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<mods:title id="E3B0CB427BB2867513DA4570F4898DE3">Redescription of Astyanax guianensis Eigenmann 1909 (Characiformes: Characidae), a poorly known and widespread fish from the Amazon, Orinoco and Guiana Shield drainages</mods:title>
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<taxonomicName id="C9E01BE64A7ADE21BCA582015A26F9FC" ID-CoL="J5S2" authority="Eigenmann 1909" authorityName="Eigenmann" authorityYear="1909" box="[151,618,1591,1617]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="569" phylum="Chordata" rank="species" species="guianensis">
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<emphasis id="3C94BC774A7ADE21BCA5820159DCF9FC" bold="true" box="[151,400,1591,1617]" italics="true" pageId="1" pageNumber="569">Astyanax guianensis</emphasis>
<bibRefCitation id="6A711D944A7ADE21BDAA82015A26F9FC" author="Eigenmann" box="[408,618,1591,1617]" pageId="1" pageNumber="569" refString="Eigenmann, C. H. (1909) Reports on the expedition to British Guiana of the Indiana University and the Carnegie Museum, 1908. Report no. 1. Some new genera and species of fishes from British Guiana. Annals of the Carnegie Museum, 6, 4 - 54." type="journal article" year="1909">Eigenmann 1909</bibRefCitation>
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<figureCitation id="96DB7CE04A7ADE21BCA5826C594DF9DF" box="[151,257,1626,1651]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1542,1564]" captionStartId-1="5.[151,250,901,923]" captionTargetBox-0="[340,1247,193,1520]" captionTargetBox-1="[346,1236,202,872]" captionTargetId-0="figure@3.[340,1248,193,1521]" captionTargetId-1="figure@5.[339,1247,193,879]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURE 1. Astyanax guianensis: (a) holotype, FMNH 51518, 41.0 mm SL, Guyana, rio Essequibo basin; (b) MZUSP 25526, 53.3. mm SL, Brazil, Pará, rio Tapajós basin; (c) MZUSP 115554, 35.7 mm SL, Brazil, Mato Grosso, rio Guaporé basin; and (d) live specimen, Brazil, Amazonas, rio Negro at Santa Isabel do Rio Negro, not preserved." captionText-1="FIGURE 2. Upper and lower jaws of Astyanax guianensis (medial view, right side), MZUSP 109674, 46.7 mm SL. Coronomeckelian bone is missing." httpUri-0="https://zenodo.org/record/234680/files/figure.png" httpUri-1="https://zenodo.org/record/234681/files/figure.png" pageId="1" pageNumber="569">Figs. 1–2</figureCitation>
, 
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<emphasis id="3C94BC774A7ADE21BCA58295593AF915" box="[151,374,1699,1720]" italics="true" pageId="1" pageNumber="569">Astyanax guianensis</emphasis>
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: 16
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at Warraputa cataract, 
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[citation].— 
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[closely allied to 
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: 350
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[identification key; literature compilation].— 
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: 274
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, plate 53, Fig. 4 [identification key; literature compilation].— 
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: 328
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[literature compilation].— 
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: 60
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: 22
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[
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: 426
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[identification key].— 
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Nijssen 
<emphasis id="3C94BC774A7ADE21B88383285CA9F89F" box="[1201,1253,1820,1843]" italics="true" pageId="1" pageNumber="569">et al</emphasis>
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: 14
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: 47
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[
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: 12
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[
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: 6
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[
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Nijssen 
<emphasis id="3C94BC774A7ADE21BDBC836D59F3F8DD" box="[398,447,1882,1904]" italics="true" pageId="1" pageNumber="569">et al</emphasis>
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[literature compilation].— 
<treatmentCitation id="8F4146744A7ADE21B8CF836C5DD7F8DD" author="Lima" box="[1277,1435,1882,1904]" page="29" pageId="1" pageNumber="569" year="2009">
<bibRefCitation id="6A711D944A7ADE21B8CF836C5D39F8DD" author="Lima" box="[1277,1397,1882,1904]" pageId="1" pageNumber="569" refString="Lima, F. C. T. (2009) Genera Incerta Sedis, family Characidae. In: Vari, R. P., Ferraris-Jr., C. J., Radosavljevic, A. &amp; Funk, V. A. (Eds.), Checklist of the freshwater fishes of the Guiana Shield. - Bulletin of the Biological Society of Washington, pp. 29 - 32." type="book chapter" year="2009">Lima 2009</bibRefCitation>
: 29
</treatmentCitation>
[
<collectingCountry id="76F720F54A7ADE21BCFE834E5954F823" box="[204,280,1912,1934]" name="Guyana" pageId="1" pageNumber="569">Guiana</collectingCountry>
Shield at Amazonas and Bolivar States, 
<collectingCountry id="76F720F54A7ADE21BE86834E5B6CF823" box="[692,800,1912,1934]" name="Venezuela" pageId="1" pageNumber="569">Venezuela</collectingCountry>
].— 
<treatmentCitation id="8F4146744A7ADE21BF71834E5C53F823" author="Lima" box="[835,1055,1912,1935]" page="109" pageId="1" pageNumber="569" year="2003">
<bibRefCitation id="6A711D944A7ADE21BF71834E5BA3F823" author="Lima" box="[835,1007,1912,1935]" pageId="1" pageNumber="569" refString="Lima, F. C. T., Malabarba, L. R., Buckup, P. A., Silva, J. F. P., Vari, R. P, Harold, A., Benine, R. C., Oyakawa, O., Pavanelli, C. S., Menezes, N. A., Lucena, C. A. S., Malabarba, M. C. S. L., Lucena, Z. M. S., Reis, R. E., Langeani, F., Casatti, L., Bertaco, V. A., Moreira, C. &amp; Lucinda, P. H. F. (2003) Genera incertae sedis in Characidae. In: Reis, R. E., Kullander, S. O. &amp; Ferraris-Jr, C. J. (Eds.), Check List of Freshwater Fishes of South and Central America. Edipucrs, Porto Alegre, pp. 106 - 169." type="book chapter" year="2003">
Lima 
<emphasis id="3C94BC774A7ADE21BFB2834C5BE0F823" box="[896,940,1912,1935]" italics="true" pageId="1" pageNumber="569">et al</emphasis>
. 2003
</bibRefCitation>
: 109
</treatmentCitation>
[questionably 
<collectingCountry id="76F720F54A7ADE21B887834E5D6DF823" box="[1205,1313,1912,1934]" name="Venezuela" pageId="1" pageNumber="569">Venezuela</collectingCountry>
].— 
<treatmentCitation id="8F4146744A7ADE21B971834E59E3F800" author="Marinho" page="46" pageId="1" pageNumber="569" year="2013">
<bibRefCitation id="6A711D944A7ADE21B971834E59C5F800" author="Marinho" pageId="1" pageNumber="569" refString="Marinho, M. M. F. &amp; Birindelli, J. L. O. (2013) Redescription of Astyanax multidens Eigenmann, 1908 (Characiformes: Characidae), a small characid of the Brazilian Amazon. Neotropical Ichthyology, 11, 45 - 54. http: // dx. doi. org / 10.1590 / S 1679 - 62252013000100005" type="journal article" year="2013">Marinho &amp; Birindelli 2013</bibRefCitation>
: 46
</treatmentCitation>
[similar to 
<taxonomicName id="C9E01BE64A7ADE21BE0683AE5AB2F800" ID-CoL="J5TM" box="[564,766,1943,1965]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="569" phylum="Chordata" rank="species" species="multidens">
<emphasis id="3C94BC774A7ADE21BE0683AE5AB2F800" box="[564,766,1943,1965]" italics="true" pageId="1" pageNumber="569">Astyanax multidens</emphasis>
</taxonomicName>
and 
<taxonomicName id="C9E01BE64A7ADE21BF0B83AE5B99F800" ID-CoL="J5RY" box="[825,981,1944,1965]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="569" phylum="Chordata" rank="species" species="guaporensis">
<emphasis id="3C94BC774A7ADE21BF0B83AE5B99F800" box="[825,981,1944,1965]" italics="true" pageId="1" pageNumber="569">A. guaporensis</emphasis>
</taxonomicName>
].—Marinho &amp; Ohara 2013: 476 [tentative identification of material from rio Beni].
