuri C.</mods:namePart>
<mods:affiliation id="5CB251173920C93A1CACF0715FA7FE9F">Laboratório de Helmintos Parasitos de Peixes, Instituto Oswaldo Cruz, FIOCRUZ. Avenida Brasil 4365, 21040 - 900 Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:namePart id="85934518895B28F510F360390273210F">Justo, Marcia Cristina N.</mods:namePart>
<mods:nameIdentifier id="93AA78B3A7B895C14C38D3B1ED94E100" type="ORCID">0000-0002-0684-3389</mods:nameIdentifier>
<mods:affiliation id="53146DE9C0A6D9C6DB31F19D4E566EF2">Laboratório de Helmintos Parasitos de Peixes, Instituto Oswaldo Cruz, FIOCRUZ. Avenida Brasil 4365, 21040 - 900 Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:namePart id="AF6B915A10B0FA763085E2C63E79B65F">Viana, Diego C.</mods:namePart>
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<mods:affiliation id="534311E9D582F5BE42BA4907B03C8508">Núcleo de Estudos Morfofisiológicos Avançados, Universidade Estadual da Região Tocantina do Maranhão. Campus Imperatriz, Rua Godofredo Viana 1300, 65901 - 480 Imperatriz, MA, Brazil.</mods:affiliation>
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<mods:namePart id="483302FF796CF4F469B116027466FF76">Cohen, Simone C.</mods:namePart>
<mods:affiliation id="BA664B4FBBA572D3D091DE7C857D4EEC">Laboratório de Helmintos Parasitos de Peixes, Instituto Oswaldo Cruz, FIOCRUZ. Avenida Brasil 4365, 21040 - 900 Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:title id="3FBF8AA52A4EACFF00FE9DEB7357DD49">Zoologia</mods:title>
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<mods:part id="26C3BDB2322784B4177E32CF5AE7E086">
<mods:date id="3549A9122CAE908C5DD425D138ABC415">2021</mods:date>
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<mods:number id="24BC1F9D9257007761E2653880A61FCA">2021-08-11</mods:number>
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<mods:number id="23088E18CE3472A36D237FD1A6818072">38</mods:number>
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<mods:classification id="E733D4A2408F46E66FE9F40634ED84E4">journal article</mods:classification>
<mods:identifier id="D76B3BFC0B173A892197A263949039BF" type="DOI">10.3897/zoologia.38.e65001</mods:identifier>
<mods:identifier id="BD184593A4630B1F9C07D14E31B718E7" type="ZooBank">A417A7BD-9459-43AD-8FAE-42151BE7D733</mods:identifier>
</mods:mods>
<treatment id="8676C93C2DF653D391B02643D6BCF6EA" ID-arpha="8676C93C-2DF6-53D3-91B0-2643D6BCF6EA" ID-zoobank="http://zoobank.org/3C18E487-2368-45B7-BB00-2A7F03AAE9AA" LSID="urn:lsid:zoobank.org:act:3C18E487-2368-45B7-BB00-2A7F03AAE9AA" LSID-ZBK="urn:lsid:zoobank.org:act:3C18E487-2368-45B7-BB00-2A7F03AAE9AA" httpUri="http://treatment.plazi.org/id/8676C93C2DF653D391B02643D6BCF6EA">
<subSubSection id="F0E18DD2C42C2909CABC3E2F98511485" type="nomenclature">
<paragraph id="BB6B4AE4E74D45E904ED83C7CC526D59">
<taxonomicName id="BFD5E18EF466B09316B1360A969DE90F" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Dactylogyridae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="172C8C3F88E339768A50AEE4FB06694F" italics="true">Urocleidoides paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B5F58FC9DB63D0A83FBE72CD324687F" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="80ABAD132EC12E174D40123B6250129C" type="description">
<paragraph id="38D6BDCE1D11C75F92D12D8799BFFE5F">
<figureCitation id="D6EB701111A66591124A8536F1C19C6C" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">Figs 1221</figureCitation>
,
<figureCitation id="BB3B4E01D7D68FED7DEC3314E4DA66FB" captionStart="Figures 3136" captionStartId="F5" captionText="Figures 3136. Light photomicrographs of Urocleidoides spp.: (31, 32) Urocleidoides boulengerellae: (31) copulatory complex; (32) vaginal sclerite; (33, 34) Urocleidoides paratriangulus: (33) copulatory complex; (34) haptor; (35, 36) Urocleidoides tocantinensis: (35) copulatory complex; (36) vagina. Scale bars: 3133, 3536 = 20 µm, 34 = 40 µm." figureDoi="10.3897/zoologia.38.e65001.figures31-36" httpUri="https://binary.pensoft.net/fig/576829">33</figureCitation>
,
<figureCitation id="694D2362BF17E27F59011D0D01A42009" captionStart="Figures 3136" captionStartId="F5" captionText="Figures 3136. Light photomicrographs of Urocleidoides spp.: (31, 32) Urocleidoides boulengerellae: (31) copulatory complex; (32) vaginal sclerite; (33, 34) Urocleidoides paratriangulus: (33) copulatory complex; (34) haptor; (35, 36) Urocleidoides tocantinensis: (35) copulatory complex; (36) vagina. Scale bars: 3133, 3536 = 20 µm, 34 = 40 µm." figureDoi="10.3897/zoologia.38.e65001.figures31-36" httpUri="https://binary.pensoft.net/fig/576829">34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELOAE" type="Type host">
