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<mods:title id="49906EEB39ECA82EAE651CA0CA109199">Taxonomic notes concerning the genus Catharsius Hope, 1837 (Scarabaeidae Scarabaeinae)</mods:title>
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<mods:namePart id="D87A61021DACED0957F827431F5F0B3B">Takano, Hitoshi</mods:namePart>
<mods:affiliation id="8E5376CBD661FE089698C702AE5F15B5">African Natural History Research Trust, Street Court, Kingsland, Leominster HR 6 9 QA, UK. &amp; Department of Zoology, Oxford University, South Parks Road, Oxford OX 1 3 PS, UK &amp; Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<treatment id="865687A5FF96FFF827974E2C25A0FB4D" ID-DOI="http://doi.org/10.5281/zenodo.5579076" ID-GBIF-Taxon="189077016" ID-Zenodo-Dep="5579076" LSID="urn:lsid:plazi:treatment:865687A5FF96FFF827974E2C25A0FB4D" httpUri="http://treatment.plazi.org/id/865687A5FF96FFF827974E2C25A0FB4D" lastPageNumber="281" pageId="1" pageNumber="281">
<subSubSection id="46E56538FF96FFF827974E2C232AFEAB" box="[151,575,281,308]" pageId="1" pageNumber="281" type="nomenclature">
<paragraph id="0E4036B3FF96FFF827974E2C232AFEAB" blockId="1.[151,575,281,308]" box="[151,575,281,308]" pageId="1" pageNumber="281">
<heading id="550881DFFF96FFF827974E2C232AFEAB" bold="true" box="[151,575,281,308]" fontSize="11" level="1" pageId="1" pageNumber="281" reason="1">
<taxonomicName id="C9FF4D30FF96FFF827974E2C232AFEAB" ID-CoL="8R7Y8" authority="Harold, 1877" authorityName="Harold" authorityYear="1877" box="[151,575,281,308]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF827974E2C232AFEAB" bold="true" box="[151,575,281,308]" pageId="1" pageNumber="281">
<emphasis id="3C8BEAA1FF96FFF827974E2C2080FEAC" bold="true" box="[151,405,281,308]" italics="true" pageId="1" pageNumber="281">Catharsius harpagus</emphasis>
<bibRefCitation id="6A6E4B42FF96FFF8269C4E2C232AFEAB" author="Harold, E. von" box="[412,575,281,307]" pageId="1" pageNumber="281" pagination="97 - 111" refId="ref4465" refString="Harold, E. von (1877) Coleopterorum species novae. Mittheilungen der Munchner entomologischen Verein, 1, 97 - 111." type="journal article" year="1877">Harold, 1877</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="46E56538FF96FFF827974E6E201FFDE9" pageId="1" pageNumber="281" type="description">
<paragraph id="0E4036B3FF96FFF827974E6E201FFDE9" blockId="1.[151,1437,347,1237]" pageId="1" pageNumber="281">
Edgar von
<bibRefCitation id="6A6E4B42FF96FFF826144E6E20A1FEED" author="Harold, E. von" box="[276,436,347,373]" pageId="1" pageNumber="281" pagination="97 - 111" refId="ref4465" refString="Harold, E. von (1877) Coleopterorum species novae. Mittheilungen der Munchner entomologischen Verein, 1, 97 - 111." type="journal article" year="1877">Harold (1877)</bibRefCitation>
described this species from a vague locality of “
<emphasis id="3C8BEAA1FF96FFF824DD4E6E2566FEED" box="[989,1139,347,373]" italics="true" pageId="1" pageNumber="281">Afric. austral</emphasis>
.” based on at least a male and a female specimen, giving a size range of
<quantity id="C9079B56FF96FFF825AB4E4A223CFE01" box="[683,809,383,409]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="1" pageNumber="281" unit="mm" value="26.0" valueMax="30.0" valueMin="22.0">2230 mm</quantity>
. Although Harold stated in his original description that the type specimens of
<taxonomicName id="C9FF4D30FF96FFF8269A4E962333FE25" authorityName="Harold" authorityYear="1877" box="[410,550,419,445]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF8269A4E962333FE25" box="[410,550,419,445]" italics="true" pageId="1" pageNumber="281">C. harpagus</emphasis>