</treatmentCitationGroup>
</paragraph>
<paragraph id="0E5F60654A7ADE21BCA583E25C1AF847" blockId="1.[151,1436,1698,2027]" box="[151,1110,2004,2027]" pageId="1" pageNumber="569">
<treatmentCitationGroup id="2EF0474B4A7ADE21BCA583E25C1AF847" box="[151,1110,2004,2027]" pageId="1" pageNumber="569">
<taxonomicName id="C9E01BE64A7ADE21BCA583E25953F847" ID-CoL="63BXC" box="[151,287,2004,2026]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="569" phylum="Chordata" rank="genus">
<emphasis id="3C94BC774A7ADE21BCA583E25953F847" box="[151,287,2004,2026]" italics="true" pageId="1" pageNumber="569">Moenkhausia</emphasis>
</taxonomicName>
sp. “ 
<taxonomicName id="C9E01BE64A7ADE21BD6683E259EDF847" ID-CoL="6RMXR" box="[340,417,2004,2026]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="569" phylum="Chordata" rank="species" species="collettii">
<emphasis id="3C94BC774A7ADE21BD6683E259EDF847" box="[340,417,2004,2026]" italics="true" pageId="1" pageNumber="569">collettii</emphasis>
</taxonomicName>
alta”.— 
<treatmentCitation id="8F4146744A7ADE21BDC683E25A9EF847" author="Lima" box="[500,722,2004,2027]" page="322" pageId="1" pageNumber="569" year="2013">
<bibRefCitation id="6A711D944A7ADE21BDC683E25AEEF847" author="Lima" box="[500,674,2004,2027]" pageId="1" pageNumber="569" refString="Lima, F. C. T., Pires, T. H. S., Ohara, W. M., Jerep, F. C., Carvalho, F. R., Marinho, M. M. F. &amp; Zuanon, J. A. S. (2013) Characidae. In: Queiroz, L. J., Torrente-Vilara, G., Ohara, W. M., Pires, T. H. S., Zuanon, J. &amp; Doria, C. R. C (Eds.), Peixes do rio Madeira. Vol. I. Dialeto, Sao Paulo, pp. 211 - 395." type="book chapter" year="2013">
Lima 
<emphasis id="3C94BC774A7ADE21BE0183E05A2AF847" box="[563,614,2004,2027]" italics="true" pageId="1" pageNumber="569">et al.</emphasis>
2013
</bibRefCitation>
: 322
</treatmentCitation>
[rio Madeira basin, brief description].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="46FA33EE4A79DE22BCA584A15B5FFDB5" pageId="2" pageNumber="570" type="diagnosis">
<paragraph id="0E5F60654A79DE22BCA584A15B5FFDB5" blockId="2.[151,1436,151,789]" pageId="2" pageNumber="570">
<emphasis id="3C94BC774A79DE22BCA584A1595DFF1D" bold="true" box="[151,273,151,176]" pageId="2" pageNumber="570">Diagnosis.</emphasis>
<taxonomicName id="C9E01BE64A79DE22BD2F84AE5A45FF02" box="[285,521,152,175]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A79DE22BD2F84AE5A45FF02" box="[285,521,152,175]" italics="true" pageId="2" pageNumber="570">Astyanax guianensis</emphasis>
</taxonomicName>
can be distinguished from all congeners, except 
<taxonomicName id="C9E01BE64A79DE22B86B84AE5D3FFF1D" authority="Regan" authorityName="Regan" box="[1113,1395,152,176]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="angustifrons">
<emphasis id="3C94BC774A79DE22B86B84AE5D45FF1D" box="[1113,1289,152,176]" italics="true" pageId="2" pageNumber="570">A. angustifrons</emphasis>
(Regan)
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B9B684AE59F7FF78" authority="Eigenmann" authorityName="Eigenmann" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="aurocaudatus">
<emphasis id="3C94BC774A79DE22B9B684AE597CFF79" italics="true" pageId="2" pageNumber="570">A. aurocaudatus</emphasis>
Eigenmann
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BDFB848A5A34FF78" authority="Dahl" authorityName="Dahl" box="[457,632,188,213]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="gisleni">
<emphasis id="3C94BC774A79DE22BDFB848A5A7AFF79" box="[457,566,188,212]" italics="true" pageId="2" pageNumber="570">A. gisleni</emphasis>
Dahl
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BEB4848A5B69FF79" box="[646,805,188,212]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="goyanensis">
<emphasis id="3C94BC774A79DE22BEB4848A5B69FF79" box="[646,805,188,212]" italics="true" pageId="2" pageNumber="570">A. goyanensis</emphasis>
</taxonomicName>
(Miranda Ribeiro), 
<taxonomicName id="C9E01BE64A79DE22B83F848A5CFBFF79" box="[1037,1207,188,212]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guaporensis">
<emphasis id="3C94BC774A79DE22B83F848A5CFBFF79" box="[1037,1207,188,212]" italics="true" pageId="2" pageNumber="570">A. guaporensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B8F5848A58BDFF55" authority="Melo &amp; Buckup" authorityName="Melo &amp; Buckup" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="henseli">
<emphasis id="3C94BC774A79DE22B8F5848A5D74FF79" box="[1223,1336,188,212]" italics="true" pageId="2" pageNumber="570">A. henseli</emphasis>
Melo &amp; Buckup
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BCCF84D65AF6FF55" authority="Gery, Planquette &amp; Le Bail" authorityName="Gery, Planquette &amp; Le Bail" box="[253,698,223,248]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="leopoldi">
<emphasis id="3C94BC774A79DE22BCCF84D65935FF55" box="[253,377,224,248]" italics="true" pageId="2" pageNumber="570">A. leopoldi</emphasis>
Géry, Planquette &amp; Le Bail
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BEF484D65B18FF55" box="[710,852,224,248]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="multidens">
<emphasis id="3C94BC774A79DE22BEF484D65B18FF55" box="[710,852,224,248]" italics="true" pageId="2" pageNumber="570">A. multidens</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BF5184D65C68FF55" authority="Meek" authorityName="Meek" box="[867,1060,223,248]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="nasutus">
<emphasis id="3C94BC774A79DE22BF5184D65B95FF55" box="[867,985,224,248]" italics="true" pageId="2" pageNumber="570">A. nasutus</emphasis>
Meek
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B80284D65D3BFF55" authority="Eigenmann" authorityName="Eigenmann" box="[1072,1399,223,248]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="nicaraguensis">
<emphasis id="3C94BC774A79DE22B80284D65CA1FF5A" box="[1072,1261,224,247]" italics="true" pageId="2" pageNumber="570">A. nicaraguensis</emphasis>
Eigenmann
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B9B684D65902FEB0" authority="Myers" authorityName="Myers" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="superbus">
<emphasis id="3C94BC774A79DE22B9B684D658B0FEB1" italics="true" pageId="2" pageNumber="570">A. superbus</emphasis>
Myers
</taxonomicName>
, and 
<taxonomicName id="C9E01BE64A79DE22BDB985325AF7FEB0" authority="Haluch &amp; Abilhoa" authorityName="Haluch &amp; Abilhoa" box="[395,699,260,285]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="totae">
<emphasis id="3C94BC774A79DE22BDB9853259AFFEB1" box="[395,483,260,284]" italics="true" pageId="2" pageNumber="570">A. totae</emphasis>
Haluch &amp; Abilhoa
</taxonomicName>
by having five to 10 maxillary teeth (vs. none to four). 
<taxonomicName id="C9E01BE64A79DE22B90785325943FE92" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A79DE22B90785325943FE92" italics="true" pageId="2" pageNumber="570">Astyanax guianensis</emphasis>
</taxonomicName>
is distinguished from the aforementioned species, except 
<taxonomicName id="C9E01BE64A79DE22BFC6851E5CE6FEED" box="[1012,1194,296,320]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="angustifrons">
<emphasis id="3C94BC774A79DE22BFC6851E5CE6FEED" box="[1012,1194,296,320]" italics="true" pageId="2" pageNumber="570">A. angustifrons</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B8F1851E5D26FE92" box="[1219,1386,296,319]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="goyanensis">
<emphasis id="3C94BC774A79DE22B8F1851E5D26FE92" box="[1219,1386,296,319]" italics="true" pageId="2" pageNumber="570">A. goyanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B9B6851E5953FEC9" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guaporensis">
<emphasis id="3C94BC774A79DE22B9B6851E5953FEC9" italics="true" pageId="2" pageNumber="570">A. guaporensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BD1F857A59E5FEC9" box="[301,425,332,356]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="leopoldi">
<emphasis id="3C94BC774A79DE22BD1F857A59E5FEC9" box="[301,425,332,356]" italics="true" pageId="2" pageNumber="570">A. leopoldi</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BD85857A5A08FEC9" box="[439,580,332,356]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="multidens">
<emphasis id="3C94BC774A79DE22BD85857A5A08FEC9" box="[439,580,332,356]" italics="true" pageId="2" pageNumber="570">A. multidens</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BE61857A5B5CFEC9" box="[595,784,332,356]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="nicaraguensis">
<emphasis id="3C94BC774A79DE22BE61857A5B5CFEC9" box="[595,784,332,356]" italics="true" pageId="2" pageNumber="570">A. nicaraguensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C9E01BE64A79DE22BF7C857A5BEAFEC9" box="[846,934,332,356]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="totae">
<emphasis id="3C94BC774A79DE22BF7C857A5BEAFEC9" box="[846,934,332,356]" italics="true" pageId="2" pageNumber="570">A. totae</emphasis>
</taxonomicName>
, by having 31 to 35 pored lateral-line scales (vs. more than 35). It can be distinguished from 
<taxonomicName id="C9E01BE64A79DE22BEF085465B21FE25" box="[706,877,368,392]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="angustifrons">
<emphasis id="3C94BC774A79DE22BEF085465B21FE25" box="[706,877,368,392]" italics="true" pageId="2" pageNumber="570">A. angustifrons</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22BF4F85465C55FE2A" box="[893,1049,368,391]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="goyanensis">
<emphasis id="3C94BC774A79DE22BF4F85465C55FE2A" box="[893,1049,368,391]" italics="true" pageId="2" pageNumber="570">A. goyanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B81885465CEAFE25" box="[1066,1190,368,392]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="leopoldi">
<emphasis id="3C94BC774A79DE22B81885465CEAFE25" box="[1066,1190,368,392]" italics="true" pageId="2" pageNumber="570">A. leopoldi</emphasis>
</taxonomicName>
, 
<taxonomicName id="C9E01BE64A79DE22B88785465D0FFE25" box="[1205,1347,368,392]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="multidens">
<emphasis id="3C94BC774A79DE22B88785465D0FFE25" box="[1205,1347,368,392]" italics="true" pageId="2" pageNumber="570">A. multidens</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C9E01BE64A79DE22B9B685465978FE01" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="nicaraguensis">
<emphasis id="3C94BC774A79DE22B9B685465978FE01" italics="true" pageId="2" pageNumber="570">A. nicaraguensis</emphasis>
</taxonomicName>
by having the middle caudal-fin rays hyaline (vs. middle caudal-fin rays dark), from 
<taxonomicName id="C9E01BE64A79DE22B8C685A25DD0FE01" box="[1268,1436,404,428]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guaporensis">
<emphasis id="3C94BC774A79DE22B8C685A25DD0FE01" box="[1268,1436,404,428]" italics="true" pageId="2" pageNumber="570">A. guaporensis</emphasis>
</taxonomicName>
by having five, rarely six (4 out of 59 examined specimens) horizontal scale row above lateral line (vs. six to seven) and the absence of a midlateral series of dark anteriorly-directed chevrons (vs. presence), and from 
<taxonomicName id="C9E01BE64A79DE22B92E85EA5D3AFE59" box="[1308,1398,476,500]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="totae">
<emphasis id="3C94BC774A79DE22B92E85EA5D3AFE59" box="[1308,1398,476,500]" italics="true" pageId="2" pageNumber="570">A. totae</emphasis>
</taxonomicName>
by presenting 21 to 25 branched anal-fin rays (vs. 15 to 18).