<paragraph id="69C700E65C831CF9D880F83C9CE1376E">
<heading id="B2BB99489D436AAD0D971E0E8554470B" reason="title">Type host.</heading>
</paragraph>
<paragraph id="A55980ACE0CE2082EED6AA7F879D3C8C">
<taxonomicName id="FAFD313196848368DCF3999D26871BD8" authority="Eigenmann &amp; Eigenmann" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" family="Curimatidae" genus="Psectrogaster" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="amazonica">
<emphasis id="464E55DF0885CB54E44C6338F390B79B" italics="true">Psectrogaster amazonica</emphasis>
Eigenmann &amp; Eigenmann
</taxonomicName>
(
<taxonomicName id="B1533B6A7562502FE8559528F16D5C71" family="Curimatidae" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="family">Curimatidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPAE" type="type locality">
<paragraph id="B5D4DD168ECE0A337DB0C9464E39DC24">
<heading id="2D095FF84ECC38E261EF39A507EF3EE5" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="97AD0BE5EEE8C09F08345E139ED7A6EB">
<collectingMunicipality id="A4D92C8145054C77EDC860A7742D087C">Tocantins River</collectingMunicipality>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENPAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.563333,-5.463889]}">
<geoCoordinate id="EB17577D488F43D736947BDF6F5DF2AE" degrees="5" minutes="27" seconds="50">5 ° 27 ' 50</geoCoordinate>
' S;
<geoCoordinate id="175582EEFBFACE8E5ABD81FA274D5FCF" degrees="47" minutes="33" seconds="48">47 ° 33 ' 48</geoCoordinate>
' W
</named-content>
), close to the municipality of
<location id="C74EA603F7BD652FDE0777D8E2A405AA" LSID="urn:lsid:plazi:treatment:8676C93C2DF653D391B02643D6BCF6EA:C74EA603F7BD652FDE0777D8E2A405AA" municipality="Tocantins River" name="Embiral" stateProvince="Maranhao">Embiral</location>
, state of
<collectingRegion id="A605607C2F3E95F30A243B6D47476409" country="Brazil" name="Maranhao">Maranhão</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESPAE" type="Other hosts and localities">
<paragraph id="8FCF6A80F296C77A9BDF11246351B0C6">
<heading id="4D059DC9A8EA0474E31A021E28789E78" reason="title">Other hosts and localities.</heading>
</paragraph>
<paragraph id="7490CD73D51F904DDE6650E6A18ADBF6">
<taxonomicName id="9CC3CDF22BE465FA934BB198C9F0B724" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" family="Curimatidae" genus="Psectrogaster" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="amazonica">
<emphasis id="62C94AC704A3F275EE61B285D82680CB" italics="true">Psectrogaster amazonica</emphasis>
</taxonomicName>
:
<collectingRegion id="65874E28FC4B57D856135B149D3B78B4" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJQAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.494750,-5.535722]}">
<geoCoordinate id="1A5F20D378CF5D6DE8A728BBFE0BF101" degrees="5" minutes="32" seconds="08.6">5 ° 32 ' 08.6</geoCoordinate>
' S;
<geoCoordinate id="5531E7617611E4B8D490B1F7D3427D24" degrees="47" minutes="29" seconds="41.1">47 ° 29 ' 41.1</geoCoordinate>
' W
</named-content>
), close to the urban perimeter of Imperatriz, state of
<collectingRegion id="98A0A919567F4F018F8DFEFF16E3B394" country="Brazil" name="Maranhao">Maranhao</collectingRegion>
.
<taxonomicName id="2BBE0323FADB1B2FE556080920B5D963" authorityName="Vari" authorityYear="1992" family="Curimatidae" genus="Cyphocharax" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="gouldingi">
<emphasis id="C5D821B65F88A18CDA212D72954B528F" italics="true">Cyphocharax gouldingi</emphasis>
</taxonomicName>
(
<taxonomicName id="31855E5568F11035A21DF016DA066902" family="Curimatidae" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="family">Curimatidae</taxonomicName>
):
<collectingRegion id="9BAE66F0975E35A615B697A70EEBD4E8" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.450208,-6.540147]}">
<geoCoordinate id="D498593F5C919BB01D9C5C8555BEAF35" degrees="6" minutes="32" seconds="24.53">6 ° 32 ' 24.53</geoCoordinate>
' S;
<geoCoordinate id="635C03788474E56A2C450FB865251815" degrees="47" minutes="27" seconds="0.75">47 ° 27 ' 0.75</geoCoordinate>
' W
</named-content>
), close to the municipalities of Aguiarnópolis and Estreito; at the mouth of the Itaueiras River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.424300,-6.499647]}">
<geoCoordinate id="32A5108BABA90394690AB1976F1D9C27" degrees="6" minutes="29" seconds="58.73">6 ° 29 ' 58.73</geoCoordinate>
' S;
<geoCoordinate id="F970F38AA1243CF43B0FB6CE6315CA1F" degrees="47" minutes="25" seconds="27.48">47 ° 25 ' 27.48</geoCoordinate>
' W
</named-content>
), in the municipality of Estreito, state of
<collectingRegion id="65AAA5233452858F7375F1AB41859AC3" country="Brazil" name="Maranhao">Maranhão</collectingRegion>
.
