210 lines
21 KiB
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210 lines
21 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1137.96306" ID-Pensoft-Pub="1313-2970-1137-75" ID-Pensoft-UUID="040858FC10C55E9BB26F5C668945EEC3" ID-ZooBank="CF82430BB0104FD48A448356992F0629" ModsDocID="1313-2970-1137-75" checkinTime="1671892175879" checkinUser="pensoft" docAuthor="Mi, Xiaoqi & Li, Shuqiang" docDate="2022" docId="86324C581C81544BB479F858B69EB31E" docLanguage="en" docName="ZooKeys 1137: 75-108" docOrigin="ZooKeys 1137" docPubDate="2022-12-22" docSource="http://dx.doi.org/10.3897/zookeys.1137.96306" docTitle="Araneus bidentatus Mi & Li 2022, sp. nov." docType="treatment" docUuid="516C5C7D-2580-4672-BF6C-A8B9F58F8674" docUuidSource="ZooBank" docVersion="1" id="040858FC10C55E9BB26F5C668945EEC3" lastPageNumber="75" masterDocId="040858FC10C55E9BB26F5C668945EEC3" masterDocTitle="On eleven new species of the orb-weaver spider genus Araneus Clerck, 1757 (Araneae, Araneidae) from Xishuangbanna, Yunnan, China" masterLastPageNumber="108" masterPageNumber="75" pageNumber="75" updateTime="1671892175879" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>On eleven new species of the orb-weaver spider genus Araneus Clerck, 1757 (Araneae, Araneidae) from Xishuangbanna, Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mi, Xiaoqi</mods:namePart>
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<mods:affiliation>College of Agriculture and Forestry Engineering and Planning, Guizhou Provincial Key Laboratory of Biodiversity Conservation and Utilization in the Fanjing Mountain Region, Tongren University, Tongren 554300, Guizhou, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1137</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>108</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1137.96306</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1137.96306</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1137-75</mods:identifier>
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<mods:identifier type="ZooBank">CF82430BB0104FD48A448356992F0629</mods:identifier>
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<mods:identifier type="Pensoft-UUID">040858FC10C55E9BB26F5C668945EEC3</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:516C5C7D-2580-4672-BF6C-A8B9F58F8674" httpUri="http://treatment.plazi.org/id/86324C581C81544BB479F858B69EB31E" lastPageNumber="75" pageId="0" pageNumber="75">
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<subSubSection pageId="0" pageNumber="75" type="nomenclature">
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName LSID="https://zoobank.org/516C5C7D-2580-4672-BF6C-A8B9F58F8674" authority="Mi & Li, 2022" authorityName="Mi & Li" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus bidentatus" order="Araneae" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="bidentatus" status="sp. nov.">Araneus bidentatus</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="75">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="description">
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<paragraph pageId="0" pageNumber="75">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus bidentatus sp. nov. A-D, G, H female paratype IZCAS-Ar 43098 E, F male holotype A epigyne, ventral view B ibid., lateral view C ibid., posterior view D vulva, posterior view E habitus, dorsal view F ibid., lateral view G ibid., dorsal view H ibid., ventral view. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1137.96306.figure3" httpUri="https://binary.pensoft.net/fig/786260" pageId="0" pageNumber="75">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Araneus bidentatus sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view E tibia of left male palp, retrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure4" httpUri="https://binary.pensoft.net/fig/786261" pageId="0" pageNumber="75">, 4</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs of Araneus spp. A-D A. arcuatus sp. nov., male holotype A leg I B leg II C leg III D leg IV E-H A. bidentatus sp. nov., male holotype E leg I F leg II G leg III H leg IV I, J A. complanatus sp. nov., female paratype Ar 43106 I leg I J leg II K, L A. cucullatus sp. nov., female paratype Ar 43110 K leg I L leg II. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1137.96306.figure18" httpUri="https://binary.pensoft.net/fig/786275" pageId="0" pageNumber="75">, 18E-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="type material">
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<paragraph pageId="0" pageNumber="75">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Holotype</emphasis>
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♂ (IZCAS-Ar43097), China: Yunnan, Xishuangbanna, Menghai County, Menghai Township, Mengweng Village, Wengnan, secondary forest (
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<geoCoordinate degrees="22" direction="north" minutes="4.94" orientation="latitude" precision="9" value="22.082333">22°4.94'N</geoCoordinate>
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,
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<geoCoordinate degrees="100" direction="east" minutes="22.03" orientation="longitude" precision="9" value="100.367165">100°22.03'E</geoCoordinate>
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, ca 1150 m), 2.VII.2013, Q.Y. Zhao & Z.G. Chen leg.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Paratypes</emphasis>
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: 1♂1♀ (IZCAS-Ar43098-43099), same data as holotype; 1♂ (IZCAS-Ar43100), Mengla County, Mohan Township, Shanggang Village, Xiaolongha, valley rainforest (