</paragraph>
</subSubSection>
<subSubSection id="46FA33EE4A79DE24BCF586125B3BF8BE" lastPageId="4" lastPageNumber="572" pageId="2" pageNumber="570" type="description">
<paragraph id="0E5F60654A79DE22BCF586125B88FCB8" blockId="2.[151,1436,151,789]" pageId="2" pageNumber="570">
<emphasis id="3C94BC774A79DE22BCF586125915FD90" bold="true" box="[199,345,548,573]" pageId="2" pageNumber="570">Description.</emphasis>
Morphometric data for 
<taxonomicName id="C9E01BE64A79DE22BE4386125B16FD91" box="[625,858,548,572]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A79DE22BE4386125B16FD91" box="[625,858,548,572]" italics="true" pageId="2" pageNumber="570">Astyanax guianensis</emphasis>
</taxonomicName>
presented in 
<tableCitation id="436255DE4A79DE22BFCA86125C03FD91" box="[1016,1103,548,573]" captionStart="TABLE 1" captionStartId="2.[151,239,832,855]" captionTargetBox="[156,1366,914,1944]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data of Astyanax guianensis. SD for Standard Deviation. The mean and SD include the values of the holotype, paratypes, and non-type specimens." httpUri="http://table.plazi.org/id/5A9F30ED4A79DE22BCA587765A8EFCDA" pageId="2" pageNumber="570" tableUuid="5A9F30ED4A79DE22BCA587765A8EFCDA">Table 1</tableCitation>
. Small sized species (largest examined specimen 
<quantity id="C918CD804A79DE22BD4E867159ACFDF2" box="[380,480,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="5.33" pageId="2" pageNumber="570" unit="mm" value="53.3">53.3 mm</quantity>
SL). Body compressed laterally, moderately elongate. Greatest body depth at dorsalfin origin. Dorsal profile of head convex from tip of snout to vertical through anterior border of anterior nostril and straight from that point to tip of supraoccipital spine. Dorsal profile of body straight to slightly convex at predorsal region, straight at dorsal-fin base, straight from end of dorsal-fin base to adipose-fin origin, and concave along caudal peduncle. Ventral profile of head and body slightly convex from anterior tip of lower lip to anal-fin origin, straight along anal-fin base, and slightly concave along caudal peduncle.
</paragraph>
<caption id="5A9F30ED4A79DE22BCA587765A8EFCDA" ID-Table-UUID="5A9F30ED4A79DE22BCA587765A8EFCDA" httpUri="http://table.plazi.org/id/5A9F30ED4A79DE22BCA587765A8EFCDA" pageId="2" pageNumber="570" targetBox="[156,1366,914,1944]" targetIsTable="true" targetPageId="2">
<paragraph id="0E5F60654A79DE22BCA587765A8EFCDA" blockId="2.[151,1436,831,887]" pageId="2" pageNumber="570">
<emphasis id="3C94BC774A79DE22BCA587765946FCFA" bold="true" box="[151,266,832,855]" pageId="2" pageNumber="570">TABLE 1.</emphasis>
Morphometric data of 
<taxonomicName id="C9E01BE64A79DE22BE3087095A90FCFA" box="[514,732,831,855]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="570" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A79DE22BE3087095A90FCFA" box="[514,732,831,855]" italics="true" pageId="2" pageNumber="570">Astyanax guianensis</emphasis>
</taxonomicName>
. SD for Standard Deviation. The mean and SD include the values of the holotype, paratypes, and non-type specimens.
</paragraph>
</caption>
<paragraph id="0E5F60654A79DE22BE0C87A45D1AF835" pageId="2" pageNumber="570">
<table id="7CE092C54A7921DFBCAE87A45D1AF835" box="[156,1366,914,1944]" gridcols="5" gridrows="24" pageId="2" pageNumber="570">
<tr id="B0D062274A7921DFBCAE87A45D1AFC05" box="[156,1366,914,936]" gridrow="0" pageId="2" pageNumber="570" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<th id="F3010B5B4A7921DFBE0C87A45B18FC05" box="[574,852,914,936]" gridcol="1" gridrow="0" pageId="2" pageNumber="570">Holotype Paratypes</th>
<th id="F3010B5B4A7921DFBF9887A45C35FC05" box="[938,1145,914,936]" gridcol="2" gridrow="0" pageId="2" pageNumber="570">Non-type specimens</th>
</tr>
<tr id="B0D062274A7921DFBCAE878A5D1AFC7F" box="[156,1366,956,978]" gridrow="1" pageId="2" pageNumber="570" rowspan-0="1">
<td id="F3010B5B4A7921DFBE0C878A5B18FC7F" box="[574,852,956,978]" gridcol="1" gridrow="1" pageId="2" pageNumber="570">n Range</td>
<td id="F3010B5B4A7921DFBF98878A5C35FC7F" box="[938,1145,956,978]" gridcol="2" gridrow="1" pageId="2" pageNumber="570">n Range</td>
<td id="F3010B5B4A7921DFB88B878A5CBFFC7F" box="[1209,1267,956,978]" gridcol="3" gridrow="1" pageId="2" pageNumber="570">Mean</td>
<td id="F3010B5B4A7921DFB904878A5D1AFC7F" box="[1334,1366,956,978]" gridcol="4" gridrow="1" pageId="2" pageNumber="570">SD</td>
</tr>
<tr id="B0D062274A7921DFBCAE87D05D1AFC51" box="[156,1366,998,1020]" gridrow="2" pageId="2" pageNumber="570" rowspan-3="1" rowspan-4="1">
<th id="F3010B5B4A7921DFBCAE87D05A4DFC51" box="[156,513,998,1020]" gridcol="0" gridrow="2" pageId="2" pageNumber="570">Standard length (mm)</th>
<td id="F3010B5B4A7921DFBE0C87D05B18FC51" box="[574,852,998,1020]" gridcol="1" gridrow="2" pageId="2" pageNumber="570">41.0 22 30.8–43.6</td>
<td id="F3010B5B4A7921DFBF9887D05C35FC51" box="[938,1145,998,1020]" gridcol="2" gridrow="2" pageId="2" pageNumber="570">43 29.1–51.4</td>
</tr>
<tr id="B0D062274A7921DFBCAE80265D1AFBFD" box="[156,1366,1040,1104]" gridrow="3" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE80265A4DFBFD" box="[156,513,1040,1104]" gridcol="0" gridrow="3" pageId="2" pageNumber="570">
<emphasis id="3C94BC774A79DE22BCAE80275911FB8A" bold="true" box="[156,349,1040,1063]" pageId="2" pageNumber="570">Percentages of SL</emphasis>
Depth at dorsal-fin origin
</th>
<td id="F3010B5B4A7921DFBE0C80265B18FBFD" box="[574,852,1040,1104]" gridcol="1" gridrow="3" pageId="2" pageNumber="570">38.1 22 33.4–39.4</td>
<td id="F3010B5B4A7921DFBF9880265C35FBFD" box="[938,1145,1040,1104]" gridcol="2" gridrow="3" pageId="2" pageNumber="570">43 28.2–39.9</td>
<td id="F3010B5B4A7921DFB88B80265CBFFBFD" box="[1209,1267,1040,1104]" gridcol="3" gridrow="3" pageId="2" pageNumber="570">35.3</td>
<td id="F3010B5B4A7921DFB90480265D1AFBFD" box="[1334,1366,1040,1104]" gridcol="4" gridrow="3" pageId="2" pageNumber="570">2.3</td>
</tr>