<taxonomicName id="AB782CBDB2779978E953DB6C7DCDD6D6" baseAuthorityName="Kner" baseAuthorityYear="1858" family="Chilodontidae" genus="Caenotropus" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="labyrinthicus">
<emphasis id="C186D2AC595E982B567B1776CFD664B2" italics="true">Caenotropus labyrinthicus</emphasis>
</taxonomicName>
(
<taxonomicName id="1AEF0705CFB3877DCD83EB590FB6774B" authorityName="Wenz" authorityYear="1938" family="Chilodontidae" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="family">Chilodontidae</taxonomicName>
): Arraias River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.136444,-12.631194]}">
<geoCoordinate id="1FCD4879B4048A7986E2CF78D8E200CB" degrees="12" minutes="37" seconds="52.3">12 ° 37 ' 52.3</geoCoordinate>
' S;
<geoCoordinate id="32E6DDE39D870F9557ED34B627246917" degrees="47" minutes="08" seconds="11.2">47 ° 08 ' 11.2</geoCoordinate>
' W
</named-content>
), close to the municipality of Babaçulândia, state of
<collectingRegion id="C5F2E9FF2FA24E13C3C513E3CED947CD" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
;
<collectingRegion id="69ADE6407FA9D9291BE54BF016228610" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.450208,-6.540147]}">
<geoCoordinate id="7E5A5BAEF017FC38696F1F2D0B654B6D" degrees="6" minutes="32" seconds="24.53">6 ° 32 ' 24.53</geoCoordinate>
' S,
<geoCoordinate id="182120C34937A0DB4FECAA894DE49F01" degrees="47" minutes="27" seconds="0.75">47 ° 27 ' 0.75</geoCoordinate>
' W
</named-content>
), close to the municipalities of Aguiarnópolis and Estreito; at the mouth of the Itaueiras River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.424300,-6.499647]}">
<geoCoordinate id="EF2CE5407397937001D524A88B03927C" degrees="6" minutes="29" seconds="58.73">6 ° 29 ' 58.73</geoCoordinate>
' S;
<geoCoordinate id="A1A5680019564DF538573E3B8EEBB566" degrees="47" minutes="25" seconds="27.48">47 ° 25 ' 27.48</geoCoordinate>
' W
</named-content>
), municipality of Estreito, state of
<collectingRegion id="4A305FA4A97C633E272976B5E4226212" country="Brazil" name="Maranhao">Maranhão</collectingRegion>
; João Aires River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.932583,-7.852944]}">
<geoCoordinate id="D747C69C389DABD9F25EC6DD056B957C" degrees="7" minutes="51" seconds="10.6">7 ° 51 ' 10.6</geoCoordinate>
' S;
<geoCoordinate id="EA540C51FC65634A8060E4BE8335FC6D" degrees="47" minutes="55" seconds="57.3">47 ° 55 ' 57.3</geoCoordinate>
' W
</named-content>
), close to the municipality of Palmeirantes, state of
<collectingRegion id="D370CE55A42881C273F2CD43A5D756DE" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
; Farinha River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBTAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.501611,-6.841806]}">
<geoCoordinate id="67766227D97149ECC91C30834FAF22C7" degrees="6" minutes="50" seconds="30.5">6 ° 50 ' 30.5</geoCoordinate>
' S;
<geoCoordinate id="9034566CEC1AA5162441BDD5526012BF" degrees="47" minutes="30" seconds="05.8">47 ° 30 ' 05.8</geoCoordinate>
' W
</named-content>
), close to the municipality of Estreito, state of
<collectingRegion id="62A532C3095639563DD454CBFBDC2E89" country="Brazil" name="Maranhao">Maranhão</collectingRegion>
,
<collectingCountry id="F9F2FDB82FFFF81CE8950EF9CA808073" name="Brazil">Brazil</collectingCountry>
.
<taxonomicName id="32859B7A9A0725BE2393E696937F1F36" authority="Jegu &amp; Santos" authorityName="Jegu &amp; Santos" family="Serrasalmidae" genus="Mylesinus" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="pauscisquamatus">
<emphasis id="033C6873A4F8556C698AD137299FA53D" italics="true">Mylesinus pauscisquamatus</emphasis>
Jégu &amp; Santos
</taxonomicName>
(
<taxonomicName id="0F551DBDE4852CDCBF3059B5FE5A7C6A" family="Serrasalmidae" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="family">Serrasalmidae</taxonomicName>
): Arraias River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZTAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.136444,-12.631194]}">
<geoCoordinate id="182252E103695AD266F31593A6307A5D" degrees="12" minutes="37" seconds="52.3">12 ° 37 ' 52.3</geoCoordinate>
' S;
<geoCoordinate id="6B1F91DA72B49A84990CE3C90F02A014" degrees="47" minutes="08" seconds="11.2">47 ° 08 ' 11.2</geoCoordinate>
' W
</named-content>
), close to the municipality of Babaçulândia, state of
<collectingRegion id="2F6AACAE74CD3A7DC740A3F8ADC7F7FE" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
; Farinha River (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBUAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.501611,-6.841806]}">
<geoCoordinate id="4FB19ACA52CAD5D3848534B3D4EC1A2B" degrees="6" minutes="50" seconds="30.5">6 ° 50 ' 30.5</geoCoordinate>
' S;
