treatments-xml/data/85/59/2A/85592A26EF79FF9BE082FB9FA045FBCA.xml

<document id="EBD70037403311BEDCA8CBC5946BC684" ID-CLB-Dataset="22399" ID-DOI="10.5852/ejt.2015.131" ID-GBIF-Dataset="1aeda346-69eb-41e6-8655-d82772c05804" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785228" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588613718640" checkinUser="carolina" docAuthor="Strandberg, Jonas &amp; Johanson, Kjell Arne" docDate="2015" docId="85592A26EF79FF9BE082FB9FA045FBCA" docLanguage="en" docName="ejt-2015-131a.pdf.imf" docOrigin="European Journal of Taxonomy 131" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dasyhelea (Dicryptoscena) gothlandica Strandberg &amp; Johanson 2015, sp. nov." docType="treatment" docUuid="5FCC2057-325D-4EFF-B0F9-0C660A747422" docUuidSource="ZooBank" docVersion="6" lastPageNumber="16" masterDocId="7960525EEF72FF94E179FFE9A422FF85" masterDocTitle="New records of Dasyhelea Kieffer, 1911 from Sweden, with descriptions of two new species (Diptera: Ceratopogonidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="12" updateTime="1698838408609" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="992A1D0DE323EFB40BAB1C686449FBDD">New records of Dasyhelea Kieffer, 1911 from Sweden, with descriptions of two new species (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="5F903DE91BA598349EB658B5DD8D8BD0">Strandberg, Jonas</mods:namePart>
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<mods:namePart id="4ABA1F888C7FC1FA2D7DAC196A83DEE3">Johanson, Kjell Arne</mods:namePart>
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<treatment id="85592A26EF79FF9BE082FB9FA045FBCA" ID-DOI="http://doi.org/10.5281/zenodo.3794739" ID-GBIF-Taxon="163887611" ID-Zenodo-Dep="3794739" LSID="urn:lsid:zoobank.org:act:5FCC2057-325D-4EFF-B0F9-0C660A747422" httpUri="http://treatment.plazi.org/id/85592A26EF79FF9BE082FB9FA045FBCA" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection id="45EAC8BBEF79FF9FE082FB9FA01BFB14" box="[507,1081,1142,1169]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="0D4F9B30EF79FF9FE082FB9FA01BFB14" blockId="11.[371,1215,1142,1240]" box="[507,1081,1142,1169]" pageId="11" pageNumber="12">
<heading id="56072C5CEF79FF9FE082FB9FA01BFB14" box="[507,1081,1142,1169]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="CAF0E0B3EF79FF9FE082FB9FA7F4FB14" ID-CoL="9HN7G" authority="Strandberg &amp; Johanson, 2015" authorityName="Strandberg &amp; Johanson" authorityYear="2015" box="[507,982,1142,1169]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gothlandica" status="sp. nov." subGenus="Dicryptoscena">
<emphasis id="3F844722EF79FF9FE082FB9FA7F4FB14" bold="true" box="[507,982,1142,1169]" pageId="11" pageNumber="12">
<emphasis id="3F844722EF79FF9FE082FB9FA65AFB14" bold="true" box="[507,632,1142,1169]" italics="true" pageId="11" pageNumber="12">Dasyhelea</emphasis>
(
<emphasis id="3F844722EF79FF9FE3F3FB9EA715FB14" bold="true" box="[650,823,1143,1169]" italics="true" pageId="11" pageNumber="12">Dicryptoscena</emphasis>
) 
<emphasis id="3F844722EF79FF9FE23EFB9FA7F4FB14" bold="true" box="[839,982,1142,1169]" italics="true" pageId="11" pageNumber="12">gothlandica</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24B7FA59EF79FF9FE2A7FB9EA01BFB14" box="[990,1081,1143,1169]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="45EAC8BBEF79FF9FE00AFB72A77EFB5D" pageId="11" pageNumber="12" type="description">
<paragraph id="0D4F9B30EF79FF9FE00AFB72A09DFB30" blockId="11.[371,1215,1142,1240]" box="[371,1215,1179,1205]" pageId="11" pageNumber="12">
<uri id="79619732EF79FF9FE00AFB72A09DFB30" box="[371,1215,1179,1205]" pageId="11" pageNumber="12">
urn:lsid:zoobank.org:act: 
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</paragraph>
<paragraph id="0D4F9B30EF79FF9FE3A1FB57A77EFB5D" blockId="11.[371,1215,1142,1240]" box="[728,860,1214,1240]" pageId="11" pageNumber="12">
<figureCitation id="95CB87B5EF79FF9FE3A1FB57A77EFB5D" box="[728,860,1214,1240]" captionStart-0="Figs 12–18" captionStart-1="Figs19–22" captionStart-2="Figs 23–29" captionStart-3="Figs 30–32" captionStartId-0="12.[189,241,1929,1955]" captionStartId-1="13.[189,241,1891,1917]" captionStartId-2="14.[189,241,1907,1933]" captionStartId-3="15.[189,241,1878,1904]" captionTargetBox-0="[253,1335,264,1898]" captionTargetBox-1="[193,1401,918,1849]" captionTargetBox-2="[190,1400,280,1872]" captionTargetBox-3="[216,1381,1167,1840]" captionTargetId-0="figure@12.[229,1358,244,1918]" captionTargetId-1="figure@13.[157,1420,909,1858]" captionTargetId-2="figure@14.[147,1433,250,1884]" captionTargetId-3="figure@15.[175,1412,1163,1847]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." captionText-1="Figs19–22. Dasyheleagothlandica sp.nov.,holotype, ♂.19–20.Ventralaspectof genitalia.21. Parameres. 22. Aedeagus. Scale bars: 19–20 = 50 μm; 21–22 = 16 μm." captionText-2="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." captionText-3="Figs 30–32. Dasyhelea gothlandica sp. nov., paratype, ♀. 30–31. Subgenital plate. 32. Seminal capsule. Scale bars = 50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3785236" figureDoi-1="http://doi.org/10.5281/zenodo.3785238" figureDoi-2="http://doi.org/10.5281/zenodo.3785240" figureDoi-3="http://doi.org/10.5281/zenodo.3785242" httpUri-0="https://zenodo.org/record/3785236/files/figure.png" httpUri-1="https://zenodo.org/record/3785238/files/figure.png" httpUri-2="https://zenodo.org/record/3785240/files/figure.png" httpUri-3="https://zenodo.org/record/3785242/files/figure.png" pageId="11" pageNumber="12">Figs 12–32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="45EAC8BBEF79FF9FE1C4FB16A542FA34" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="0D4F9B30EF79FF9FE1C4FB16A514FA9C" blockId="11.[189,310,1279,1305]" box="[189,310,1279,1305]" pageId="11" pageNumber="12">