</taxonomicName>
were housed in ZMHB, the type series was found in the MNHN collections. This series consists of
<specimenCount id="18F9FD3AFF96FFF826A14EF22302FE79" box="[417,535,455,481]" pageId="1" pageNumber="281" type="male">two males</specimenCount>
and
<specimenCount id="18F9FD3AFF96FFF825544EF223C0FE79" box="[596,725,455,481]" pageId="1" pageNumber="281" type="female">one female</specimenCount>
, the larger male with a typical Harold type label (
<figureCitation id="96C42A36FF96FFF8221F4EF2249AFE79" box="[1311,1423,455,481]" captionStart="FIGURES 19" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 19. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="1" pageNumber="281">Figs. 12</figureCitation>
). Rather surprisingly, the
<specimenCount id="18F9FD3AFF96FFF826A64EDE230DFD9D" box="[422,536,491,517]" pageId="1" pageNumber="281" type="male">two males</specimenCount>
pertain to the Asian
<taxonomicName id="C9FF4D30FF96FFF824044EDE25C0FD9D" authority="Lansberge, 1874" authorityName="Lansberge" authorityYear="1874" box="[772,1237,491,517]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="birmanensis">
<emphasis id="3C8BEAA1FF96FFF824044EDE2505FD9D" box="[772,1040,491,517]" italics="true" pageId="1" pageNumber="281">Catharsius birmanensis</emphasis>
<bibRefCitation id="6A6E4B42FF96FFF823174EDE25C0FD9D" author="Lansberge, G. W. van" box="[1047,1237,491,517]" pageId="1" pageNumber="281" pagination="4 - 12" refId="ref4723" refString="Lansberge, G. W. van (1874) Description de quelque Lamellicornes Coprophages. Coleopterologische Hefte, 12 (2), 4 - 12." type="journal article" year="1874">Lansberge, 1874</bibRefCitation>
</taxonomicName>
, while the female specimen is a West African
<taxonomicName id="C9FF4D30FF96FFF826CD4D3A2244FDB1" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[461,849,527,553]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="phidias">
<emphasis id="3C8BEAA1FF96FFF826CD4D3A23B4FDB1" box="[461,673,527,553]" italics="true" pageId="1" pageNumber="281">Catharsius phidias</emphasis>
(Olivier, 1789)
</taxonomicName>
. In order to stabilise the taxonomy of this species by fixing the published name to a single specimen, the larger male with the following label data is here
<emphasis id="3C8BEAA1FF96FFF823F14D072017FDE9" bold="true" pageId="1" pageNumber="281">
designated the
<typeStatus id="D1448811FF96FFF827974D622017FDE9" box="[151,258,599,625]" pageId="1" pageNumber="281" type="lectotype">lectotype</typeStatus>
</emphasis>
:
</paragraph>
</subSubSection>
<subSubSection id="46E56538FF96FFF827C74D4E25A0FB4D" pageId="1" pageNumber="281" type="description">
<paragraph id="0E4036B3FF96FFF827C74D4E2467FD45" blockId="1.[151,1437,347,1237]" pageId="1" pageNumber="281">
“Afr. austr. / C. / Harpagus / Typ. Harold [cream-colored card with red border; handwritten] // Ex. Musaeo / E. Harold [cream-colored card with black border] // MUSEUM PARIS / ex coll. / OBERTHÜR / 1953 //
<typeStatus id="D1448811FF96FFF823FD4DAA2488FD21" box="[1277,1437,671,697]" pageId="1" pageNumber="281" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="C9FF4D30FF96FFF827A64DF62046FD45" authorityName="Hope" authorityYear="1837" box="[166,339,707,733]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="genus">CATHARSIUS</taxonomicName>
/ harpagus / Harold / det. H Takano 2017 [white card with red border; partially handwritten]”.