<geoCoordinate id="DE389BACC51562E88D36686C9240FCA2" degrees="47" minutes="30" seconds="05.8">47 ° 30 ' 05.8</geoCoordinate>
' W
</named-content>
), close to the municipality of Estreito, state of
<collectingRegion id="ABB6023CCDC4ACE3E31493937BE207A0" country="Brazil" name="Maranhao">Maranhão</collectingRegion>
,
<collectingCountry id="620153DB819BA73C240D8DCD3EB30254" name="Brazil">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGUAE" type="Infestation parameters">
<paragraph id="2B58605745E376B0F55BAC5221F5FEC6">
<heading id="3AE95C92122AD3B8EF8D2C630957D94D" reason="title">Infestation parameters.</heading>
</paragraph>
<paragraph id="F777B6164951A01F035FEAB57C65DC93">
<taxonomicName id="1DFFF7FCC8EAF33F8285C20AB9167C07" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" family="Curimatidae" genus="Psectrogaster" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="amazonica">
<emphasis id="51877344BCE83CE55A964BE398F64267" italics="true">Psectrogaster amazonica</emphasis>
</taxonomicName>
: total number of hosts: 97; prevalence: 11.3 %; total number of parasites: 37, mean intensity: 3.1 ± 2.5, range of infection: 19;
<taxonomicName id="D4993A94EA813CC0420A770C4B504093" baseAuthorityName="Kner" baseAuthorityYear="1858" family="Chilodontidae" genus="Caenotropus" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="labyrinthicus">
<emphasis id="ED93FA0E09A20BCA1150ED2425800BE3" italics="true">Caenotropus labyrinthicus</emphasis>
</taxonomicName>
: total number of hosts: 8; number of infected hosts: 3; total number of parasites: 4;
<taxonomicName id="9A48CF81E71A6E0E4C75704B23573BCB" authorityName="Vari" authorityYear="1992" family="Curimatidae" genus="Cyphocharax" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="gouldingi">
<emphasis id="B57761D530B1D1B2BFE63546F34103CE" italics="true">Cyphocharax gouldingi</emphasis>
</taxonomicName>
: total number of hosts: 8; number of infected hosts: 1; total number of parasites: 2;
<taxonomicName id="F96EF183AF2647F291DF7C083E4F7D88" authorityName="Jegu &amp; Santos" authorityYear="1988" family="Serrasalmidae" genus="Mylesinus" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="paucisquamatus">
<emphasis id="937E0186E1E01EDD06772D49A0950F0A" italics="true">Mylesinus paucisquamatus</emphasis>
</taxonomicName>
: total number of hosts: 7; number of infected hosts: 1; total number of parasites: 10.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2VAE" type="material">
<paragraph id="7C516F1FCA55398FD2B3D49EA7C7CBDF">
<heading id="BF0C682673F00119A6F3E0CFA0D7E0A1" reason="title">Deposited material.</heading>
</paragraph>
<paragraph id="A42B7E3F673C6DE5F06533F4AA823549">
<materialsCitation id="1C4966F34EBCDE31AB7B9ECC83E95C82" collectionCode="CHIOC" location="Embiral" municipality="Tocantins River" specimenCode="39567 a" specimenCount="1" stateProvince="Maranhao" typeStatus="holotype">
<typeStatus id="EC70BAA4A75778776FFDBAFE106D6804" type="holotype">Holotype</typeStatus>
:
<taxonomicName id="B63A08574C057FB4C0176844EECEA781" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" family="Curimatidae" genus="Psectrogaster" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="amazonica">
<emphasis id="24FA589B9C09A84C6FDCB18AEA14B30D" italics="true">
<originalDetermination id="C0A45F85B230005A6662BC08E46E9AF5">Psectrogaster amazonica</originalDetermination>
</emphasis>
</taxonomicName>
:
<collectionCode id="1EA81091EF34A63970B0146626B996C7" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" type="">CHIOC</collectionCode>
:
<specimenCode id="4E287D4CF7B007E4571B4358214D9428">39567 a</specimenCode>
</materialsCitation>
;
<materialsCitation id="4A537C26ADF8BD130EC424B656F26B31" collectionCode="CHIOC, INPA" location="Embiral" municipality="Tocantins River" specimenCode="39567 b, 39571, 39572, 39573, 39574, 836, 837" specimenCount="1" stateProvince="Maranhao" typeStatus="paratype">
<typeStatus id="F918830487E8A343CEE188C4CC6651DB" type="paratype">paratypes</typeStatus>
<collectionCode id="7F0A6833EF507C42EE19E27F064F2C13" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" type="">CHIOC</collectionCode>
<specimenCode id="CDA1B4B981A65BA824BBC14EC0644540">39567 b</specimenCode>
,
<specimenCode id="048934AB2DFE898BD8089809EEF38AA2">39571</specimenCode>
,
<specimenCode id="75D35B9602CBECC58F8B25713D59C9AE">39572</specimenCode>
,
<specimenCode id="F6FB1E190D6AFBC3D5670A34B0663B7E">39573</specimenCode>
,
<specimenCode id="3DB12BE25524B39CFA7AF7811730CF04">39574</specimenCode>
,
<collectionCode id="15DF7B6127CAD25168998D31E62EBD79" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" type="">INPA</collectionCode>
<specimenCode id="A4E73DAFFAE8E1648EE21AAE9F09B02E">836</specimenCode>
,
<specimenCode id="23A1B7471F40D1E58BF6C7127E6A29C8">837</specimenCode>
</materialsCitation>
.