<heading id="56072C5CEF79FF9FE1C4FB16A514FA9C" bold="true" box="[189,310,1279,1305]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="3F844722EF79FF9FE1C4FB16A514FA9C" bold="true" box="[189,310,1279,1305]" pageId="11" pageNumber="12">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="0D4F9B30EF79FF9FE1C4FAC7A542FA34" blockId="11.[189,1399,1326,1457]" pageId="11" pageNumber="12">The male genitalia of this species are characteristic in having the posterior margin of sternite 9 with a pair of processes apically separated by a shallow furrow. Gonostyli short, tapering to an indented sharp point. The aedeagus has broad apicolateral processes, and the posterior process of the paramere has fine hairs apically.</paragraph>
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<subSubSection id="45EAC8BBEF79FF9FE1C4FA30A66EF9C0" pageId="11" pageNumber="12" type="etymology">
<paragraph id="0D4F9B30EF79FF9FE1C4FA30A566FA76" blockId="11.[189,324,1497,1523]" box="[189,324,1497,1523]" pageId="11" pageNumber="12">
<heading id="56072C5CEF79FF9FE1C4FA30A566FA76" bold="true" box="[189,324,1497,1523]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="3F844722EF79FF9FE1C4FA30A566FA76" bold="true" box="[189,324,1497,1523]" pageId="11" pageNumber="12">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="0D4F9B30EF79FF9FE1C4F9EEA66EF9C0" blockId="11.[189,1398,1543,1605]" pageId="11" pageNumber="12">
<emphasis id="3F844722EF79FF9FE1C4F9EEA572F9A4" box="[189,336,1543,1569]" italics="true" pageId="11" pageNumber="12">Gothlandica</emphasis>
refers to 
<collectingRegion id="CF3455D2EF79FF9FE0BAF9EEA607F9A4" box="[451,549,1543,1569]" country="Sweden" name="Gotland" pageId="11" pageNumber="12">Gotland</collectingRegion>
, a calcareous island located in the Baltic Sea east of mainland 
<collectingCountry id="75E7DBA0EF79FF9FE46BF9EEA150F9A4" box="[1298,1394,1543,1569]" name="Sweden" pageId="11" pageNumber="12">Sweden</collectingCountry>
, where the 
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was collected.
</paragraph>
</subSubSection>
<subSubSection id="45EAC8BBEF79FF9FE1C4F985A52EF84C" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="0D4F9B30EF79FF9FE1C4F985A54EF903" blockId="11.[189,364,1644,1670]" box="[189,364,1644,1670]" pageId="11" pageNumber="12">
<heading id="56072C5CEF79FF9FE1C4F985A54EF903" bold="true" box="[189,364,1644,1670]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="3F844722EF79FF9FE1C4F985A54EF903" bold="true" box="[189,364,1644,1670]" pageId="11" pageNumber="12">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="BD98916DEF79FF9FE193F972A707F97E" ID-GBIF-Occurrence="2610058608" collectingDate="2011-05-28" collectingDateMax="2011-06-12" collectingDateMin="2011-05-28" collectingMethod="Malaise trap" collectionCode="NHRS" collectorName="B. Eklund" country="Sweden" elevation="9" latitude="57.899384" location="Bastetrask" longLatPrecision="1" longitude="18.901083" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Gotland" typeStatus="holotype">
<paragraph id="0D4F9B30EF79FF9FE193F972A57EF930" blockId="11.[189,1399,1691,1787]" box="[234,348,1691,1717]" pageId="11" pageNumber="12">
<emphasis id="3F844722EF79FF9FE193F972A57EF930" bold="true" box="[234,348,1691,1717]" pageId="11" pageNumber="12">
<typeStatus id="D24B2592EF79FF9FE193F972A57EF930" box="[234,348,1691,1717]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="0D4F9B30EF79FF9FE1C4F957A707F97E" blockId="11.[189,1399,1691,1787]" pageId="11" pageNumber="12">
<collectingCountry id="75E7DBA0EF79FF9FE1C4F957A51EF95D" box="[189,316,1726,1752]" name="Sweden" pageId="11" pageNumber="12">SWEDEN</collectingCountry>
: 
<specimenCount id="1BF650B9EF79FF9FE03FF957A57EF95D" box="[326,348,1726,1752]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
, 
<collectingRegion id="CF3455D2EF79FF9FE013F957A5EEF95D" box="[362,460,1726,1752]" country="Sweden" name="Gotland" pageId="11" pageNumber="12">Gotland</collectingRegion>
, 
<location id="082FCDEBEF79FF9FE0AEF957A676F95D" LSID="urn:lsid:plazi:treatment:85592A26EF79FF9BE082FB9FA045FBCA:082FCDEBEF79FF9FE0AEF957A676F95D" box="[471,596,1726,1752]" country="Sweden" latitude="57.899384" longLatPrecision="1" longitude="18.901083" name="Bastetrask" pageId="11" pageNumber="12" stateProvince="Gotland">Bästeträsk</location>
, 