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<geoCoordinate degrees="21" direction="north" minutes="24.25" orientation="latitude" precision="9" value="21.404167">21°24.25'N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="36.32" orientation="longitude" precision="9" value="101.60533">101°36.32'E</geoCoordinate>
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, ca 760 m), 15.VI.2013, Q.Y. Zhao & Z.G. Chen leg.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="etymology">
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<paragraph pageId="0" pageNumber="75">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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The specific name is a combination of the Latin prefix
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<normalizedToken originalValue="“bi-”">"bi-"</normalizedToken>
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and
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<normalizedToken originalValue="“dentatus”">"dentatus"</normalizedToken>
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(two toothed), referring to the two heavily sclerotized denticulate protuberances on the tibia of the male palp.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="diagnosis">
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<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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The new species resembles
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="A. bidentatoides" order="Araneae" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="bidentatoides">
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<emphasis italics="true" pageId="0" pageNumber="75">A. bidentatoides</emphasis>
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</taxonomicName>
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sp. nov. in appearance, but differs in the following: 1) copulatory openings located at lateral ends of the scape groove vs at anterolateral base of the scape (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Araneus bidentatoides sp. nov. A-C, F, G female paratype IZCAS-Ar 43102 D, E male holotype A epigyne, ventral view B ibid., posterior view C vulva, posterior view D habitus, dorsal view E ibid., lateral view F ibid., dorsal view G ibid., ventral view. Scale bars: 0.1 mm (A-C); 1 mm (D-G)." figureDoi="10.3897/zookeys.1137.96306.figure5" httpUri="https://binary.pensoft.net/fig/786262" pageId="0" pageNumber="75">5A</figureCitation>
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); 2) copulatory ducts not expanded vs expanded at their origin (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Araneus bidentatoides sp. nov. A-C, F, G female paratype IZCAS-Ar 43102 D, E male holotype A epigyne, ventral view B ibid., posterior view C vulva, posterior view D habitus, dorsal view E ibid., lateral view F ibid., dorsal view G ibid., ventral view. Scale bars: 0.1 mm (A-C); 1 mm (D-G)." figureDoi="10.3897/zookeys.1137.96306.figure5" httpUri="https://binary.pensoft.net/fig/786262" pageId="0" pageNumber="75">5C</figureCitation>
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); 3) embolus slender, a bit more slender than the patellar bristle vs embolus stout, several times bigger than the patellar bristle (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Araneus bidentatoides sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure6" httpUri="https://binary.pensoft.net/fig/786263" pageId="0" pageNumber="75">6A, C, D</figureCitation>
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); 4) distal end of the terminal apophysis not tapered to a long tip vs tapered (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Araneus bidentatoides sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure6" httpUri="https://binary.pensoft.net/fig/786263" pageId="0" pageNumber="75">6</figureCitation>
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); 5) median apophysis slightly curved vs curved about 90° (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Araneus bidentatoides sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure6" httpUri="https://binary.pensoft.net/fig/786263" pageId="0" pageNumber="75">6A</figureCitation>
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); 6) tibia with 2 heavily sclerotized denticulate protuberances vs protuberances absent (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Araneus bidentatoides sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure6" httpUri="https://binary.pensoft.net/fig/786263" pageId="0" pageNumber="75">6A, B</figureCitation>
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); and 7) fovea region and sides of thoracic region paler than thoracic region vs unicolor (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Araneus bidentatoides sp. nov. A-C, F, G female paratype IZCAS-Ar 43102 D, E male holotype A epigyne, ventral view B ibid., posterior view C vulva, posterior view D habitus, dorsal view E ibid., lateral view F ibid., dorsal view G ibid., ventral view. Scale bars: 0.1 mm (A-C); 1 mm (D-G)." figureDoi="10.3897/zookeys.1137.96306.figure5" httpUri="https://binary.pensoft.net/fig/786262" pageId="0" pageNumber="75">5D-G</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="description">
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<paragraph pageId="0" pageNumber="75">Description.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Male</emphasis>
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(holotype, Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus bidentatus sp. nov. A-D, G, H female paratype IZCAS-Ar 43098 E, F male holotype A epigyne, ventral view B ibid., lateral view C ibid., posterior view D vulva, posterior view E habitus, dorsal view F ibid., lateral view G ibid., dorsal view H ibid., ventral view. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1137.96306.figure3" httpUri="https://binary.pensoft.net/fig/786260" pageId="0" pageNumber="75">3E, F</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Araneus bidentatus sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view E tibia of left male palp, retrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure4" httpUri="https://binary.pensoft.net/fig/786261" pageId="0" pageNumber="75">4</figureCitation>