<materialsCitation id="06F3E165E5CF5A2A7D1163C9CDEFE8A4" collectionCode="CHIOC" specimenCode="39568 a-b" specimenCount="1">
<taxonomicName id="880E321C444105E9F48F72B3053432A6" authorityName="Vari" authorityYear="1992" family="Curimatidae" genus="Cyphocharax" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="gouldingi">
<emphasis id="26A7D6EBFA453F584CD85230BCE9A1DE" italics="true">
<originalDetermination id="7F40113930B3E8466B650904084361E2">Cyphocharax gouldingi</originalDetermination>
</emphasis>
</taxonomicName>
:
<collectionCode id="C055D7F9767F510B3C716CBC0D0A39F1" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" type="">CHIOC</collectionCode>
:
<specimenCode id="353D00061E81DE3F15110521AA41E6C5">39568 a-b</specimenCode>
</materialsCitation>
.
<materialsCitation id="8AEE143F8C0DF2A63087ED9356E72967" collectionCode="CHIOC" specimenCode="39570" specimenCount="1">
<taxonomicName id="7D540D08B1531ADEE78E4C0B6F5C5A0F" baseAuthorityName="Kner" baseAuthorityYear="1858" family="Chilodontidae" genus="Caenotropus" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="labyrinthicus">
<emphasis id="D61EC69A4229118143993B0CA30F9876" italics="true">
<originalDetermination id="C185E9201F999A34ABC0E9A8F7861196">Caenotropus labyrinthicus</originalDetermination>
</emphasis>
</taxonomicName>
:
<collectionCode id="3B81ADC6CD46873D446F1EACE2106073" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" type="">CHIOC</collectionCode>
:
<specimenCode id="B518898981DA896C6ABB68E786009C21">39570</specimenCode>
</materialsCitation>
,
<materialsCitation id="58A35924FDD5B87C045E8159FA965D53" collectionCode="CHIOC, INPA" specimenCode="39569 a-c, 835" specimenCount="1">
<taxonomicName id="BC2D38ABD9B9433D0D065EA621DAC537" authorityName="Jegu &amp; Santos" authorityYear="1988" family="Serrasalmidae" genus="Mylesinus" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="paucisquamatus">
<emphasis id="17340FF63FBDADFEA08CE0AC741B3821" italics="true">
<originalDetermination id="5811AE048A24D8A1CD73BB2F3E7F7004">Mylesinus paucisquamatus</originalDetermination>
</emphasis>
</taxonomicName>
:
<collectionCode id="A3553A92E9A806A7C3CEDEEE8EBBA9B8" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" type="">CHIOC</collectionCode>
:
<specimenCode id="85FF4BE6CF1A647D804452E90A9AAD56">39569 a-c</specimenCode>
,
<collectionCode id="12581864024C7E7733B95EBB9A9B9139" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" type="">INPA</collectionCode>
<specimenCode id="E46410496C800C36D3E17D2A4192A3C2">835</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENXAE" type="description">
<paragraph id="19F0F897F901147BA527BF0BC51020C6">
<heading id="BD62FE7C7BB6C572BDEF742591855F08" reason="title">Description.</heading>
</paragraph>
<paragraph id="36FD4CF2FC70161FBBC30E0CD1A35CFA">
Based on
<specimenCount id="06A74FD81D884559E2B1BB59213CE733" count="51" type="generic">51 specimens</specimenCount>
mounted in Hoyers' medium: Body fusiform, robust 195402 (290; n = 15) long by 65132 (103; n = 15) wide. Cephalic region with cephalic lobes poorly developed; four bilateral pairs of head organs. Eyespots absent; accessory chromatic granules dispersed in the cephalic region. Mouth subterminal, midventral; pharynx spherical, esophagus short; two intestinal caeca, confluent posteriorly to testes. Haptor sub-hexagonal, 60105 (83; n = 15) wide (Fig.
<figureCitation id="6A93E3AEA44B5F82DAE6CDF6AE165FA0" captionStart="Figures 3136" captionStartId="F5" captionText="Figures 3136. Light photomicrographs of Urocleidoides spp.: (31, 32) Urocleidoides boulengerellae: (31) copulatory complex; (32) vaginal sclerite; (33, 34) Urocleidoides paratriangulus: (33) copulatory complex; (34) haptor; (35, 36) Urocleidoides tocantinensis: (35) copulatory complex; (36) vagina. Scale bars: 3133, 3536 = 20 µm, 34 = 40 µm." figureDoi="10.3897/zoologia.38.e65001.figures31-36" httpUri="https://binary.pensoft.net/fig/576829">34</figureCitation>
). Ventral anchor with superficial root elongated, round deep root knob-like, straight shaft and short point, 60105 (83; n = 15) long, base 1331 (22; n = 38) (Fig.
<figureCitation id="1E2B06C601641D4636F68397DC74EEA6" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">18</figureCitation>
). Dorsal anchor with well-developed elongated superficial root and inconspicuous deep root, short shaft and long point, 2650 (36; n = 27) in length and base 1530 (22; n = 28) (Fig.
<figureCitation id="84F70F50BE82B04A4ADC9A0D7C6176C7" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">19</figureCitation>
). Ventral bar straight, with an anteromedian groove and rounded extremities, anteriorly directed, 2445 (33; n = 15) long (Fig.