<geoCoordinate id="68C4FDF7EF79FF9FE326F957A724F95D" box="[607,774,1726,1752]" degrees="57.899383" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="57.899384">57.899383° N</geoCoordinate>
, 
<geoCoordinate id="68C4FDF7EF79FF9FE268F957A796F95D" box="[785,948,1726,1752]" degrees="18.901083" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="18.901083">18.901083° E</geoCoordinate>
, 
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<elevation id="86DD7C03EF79FF9FE2B9F957A02DF95D" box="[960,1039,1726,1752]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="12" unit="m" value="9.0">9 masl</elevation>
</quantity>
, 
<collectingMethod id="D4B1E327EF79FF9FE562F957A08DF95D" box="[1051,1199,1726,1752]" pageId="11" pageNumber="12">Malaise trap</collectingMethod>
, 
<date id="794EBDF0EF79FF9FE5C3F957A4DBF97E" pageId="11" pageNumber="12" value="2011-05-28" valueMax="2011-06-12" valueMin="2011-05-28">
<collectingDate id="690A4418EF79FF9FE5C3F957A4DBF97E" pageId="11" pageNumber="12" value="2011-05-28" valueMax="2011-06-12" valueMin="2011-05-28">28 May–12 Jun. 2011</collectingDate>
</date>
, leg. 
<collectorName id="A005FEE6EF79FF9FE041F908A58DF97E" box="[312,431,1761,1787]" pageId="11" pageNumber="12">B. Eklund</collectorName>
(
<collectionCode id="6BE103F5EF79FF9FE0C7F908A632F97E" box="[446,528,1761,1787]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="11" pageNumber="12">NHRS</collectionCode>
) (DNA voucher: 4BA).
</paragraph>
</materialsCitation>
<paragraph id="0D4F9B30EF79FF9FE193F8CBA545F8B9" blockId="11.[189,1399,1826,1993]" box="[234,359,1826,1852]" pageId="11" pageNumber="12">
<emphasis id="3F844722EF79FF9FE193F8CBA545F8B9" bold="true" box="[234,359,1826,1852]" pageId="11" pageNumber="12">
<typeStatus id="D24B2592EF79FF9FE193F8CBA545F8B9" box="[234,359,1826,1852]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="0D4F9B30EF79FF9FE1C4F8AFA52EF84C" blockId="11.[189,1399,1826,1993]" pageId="11" pageNumber="12">
<collectingCountry id="75E7DBA0EF79FF9FE1C4F8AFA51EF8E5" box="[189,316,1862,1888]" name="Sweden" pageId="11" pageNumber="12">SWEDEN</collectingCountry>
: 
<specimenCount id="1BF650B9EF79FF9FE031F8AFA5A9F8E5" box="[328,395,1862,1888]" pageId="11" pageNumber="12" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="1BF650B9EF79FF9FE0E2F8AFA5FDF8E5" box="[411,479,1862,1888]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
, same data as 
<typeStatus id="D24B2592EF79FF9FE3F7F8AFA6DAF8E5" box="[654,760,1862,1888]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
; 
<specimenCount id="1BF650B9EF79FF9FE27FF8AFA711F8E5" box="[774,819,1862,1888]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="CF3455D2EF79FF9FE23AF8AFA787F8E5" box="[835,933,1862,1888]" country="Sweden" name="Gotland" pageId="11" pageNumber="12">Gotland</collectingRegion>
, Horsan, 
<geoCoordinate id="68C4FDF7EF79FF9FE56FF8AFA09DF8E5" box="[1046,1215,1862,1888]" degrees="57.865883" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="57.865883">57.865883° N</geoCoordinate>
, 
<geoCoordinate id="68C4FDF7EF79FF9FE5B5F8AFA153F8E5" box="[1228,1393,1862,1888]" degrees="18.846350" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="18.84635">18.846350° E</geoCoordinate>
, 
<quantity id="CA0836D5EF79FF9FE1C4F880A530F806" box="[189,274,1897,1923]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="12" unit="m" value="4.0">4 masl</quantity>
, Malaise trap, 
<date id="794EBDF0EF79FF9FE0B6F880A6BAF806" box="[463,664,1897,1923]" pageId="11" pageNumber="12" value="2011-07-10" valueMax="2011-07-25" valueMin="2011-07-10">10–25 Jul. 2011</date>
, leg. B. Eklund (
<collectionCode id="6BE103F5EF79FF9FE20AF880A7EAF806" box="[883,968,1897,1923]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="11" pageNumber="12">NHRS</collectionCode>
); 
<specimenCount id="1BF650B9EF79FF9FE298F880A031F806" box="[993,1043,1897,1923]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
, 
<specimenCount id="1BF650B9EF79FF9FE55EF880A04CF806" box="[1063,1134,1897,1923]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
, 
<collectingRegion id="CF3455D2EF79FF9FE5FBF880A0C6F806" box="[1154,1252,1897,1923]" country="Sweden" name="Gotland" pageId="11" pageNumber="12">Gotland</collectingRegion>
, Bästeträsk, 
<geoCoordinate id="68C4FDF7EF79FF9FE1C4F865A546F823" box="[189,356,1932,1958]" degrees="57.899383" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="57.899384">57.899383° N</geoCoordinate>
, 
<geoCoordinate id="68C4FDF7EF79FF9FE016F865A630F823" box="[367,530,1932,1958]" degrees="18.901083" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="18.901083">18.901083° E</geoCoordinate>
, 
<quantity id="CA0836D5EF79FF9FE367F865A64FF823" box="[542,621,1932,1958]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="12" unit="m" value="9.0">9 masl</quantity>
, sweep net, 
<date id="794EBDF0EF79FF9FE383F865A75EF823" box="[762,892,1932,1958]" pageId="11" pageNumber="12" value="2011-07-03">3 Jul. 2011</date>
, leg. J. Strandberg (
<collectionCode id="6BE103F5EF79FF9FE518F865A091F823" box="[1121,1203,1932,1958]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="11" pageNumber="12">NHRS</collectionCode>
) (DNA voucher: 6AW).