<tr id="B0D062274A7921DFBCAE80525D1AFBD7" box="[156,1366,1124,1146]" gridrow="4" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE80525A4DFBD7" box="[156,513,1124,1146]" gridcol="0" gridrow="4" pageId="2" pageNumber="570">Snout to dorsal-fin origin</th>
<td id="F3010B5B4A7921DFBE0C80525B18FBD7" box="[574,852,1124,1146]" gridcol="1" gridrow="4" pageId="2" pageNumber="570">49.8 22 46.1–51.6</td>
<td id="F3010B5B4A7921DFBF9880525C35FBD7" box="[938,1145,1124,1146]" gridcol="2" gridrow="4" pageId="2" pageNumber="570">43 46.8–51.3</td>
<td id="F3010B5B4A7921DFB88B80525CBFFBD7" box="[1209,1267,1124,1146]" gridcol="3" gridrow="4" pageId="2" pageNumber="570">48.5</td>
<td id="F3010B5B4A7921DFB90480525D1AFBD7" box="[1334,1366,1124,1146]" gridcol="4" gridrow="4" pageId="2" pageNumber="570">1.1</td>
</tr>
<tr id="B0D062274A7921DFBCAE80B85D1AFB09" box="[156,1366,1166,1188]" gridrow="5" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE80B85A4DFB09" box="[156,513,1166,1188]" gridcol="0" gridrow="5" pageId="2" pageNumber="570">Snout to pectoral-fin origin</th>
<td id="F3010B5B4A7921DFBE0C80B85B18FB09" box="[574,852,1166,1188]" gridcol="1" gridrow="5" pageId="2" pageNumber="570">27.3 21 25.1–29.3</td>
<td id="F3010B5B4A7921DFBF9880B85C35FB09" box="[938,1145,1166,1188]" gridcol="2" gridrow="5" pageId="2" pageNumber="570">43 25.5–31.0</td>
<td id="F3010B5B4A7921DFB88B80B85CBFFB09" box="[1209,1267,1166,1188]" gridcol="3" gridrow="5" pageId="2" pageNumber="570">27.6</td>
<td id="F3010B5B4A7921DFB90480B85D1AFB09" box="[1334,1366,1166,1188]" gridcol="4" gridrow="5" pageId="2" pageNumber="570">1.1</td>
</tr>
<tr id="B0D062274A7921DFBCAE808E5D1AFB63" box="[156,1366,1208,1230]" gridrow="6" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE808E5A4DFB63" box="[156,513,1208,1230]" gridcol="0" gridrow="6" pageId="2" pageNumber="570">Snout to pelvic-fin origin</th>
<td id="F3010B5B4A7921DFBE0C808E5B18FB63" box="[574,852,1208,1230]" gridcol="1" gridrow="6" pageId="2" pageNumber="570">48.1 22 45.9–49.7</td>
<td id="F3010B5B4A7921DFBF98808E5C35FB63" box="[938,1145,1208,1230]" gridcol="2" gridrow="6" pageId="2" pageNumber="570">43 44.2–49.7</td>
<td id="F3010B5B4A7921DFB88B808E5CBFFB63" box="[1209,1267,1208,1230]" gridcol="3" gridrow="6" pageId="2" pageNumber="570">47.1</td>
<td id="F3010B5B4A7921DFB904808E5D1AFB63" box="[1334,1366,1208,1230]" gridcol="4" gridrow="6" pageId="2" pageNumber="570">1.1</td>
</tr>
<tr id="B0D062274A7921DFBCAE80D45D1AFB55" box="[156,1366,1250,1272]" gridrow="7" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE80D45A4DFB55" box="[156,513,1250,1272]" gridcol="0" gridrow="7" pageId="2" pageNumber="570">Snout to anal-fin origin</th>
<td id="F3010B5B4A7921DFBE0C80D45B18FB55" box="[574,852,1250,1272]" gridcol="1" gridrow="7" pageId="2" pageNumber="570">63.3 22 61.3–64.7</td>
<td id="F3010B5B4A7921DFBF9880D45C35FB55" box="[938,1145,1250,1272]" gridcol="2" gridrow="7" pageId="2" pageNumber="570">43 59.8–67.0</td>
<td id="F3010B5B4A7921DFB88B80D45CBFFB55" box="[1209,1267,1250,1272]" gridcol="3" gridrow="7" pageId="2" pageNumber="570">62.8</td>
<td id="F3010B5B4A7921DFB90480D45D1AFB55" box="[1334,1366,1250,1272]" gridcol="4" gridrow="7" pageId="2" pageNumber="570">1.6</td>
</tr>
<tr id="B0D062274A7921DFBCAE813A5D1AFA8F" box="[156,1366,1292,1314]" gridrow="8" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE813A5A4DFA8F" box="[156,513,1292,1314]" gridcol="0" gridrow="8" pageId="2" pageNumber="570">Caudal-peduncle depth</th>
<td id="F3010B5B4A7921DFBE0C813A5B18FA8F" box="[574,852,1292,1314]" gridcol="1" gridrow="8" pageId="2" pageNumber="570">10.4 22 8.6–10.5</td>
<td id="F3010B5B4A7921DFBF98813A5C35FA8F" box="[938,1145,1292,1314]" gridcol="2" gridrow="8" pageId="2" pageNumber="570">43 8.3–10.9</td>
<td id="F3010B5B4A7921DFB88B813A5CBFFA8F" box="[1209,1267,1292,1314]" gridcol="3" gridrow="8" pageId="2" pageNumber="570">9.6</td>
<td id="F3010B5B4A7921DFB904813A5D1AFA8F" box="[1334,1366,1292,1314]" gridcol="4" gridrow="8" pageId="2" pageNumber="570">0.6</td>
</tr>
<tr id="B0D062274A7921DFBCAE81005D1AFAE1" box="[156,1366,1334,1356]" gridrow="9" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE81005A4DFAE1" box="[156,513,1334,1356]" gridcol="0" gridrow="9" pageId="2" pageNumber="570">Caudal-peduncle length</th>
<td id="F3010B5B4A7921DFBE0C81005B18FAE1" box="[574,852,1334,1356]" gridcol="1" gridrow="9" pageId="2" pageNumber="570">11.3 22 8.9–12.6</td>
<td id="F3010B5B4A7921DFBF9881005C35FAE1" box="[938,1145,1334,1356]" gridcol="2" gridrow="9" pageId="2" pageNumber="570">43 6.9–12.6</td>
<td id="F3010B5B4A7921DFB88B81005CBFFAE1" box="[1209,1267,1334,1356]" gridcol="3" gridrow="9" pageId="2" pageNumber="570">10.1</td>
<td id="F3010B5B4A7921DFB90481005D1AFAE1" box="[1334,1366,1334,1356]" gridcol="4" gridrow="9" pageId="2" pageNumber="570">1.0</td>
</tr>
<tr id="B0D062274A7921DFBCAE81565D1AFADB" box="[156,1366,1376,1398]" gridrow="10" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE81565A4DFADB" box="[156,513,1376,1398]" gridcol="0" gridrow="10" pageId="2" pageNumber="570">Pectoral-fin length</th>
<td id="F3010B5B4A7921DFBE0C81565B18FADB" box="[574,852,1376,1398]" gridcol="1" gridrow="10" pageId="2" pageNumber="570">14.4 22 20.8–26.5</td>
<td id="F3010B5B4A7921DFBF9881565C35FADB" box="[938,1145,1376,1398]" gridcol="2" gridrow="10" pageId="2" pageNumber="570">43 19.2–26.0</td>
<td id="F3010B5B4A7921DFB88B81565CBFFADB" box="[1209,1267,1376,1398]" gridcol="3" gridrow="10" pageId="2" pageNumber="570">22.8</td>
<td id="F3010B5B4A7921DFB90481565D1AFADB" box="[1334,1366,1376,1398]" gridcol="4" gridrow="10" pageId="2" pageNumber="570">1.7</td>
</tr>
<tr id="B0D062274A7921DFBCAE81BC5D1AFA0D" box="[156,1366,1418,1440]" gridrow="11" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE81BC5A4DFA0D" box="[156,513,1418,1440]" gridcol="0" gridrow="11" pageId="2" pageNumber="570">Pelvic-fin length</th>
<td id="F3010B5B4A7921DFBE0C81BC5B18FA0D" box="[574,852,1418,1440]" gridcol="1" gridrow="11" pageId="2" pageNumber="570">18.0 22 15.9–19.5</td>
<td id="F3010B5B4A7921DFBF9881BC5C35FA0D" box="[938,1145,1418,1440]" gridcol="2" gridrow="11" pageId="2" pageNumber="570">43 16.0–20.7</td>