<figureCitation id="4A6E959186F5D1A9817BD9111C7737BC" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">17</figureCitation>
). Dorsal bar V-shaped, postero-median projection 1938 (24; n = 12) long, with two variations: a slender or a more robust projection with lateral allae, 3347 (40; n = 11) long (Figs
<figureCitation id="23AE30C61DD5209A17F1CB964878D2A3" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">15</figureCitation>
,
<figureCitation id="DB27B55579E1579A4C6F709972D9650E" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">16</figureCitation>
). Seven pairs of robust marginal hooks, five ventral and two dorsal, similar in shape, pairs 1 and 5 reduced in size, presenting curved point, dilated shank, composed of one subunit, protruded thumb, filamentous hook loop about 1 / 6 shank length (Figs
<figureCitation id="5536831B89DEFA8B8B0CC3209A390447" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">20</figureCitation>
,
<figureCitation id="DBB952DB6DCBC146A8696FC96483203F" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">21</figureCitation>
), pair 1, 720 (13; n = 30) long; pair 2, 1851 (35; n = 23) long; pair 3, 1737 (28; n = 20) long; pair 4, 1527 (21; n = 19) long; pair 5, 914 (11; n = 17) long, pair 6, 1442 (29; n = 22) long; pair 7, 643 (21; n = 21) long. Copulatory complex consists of male copulatory organ (MCO) and accessory piece, which serve as guide to MCO. Male copulatory organ about two counterclockwise rings, the proximal ring has a smaller diameter than the distal one, 54100 (75; n = 18) long. Accessory piece not connected to MCO, situated at distal portion of the organ, consisting of a shaft with medial portion enlarged, with distal groove supporting MCO, 2138 (28; n = 10) long (Figs
<figureCitation id="0D13A75B16602371CD83ECD5BACF71E2" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">12</figureCitation>
,
<figureCitation id="2D44631157704F343861CE23E462A95E" captionStart="Figures 3136" captionStartId="F5" captionText="Figures 3136. Light photomicrographs of Urocleidoides spp.: (31, 32) Urocleidoides boulengerellae: (31) copulatory complex; (32) vaginal sclerite; (33, 34) Urocleidoides paratriangulus: (33) copulatory complex; (34) haptor; (35, 36) Urocleidoides tocantinensis: (35) copulatory complex; (36) vagina. Scale bars: 3133, 3536 = 20 µm, 34 = 40 µm." figureDoi="10.3897/zoologia.38.e65001.figures31-36" httpUri="https://binary.pensoft.net/fig/576829">33</figureCitation>
). Vagina ventral, funnel-shaped, consisting of a sclerotized tube, opening dorsally (Fig.
<figureCitation id="8370DC72EC0719D5D8B18F6CDF305F8A" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">14</figureCitation>
). Vaginal sclerite composed of a long rod, longitudinally grooved, distal extremity hooked and subterminal short projection, 1832 (23; n = 13) long (Fig.
<figureCitation id="DA48FA8D928FB6242DF94AF4F88E6AA3" captionStart="Figures 1221" captionStartId="F3" captionText="Figures 1221. Urocleidoides paratriangulus sp. nov. from Psectogaster amazonica from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm." figureDoi="10.3897/zoologia.38.e65001.figures12-21" httpUri="https://binary.pensoft.net/fig/576827">13</figureCitation>
). Eggs, Mehlis' glands and ootype not observed. Vitellaria distributed throughout the body, absent in the region of reproductive organs.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDZAE" type="etymology">
<paragraph id="2CF541094A96BDDD10A7F88BA5BE2354">
<heading id="DEB8336905229E4BAA08B0C7253A214A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A55F629A349B5C50265A0596A80F7BBA">
The specific name refers to the similarity with
<taxonomicName id="A9EFCFE3FD15F61E16CE805C5BA0FA03" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="717065BBEC329E4876B88BEA4A52B362" italics="true">Urocleidoides triangulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUZAE" type="remarks">
<paragraph id="78958A9EBBFF6728756E85B2442E8190">
<heading id="26FFD48A52B9959F9451F1877B6FAC4A" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="EFB2AD8318CE2D8DF43DA7FD63BC0B6C">
<taxonomicName id="D3A4984A0FDF8C21117244BD7E2C1564" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="C02FD7E4BF8848B96529DC0BBACFFBEB" italics="true">Urocleidoides paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CFEFFF4411A1AC5CB493F04A2B10A31" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="08713DD27C4C6DC26AA4755B28EACD4F" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="0A5CA2B5695BB2C1FBE2A98A15A126D5" italics="true">U. triangulus</emphasis>
</taxonomicName>
considering the shape of the anchors and bars. The new species differs from
<taxonomicName id="8443DF02BC839C0820E5E75AF33BCA05" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="7F5D4D9C5EBBFE9D3E674C69DC92EF60" italics="true">U. triangulus</emphasis>