</paragraph>
</subSubSection>
<caption id="598FCBB8EF7EFF98E1C4F860A6F6F86C" ID-DOI="http://doi.org/10.5281/zenodo.3785236" ID-Zenodo-Dep="3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,241,1929,1955]" targetBox="[253,1335,264,1898]" targetPageId="12">
<paragraph id="0D4F9B30EF7EFF98E1C4F860A6F6F86C" blockId="12.[189,1399,1929,2025]" pageId="12" pageNumber="13">
<emphasis id="3F844722EF7EFF98E1C4F860A565F826" bold="true" box="[189,327,1929,1955]" pageId="12" pageNumber="13">Figs 12–18.</emphasis>
<taxonomicName id="CAF0E0B3EF7EFF98E037F860A67EF826" authority="Strandberg &amp; Johanson, 2015" authorityName="Strandberg &amp; Johanson" authorityYear="2015" box="[334,604,1929,1955]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gothlandica" status="sp. nov.">
<emphasis id="3F844722EF7EFF98E037F860A67EF826" box="[334,604,1929,1955]" italics="true" pageId="12" pageNumber="13">Dasyhelea gothlandica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24B7FA59EF7EFF98E31BF860A69FF826" box="[610,701,1929,1955]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂. 
<emphasis id="3F844722EF7EFF98E226F860A75EF826" bold="true" box="[863,892,1929,1955]" pageId="12" pageNumber="13">12</emphasis>
. Head. 
<emphasis id="3F844722EF7EFF98E2ACF860A7D0F826" bold="true" box="[981,1010,1929,1955]" pageId="12" pageNumber="13">13</emphasis>
. Distal flagellomeres. 
<emphasis id="3F844722EF7EFF98E58EF860A136F826" bold="true" box="[1271,1300,1929,1955]" pageId="12" pageNumber="13">14</emphasis>
. Frontal sclerite. 
<emphasis id="3F844722EF7EFF98E058F845A51CF843" bold="true" box="[289,318,1964,1990]" pageId="12" pageNumber="13">15</emphasis>
. Clypeus. 
<emphasis id="3F844722EF7EFF98E0C7F845A5F9F843" bold="true" box="[446,475,1964,1990]" pageId="12" pageNumber="13">16</emphasis>
. Palp. 
<emphasis id="3F844722EF7EFF98E356F845A66EF843" bold="true" box="[559,588,1964,1990]" pageId="12" pageNumber="13">17</emphasis>
. Wing. 
<emphasis id="3F844722EF7EFF98E3D2F845A6EAF843" bold="true" box="[683,712,1964,1990]" pageId="12" pageNumber="13">18</emphasis>
. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm.
</paragraph>
</caption>
<subSubSection id="45EAC8BBEF7FFF9BE1C4FEE0A55BFD45" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="0D4F9B30EF7FFF99E1C4FEE0A56CFEA6" blockId="13.[189,334,265,291]" box="[189,334,265,291]" pageId="13" pageNumber="14">
<heading id="56072C5CEF7FFF99E1C4FEE0A56CFEA6" bold="true" box="[189,334,265,291]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="3F844722EF7FFF99E1C4FEE0A56CFEA6" bold="true" box="[189,334,265,291]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="0D4F9B30EF7FFF99E193FED1A508FED7" blockId="13.[189,1399,312,478]" box="[234,298,312,338]" pageId="13" pageNumber="14">
<emphasis id="3F844722EF7FFF99E193FED1A508FED7" bold="true" box="[234,298,312,338]" pageId="13" pageNumber="14">Male</emphasis>
</paragraph>
<paragraph id="0D4F9B30EF7FFF99E1C4FEB2A5CCFE5B" blockId="13.[189,1399,312,478]" pageId="13" pageNumber="14">
HEAD (
<figureCitation id="95CB87B5EF7FFF99E072FEB2A5B1FEF0" box="[267,403,347,373]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Figs 12–16</figureCitation>
). Antennal flagellum length 727 μm (687–727 μm, n = 4); AR 1.06 (0.97–1.19, n = 4) (
<figureCitation id="95CB87B5EF7FFF99E1BFFE97A53CFE1D" box="[198,286,382,408]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Fig. 13</figureCitation>
). Frontal sclerite elliptical (
<figureCitation id="95CB87B5EF7FFF99E313FE97A6E6FE1D" box="[618,708,382,408]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Fig. 14</figureCitation>
). Clypeus entire, with 8 (6–8, n = 4) long setae (
<figureCitation id="95CB87B5EF7FFF99E476FE97A14AFE1D" box="[1295,1384,382,408]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Fig. 15</figureCitation>
). Palp segment 3 slender, 90 μm (85–91 μm, n = 4); PR (III) 6.6 (6.3, n = 4) (
<figureCitation id="95CB87B5EF7FFF99E569FE48A045FE3E" box="[1040,1127,417,443]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
); first half with hyaline sensillae on inner surface.
