119 lines
10 KiB
XML
119 lines
10 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno" docDate="2015" docId="8664E66D8CF84C710BD0B2555D67CB20" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Acmella nana Vermeulen, Liew & Schilthuizen, sp. n." docType="treatment" docUuid="A2287C13-AE75-4E08-B3D9-BC093116E64A" docUuidSource="ZooBank" docVersion="10" lastPageNumber="8" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="7" updateTime="1668160884494" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>531</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>139</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
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<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="144057470" LSID="urn:lsid:zoobank.org:act:A2287C13-AE75-4E08-B3D9-BC093116E64A" httpUri="http://treatment.plazi.org/id/8664E66D8CF84C710BD0B2555D67CB20" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection pageId="6" pageNumber="7" type="multiple">
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<paragraph pageId="6" pageNumber="7">
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<pageBreakToken pageId="6" pageNumber="7" start="start">Taxon</pageBreakToken>
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classification Animalia Littorinimorpha Assimineidae
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="nomenclature">
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<paragraph pageId="6" pageNumber="7">
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<taxonomicName LSID="http://zoobank.org/A2287C13-AE75-4E08-B3D9-BC093116E64A" authority="Vermeulen, Liew & Schilthuizen" class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella nana" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="nana">Acmella nana Vermeulen, Liew & Schilthuizen</taxonomicName>
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<taxonomicNameLabel pageId="6" pageNumber="7">sp. n.</taxonomicNameLabel>
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Figure 5
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="reference_group">
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<paragraph pageId="7" pageNumber="8">
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella nana" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="nana">
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<pageBreakToken pageId="7" pageNumber="8" start="start">Acmella</pageBreakToken>
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nana
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</taxonomicName>
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nomen nudum:
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<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="7" pageNumber="8" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
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: 2761-2762.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="holotype">
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<paragraph pageId="7" pageNumber="8">Holotype. Malaysia, Sarawak, 4th Division, Niah Caves, South side of limestone area, West side of quarry, soil-filled crevice opened in quarry (leg J.J. Vermeulen, RMNH.5003950).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="materials_examined">
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<paragraph pageId="7" pageNumber="8">Examined material from Sabah.</paragraph>
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<paragraph pageId="7" pageNumber="8">Interior Province. Pinangah valley, Batu Urun (= Bukit Sinobang) (leg. J.J. Vermeulen, V 7978). Sepulut valley, Gua Pungiton (leg. J.J. Vermeulen & M. Schilthuizen, V 8075, BOR/MOL 529); Gua Sanaron (leg. J.J. Vermeulen & M. Schilthuizen, V 8063). Sandakan Province. Kinabatangan valley, Gomantong Hill 30 km South of Sandakan (leg. J.J. Vermeulen & H. Duistermaat, V 13268).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="description">
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<paragraph pageId="7" pageNumber="8">Description.</paragraph>
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<paragraph pageId="7" pageNumber="8">Shell minute (one of the smallest Borneo snail species), thin, translucent, white. Surface shiny. Spire conical with almost flat to distinctly convex sides, apex broadly obtuse, whorls moderately convex, sometimes slightly shouldered. Sculpture. Spiral sculpture predominant. Radial sculpture: scattered, inconspicuous growth lines only, grading into somewhat coarser riblets at irregular intervals. Spiral sculpture: rather conspicuous, densely placed and regularly spaced, fine, low and wide threads with only very narrow grooves in between. Aperture obliquely ovate in outline, with a slightly concave to slightly convex parietal side, transition from parietal to basal side rounded to obtusely angular. Umbilicus open, narrow. Dimensions. Height 0.60-0.79 mm; width 0.50-0.60 mm; h/w 1.00-1.35; number of whorls 2 7/8-3 7/8; height aperture 0.30-0.37 mm; width aperture 0.26-0.30 mm.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="habitat in sabah and distribution">
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<paragraph pageId="7" pageNumber="8">Habitat in Sabah and distribution.</paragraph>
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<paragraph pageId="7" pageNumber="8">(Disturbed) primary forest on limestone bedrock. Alt. 0-500 m. Sabah: Interior Province, South part; East coast: Kinabatangan River valley, 1 locality. Also in Sarawak and East Kalimantan. Endemic to Borneo.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="cross diagnosis">
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<paragraph pageId="7" pageNumber="8">Cross diagnosis.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Most similar to
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella caelata" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="caelata">Acmella caelata</taxonomicName>
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Vermeulen & Junau, 2007, from Sarawak. This species is consistently wider (c. 0.7 mm in
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella caelata" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="caelata">Acmella caelata</taxonomicName>
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, 0.50-0.60 mm in
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella nana" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="nana">Acmella nana</taxonomicName>
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). If specimens with the same number of whorls are compared, the whorls of
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella caelata" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="caelata">Acmella caelata</taxonomicName>
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are less depressed (at 3 1/8-3 3/8 whorl
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella caelata" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="caelata">Acmella caelata</taxonomicName>
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is 0.80-0.95 mm high,
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella nana" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="nana">Acmella nana</taxonomicName>
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only 0.60-0.75 mm high).
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</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Also similar to
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella ovoidea" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="ovoidea">Acmella ovoidea</taxonomicName>
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but smaller, with a narrower spire than juvenile
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<taxonomicName class="Gastropoda" family="Assimineidae" genus="Acmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acmella ovoidea" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="ovoidea">Acmella ovoidea</taxonomicName>
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of the same size. It also has coarser spiral sculpture.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="etymology">
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<paragraph pageId="7" pageNumber="8">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="8">The name refers to the size [nanus (L.) = dwarf].</paragraph>
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</subSubSection>
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</treatment>
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</document> |