<td id="F3010B5B4A7921DFB88B81BC5CBFFA0D" box="[1209,1267,1418,1440]" gridcol="3" gridrow="11" pageId="2" pageNumber="570">18.2</td>
<td id="F3010B5B4A7921DFB90481BC5D1AFA0D" box="[1334,1366,1418,1440]" gridcol="4" gridrow="11" pageId="2" pageNumber="570">1.0</td>
</tr>
<tr id="B0D062274A7921DFBCAE81825D1AFA67" box="[156,1366,1460,1482]" gridrow="12" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE81825A4DFA67" box="[156,513,1460,1482]" gridcol="0" gridrow="12" pageId="2" pageNumber="570">Dorsal-fin base length</th>
<td id="F3010B5B4A7921DFBE0C81825B18FA67" box="[574,852,1460,1482]" gridcol="1" gridrow="12" pageId="2" pageNumber="570">13.5 22 13.6–16.1</td>
<td id="F3010B5B4A7921DFBF9881825C35FA67" box="[938,1145,1460,1482]" gridcol="2" gridrow="12" pageId="2" pageNumber="570">43 13.4–16.2</td>
<td id="F3010B5B4A7921DFB88B81825CBFFA67" box="[1209,1267,1460,1482]" gridcol="3" gridrow="12" pageId="2" pageNumber="570">14.9</td>
<td id="F3010B5B4A7921DFB90481825D1AFA67" box="[1334,1366,1460,1482]" gridcol="4" gridrow="12" pageId="2" pageNumber="570">0.7</td>
</tr>
<tr id="B0D062274A7921DFBCAE81E85D1AFA59" box="[156,1366,1502,1524]" gridrow="13" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE81E85A4DFA59" box="[156,513,1502,1524]" gridcol="0" gridrow="13" pageId="2" pageNumber="570">Dorsal-fin length</th>
<td id="F3010B5B4A7921DFBE0C81E85B18FA59" box="[574,852,1502,1524]" gridcol="1" gridrow="13" pageId="2" pageNumber="570">– 22 30.4–34.1</td>
<td id="F3010B5B4A7921DFBF9881E85C35FA59" box="[938,1145,1502,1524]" gridcol="2" gridrow="13" pageId="2" pageNumber="570">43 29.0–35.3</td>
<td id="F3010B5B4A7921DFB88B81E85CBFFA59" box="[1209,1267,1502,1524]" gridcol="3" gridrow="13" pageId="2" pageNumber="570">32.3</td>
<td id="F3010B5B4A7921DFB90481E85D1AFA59" box="[1334,1366,1502,1524]" gridcol="4" gridrow="13" pageId="2" pageNumber="570">1.3</td>
</tr>
<tr id="B0D062274A7921DFBCAE823E5D1AF9B3" box="[156,1366,1544,1566]" gridrow="14" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE823E5A4DF9B3" box="[156,513,1544,1566]" gridcol="0" gridrow="14" pageId="2" pageNumber="570">Anal-fin base length</th>
<td id="F3010B5B4A7921DFBE0C823E5B18F9B3" box="[574,852,1544,1566]" gridcol="1" gridrow="14" pageId="2" pageNumber="570">31.5 22 28.7–34.6</td>
<td id="F3010B5B4A7921DFBF98823E5C35F9B3" box="[938,1145,1544,1566]" gridcol="2" gridrow="14" pageId="2" pageNumber="570">43 28.8–36.2</td>
<td id="F3010B5B4A7921DFB88B823E5CBFF9B3" box="[1209,1267,1544,1566]" gridcol="3" gridrow="14" pageId="2" pageNumber="570">32.2</td>
<td id="F3010B5B4A7921DFB904823E5D1AF9B3" box="[1334,1366,1544,1566]" gridcol="4" gridrow="14" pageId="2" pageNumber="570">1.6</td>
</tr>
<tr id="B0D062274A7921DFBCAE82045D1AF9E5" box="[156,1366,1586,1608]" gridrow="15" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE82045A4DF9E5" box="[156,513,1586,1608]" gridcol="0" gridrow="15" pageId="2" pageNumber="570">Anal-fin lobe length</th>
<td id="F3010B5B4A7921DFBE0C82045B18F9E5" box="[574,852,1586,1608]" gridcol="1" gridrow="15" pageId="2" pageNumber="570">20.0 17 18.7–25.2</td>
<td id="F3010B5B4A7921DFBF9882045C35F9E5" box="[938,1145,1586,1608]" gridcol="2" gridrow="15" pageId="2" pageNumber="570">43 15.8–25.1</td>
<td id="F3010B5B4A7921DFB88B82045CBFF9E5" box="[1209,1267,1586,1608]" gridcol="3" gridrow="15" pageId="2" pageNumber="570">22.1</td>
<td id="F3010B5B4A7921DFB90482045D1AF9E5" box="[1334,1366,1586,1608]" gridcol="4" gridrow="15" pageId="2" pageNumber="570">1.8</td>
</tr>
<tr id="B0D062274A7921DFBCAE826A5D1AF9DF" box="[156,1366,1628,1650]" gridrow="16" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE826A5A4DF9DF" box="[156,513,1628,1650]" gridcol="0" gridrow="16" pageId="2" pageNumber="570">Eye to dorsal-fin origin</th>
<td id="F3010B5B4A7921DFBE0C826A5B18F9DF" box="[574,852,1628,1650]" gridcol="1" gridrow="16" pageId="2" pageNumber="570">35.3 22 31.1–35.3</td>
<td id="F3010B5B4A7921DFBF98826A5C35F9DF" box="[938,1145,1628,1650]" gridcol="2" gridrow="16" pageId="2" pageNumber="570">43 30.4–35.3</td>
<td id="F3010B5B4A7921DFB88B826A5CBFF9DF" box="[1209,1267,1628,1650]" gridcol="3" gridrow="16" pageId="2" pageNumber="570">33.2</td>
<td id="F3010B5B4A7921DFB904826A5D1AF9DF" box="[1334,1366,1628,1650]" gridcol="4" gridrow="16" pageId="2" pageNumber="570">1.1</td>
</tr>
<tr id="B0D062274A7921DFBCAE82B05D1AF931" box="[156,1366,1670,1692]" gridrow="17" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE82B05A4DF931" box="[156,513,1670,1692]" gridcol="0" gridrow="17" pageId="2" pageNumber="570">Dorsal-fin origin to caudal-fin base</th>
<td id="F3010B5B4A7921DFBE0C82B05B18F931" box="[574,852,1670,1692]" gridcol="1" gridrow="17" pageId="2" pageNumber="570">53.9 21 53.4–59.5</td>
<td id="F3010B5B4A7921DFBF9882B05C35F931" box="[938,1145,1670,1692]" gridcol="2" gridrow="17" pageId="2" pageNumber="570">43 53.5–59.9</td>
<td id="F3010B5B4A7921DFB88B82B05CBFF931" box="[1209,1267,1670,1692]" gridcol="3" gridrow="17" pageId="2" pageNumber="570">55.7</td>
<td id="F3010B5B4A7921DFB90482B05D1AF931" box="[1334,1366,1670,1692]" gridcol="4" gridrow="17" pageId="2" pageNumber="570">1.3</td>
</tr>
<tr id="B0D062274A7921DFBCAE82865D1AF96B" box="[156,1366,1712,1734]" gridrow="18" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE82865A4DF96B" box="[156,513,1712,1734]" gridcol="0" gridrow="18" pageId="2" pageNumber="570">Head length</th>
<td id="F3010B5B4A7921DFBE0C82865B18F96B" box="[574,852,1712,1734]" gridcol="1" gridrow="18" pageId="2" pageNumber="570">27.0 21 24.9–29.4</td>
<td id="F3010B5B4A7921DFBF9882865C35F96B" box="[938,1145,1712,1734]" gridcol="2" gridrow="18" pageId="2" pageNumber="570">43 25.4–30.3</td>
<td id="F3010B5B4A7921DFB88B82865CBFF96B" box="[1209,1267,1712,1734]" gridcol="3" gridrow="18" pageId="2" pageNumber="570">27.4</td>
<td id="F3010B5B4A7921DFB90482865D1AF96B" box="[1334,1366,1712,1734]" gridcol="4" gridrow="18" pageId="2" pageNumber="570">1.2</td>
</tr>