</paragraph>
<paragraph id="0E4036B3FF96FFF827C74DD2243FFC0A" blockId="1.[151,1437,347,1237]" pageId="1" pageNumber="281">
Louis
<bibRefCitation id="6A6E4B42FF96FFF826104DD220C1FC99" author="Peringuey, L." box="[272,468,743,769]" pageId="1" pageNumber="281" pagination="1 - 563" refId="ref4905" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae &amp; Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey (1901)</bibRefCitation>
synonymised
<taxonomicName id="C9FF4D30FF96FFF8257A4DD22213FC99" authorityName="Harold" authorityYear="1877" box="[634,774,743,769]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF8257A4DD22213FC99" box="[634,774,743,769]" italics="true" pageId="1" pageNumber="281">C. harpagus</emphasis>
</taxonomicName>
under
<taxonomicName id="C9FF4D30FF96FFF824574DD225FBFC99" authority="Boheman, 1857" authorityName="Boheman" authorityYear="1857" box="[855,1262,742,769]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF824574DD22527FC99" box="[855,1074,743,769]" italics="true" pageId="1" pageNumber="281">Catharsius laticeps</emphasis>
<bibRefCitation id="6A6E4B42FF96FFF8233B4DD225FBFC99" author="Boheman, C. H." box="[1083,1262,742,769]" pageId="1" pageNumber="281" refId="ref3963" refString="Boheman, C. H. (1857) Insecta Caffraria annis 1838 - 1845 a J. A. Wahlberg collecta amici auxilios sultus descripsit. Pars II. Coleoptera (Scarabaeidae). Officina Norstedtiana, Holmiae, 395 pp." type="book" year="1857">Boheman, 1857</bibRefCitation>
</taxonomicName>
without giving any justification.
<taxonomicName id="C9FF4D30FF96FFF826584C3E2325FCBD" authorityName="Boheman" authorityYear="1857" box="[344,560,779,805]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF826584C3E2325FCBD" box="[344,560,779,805]" italics="true" pageId="1" pageNumber="281">Catharsius laticeps</emphasis>
</taxonomicName>
was described from a female specimen by Boheman and Péringuey incorrectly associated with it a male which belonged to a different species. The male specimen he described and figured as
<taxonomicName id="C9FF4D30FF96FFF822834C1A21FBFCF5" authorityName="Boheman" authorityYear="1857" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF822834C1A21FBFCF5" italics="true" pageId="1" pageNumber="281">C. laticeps</emphasis>
</taxonomicName>
(
<figureCitation id="96C42A36FF96FFF827FD4C66202AFCF5" box="[253,319,851,877]" captionStart="FIGURES 19" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 19. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="1" pageNumber="281">Fig. 3</figureCitation>
) is unmistakably
<taxonomicName id="C9FF4D30FF96FFF825054C6623F6FCF5" authority="Harold" authorityName="Harold" authorityYear="1877" box="[517,739,851,877]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF825054C66239AFCF5" box="[517,655,851,877]" italics="true" pageId="1" pageNumber="281">C. harpagus</emphasis>
Harold
</taxonomicName>
and so, despite the incorrect assumption of this being the male of
<taxonomicName id="C9FF4D30FF96FFF827B64C422038FC09" authorityName="Boheman" authorityYear="1857" box="[182,301,887,913]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF827B64C422038FC09" box="[182,301,887,913]" italics="true" pageId="1" pageNumber="281">C. laticeps</emphasis>
</taxonomicName>
, he correctly synonymised
<taxonomicName id="C9FF4D30FF96FFF825624C4223F8FC09" authorityName="Harold" authorityYear="1877" box="[610,749,887,913]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF825624C4223F8FC09" box="[610,749,887,913]" italics="true" pageId="1" pageNumber="281">C. harpagus</emphasis>
</taxonomicName>
under
<taxonomicName id="C9FF4D30FF96FFF8243A4C422565FC09" authority="sensu Peringuey" authorityName="Peringuey" box="[826,1136,887,913]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF8243A4C4222A7FC09" box="[826,946,887,913]" italics="true" pageId="1" pageNumber="281">C. laticeps</emphasis>
sensu Péringuey
</taxonomicName>
(nec Boheman).