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,
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs of Araneus spp. A-D A. arcuatus sp. nov., male holotype A leg I B leg II C leg III D leg IV E-H A. bidentatus sp. nov., male holotype E leg I F leg II G leg III H leg IV I, J A. complanatus sp. nov., female paratype Ar 43106 I leg I J leg II K, L A. cucullatus sp. nov., female paratype Ar 43110 K leg I L leg II. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1137.96306.figure18" httpUri="https://binary.pensoft.net/fig/786275" pageId="0" pageNumber="75">18E-H</figureCitation>
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). Total length 2.70. Carapace 1.60 long, 1.20 wide. Abdomen 1.50 long, 0.95 wide. Clypeus 0.13 high. Eye sizes and interdistances: AME 0.13, ALE 0.10, PME 0.13, PLE 0.13, AME-AME 0.13, AME-ALE 0.10, PME-PME 0.13, PME-PLE 0.20, MOA length 0.23, anterior width 0.35, posterior width 0.35. Leg measurements: I 5.45 (1.65, 2.00, 1.15, 0.65), II 4.50 (1.45, 1.55, 0.95, 0.55), III 3.05 (1.00, 1.05, 0.55, 0.45), IV 4.15 (1.40, 1.40, 0.85, 0.50). Carapace pear-shaped, grayish black with yellow patches around fovea and on lateral edges of thoracic region, ALEs, PMEs and PLEs with black base, with pale setae, cervical groove inconspicuous. Chelicerae yellowish brown, 5 promarginal teeth, 3 retromarginal teeth. Endites and labium dark brown at base, paler distally, endites square, labium triangular. Sternum cordiform, dark brown with dark setae. Legs yellow with grayish brown annuli, tibia I with 9 macrosetae, tibia II with 11 macrosetae, tibia III with 7 macrosetae, tibia IV with 8 macrosetae. Abdomen elliptical, about 1.25 times longer than wide, covered with gray setae, dorsum yellow with white spot anteriorly and irregular black markings; venter yellow with dark brown patches. Spinnerets yellowish brown.
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</paragraph>
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<caption doi="10.3897/zookeys.1137.96306.figure3" httpUri="https://binary.pensoft.net/fig/786260" pageId="0" pageNumber="75" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 3.</emphasis>
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<taxonomicName authorityName="Mi & Li" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus bidentatus" order="Araneae" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="bidentatus">
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<emphasis italics="true" pageId="0" pageNumber="75">Araneus bidentatus</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="75">A-D, G, H</emphasis>
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female paratype IZCAS-Ar43098
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<emphasis bold="true" pageId="0" pageNumber="75">E, F</emphasis>
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male holotype
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<emphasis bold="true" pageId="0" pageNumber="75">A</emphasis>
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epigyne, ventral view
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<emphasis bold="true" pageId="0" pageNumber="75">B</emphasis>
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ibid., lateral view
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<emphasis bold="true" pageId="0" pageNumber="75">C</emphasis>
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ibid., posterior view
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<emphasis bold="true" pageId="0" pageNumber="75">D</emphasis>
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vulva, posterior view
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<emphasis bold="true" pageId="0" pageNumber="75">E</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="75">F</emphasis>
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ibid., lateral view
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<emphasis bold="true" pageId="0" pageNumber="75">G</emphasis>
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ibid., dorsal view
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<emphasis bold="true" pageId="0" pageNumber="75">H</emphasis>
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ibid., ventral view. Scale bars: 0.1 mm (
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<emphasis bold="true" pageId="0" pageNumber="75">A-D</emphasis>
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); 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="75">E-H</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Palp</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Araneus bidentatus sp. nov. male holotype A male palp, prolateral view B ibid., retrolateral view C ibid., ventral view D ibid., apical view E tibia of left male palp, retrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1137.96306.figure4" httpUri="https://binary.pensoft.net/fig/786261" pageId="0" pageNumber="75">4</figureCitation>
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): patella with 2 bristles; tibia with 2 heavily sclerotized denticulate protuberances (fig. 4B, E); paracymbium widened at base, with a finger-like tip; median apophysis shorter than the conductor, tapered to a pointed tip; embolus slender, slightly curved; conductor very wide, semicircular in prolateral view; terminal apophysis extremely large, heavily sclerotized, with 2 widely separated protuberances in apical view.