</taxonomicName>
by the projection of the dorsal bar (prominent postero-median process in
<taxonomicName id="4224A62CE096747D32B2165CB4DEEFAF" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="2B59676FC28AE5241DB1722D02E9AF23" italics="true">U. triangulus</emphasis>
</taxonomicName>
and slender or a more robust projection with lateral allae in the new species); the number of rings of MCO (
<quantity id="C4B6FC089CA568175BD9EED2D578D4E5" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.62" metricValueMin="6.35" unit="in" value="2.75" valueMax="3.0" valueMin="2.5">2.5 to 3 in</quantity>
<taxonomicName id="284855E04CF90F7FB10D41B6BBF08094" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="109947B32E81DD64F68A1EF9FF998122" italics="true">U. triangulus</emphasis>
</taxonomicName>
and about two in the new species); and by the reduced size of hooks (pairs 1, 5 and 7 reduced in size in
<taxonomicName id="5EA181B8A4866D67BA6592287AD3383C" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="03942998E795D0054C488E20870A552C" italics="true">U. triangulus</emphasis>
</taxonomicName>
and 1 and 5 only in
<taxonomicName id="C638766D07A407EBD9F7035563ECF2B7" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="94749967DEA0064827A44522D3BC7F42" italics="true">U. paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B44C3C74EDC4DDB21DE0E99EE28DAB" rank="species">sp. nov.</taxonomicNameLabel>
). Only five species of
<taxonomicName id="4BF8EE7A796FBDD89ADF968D0EDC6E29" authorityName="Mizelle &amp; Price" authorityYear="1964" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="genus">
<emphasis id="54BD67531AEA73B735DE3FFA95D74B68" italics="true">Urocleidoides</emphasis>
</taxonomicName>
possess a medial projection on the posterior margin of the dorsal bar:
<taxonomicName id="64D7200F5A81295E757E1581C18F5BFB" authority="Molnar, Henek &amp; Fernando, 1974" authorityName="Molnar, Henek &amp; Fernando" authorityYear="1974" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="curimatae">
<emphasis id="8EF20BD1E091B75F3DA24D1E2BCB85A5" italics="true">Urocleidoides curimatae</emphasis>
Molnar, Henek &amp; Fernando, 1974
</taxonomicName>
;
<taxonomicName id="F69EF4D4BBF21098F42B3BAF134175A5" authorityName="Mendoza-Franco &amp; Reina" authorityYear="2008" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="neotropicalis">
<emphasis id="37792313E908DB331D471FC45ACA8481" italics="true">U. neotropicalis</emphasis>
</taxonomicName>
;
<taxonomicName id="C46CDAD0CA4997CC38A1FFD1740A886A" authorityName="Mendoza-Franco &amp; Reina" authorityYear="2008" family="Ctenoluciidae" genus="Urocelidoides" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="piriatiu">
<emphasis id="24E7BB4E3492F43291153E2CFB9CD232" italics="true">U. piriatiu</emphasis>
</taxonomicName>
;
<taxonomicName id="8C499D17A798373BFB1B5342CCCF5A6A" authority="Zago, Yamada, Yamada, Franceschini, Bongiovani &amp; Silva, 2020" authorityName="Zago, Yamada, Yamada, Franceschini, Bongiovani &amp; Silva" authorityYear="2020" family="Serrasalmidae" genus="Urocleidoide" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="tenuis">
<emphasis id="D5F461CF5FF861726FC62FA824A2EBC2" italics="true">Urocleidoide tenuis</emphasis>
<bibRefCitation id="CB23058493E7D29A69320A4CB3958768" DOI="10.1007/s00436-020-06831-z" author="Zago AC &amp; Yamada FH &amp; Yamada POF &amp; Franceschini L. Bongiovani MF &amp; Silva RJ" refId="B34" refString="Zago AC, Yamada FH, Yamada POF, Franceschini L. Bongiovani MF, Silva RJ (2020) Seven new species of Urocleidoides (Monogenea: Dactylogyridae) from Brazilian fishes supported by morphological and molecular data. Parasitology Research, https: // doi. org / 10.1007 / s 00436 - 020 - 06831 - z" year="2020">Zago, Yamada, Yamada, Franceschini, Bongiovani &amp; Silva, 2020</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="9E15C1A738E5AFC417CD4D13701392A7" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="boulengerellae" status="sp. nov.">
<emphasis id="5EC0375A75DCE2B3F78296FFF5506AE4" italics="true">U. boulengerellae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C37379B76C282CD55D39C85535B0B132" rank="species">sp. nov.</taxonomicNameLabel>
The medial projection is slenderer than the bar arms in the five species, whereas in
<taxonomicName id="2EE0A4EC63F133680C53BAC95C976585" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="C480C578506377E219B42E24D54ED9CF" italics="true">U. triangulus</emphasis>
</taxonomicName>
and in some specimens of
<taxonomicName id="5EF7F7750B03C8D8B4FA977BC3D6FF31" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="DC1DFB42DD68FE0882F249C5207690CD" italics="true">U. paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7450FB3A92986272F2DBBBAE9D3BD26" rank="species">sp. nov.</taxonomicNameLabel>
, the projection is thicker than the arms.