</paragraph>
<paragraph id="0D4F9B30EF7FFF99E1C4FDEFA6E7FDE3" blockId="13.[189,1398,518,614]" pageId="13" pageNumber="14">
THORAX (
<figureCitation id="95CB87B5EF7FFF99E051FDEFA593FDA5" box="[296,433,518,544]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Figs 17–18</figureCitation>
). Scutellum pale, with 10 bristles (9–10, n = 4). Wing length 1.15 (
<quantity id="CA0836D5EF7FFF99E5BAFDEFA151FDA5" box="[1219,1395,518,544]" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.17" metricValueMin="1.08" pageId="13" pageNumber="14" unit="mm" value="1.125" valueMax="1.17" valueMin="1.08">1.08–1.17 mm</quantity>
, n = 4); CR 0.45 (0.46–048, n = 4); macrotrichia numerous (
<figureCitation id="95CB87B5EF7FFF99E208FDC0A7E5FDC6" box="[881,967,553,579]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
); first radial cell slit-like, second cell open (
<figureCitation id="95CB87B5EF7FFF99E07FFDA5A57FFDE3" box="[262,349,588,614]" captionStart="Figs 12–18" captionStartId="12.[189,241,1929,1955]" captionTargetBox="[253,1335,264,1898]" captionTargetId="figure@12.[229,1358,244,1918]" captionTargetPageId="12" captionText="Figs 12–18. Dasyhelea gothlandica sp. nov., holotype, ♂. 12. Head. 13. Distal flagellomeres. 14. Frontal sclerite. 15. Clypeus. 16. Palp. 17. Wing. 18. First and second radial cell of wing. Scale bars: 12, 17 = 100 μm; 14, 18 = 50 μm; 13, 15–16 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785236" httpUri="https://zenodo.org/record/3785236/files/figure.png" pageId="13" pageNumber="14">Fig. 18</figureCitation>
). Legs light brown, tarsi paler.
</paragraph>
<paragraph id="0D4F9B30EF7FFF99E1C4FD67A542FCDD" blockId="13.[189,1399,654,856]" pageId="13" pageNumber="14">
GENITALIA (
<figureCitation id="95CB87B5EF7FFF99E038FD67A5EEFD2D" box="[321,460,654,680]" captionStart="Figs19–22" captionStartId="13.[189,241,1891,1917]" captionTargetBox="[193,1401,918,1849]" captionTargetId="figure@13.[157,1420,909,1858]" captionTargetPageId="13" captionText="Figs19–22. Dasyheleagothlandica sp.nov.,holotype, ♂.19–20.Ventralaspectof genitalia.21. Parameres. 22. Aedeagus. Scale bars: 19–20 = 50 μm; 21–22 = 16 μm." figureDoi="http://doi.org/10.5281/zenodo.3785238" httpUri="https://zenodo.org/record/3785238/files/figure.png" pageId="13" pageNumber="14">Figs 19–22</figureCitation>
). Apicolateral process of tergite 9 short. Posterior margin of sternite 9 apically divided into pair of obtuse projections. Gonostylus short, tapering to indented sharp point, covered by small setae, two long bristles at midpoint present (
<figureCitation id="95CB87B5EF7FFF99E245FD3DA7E5FD6B" box="[828,967,724,750]" captionStart="Figs19–22" captionStartId="13.[189,241,1891,1917]" captionTargetBox="[193,1401,918,1849]" captionTargetId="figure@13.[157,1420,909,1858]" captionTargetPageId="13" captionText="Figs19–22. Dasyheleagothlandica sp.nov.,holotype, ♂.19–20.Ventralaspectof genitalia.21. Parameres. 22. Aedeagus. Scale bars: 19–20 = 50 μm; 21–22 = 16 μm." figureDoi="http://doi.org/10.5281/zenodo.3785238" httpUri="https://zenodo.org/record/3785238/files/figure.png" pageId="13" pageNumber="14">Figs 19–20</figureCitation>
). Parameres asymmetrical; posterior process fused with right arm; long and slender, distinctly bent apically, with fine apical hairs (
<figureCitation id="95CB87B5EF7FFF99E468FD1EA14BFC94" box="[1297,1385,759,785]" captionStart="Figs19–22" captionStartId="13.[189,241,1891,1917]" captionTargetBox="[193,1401,918,1849]" captionTargetId="figure@13.[157,1420,909,1858]" captionTargetPageId="13" captionText="Figs19–22. Dasyheleagothlandica sp.nov.,holotype, ♂.19–20.Ventralaspectof genitalia.21. Parameres. 22. Aedeagus. Scale bars: 19–20 = 50 μm; 21–22 = 16 μm." figureDoi="http://doi.org/10.5281/zenodo.3785238" httpUri="https://zenodo.org/record/3785238/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
). Aedeagus low arched with apicolateral processes broad, somewhat folded, extended laterally in distal part (
<figureCitation id="95CB87B5EF7FFF99E182FCD7A570FCDD" box="[251,338,830,856]" captionStart="Figs19–22" captionStartId="13.[189,241,1891,1917]" captionTargetBox="[193,1401,918,1849]" captionTargetId="figure@13.[157,1420,909,1858]" captionTargetPageId="13" captionText="Figs19–22. Dasyheleagothlandica sp.nov.,holotype, ♂.19–20.Ventralaspectof genitalia.21. Parameres. 22. Aedeagus. Scale bars: 19–20 = 50 μm; 21–22 = 16 μm." figureDoi="http://doi.org/10.5281/zenodo.3785238" httpUri="https://zenodo.org/record/3785238/files/figure.png" pageId="13" pageNumber="14">Fig. 22</figureCitation>
).