<tr id="B0D062274A7921DFBCAE82EC5D1AF95D" box="[156,1366,1754,1776]" gridrow="19" pageId="2" pageNumber="570" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="F3010B5B4A7921DFBCAE82EC5A4DF95D" box="[156,513,1754,1776]" gridcol="0" gridrow="19" pageId="2" pageNumber="570">
<emphasis id="3C94BC774A79DE22BCAE82ED592EF95D" bold="true" box="[156,354,1754,1776]" pageId="2" pageNumber="570">Percentages of HL</emphasis>
</th>
</tr>
<tr id="B0D062274A7921DFBCAE83325D1AF8B7" box="[156,1366,1796,1818]" gridrow="20" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE83325A4DF8B7" box="[156,513,1796,1818]" gridcol="0" gridrow="20" pageId="2" pageNumber="570">Horizontal eye diameter</th>
<td id="F3010B5B4A7921DFBE0C83325B18F8B7" box="[574,852,1796,1818]" gridcol="1" gridrow="20" pageId="2" pageNumber="570">41.7 19 41.9–47.1</td>
<td id="F3010B5B4A7921DFBF9883325C35F8B7" box="[938,1145,1796,1818]" gridcol="2" gridrow="20" pageId="2" pageNumber="570">41 33.6–43.6</td>
<td id="F3010B5B4A7921DFB88B83325CBFF8B7" box="[1209,1267,1796,1818]" gridcol="3" gridrow="20" pageId="2" pageNumber="570">40.9</td>
<td id="F3010B5B4A7921DFB90483325D1AF8B7" box="[1334,1366,1796,1818]" gridcol="4" gridrow="20" pageId="2" pageNumber="570">4.0</td>
</tr>
<tr id="B0D062274A7921DFBCAE83185D1AF8E9" box="[156,1366,1838,1860]" gridrow="21" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE83185A4DF8E9" box="[156,513,1838,1860]" gridcol="0" gridrow="21" pageId="2" pageNumber="570">Snout length</th>
<td id="F3010B5B4A7921DFBE0C83185B18F8E9" box="[574,852,1838,1860]" gridcol="1" gridrow="21" pageId="2" pageNumber="570">24.8 21 22.0–29.7</td>
<td id="F3010B5B4A7921DFBF9883185C35F8E9" box="[938,1145,1838,1860]" gridcol="2" gridrow="21" pageId="2" pageNumber="570">43 20.2–31.3</td>
<td id="F3010B5B4A7921DFB88B83185CBFF8E9" box="[1209,1267,1838,1860]" gridcol="3" gridrow="21" pageId="2" pageNumber="570">26.1</td>
<td id="F3010B5B4A7921DFB90483185D1AF8E9" box="[1334,1366,1838,1860]" gridcol="4" gridrow="21" pageId="2" pageNumber="570">2.1</td>
</tr>
<tr id="B0D062274A7921DFBCAE836E5D1AF8C3" box="[156,1366,1880,1902]" gridrow="22" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE836E5A4DF8C3" box="[156,513,1880,1902]" gridcol="0" gridrow="22" pageId="2" pageNumber="570">Interorbital width</th>
<td id="F3010B5B4A7921DFBE0C836E5B18F8C3" box="[574,852,1880,1902]" gridcol="1" gridrow="22" pageId="2" pageNumber="570">30.4 21 27.5–33.2</td>
<td id="F3010B5B4A7921DFBF98836E5C35F8C3" box="[938,1145,1880,1902]" gridcol="2" gridrow="22" pageId="2" pageNumber="570">43 26.7–33.8</td>
<td id="F3010B5B4A7921DFB88B836E5CBFF8C3" box="[1209,1267,1880,1902]" gridcol="3" gridrow="22" pageId="2" pageNumber="570">30.6</td>
<td id="F3010B5B4A7921DFB904836E5D1AF8C3" box="[1334,1366,1880,1902]" gridcol="4" gridrow="22" pageId="2" pageNumber="570">1.7</td>
</tr>
<tr id="B0D062274A7921DFBCAE83B45D1AF835" box="[156,1366,1922,1944]" gridrow="23" pageId="2" pageNumber="570">
<th id="F3010B5B4A7921DFBCAE83B45A4DF835" box="[156,513,1922,1944]" gridcol="0" gridrow="23" pageId="2" pageNumber="570">Upper jaw length</th>
<td id="F3010B5B4A7921DFBE0C83B45B18F835" box="[574,852,1922,1944]" gridcol="1" gridrow="23" pageId="2" pageNumber="570">43.6 21 36.1–46.0</td>
<td id="F3010B5B4A7921DFBF9883B45C35F835" box="[938,1145,1922,1944]" gridcol="2" gridrow="23" pageId="2" pageNumber="570">43 36.4–46.9</td>
<td id="F3010B5B4A7921DFB88B83B45CBFF835" box="[1209,1267,1922,1944]" gridcol="3" gridrow="23" pageId="2" pageNumber="570">41.2</td>
<td id="F3010B5B4A7921DFB90483B45D1AF835" box="[1334,1366,1922,1944]" gridcol="4" gridrow="23" pageId="2" pageNumber="570">2.5</td>
</tr>
</table>
</paragraph>
<caption id="5A9F30ED4A78DE23BCA582305C72F9F7" httpUri="https://zenodo.org/record/234680/files/figure.png" pageId="3" pageNumber="571" targetBox="[340,1247,193,1520]" targetPageId="3">
<paragraph id="0E5F60654A78DE23BCA582305C72F9F7" blockId="3.[151,1436,1542,1626]" pageId="3" pageNumber="571">
<emphasis id="3C94BC774A78DE23BCA58230595FF9B1" bold="true" box="[151,275,1542,1564]" pageId="3" pageNumber="571">FIGURE 1.</emphasis>
<taxonomicName id="C9E01BE64A78DE23BD28823159A5F9B0" box="[282,489,1543,1565]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="571" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A78DE23BD28823159A5F9B0" box="[282,489,1543,1565]" italics="true" pageId="3" pageNumber="571">Astyanax guianensis</emphasis>
</taxonomicName>
: (a) holotype, FMNH 51518, 41.0 mm SL, Guyana, rio Essequibo basin; (b) MZUSP 25526, 53.3. mm SL, Brazil, Pará, rio Tapajós basin; (c) MZUSP 115554, 35.7 mm SL, Brazil, Mato Grosso, rio Guaporé basin; and (d) live specimen, Brazil, Amazonas, rio Negro at Santa Isabel do Rio Negro, not preserved.
</paragraph>
</caption>
<paragraph id="0E5F60654A78DE23BCF582B459EDF83B" blockId="3.[151,1437,1665,2014]" pageId="3" pageNumber="571">
Jaws aligned vertically, mouth terminal. Premaxillary teeth in two rows. Outer row with 2(4), 3(32), 4(30) or 5*(3) tricuspid teeth. Inner row with 5*(66) or 6(3) teeth, 4 of which pentacuspid, and lateralmost one tricuspid. Maxilla with 5(17), 6*(22), 7(22), 8(5), 9(3) or 10(1) uni- to tetracuspid teeth. Dentary with 4(26) large pentacuspid teeth, followed by one or two smaller tricuspid and one series of 5 to 12 small uni- to tricuspid teeth. Central median cusp in all teeth longer than lateral cusps (
<figureCitation id="96DB7CE04A78DE23BF1283275B29F887" box="[800,869,1809,1834]" captionStart="FIGURE 2" captionStartId="5.[151,250,901,923]" captionTargetBox="[346,1236,202,872]" captionTargetId="figure@5.[339,1247,193,879]" captionTargetPageId="5" captionText="FIGURE 2. Upper and lower jaws of Astyanax guianensis (medial view, right side), MZUSP 109674, 46.7 mm SL. Coronomeckelian bone is missing." httpUri="https://zenodo.org/record/234681/files/figure.png" pageId="3" pageNumber="571">Fig. 2</figureCitation>
). Branchiostegal rays 4. First gill arch with 2(3) or 3(3) rakers on hypobranchial, none (3) or 1(3) raker on intermediate cartilage, 10(2) or 11(4) rakers on ceratobranchial, 1(6) raker on intermediate cartilage, and 7(3) or 8(3) rakers on epibranchial. Each gill raker bearing small denticles.
