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<document id="CB128D22B263FB0BD1DEC6DC9BD20C14" ID-CLB-Dataset="31186" ID-DOI="10.11646/zootaxa.4363.3.9" ID-GBIF-Dataset="0313c9b7-5656-4c15-8218-3bea41391dbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1108254" ID-ZooBank="15B32E25-7BCD-4AA3-9610-9B6E9AD5500B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513065121575" checkinUser="plazi" docAuthor="Sankaran, Pradeep M. &amp; Sebastian, Pothalil A." docDate="2017" docId="854687F2FF9CFFDF7DC8FA11FF00AEC1" docLanguage="en" docName="zootaxa.4363.3.9.pdf" docOrigin="Zootaxa 4363 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anaxibia folia Sankaran &amp; Sebastian 2017, spec. nov." docType="treatment" docVersion="7" lastPageNumber="444" masterDocId="797FFF8AFF9CFFDC7D5FFFE4FFDCA843" masterDocTitle="Anaxibia folia spec. nov. - a new litter-dwelling dictynid spider from India (Araneae: Dictynidae: Dictyninae)" masterLastPageNumber="444" masterPageNumber="441" pageNumber="441" updateTime="1698509457369" updateUser="ExternalLinkService">
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<mods:title id="B4A16D52AFEDC7FD8C880002CD4F7371">Anaxibia folia spec. nov. - a new litter-dwelling dictynid spider from India (Araneae: Dictynidae: Dictyninae)</mods:title>
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<mods:name id="EF45677EB699C436631C9B9ACEC70184" type="personal">
<mods:role id="8A90DC9B8DF60878DBF746B11C6F5B92">
<mods:roleTerm id="E29AFBEBECA06AA609D4D9907B6E8D0A">Author</mods:roleTerm>
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<mods:namePart id="62CC7F3263ED38F34DA3D9E3C40D83B4">Sankaran, Pradeep M.</mods:namePart>
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<mods:namePart id="CA48B6C0F640DA3C10078157BD486499">Sebastian, Pothalil A.</mods:namePart>
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<mods:typeOfResource id="070F70084FFA50A0B958D63C9EC5F766">text</mods:typeOfResource>
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<mods:title id="48E0435B6799C3FBB6763AC4DD5547BA">Zootaxa</mods:title>
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<mods:part id="EB8AAAF96A3B5199212A69B3C2FF2841">
<mods:date id="57A525BE5F8BD2594B7EAC223E75EB95">2017</mods:date>
<mods:detail id="6D8B165E9F02C4F2D083462FB94E43D7" type="pubDate">
<mods:number id="EE6C9717B47EEE653E47925D6C593365">2017-12-12</mods:number>
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<mods:number id="2D6736ACB470E7B51469D31D5280E033">4363</mods:number>
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<treatment id="854687F2FF9CFFDF7DC8FA11FF00AEC1" ID-DOI="http://doi.org/10.5281/zenodo.5999298" ID-GBIF-Taxon="137678958" ID-Zenodo-Dep="5999298" LSID="urn:lsid:plazi:treatment:854687F2FF9CFFDF7DC8FA11FF00AEC1" httpUri="http://treatment.plazi.org/id/854687F2FF9CFFDF7DC8FA11FF00AEC1" lastPageId="3" lastPageNumber="444" pageId="0" pageNumber="441">
<subSubSection id="45F5656FFF9CFFDC7DC8FA11FD9AAE53" box="[151,582,1525,1552]" pageId="0" pageNumber="441" type="nomenclature">
<paragraph id="0D5036E4FF9CFFDC7DC8FA11FD9AAE53" blockId="0.[151,1436,1388,2037]" box="[151,582,1525,1552]" pageId="0" pageNumber="441">