</paragraph>
<paragraph id="0E4036B3FF96FFF827C74CAE23D7FBB9" blockId="1.[151,1437,347,1237]" pageId="1" pageNumber="281">
Maria Corinta
<bibRefCitation id="6A6E4B42FF96FFF826694CAE2307FC2D" author="Ferreira, M. C." box="[361,530,923,949]" pageId="1" pageNumber="281" pagination="5 - 6" refId="ref4166" refString="Ferreira, M. C. (1971) Contribuicao para o estudo dos Escarabideos Africanos. Descricao de uma especie nova de Catharsius s. str. Novos Taxa Entomologicos, 96, 5 - 6." type="journal article" year="1971">Ferreira (1971)</bibRefCitation>
re-examined the very male specimen upon which Péringuey based his description and correctly stated that this is not a male of
<taxonomicName id="C9FF4D30FF96FFF825904C8A2212FC41" box="[656,775,959,985]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF825904C8A2212FC41" box="[656,775,959,985]" italics="true" pageId="1" pageNumber="281">C. laticeps</emphasis>
</taxonomicName>
nor is
<taxonomicName id="C9FF4D30FF96FFF824554C8A2590FC41" authority="sensu Ferreira" authorityName="sensu Ferreira" authorityYear="1960" box="[853,1157,959,985]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF824554C8A22F5FC41" box="[853,992,959,985]" italics="true" pageId="1" pageNumber="281">C. harpagus</emphasis>
sensu Ferreira
</taxonomicName>
(nec Harold) a synonym of
<taxonomicName id="C9FF4D30FF96FFF827B44CD6203EFC65" box="[180,299,995,1021]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3C8BEAA1FF96FFF827B44CD6203EFC65" box="[180,299,995,1021]" italics="true" pageId="1" pageNumber="281">C. laticeps</emphasis>
</taxonomicName>
. She proceeded to describe this specimen as
<taxonomicName id="C9FF4D30FF96FFF8241C4CD6255FFC65" authorityName="Ferreira" authorityYear="1971" box="[796,1098,995,1021]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="parafastidiosus">
<emphasis id="3C8BEAA1FF96FFF8241C4CD6255FFC65" box="[796,1098,995,1021]" italics="true" pageId="1" pageNumber="281">Catharsius parafastidiosus</emphasis>
</taxonomicName>
. The
<typeStatus id="D1448811FF96FFF823874CD625FCFC65" box="[1159,1257,995,1021]" pageId="1" pageNumber="281" type="holotype">holotype</typeStatus>
in SANC (
<figureCitation id="96C42A36FF96FFF822644CD621D6FBB9" captionStart="FIGURES 19" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 19. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="1" pageNumber="281">Figs. 45</figureCitation>
) with the following label data was examined:
</paragraph>
<paragraph id="0E4036B3FF96FFF827C74B1E200CFB29" blockId="1.[151,1437,347,1237]" pageId="1" pageNumber="281">
<materialsCitation id="BE973CEEFF96FFF827D14B1E2000FB29" ID-GBIF-Occurrence="3387267306" collectionCode="SANC" country="South Africa" determinerName="M. C. Ferreira" location="Pretoria" pageId="1" pageNumber="281" specimenCount="3" specimenCount-male="3" typeStatus="holotype">
no 3 /
<taxonomicName id="C9FF4D30FF96FFF826154B1E20AEFBDD" box="[277,443,1067,1093]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="laticeps">Copris laticeps</taxonomicName>
/