</paragraph>
<caption id="598FCBB8EF7FFF99E1C4F88AA753F825" ID-DOI="http://doi.org/10.5281/zenodo.3785238" ID-Zenodo-Dep="3785238" httpUri="https://zenodo.org/record/3785238/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,241,1891,1917]" targetBox="[193,1401,918,1849]" targetPageId="13">
<paragraph id="0D4F9B30EF7FFF99E1C4F88AA753F825" blockId="13.[189,1399,1891,1952]" pageId="13" pageNumber="14">
<emphasis id="3F844722EF7FFF99E1C4F88AA566F8F8" bold="true" box="[189,324,1891,1917]" pageId="13" pageNumber="14">Figs19–22.</emphasis>
<taxonomicName id="CAF0E0B3EF7FFF99E031F88AA671F8F8" authority="Strandberg &amp; Johanson, 2015" authorityName="Strandberg &amp; Johanson" authorityYear="2015" box="[328,595,1891,1917]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gothlandica" status="sp. nov.">
<emphasis id="3F844722EF7FFF99E031F88AA671F8F8" box="[328,595,1891,1917]" italics="true" pageId="13" pageNumber="14">Dasyhelea gothlandica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24B7FA59EF7FFF99E32EF88AA692F8F8" box="[599,688,1891,1917]" pageId="13" pageNumber="14" rank="species">sp.nov.</taxonomicNameLabel>
,holotype, ♂. 
<emphasis id="3F844722EF7FFF99E230F88AA7B1F8F8" bold="true" box="[841,915,1891,1917]" pageId="13" pageNumber="14">19–20</emphasis>
.Ventral aspect of genitalia. 
<emphasis id="3F844722EF7FFF99E5B4F88AA0C8F8F8" bold="true" box="[1229,1258,1891,1917]" pageId="13" pageNumber="14">21</emphasis>
. Parameres. 
<emphasis id="3F844722EF7FFF99E1C4F86FA4F8F825" bold="true" box="[189,218,1926,1952]" pageId="13" pageNumber="14">22</emphasis>
. Aedeagus. Scale bars: 19–20 = 50 μm; 21–22 = 16 μm.
</paragraph>
</caption>
<caption id="598FCBB8EF7CFF9AE1C4F89AA64BF856" ID-DOI="http://doi.org/10.5281/zenodo.3785240" ID-Zenodo-Dep="3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,241,1907,1933]" targetBox="[190,1400,280,1872]" targetPageId="14">
<paragraph id="0D4F9B30EF7CFF9AE1C4F89AA64BF856" blockId="14.[189,1399,1907,2003]" pageId="14" pageNumber="15">
<emphasis id="3F844722EF7CFF9AE1C4F89AA567F808" bold="true" box="[189,325,1907,1933]" pageId="14" pageNumber="15">Figs 23–29.</emphasis>
<taxonomicName id="CAF0E0B3EF7CFF9AE030F89AA677F808" authority="Strandberg &amp; Johanson, 2015" authorityName="Strandberg &amp; Johanson" authorityYear="2015" box="[329,597,1907,1933]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gothlandica" status="sp. nov.">
<emphasis id="3F844722EF7CFF9AE030F89AA677F808" box="[329,597,1907,1933]" italics="true" pageId="14" pageNumber="15">Dasyhelea gothlandica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24B7FA59EF7CFF9AE320F89AA691F808" box="[601,691,1907,1933]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
,paratype, ♀. 
<emphasis id="3F844722EF7CFF9AE234F89AA748F808" bold="true" box="[845,874,1907,1933]" pageId="14" pageNumber="15">23</emphasis>
. Head. 
<emphasis id="3F844722EF7CFF9AE2C6F89AA7FEF808" bold="true" box="[959,988,1907,1933]" pageId="14" pageNumber="15">24</emphasis>
. Flagellomeres. 
<emphasis id="3F844722EF7CFF9AE5E0F89AA094F808" bold="true" box="[1177,1206,1907,1933]" pageId="14" pageNumber="15">25</emphasis>
. Frontal sclerite. 
<emphasis id="3F844722EF7CFF9AE1C4F87FA4F8F835" bold="true" box="[189,218,1942,1968]" pageId="14" pageNumber="15">26</emphasis>
. Clypeus. 
<emphasis id="3F844722EF7CFF9AE021F87FA557F835" bold="true" box="[344,373,1942,1968]" pageId="14" pageNumber="15">27</emphasis>
. Palp. 
<emphasis id="3F844722EF7CFF9AE0B1F87FA5C7F835" bold="true" box="[456,485,1942,1968]" pageId="14" pageNumber="15">28</emphasis>
. Wing. 
<emphasis id="3F844722EF7CFF9AE33AF87FA642F835" bold="true" box="[579,608,1942,1968]" pageId="14" pageNumber="15">29</emphasis>
. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm.