<taxonomicName id="0B8EDE19F106FABA2AA1909A8AF4DD93" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus">
<emphasis id="8DF7743E8D1FECDB15BAFA4B8F2BA589" italics="true">Urocleidoides paratriangulus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="7FC766D68F2F066DC332A4BDBBC6A2C2" authorityName="Molnar, Henek &amp; Fernando" authorityYear="1974" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="curimatae">
<emphasis id="30AD65994653DC9724A9DCBA4705E638" italics="true">U. curimatae</emphasis>
</taxonomicName>
and
<taxonomicName id="F7D6190AB7010918905139F858232A5C" authorityName="Zago, Yamada, Yamada, Franceschini, Bongiovani &amp; Silva" authorityYear="2020" family="Serrasalmidae" genus="Urocleidoide" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="tenuis">
<emphasis id="79FD0613AD32EB5EDFF06FC1DF0FD06E" italics="true">U. tenuis</emphasis>
</taxonomicName>
also in terms of the triangular shape of the ventral anchor, but it can be differentiated by the number of coils and the length of the MCO (3 coils in
<taxonomicName id="7D9584FF9CD27521C2E52FEC8A01A0FF" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="D279BA10A52A99D0838209D3C4E780E7" italics="true">U. paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E90D89217FEA329DA2A62673D2820115" rank="species">sp. nov.</taxonomicNameLabel>
vs 1.5 coils in
<taxonomicName id="D80B8370CBA7CF81DC1B2EDF6AC6207C" authorityName="Molnar, Henek &amp; Fernando" authorityYear="1974" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="curimatae">
<emphasis id="666AD10A628B73958E132B10DF88FDD9" italics="true">U. curimatae</emphasis>
</taxonomicName>
and
<quantity id="67B87AF19ABFA6094EB77E08C981F15B" metricMagnitude="-1" metricUnit="m" metricValue="1.905" unit="in" value="7.5">7.5 in</quantity>
<taxonomicName id="DFDE0776613F8F2CFBF4181D86D08FDE" authorityName="Zago, Yamada, Yamada, Franceschini, Bongiovani &amp; Silva" authorityYear="2020" family="Serrasalmidae" genus="Urocleidoide" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="tenuis">
<emphasis id="259E296362F39225C38FE4AC86886F2A" italics="true">U. tenuis</emphasis>
</taxonomicName>
), as well as through the morphology of the accessory piece (curved shaft in
<taxonomicName id="15948611FC1D49CD0F963FC49EAA8524" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="675405632756A5F346387A8DB584BB81" italics="true">U. paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA3B644FC1F705794482559980D06667" rank="species">sp. nov.</taxonomicNameLabel>
, straight in
<taxonomicName id="067F3BB7DA3D64B2CCF0225712211931" authorityName="Molnar, Henek &amp; Fernando" authorityYear="1974" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="curimatae">
<emphasis id="C11441070B759CC35B50AD751C3971B6" italics="true">U. curimatae</emphasis>
</taxonomicName>
and pincer-shaped in
<taxonomicName id="86ABBED79E06E39C3552D8D70125C0EF" authorityName="Zago, Yamada, Yamada, Franceschini, Bongiovani &amp; Silva" authorityYear="2020" family="Serrasalmidae" genus="Urocleidoide" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="tenuis">
<emphasis id="7DDAE9C17618D779DBE5B76EDA3AE011" italics="true">U. tenuis</emphasis>
</taxonomicName>
). During a study carried out in the Guandu River by the laboratory team, specimens of
<taxonomicName id="53739EB971767BDE3B12B3A25A4137E3" authorityName="Rossin &amp; Timi" authorityYear="2016" baseAuthorityName="Suriano" baseAuthorityYear="1981" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="triangulus">
<emphasis id="BBF420B7DCDC0153C463B0C011F0E460" italics="true">U. triangulus</emphasis>
</taxonomicName>
were found parasitizing
<taxonomicName id="A1BD252E39F85298B6B3FEEF730FBEB4" authority="(Quoy &amp; Gaimard, 1824)" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" family="Curimatidae" genus="Cyphocharax" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="gilbert">
<emphasis id="6A8951DD9E0DEFA81ABDF86664F90987" italics="true">Cyphocharax gilbert</emphasis>
(Quoy &amp; Gaimard, 1824)
</taxonomicName>
. The morphometry of the specimens studied herein were used to make comparisons with those of
<taxonomicName id="6C7599FE9DA30AC8158BFB3C9DE21C76" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="60885FA934623EB4D2B48FA52ECD070A" italics="true">U. paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CEFD5857E61F878C7E313B181C07C44" rank="species">sp. nov.</taxonomicNameLabel>
This was the first record in this host in
<collectingCountry id="3F8E28FE33F28599BDF076A4F5A91C91" name="Brazil">Brazil</collectingCountry>
.
</paragraph>
<caption id="B8C5E1965B46DD97426FD0F031F4981D" ID-DOI="10.3897/zoologia.38.e65001.figures12-21" ID-arpha="64A3C5ED-8986-50D7-8028-EE0EF5D93FAD" ID-zenodo_dep_id="5219829" httpUri="https://binary.pensoft.net/fig/576827" startId="F3">
<paragraph id="F86F4D586962DA020DD83F6F17CA580B">
<label id="DD4A486328805FB48E702F5F3995C194">Figures 1221.</label>
</paragraph>
<paragraph id="45292EF0A38F4C1EDF57FB32DD35C341">
<taxonomicName id="9181E3FC6D0A558722129E2FE8065C30" authorityName="de Freitas &amp; Bezerra &amp; Meneses &amp; Justo &amp; Viana &amp; Cohen" authorityYear="2021" class="Monogenea" family="Ancyrocephalidae" genus="Urocleidoides" kingdom="Animalia" order="Dactylogyridea" phylum="Platyhelminthes" rank="species" species="paratriangulus" status="sp. nov.">
<emphasis id="F2B768A5EACF44A94C614E323DF72FA3" italics="true">Urocleidoides paratriangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="951B924668E5E0B594C2E7AA08E60130" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="8FD4F34EF8EC779FE1F2861DFCDE5908" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" family="Serrasalmidae" genus="Psectogaster" kingdom="Animalia" order="Characiformes" phylum="Chordata" rank="species" species="amazonica">
<emphasis id="814FF1DE17BC3299A51F8FFFC85DF172" italics="true">Psectogaster amazonica</emphasis>
</taxonomicName>
from Tocantins River: (12) copulatory complex, ventral view; (13) vaginal sclerite; (14) vagina; (15, 16) dorsal bar; (17) ventral bar; (18) ventral anchor; (19) dorsal anchor, (20) hook pairs 1, 5; (21) hook pairs 24, 6, 7. Scale bars: 12 = 10 µm, 1314 = 10 µm, 1517 = 20 µm, 1819 = 20 µm, 20 = 5 µm, 21 = 10 µm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>