<emphasis id="3F9BEAF6FF9CFFDC7DC8FA11FD9AAE53" bold="true" box="[151,582,1525,1552]" pageId="0" pageNumber="441">
<taxonomicName id="CAEF4D67FF9CFFDC7DC8FA11FE98AE4C" ID-CoL="DK2P" authority="Sankaran &amp; Sebastian, 2017" authorityName="Sankaran &amp; Sebastian" authorityYear="2017" box="[151,324,1525,1551]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="folia" status="sp. nov.">
<emphasis id="3F9BEAF6FF9CFFDC7DC8FA11FE98AE4C" bold="true" box="[151,324,1525,1551]" italics="true" pageId="0" pageNumber="441">Anaxibia folia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24A8578DFF9CFFDC7C13FA12FE1FAE53" box="[332,451,1526,1552]" pageId="0" pageNumber="441" rank="species">spec. nov.</taxonomicNameLabel>
(
<figureCitation id="95D42A61FF9CFFDC7C8CFA12FDE0AE53" box="[467,572,1526,1552]" captionStart-0="FIGURES 1" captionStart-1="FIGURES 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,258,1573,1595]" captionStartId-1="2.[151,258,1850,1872]" captionStartId-2="3.[151,244,994,1016]" captionTargetBox-0="[161,1428,307,1541]" captionTargetBox-1="[212,1385,657,1829]" captionTargetBox-2="[163,1424,189,976]" captionTargetId-0="figure@1.[151,1436,295,1551]" captionTargetId-1="figure@2.[196,1401,637,1841]" captionTargetId-2="figure@3.[151,1436,180,989]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." captionText-1="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." captionText-2="FIGURE 3. Anaxibiafolia spec. nov., type locality." httpUri-0="https://zenodo.org/record/1108256/files/figure.png" httpUri-1="https://zenodo.org/record/1108258/files/figure.png" httpUri-2="https://zenodo.org/record/1108260/files/figure.png" pageId="0" pageNumber="441">
Figs 1
<docIssue id="498C490BFF9CFFDC7F72FA12FDE0AE53" box="[557,572,1526,1552]" pageId="0" pageNumber="441">3</docIssue>
</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="45F5656FFF9CFFDC7DC8F9FDFC5FAE3B" pageId="0" pageNumber="441" type="materials_examined">
<paragraph id="0D5036E4FF9CFFDC7DC8F9FDFC5FAE3B" blockId="0.[151,1436,1388,2037]" pageId="0" pageNumber="441">
<materialsCitation id="BD873CB9FF9CFFDC7DC8F9FDFB21AE16" ID-GBIF-Occurrence="1705249402" collectingDate="2013-05-16" collectionCode="ADSH" collectorName="P. M. Sankaran" country="India" county="Male" elevation="20" latitude="11.26838" location="Nilambur" longLatPrecision="1" longitude="76.20617" municipality="Malappuram" pageId="0" pageNumber="441" specimenCode="ADSH2831" specimenCount="2" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="3F9BEAF6FF9CFFDC7DC8F9FDFE0FAE73" bold="true" box="[151,467,1561,1584]" pageId="0" pageNumber="441">
<typeStatus id="D2548846FF9CFFDC7DC8F9FDFF13AE73" box="[151,207,1561,1584]" pageId="0" pageNumber="441">Type</typeStatus>
material:
<typeStatus id="D2548846FF9CFFDC7C3CF9FDFE13AE73" box="[355,463,1561,1584]" pageId="0" pageNumber="441" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCounty id="E4314E68FF9CFFDC7CB6F9FDFDFCAE73" box="[489,544,1561,1584]" pageId="0" pageNumber="441">Male</collectingCounty>
(