</paragraph>
<paragraph id="0E5F60654A78DE24BCF583975AF8FE15" blockId="3.[151,1437,1665,2014]" lastBlockId="4.[151,1437,151,1990]" lastPageId="4" lastPageNumber="572" pageId="3" pageNumber="571">Supraneurals 3(3) or 4*(4), upper portion with bony lamellae. Dorsal-fin rays ii,9*(65), first unbranched ray about one-half length of second unbranched ray. First dorsal-fin pterygiophore inserted posterior to neural spine of 7th(1) or 8th*(6) vertebrae. Dorsal-fin origin slightly posterior to vertical through pelvic-fin origin. Adipose-fin origin at vertical through base of 18th or 19th branched anal-fin rays. Pectoral-fin rays i(64), 12(22), 13(37) or 14(4). Tip of adpressed pectoral fin extending beyond pelvic-fin origin. Pelvic-fin rays i*(65), 6(1) or 7*(64). Tip of adpressed pelvic fin extending beyond anal-fin origin. Anal-fin rays iv*(7), 21(8), 22(6), 23*(18), 24(27) or 25(5). First anal-fin pterygiophore inserted posterior of haemal spine of 15th*(7) vertebra. Anal-fin origin located at vertical through base of 5th or 6th branched dorsal-fin ray. Principal caudal-fin rays i,9*(57) at upper and i,8*(57) at lower lobe. Dorsal procurrent caudal-fin rays 11(1), 12(4) or 13(1) and ventral procurrent caudal-fin rays 9(2), 10(2) or 11(2). Caudal fin forked, lobes pointed.</paragraph>
<paragraph id="0E5F60654A7FDE24BCF585F55A1EFD19" blockId="4.[151,1437,151,1990]" pageId="4" pageNumber="572">Lateral line straight to slightly curved ventrally, with 31(3), 32(7), 33(17), 34(17) or 35*(16) perforated scales from supracleithrum to caudal-fin base. Longitudinal scale rows between dorsal-fin origin and lateral line 5*(55) or 6(4). Longitudinal scale rows between lateral line and pelvic-fin origin 4*(44) or 5(10). Predorsal scales 8(14) or 9(51) in one aligned series. Single row of 4–8 scales overlying base of anteriormost anal-fin rays. Scale rows around caudal peduncle 14*(50). Scales usually at base of caudal fin, not extending over lobes; some specimens with scales along basal one-sixth of lower lobe. Abdominal vertebrae 14*(5) or 15(2), caudal vertebrae 18(3) or 19*(4). Total vertebrae 32(1) or 33*(6).</paragraph>
<paragraph id="0E5F60654A7FDE24BCF58689591FFA93" blockId="4.[151,1437,151,1990]" pageId="4" pageNumber="572">
<emphasis id="3C94BC774A7FDE24BCF5868959DAFD75" bold="true" box="[199,406,703,728]" pageId="4" pageNumber="572">Color in alcohol.</emphasis>
General body color pale yellow (
<figureCitation id="96DB7CE04A7FDE24BF1586895B39FD7A" box="[807,885,703,728]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1564]" captionTargetBox="[340,1247,193,1520]" captionTargetId="figure@3.[340,1248,193,1521]" captionTargetPageId="3" captionText="FIGURE 1. Astyanax guianensis: (a) holotype, FMNH 51518, 41.0 mm SL, Guyana, rio Essequibo basin; (b) MZUSP 25526, 53.3. mm SL, Brazil, Pará, rio Tapajós basin; (c) MZUSP 115554, 35.7 mm SL, Brazil, Mato Grosso, rio Guaporé basin; and (d) live specimen, Brazil, Amazonas, rio Negro at Santa Isabel do Rio Negro, not preserved." httpUri="https://zenodo.org/record/234680/files/figure.png" pageId="4" pageNumber="572">Figs. 1</figureCitation>
a–c). Upper and lower lips, anterior portion of maxilla, snout, top of head, dorsal portion of opercle and pectoral girdle with scattered small dark chromatophores. Some specimens with dark chromatophores scattered on third infraorbital. Middorsal horizontal scale rows bordered by dark pigmentation, forming slightly reticulated pattern. More pigmented specimens with faint dark chromatophores bordering all scales, except on ventral portion of body. Scattered dark chromatophores above analfin base. More pigmented specimens with dark chromatophores concentrated along margins of myosepta of posterior half of lower portion of body, between lateral line and anal-fin base. Humeral blotch vertically oriented, extending over one scale row ventral to, and three scale rows dorsal to lateral line, and over three scales horizontally. Portion of humeral blotch located dorsal to lateral line often interrupted by clear horizontal stripe. Humeral blotch followed posteriorly by clear area and then by pigmented area progressively fading posteriorly as a longitudinal stripe, from vertical through 7th or 8th lateral-line scale to caudal peduncle. Some specimens retaining guanine over longitudinal stripe. Dark pigmentation along horizontal septum extending approximately from vertical through dorsal-fin origin to caudal peduncle, falling short of caudal-fin base. Faint pigmented area at caudal peduncle. First unbranched dorsal-fin ray with dark chromatophores, remaining dorsal-fin rays hyaline. Distal half of interadial membranes of all dorsal-fin rays with scattered dark chromatophores. Pectoral, pelvic, caudal and adipose fins with few scattered dark chromatophores. Distal portion of interadial membranes of branched anal-fin rays with concentration of dark chromatophores; proximal portion with few scattered dark chromatophores.
</paragraph>
<paragraph id="0E5F60654A7FDE24BCF5817C5AA8FA63" blockId="4.[151,1437,151,1990]" pageId="4" pageNumber="572">
<emphasis id="3C94BC774A7FDE24BCF5817C5910FACE" bold="true" box="[199,348,1354,1379]" pageId="4" pageNumber="572">Color in life.</emphasis>
General body coloration silver to pale (
<figureCitation id="96DB7CE04A7FDE24BF24817C5B19FACE" box="[790,853,1354,1379]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1564]" captionTargetBox="[340,1247,193,1520]" captionTargetId="figure@3.[340,1248,193,1521]" captionTargetPageId="3" captionText="FIGURE 1. Astyanax guianensis: (a) holotype, FMNH 51518, 41.0 mm SL, Guyana, rio Essequibo basin; (b) MZUSP 25526, 53.3. mm SL, Brazil, Pará, rio Tapajós basin; (c) MZUSP 115554, 35.7 mm SL, Brazil, Mato Grosso, rio Guaporé basin; and (d) live specimen, Brazil, Amazonas, rio Negro at Santa Isabel do Rio Negro, not preserved." httpUri="https://zenodo.org/record/234680/files/figure.png" pageId="4" pageNumber="572">Fig. 1</figureCitation>
d). Dorsal portions of head and eye, middle portions of dorsal and anal fins (especially anteriorly), adipose fin, and base of upper and lower caudal-fin lobes yellow to orange. Humeral blotch and longitudinal stripe conspicuous in life. Tip of the three longest pelvic-fin rays white. Tip of first six dorsal- and anal-fin rays milky white.
</paragraph>
<paragraph id="0E5F60654A7FDE24BCF581EC5BE6FA5E" blockId="4.[151,1437,151,1990]" box="[199,938,1498,1523]" pageId="4" pageNumber="572">
<emphasis id="3C94BC774A7FDE24BCF581EC59FBFA5E" bold="true" box="[199,439,1498,1523]" pageId="4" pageNumber="572">Sexual dimorphism.</emphasis>
No sexually dimorphic characters observed.
</paragraph>
<paragraph id="0E5F60654A7FDE24BCF581CB5AF4F997" blockId="4.[151,1437,151,1990]" pageId="4" pageNumber="572">
<emphasis id="3C94BC774A7FDE24BCF581CB59A2F9BB" bold="true" box="[199,494,1533,1558]" pageId="4" pageNumber="572">Geographic distribution.</emphasis>
<taxonomicName id="C9E01BE64A7FDE24BDC681C85A95F9BB" box="[500,729,1534,1558]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="572" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7FDE24BDC681C85A95F9BB" box="[500,729,1534,1558]" italics="true" pageId="4" pageNumber="572">Astyanax guianensis</emphasis>
</taxonomicName>
is widespread throughout the 
<collectingCountry id="76F720F54A7FDE24B81B81CB5C36F9BB" box="[1065,1146,1533,1558]" name="Guyana" pageId="4" pageNumber="572">Guiana</collectingCountry>
Shield drainages and also in the Amazon and Orinoco river basins (
<figureCitation id="96DB7CE04A7FDE24BE5482145AE6F997" box="[614,682,1570,1595]" captionStart="FIGURE 3" captionStartId="5.[151,250,1979,2001]" captionTargetBox="[169,1418,1010,1951]" captionTargetId="figure@5.[161,1421,1003,1960]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Astyanax guianensis in the Guiana Shield drainages, Amazon, and Orinoco basins. Type locality is represented by the star." httpUri="https://zenodo.org/record/234682/files/figure.png" pageId="4" pageNumber="572">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="0E5F60654A7FDE24BCF582735B3BF8BE" blockId="4.[151,1437,151,1990]" pageId="4" pageNumber="572">
<emphasis id="3C94BC774A7FDE24BCF582735A69F9F3" bold="true" box="[199,549,1605,1630]" pageId="4" pageNumber="572">Habitat and ecological notes.</emphasis>
<taxonomicName id="C9E01BE64A7FDE24BE1C82705B5AF9F3" box="[558,790,1606,1630]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="572" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7FDE24BE1C82705B5AF9F3" box="[558,790,1606,1630]" italics="true" pageId="4" pageNumber="572">Astyanax guianensis</emphasis>
</taxonomicName>
was collected in clear water streams, running over sand and silt bottoms, at altitudes ranging from 
<quantity id="C918CD804A7FDE24BEBF825C5B62F92F" box="[653,814,1642,1666]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" metricValueMax="2.4" metricValueMin="2.04" pageId="4" pageNumber="572" unit="m" value="222.0" valueMax="240.0" valueMin="204.0">204 to 240 m</quantity>
above sea level, in habitats characterized by slow to moderate water current, less than one meter deep. The riparian vegetation was mainly composed by grass, shrubs, and trees. The analysis of stomach contents of five specimens of 
<taxonomicName id="C9E01BE64A7FDE24BF9282845C70F967" box="[928,1084,1714,1738]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="572" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7FDE24BF9282845C70F967" box="[928,1084,1714,1738]" italics="true" pageId="4" pageNumber="572">A. guianensis</emphasis>
</taxonomicName>
(MZUSP 97149 and MZUSP 109674) revealed the presence of organic debris, aquatic insects (caddisflies larvae, chironomids pupae, and Odonota nymph), and fragments of other unidentified arthropods.
</paragraph>
</subSubSection>
<subSubSection id="46FA33EE4A7FDE26BCF5832B5988FDBA" lastPageId="6" lastPageNumber="574" pageId="4" pageNumber="572" type="discussion">
<paragraph id="0E5F60654A7FDE24BCF5832B5B96F86B" blockId="4.[151,1437,151,1990]" pageId="4" pageNumber="572">
<emphasis id="3C94BC774A7FDE24BCF5832B5977F89B" bold="true" box="[199,315,1821,1846]" pageId="4" pageNumber="572">Remarks.</emphasis>
Most features described by 
<bibRefCitation id="6A711D944A7FDE24BEB6832B5B14F89B" author="Eigenmann" box="[644,856,1821,1846]" pageId="4" pageNumber="572" refString="Eigenmann, C. H. (1909) Reports on the expedition to British Guiana of the Indiana University and the Carnegie Museum, 1908. Report no. 1. Some new genera and species of fishes from British Guiana. Annals of the Carnegie Museum, 6, 4 - 54." type="journal article" year="1909">Eigenmann (1909)</bibRefCitation>
in the original description of 
<taxonomicName id="C9E01BE64A7FDE24B88683285DD0F89B" box="[1204,1436,1822,1846]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="572" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7FDE24B88683285DD0F89B" box="[1204,1436,1822,1846]" italics="true" pageId="4" pageNumber="572">Astyanax guianensis</emphasis>
</taxonomicName>
matches those observed by us. There are only two exceptions. 