</paragraph>
</caption>
<paragraph id="0D4F9B30EF7DFF9BE193FEE0A567FEA6" blockId="15.[189,1399,265,432]" box="[234,325,265,291]" pageId="15" pageNumber="16">
<emphasis id="3F844722EF7DFF9BE193FEE0A567FEA6" bold="true" box="[234,325,265,291]" pageId="15" pageNumber="16">Female</emphasis>
</paragraph>
<paragraph id="0D4F9B30EF7DFF9BE1C4FEC5A6FEFE35" blockId="15.[189,1399,265,432]" pageId="15" pageNumber="16">
HEAD (
<figureCitation id="95CB87B5EF7DFF9BE075FEC5A5B4FEC3" box="[268,406,300,326]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Figs 23–27</figureCitation>
). Antennal flagellum length 545 μm (444–525, n = 3); distal flagellomeres elongate; AR 0.96 (0.86–1.0, n = 3) (
<figureCitation id="95CB87B5EF7DFF9BE086FEA6A67AFEEC" box="[511,600,335,361]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Fig. 24</figureCitation>
). Clypeus entire, with 8 (n = 3) long setae (
<figureCitation id="95CB87B5EF7DFF9BE521FEA6A092FEEC" box="[1112,1200,335,361]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Fig. 25</figureCitation>
). Frontal sclerite elliptical (
<figureCitation id="95CB87B5EF7DFF9BE04DFE9AA5AEFE08" box="[308,396,371,397]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Fig. 26</figureCitation>
). Palp segment 3 slender, 80 μm (80 μm, n = 4); PR (III) 5.6 (5.3, n = 3) (
<figureCitation id="95CB87B5EF7DFF9BE5ACFE9AA10FFE08" box="[1237,1325,371,397]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Fig. 27</figureCitation>
); with hyaline sensillae at basal part of inner surface.
</paragraph>
<paragraph id="0D4F9B30EF7DFF9BE1C4FE3EA5D3FDBD" blockId="15.[189,1399,471,568]" pageId="15" pageNumber="16">
THORAX (
<figureCitation id="95CB87B5EF7DFF9BE050FE3EA597FE74" box="[297,437,471,497]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Figs 28–29</figureCitation>
). Scutellum pale, with 8 bristles (9–10, n = 3). Wing length 
<quantity id="CA0836D5EF7DFF9BE5F2FE3EA0C5FE74" box="[1163,1255,471,497]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="1.1">1.1 mm</quantity>
(1.05–1.13, n = 3); CR 0.46 (0.44–0.47, n = 3) (
<figureCitation id="95CB87B5EF7DFF9BE3FEFE12A6C1FD90" box="[647,739,507,533]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Fig. 28</figureCitation>
). First radial cell slit-like, second cell open but small (
<figureCitation id="95CB87B5EF7DFF9BE1BFFDF7A53CFDBD" box="[198,286,542,568]" captionStart="Figs 23–29" captionStartId="14.[189,241,1907,1933]" captionTargetBox="[190,1400,280,1872]" captionTargetId="figure@14.[147,1433,250,1884]" captionTargetPageId="14" captionText="Figs 23–29. Dasyhelea gothlandica sp. nov.,paratype, ♀. 23. Head. 24. Flagellomeres. 25. Frontal sclerite. 26. Clypeus. 27. Palp. 28. Wing. 29. First and second radial cell of wing. Scale bars: 23, 28 = 100 μm; 25–26, 29 = 50 μm; 24, 27 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.3785240" httpUri="https://zenodo.org/record/3785240/files/figure.png" pageId="15" pageNumber="16">Fig. 29</figureCitation>
). Legs as in male.
</paragraph>
<paragraph id="0D4F9B30EF7DFF9BE1C4FDB6A55BFD45" blockId="15.[189,1399,607,704]" pageId="15" pageNumber="16">
GENITALIA (
<figureCitation id="95CB87B5EF7DFF9BE031FDB6A5FBFDFC" box="[328,473,607,633]" captionStart="Figs 30–32" captionStartId="15.[189,241,1878,1904]" captionTargetBox="[216,1381,1167,1840]" captionTargetId="figure@15.[175,1412,1163,1847]" captionTargetPageId="15" captionText="Figs 30–32. Dasyhelea gothlandica sp. nov., paratype, ♀. 30–31. Subgenital plate. 32. Seminal capsule. Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3785242" httpUri="https://zenodo.org/record/3785242/files/figure.png" pageId="15" pageNumber="16">Figs 30–32</figureCitation>
). Subgenital plate with notum separated from ramus, slightly fragmented (
<figureCitation id="95CB87B5EF7DFF9BE1BCFD6BA56EFD19" box="[197,332,642,668]" captionStart="Figs 30–32" captionStartId="15.[189,241,1878,1904]" captionTargetBox="[216,1381,1167,1840]" captionTargetId="figure@15.[175,1412,1163,1847]" captionTargetPageId="15" captionText="Figs 30–32. Dasyhelea gothlandica sp. nov., paratype, ♀. 30–31. Subgenital plate. 32. Seminal capsule. Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3785242" httpUri="https://zenodo.org/record/3785242/files/figure.png" pageId="15" pageNumber="16">Figs 30–31</figureCitation>
). Spermatheca single; ovoid-shaped with short neck; length × width: 60 × 40 μm (60 × 40 μm, n = 3) (