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</paragraph>
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<caption doi="10.3897/zookeys.1137.96306.figure4" httpUri="https://binary.pensoft.net/fig/786261" pageId="0" pageNumber="75" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 4.</emphasis>
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<taxonomicName authorityName="Mi & Li" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus bidentatus" order="Araneae" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="bidentatus">
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<emphasis italics="true" pageId="0" pageNumber="75">Araneus bidentatus</emphasis>
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</taxonomicName>
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sp. nov. male holotype
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<emphasis bold="true" pageId="0" pageNumber="75">A</emphasis>
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male palp, prolateral view
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<emphasis bold="true" pageId="0" pageNumber="75">B</emphasis>
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ibid., retrolateral view
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<emphasis bold="true" pageId="0" pageNumber="75">C</emphasis>
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ibid., ventral view
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<emphasis bold="true" pageId="0" pageNumber="75">D</emphasis>
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ibid., apical view
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<emphasis bold="true" pageId="0" pageNumber="75">E</emphasis>
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tibia of left male palp, retrolateral view. Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Female</emphasis>
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(paratype IZCAS-Ar43098, Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus bidentatus sp. nov. A-D, G, H female paratype IZCAS-Ar 43098 E, F male holotype A epigyne, ventral view B ibid., lateral view C ibid., posterior view D vulva, posterior view E habitus, dorsal view F ibid., lateral view G ibid., dorsal view H ibid., ventral view. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1137.96306.figure3" httpUri="https://binary.pensoft.net/fig/786260" pageId="0" pageNumber="75">3A-D, G, H</figureCitation>
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). Total length 3.25. Carapace 1.55 long, 1.15 wide. Abdomen 2.35 long, 1.90 wide. Clypeus 0.08 high. Eye sizes and interdistances: AME 0.13, ALE 0.08, PME 0.13, PLE 0.10, AME-AME 0.13, AME-ALE 0.18, PME-PME 0.15, PME-PLE 0.25, MOA length 0.33, anterior width 0.35, posterior width 0.33. Habitus similar to that of male but with a pair of low anterolateral humps and a little paler.
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</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Epigyne</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus bidentatus sp. nov. A-D, G, H female paratype IZCAS-Ar 43098 E, F male holotype A epigyne, ventral view B ibid., lateral view C ibid., posterior view D vulva, posterior view E habitus, dorsal view F ibid., lateral view G ibid., dorsal view H ibid., ventral view. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1137.96306.figure3" httpUri="https://binary.pensoft.net/fig/786260" pageId="0" pageNumber="75">3A-D</figureCitation>
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): about 1.5 times wider than long; scape triangular, very short, about 5 times wider than long, distally with transverse groove; copulatory openings on the ends of the groove; copulatory ducts longer than the spermatheca diameter, curved; spermathecae globular, touching each other.
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</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">
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<emphasis italics="true" pageId="0" pageNumber="75">Variation</emphasis>
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.
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</emphasis>
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Total length: ♂♂ 2.70-3.25.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="distribution">
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<paragraph pageId="0" pageNumber="75">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="75">Known only from type localities (Yunnan, China).</paragraph>
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</subSubSection>
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</treatment>
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</document> |