<specimenCount id="18F9FD3AFF96FFF826CD4B1E20F7FBDD" box="[461,482,1067,1093]" pageId="1" pageNumber="281" type="male"></specimenCount>
[cream-colored paper; handwritten in Péringueys hand] // HOLOTIPO
<specimenCount id="18F9FD3AFF96FFF823FF4B1E2401FBDD" box="[1279,1300,1067,1093]" pageId="1" pageNumber="281" type="male"></specimenCount>
/
<taxonomicName id="C9FF4D30FF96FFF822264B1E2489FBDD" authorityName="Hope" authorityYear="1837" box="[1318,1436,1067,1093]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="genus">Catharsius</taxonomicName>
parafas- / tidiosus n.sp. /
<determinerName id="FB9CDA77FF96FFF826B94B7A2341FBF1" box="[441,596,1103,1129]" pageId="1" pageNumber="281">M.C. Ferreira</determinerName>
det., 1971 [partially handwritten in Ferreiras hand] //
<typeStatus id="D1448811FF96FFF823C74B7A2478FBF1" box="[1223,1389,1103,1129]" pageId="1" pageNumber="281" type="holotype">HOLOTYPUS</typeStatus>
/
<specimenCount id="18F9FD3AFF96FFF822874B7A2489FBF1" box="[1415,1436,1103,1129]" pageId="1" pageNumber="281" type="male"></specimenCount>
[red card] // NATIONAL COLL. / OF INSECTS (
<collectionCode id="68EEAE76FF96FFF825D84B462233FB15" box="[728,806,1139,1165]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="1" pageNumber="281" type="Museum">SANC</collectionCode>
) /
<location id="0B206068FF96FFF824484B4622B1FB15" LSID="urn:lsid:plazi:treatment:865687A5FF96FFF827974E2C25A0FB4D:0B206068FF96FFF824484B4622B1FB15" box="[840,932,1139,1165]" country="South Africa" name="Pretoria" pageId="1" pageNumber="281">Pretoria</location>
,
<collectingCountry id="76E87623FF96FFF824B14B462557FB15" box="[945,1090,1139,1165]" name="South Africa" pageId="1" pageNumber="281">South Africa</collectingCountry>
[black border] // TYPH00479 [red card]”
</materialsCitation>
.
</paragraph>
<paragraph id="0E4036B3FF96FFF827C74B8E25A0FB4D" blockId="1.[151,1437,347,1237]" box="[199,1205,1211,1237]" pageId="1" pageNumber="281">
Both
<taxonomicName id="C9FF4D30FF96FFF826054B8E209AFB4D" authorityName="sensu Ferreira" authorityYear="1960" box="[261,399,1211,1237]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="harpagus">
<emphasis id="3C8BEAA1FF96FFF826054B8E209AFB4D" box="[261,399,1211,1237]" italics="true" pageId="1" pageNumber="281">C. harpagus</emphasis>
</taxonomicName>
and
<taxonomicName id="C9FF4D30FF96FFF826C64B8E2386FB4D" authorityName="Ferreira" authorityYear="1971" box="[454,659,1211,1237]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="parafastidiosus">
<emphasis id="3C8BEAA1FF96FFF826C64B8E2386FB4D" box="[454,659,1211,1237]" italics="true" pageId="1" pageNumber="281">C. parafastidiosus</emphasis>
</taxonomicName>
are here synonymised with
<taxonomicName id="C9FF4D30FF96FFF824CE4B8E256DFB4D" authorityName="Lansberge" authorityYear="1874" box="[974,1144,1211,1237]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="281" phylum="Arthropoda" rank="species" species="birmanensis">
<emphasis id="3C8BEAA1FF96FFF824CE4B8E256DFB4D" box="[974,1144,1211,1237]" italics="true" pageId="1" pageNumber="281">C. birmanensis</emphasis>
</taxonomicName>
thus:
</paragraph>
</subSubSection>
</treatment>
</document>