<figureCitation id="95CB87B5EF7DFF9BE06AFD4FA549FD45" box="[275,363,678,704]" captionStart="Figs 30–32" captionStartId="15.[189,241,1878,1904]" captionTargetBox="[216,1381,1167,1840]" captionTargetId="figure@15.[175,1412,1163,1847]" captionTargetPageId="15" captionText="Figs 30–32. Dasyhelea gothlandica sp. nov., paratype, ♀. 30–31. Subgenital plate. 32. Seminal capsule. Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3785242" httpUri="https://zenodo.org/record/3785242/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="45EAC8BBEF7DFF9BE1C4FD0EA79CFC1C" pageId="15" pageNumber="16" type="discussion">
<paragraph id="0D4F9B30EF7DFF9BE1C4FD0EA50DFC84" blockId="15.[189,303,743,769]" box="[189,303,743,769]" pageId="15" pageNumber="16">
<heading id="56072C5CEF7DFF9BE1C4FD0EA50DFC84" bold="true" box="[189,303,743,769]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="3F844722EF7DFF9BE1C4FD0EA50DFC84" bold="true" box="[189,303,743,769]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="0D4F9B30EF7DFF9BE1C4FCFFA79CFC1C" blockId="15.[189,1399,790,921]" pageId="15" pageNumber="16">
This species is a member of the subgenus 
<taxonomicName id="CAF0E0B3EF7DFF9BE3CDFCFEA77DFCB5" authorityName="Enderlein" authorityYear="1936" box="[692,863,791,816]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subGenus" subGenus="Dicryptoscena">
<emphasis id="3F844722EF7DFF9BE3CDFCFEA77DFCB5" box="[692,863,791,816]" italics="true" pageId="15" pageNumber="16">Dicryptoscena</emphasis>
</taxonomicName>
, and the male is similar to that of 
<taxonomicName id="CAF0E0B3EF7DFF9BE582FCFFA63BFCD6" authority="Santos Abreu, 1918" authorityName="Santos Abreu" authorityYear="1918" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="albidipes">
<emphasis id="3F844722EF7DFF9BE582FCFFA50BFCD6" italics="true" pageId="15" pageNumber="16">Dasyhelea albidipes</emphasis>
Santos Abreu, 
<quantity id="CA0836D5EF7DFF9BE0A7FCD0A63BFCD6" box="[478,537,825,851]" metricMagnitude="1" metricUnit="m" metricValue="4.87172" pageId="15" pageNumber="16" unit="in" value="1918.0">1918</quantity>
</taxonomicName>
in the morphology of the genitalia, particularly by the presence of a pair of processes on the posterior margin of sternite 9. 
<taxonomicName id="CAF0E0B3EF7DFF9BE269FCB4A7E2FCF3" authority="Strandberg &amp; Johanson, 2015" authorityName="Strandberg &amp; Johanson" authorityYear="2015" box="[784,960,860,886]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gothlandica" status="sp. nov.">
<emphasis id="3F844722EF7DFF9BE269FCB4A7E2FCF3" box="[784,960,860,886]" italics="true" pageId="15" pageNumber="16">D. gothlandica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24B7FA59EF7DFF9BE2B1FCB5A006FCF3" box="[968,1060,860,886]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="CAF0E0B3EF7DFF9BE5B8FCB4A173FCF3" authorityName="Santos Abreu" authorityYear="1918" box="[1217,1361,860,886]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="albidipes">
<emphasis id="3F844722EF7DFF9BE5B8FCB4A173FCF3" box="[1217,1361,860,886]" italics="true" pageId="15" pageNumber="16">D. albidipes</emphasis>
</taxonomicName>
by the broadly shaped apicolateral processes and the short gonostyli.
</paragraph>
</subSubSection>
<subSubSection id="45EAC8BBEF7DFF9BE1C4FC28A045FBCA" pageId="15" pageNumber="16" type="biology_ecology">
<paragraph id="0D4F9B30EF7DFF9BE1C4FC28A53EFC5E" blockId="15.[189,284,961,987]" box="[189,284,961,987]" pageId="15" pageNumber="16">
<heading id="56072C5CEF7DFF9BE1C4FC28A53EFC5E" bold="true" box="[189,284,961,987]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="3F844722EF7DFF9BE1C4FC28A53EFC5E" bold="true" box="[189,284,961,987]" pageId="15" pageNumber="16">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="0D4F9B30EF7DFF9BE1C4FC06A045FBCA" blockId="15.[189,1399,1007,1103]" pageId="15" pageNumber="16">
The 
<typeStatus id="D24B2592EF7DFF9BE18CFC06A57EFB8C" box="[245,348,1007,1033]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
was collected in a Malaise trap next to Lake Bästeträsk, with sourrounding vegetation consisting of 
<taxonomicName id="CAF0E0B3EF7DFF9BE025FBFDA6AAFBA9" authority="Linnaeus" authorityName="Linnaeus" box="[348,648,1042,1069]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="3F844722EF7DFF9BE025FBFDA636FBA9" box="[348,532,1042,1069]" italics="true" pageId="15" pageNumber="16">Pinus sylvestris</emphasis>
Linnaeus
</taxonomicName>
and small shrubs. One of the male 
<typeStatus id="D24B2592EF7DFF9BE552FBFBA0B9FBA9" box="[1067,1179,1042,1068]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
was collected in a Malaise trap next to Lake Horsan, approximatley 
<quantity id="CA0836D5EF7DFF9BE27DFBDCA71DFBCA" box="[772,831,1077,1103]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="5.0">5 km</quantity>
south-east of Bästeträsk.
</paragraph>
</subSubSection>
</treatment>
</document>