<bibRefCitation id="6A711D944A7FDE24BFA683745C20F8F7" author="Eigenmann" box="[916,1132,1858,1883]" pageId="4" pageNumber="572" refString="Eigenmann, C. H. (1909) Reports on the expedition to British Guiana of the Indiana University and the Carnegie Museum, 1908. Report no. 1. Some new genera and species of fishes from British Guiana. Annals of the Carnegie Museum, 6, 4 - 54." type="journal article" year="1909">Eigenmann (1909)</bibRefCitation>
mentioned four to seven maxillary teeth. In 70 examined specimens, five to 10 tiny teeth were counted in the maxilla, none had four. 
<bibRefCitation id="6A711D944A7FDE24BCA583BC5925F80F" author="Eigenmann" box="[151,361,1930,1955]" pageId="4" pageNumber="572" refString="Eigenmann, C. H. (1909) Reports on the expedition to British Guiana of the Indiana University and the Carnegie Museum, 1908. Report no. 1. Some new genera and species of fishes from British Guiana. Annals of the Carnegie Museum, 6, 4 - 54." type="journal article" year="1909">Eigenmann (1909)</bibRefCitation>
also mentioned the presence of 34 to 35 (rarely 36) scales on the lateral series. In 60 examined specimens, 31 to 35 scales were counted on the lateral series, none had 36.
</paragraph>
<caption id="5A9F30ED4A7EDE25BCA587B359BAFC14" httpUri="https://zenodo.org/record/234681/files/figure.png" pageId="5" pageNumber="573" targetBox="[346,1236,202,872]" targetPageId="5">
<paragraph id="0E5F60654A7EDE25BCA587B359BAFC14" blockId="5.[151,1436,900,953]" pageId="5" pageNumber="573">
<emphasis id="3C94BC774A7EDE25BCA587B35955FC37" bold="true" box="[151,281,901,923]" pageId="5" pageNumber="573">FIGURE 2.</emphasis>
Upper and lower jaws of 
<taxonomicName id="C9E01BE64A7EDE25BE7A87B35B52FC36" box="[584,798,901,923]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="573" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7EDE25BE7A87B35B52FC36" box="[584,798,901,923]" italics="true" pageId="5" pageNumber="573">Astyanax guianensis</emphasis>
</taxonomicName>
(medial view, right side), MZUSP 109674, 46.7 mm SL. Coronomeckelian bone is missing.
</paragraph>
</caption>
<caption id="5A9F30ED4A7EDE25BCA5838D59CDF85D" httpUri="https://zenodo.org/record/234682/files/figure.png" pageId="5" pageNumber="573" targetBox="[169,1418,1010,1951]" targetPageId="5">
<paragraph id="0E5F60654A7EDE25BCA5838D59CDF85D" blockId="5.[151,1436,1979,2032]" pageId="5" pageNumber="573">
<emphasis id="3C94BC774A7EDE25BCA5838D595EF87D" bold="true" box="[151,274,1979,2001]" pageId="5" pageNumber="573">FIGURE 3.</emphasis>
Distribution of 
<taxonomicName id="C9E01BE64A7EDE25BD87838A5AC8F87C" box="[437,644,1980,2001]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="573" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7EDE25BD87838A5AC8F87C" box="[437,644,1980,2001]" italics="true" pageId="5" pageNumber="573">Astyanax guianensis</emphasis>
</taxonomicName>
in the Guiana Shield drainages, Amazon, and Orinoco basins. Type locality is represented by the star.
</paragraph>
</caption>
<paragraph id="0E5F60654A7DDE26BCF584AE5D47FEB0" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="574">
In the label corresponding to lot CAS 68733 (ex IU 12078), the word cotype, is written in reference to one specimen of 
<taxonomicName id="C9E01BE64A7DDE26BD14848A5A40FF79" box="[294,524,188,212]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7DDE26BD14848A5A40FF79" box="[294,524,188,212]" italics="true" pageId="6" pageNumber="574">Astyanax guianensis</emphasis>
</taxonomicName>
collected by Eigenmann at 
<taxonomicName id="C9E01BE64A7DDE26BF71848A5BF8FF78" box="[835,948,188,213]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="577" phylum="Chordata" rank="species" species="rupununi">Rupununi</taxonomicName>
, near Rockstone. A second label was found inside the jar, stating that this specimen was not cited in the original description of 
<taxonomicName id="C9E01BE64A7DDE26B80A84D65C83FF5A" box="[1080,1231,224,247]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7DDE26B80A84D65C83FF5A" box="[1080,1231,224,247]" italics="true" pageId="6" pageNumber="574">A. guianensis</emphasis>
</taxonomicName>
and should not be considered as cotype. We agree with this remark and thus will not consider this specimen as 
<typeStatus id="D15BDEC74A7DDE26B89185335D4BFEB0" box="[1187,1287,261,285]" pageId="6" pageNumber="574" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="0E5F60654A7DDE26BCF5851E59CFFE01" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="574">
<bibRefCitation id="6A711D944A7DDE26BCF5851E593DFEED" author="Pearson" box="[199,369,296,320]" pageId="6" pageNumber="574" refString="Pearson, N. E. (1924) The fishes of the eastern slope of the Andes. I. The fishes of the rio Beni basin, Bolivia, collected by the Mulford Expedition. Indiana University Studies, 11, 1 - 83." type="journal article" year="1924">Pearson (1924)</bibRefCitation>
mentioned 
<specimenCount id="18E6ABEC4A7DDE26BDC4851E5AC2FEED" box="[502,654,295,320]" pageId="6" pageNumber="574" type="generic">12 specimens</specimenCount>
of 
<taxonomicName id="C9E01BE64A7DDE26BE81851E5BE5FE92" box="[691,937,296,319]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guaporensis">
<emphasis id="3C94BC774A7DDE26BE81851E5BE5FE92" box="[691,937,296,319]" italics="true" pageId="6" pageNumber="574">Astyanax guaporensis</emphasis>
</taxonomicName>
from lagoons near Reyes [= pampa town 
<quantity id="C918CD804A7DDE26B9B2851E58F9FEC8" metricMagnitude="4" metricUnit="m" metricValue="3.8624256" pageId="6" pageNumber="574" unit="mi" value="24.0">24 mi</quantity>
NE of Rurrenabaque], rio Beni, 
<collectingCountry id="76F720F54A7DDE26BE04857A5AC7FEC8" box="[566,651,332,357]" name="Bolivia" pageId="6" pageNumber="574">Bolivia</collectingCountry>
, identified as 
<taxonomicName id="C9E01BE64A7DDE26BF03857A5C0DFEC9" box="[817,1089,332,357]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7DDE26BF03857A5BDBFECE" box="[817,919,332,355]" italics="true" pageId="6" pageNumber="574">Astyanax</emphasis>
cf. 
<emphasis id="3C94BC774A7DDE26BFFB857B5C0DFEC9" box="[969,1089,333,356]" italics="true" pageId="6" pageNumber="574">guianensis</emphasis>
</taxonomicName>
by Marinho &amp; Ohara (2013). Examination of this material (CAS 68730 [ex IU 17322]) confirmed the identification of the specimens as 
<taxonomicName id="C9E01BE64A7DDE26BCA585A25931FE01" box="[151,381,404,428]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7DDE26BCA585A25931FE01" box="[151,381,404,428]" italics="true" pageId="6" pageNumber="574">Astyanax guianensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0E5F60654A7DDE26BCF585815988FDBA" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="574">
<bibRefCitation id="6A711D944A7DDE26BCF585815993FE7D" author="Lowe-McConnell" box="[199,479,439,464]" pageId="6" pageNumber="574" refString="Lowe-McConnell, R. H. (1991) Natural history of fishes in Araguaia and Xingu Amazonian tributaries, Serra do Roncador, Mato Grosso, Brazil. Icthyological Exploration of Freshwater, 2, 63 - 82." type="journal article" year="1991">Lowe-McConnell (1991)</bibRefCitation>
cited 
<taxonomicName id="C9E01BE64A7DDE26BE16858E5B47FE62" box="[548,779,440,463]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7DDE26BE16858E5B47FE62" box="[548,779,440,463]" italics="true" pageId="6" pageNumber="574">Astyanax guianensis</emphasis>
</taxonomicName>
from rio das Mortes, rio Araguaia basin. These specimens could not be analyzed herein. However, extensive material from that drainage was examined and did not yield any specimen of 
<taxonomicName id="C9E01BE64A7DDE26BD14863659F1FDBA" box="[294,445,512,535]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="574" phylum="Chordata" rank="species" species="guianensis">
<emphasis id="3C94BC774A7DDE26BD14863659F1FDBA" box="[294,445,512,535]" italics="true" pageId="6" pageNumber="574">A. guianensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>