<specimenCode id="5D499E9FFF9CFFDC7F63F9FDFD63AE73" box="[572,703,1561,1584]" collectionCode="ADSH" country="India" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="0" pageNumber="441">ADSH2831</specimenCode>
A),
<emphasis id="3F9BEAF6FF9CFFDC7FABF9FDFC68AE73" bold="true" box="[756,948,1560,1584]" pageId="0" pageNumber="441">
<collectingCountry id="75F87674FF9CFFDC7FABF9FDFC9BAE73" box="[756,839,1561,1584]" name="India" pageId="0" pageNumber="441">INDIA</collectingCountry>
:
<collectingRegion id="CF2BF806FF9CFFDC7E3EF9FCFC77AE73" box="[865,939,1560,1584]" country="India" name="Kerala" pageId="0" pageNumber="441">
<emphasis id="3F9BEAF6FF9CFFDC7E3EF9FCFC77AE73" bold="true" box="[865,939,1560,1584]" italics="true" pageId="0" pageNumber="441">Kerala</emphasis>
</collectingRegion>
:
</emphasis>
<collectingMunicipality id="ED34AC9EFF9CFFDC7E96F9FDFB88AE73" box="[969,1108,1561,1584]" pageId="0" pageNumber="441">Malappuram</collectingMunicipality>
,
<location id="0830603FFF9CFFDC7932F9FDFB09AE73" LSID="urn:lsid:plazi:treatment:854687F2FF9CFFDF7DC8FA11FF00AEC1:0830603FFF9CFFDC7932F9FDFB09AE73" box="[1133,1237,1561,1584]" country="India" county="Male" latitude="11.26838" longLatPrecision="1" longitude="76.20617" municipality="Malappuram" name="Nilambur" pageId="0" pageNumber="441" stateProvince="Kerala">Nilambur</location>
,
<location id="0830603FFF9CFFDC79B0F9FDFA44AE73" LSID="urn:lsid:plazi:treatment:854687F2FF9CFFDF7DC8FA11FF00AEC1:0830603FFF9CFFDC79B0F9FDFA44AE73" box="[1263,1432,1561,1584]" country="India" county="Male" latitude="11.26838" longLatPrecision="1" longitude="76.20617" municipality="Malappuram" name="Canolly's Plot" pageId="0" pageNumber="441" stateProvince="Kerala">Canollys Plot</location>
,
<geoCoordinate id="68DB5023FF9CFFDC7DC8F9DAFEEFAE16" box="[151,307,1596,1621]" direction="north" orientation="latitude" pageId="0" pageNumber="441" precision="1" value="11.26838">11o16'06.17''N</geoCoordinate>
,
<geoCoordinate id="68DB5023FF9CFFDC7C1FF9DAFE06AE16" box="[320,474,1596,1621]" direction="east" orientation="longitude" pageId="0" pageNumber="441" precision="1" value="76.20617">76o12'22.21''E</geoCoordinate>
,
<elevation id="86C2D1D7FF9CFFDC7CB8F9DAFDFCAE16" box="[487,544,1598,1621]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="441" unit="m" value="20.0">
<quantity id="CA179B01FF9CFFDC7CB8F9DAFDFCAE16" box="[487,544,1598,1621]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="441" unit="m" value="20.0">20 m</quantity>
</elevation>
alt.,
<collectingDate id="6915E9CCFF9CFFDC7F04F9DAFD2DAE16" box="[603,753,1598,1621]" pageId="0" pageNumber="441" value="2013-05-16">
<date id="79511024FF9CFFDC7F04F9DAFD2DAE16" box="[603,753,1598,1621]" pageId="0" pageNumber="441" value="2013-05-16">16 May 2013</date>
</collectingDate>
,
<collectorName id="A01A5332FF9CFFDC7FA0F9DAFC40AE16" box="[767,924,1598,1621]" pageId="0" pageNumber="441">P.M. Sankaran</collectorName>
leg., from forest litter, by hand
</materialsCitation>
.
<materialsCitation id="BD873CB9FF9CFFDC7855F9DAFCA3AE3B" ID-GBIF-Occurrence="1705249403" collectingDate="2013-05-16" collectionCode="ADSH" collectorName="P. M. Sankaran" country="India" county="Male" elevation="20" latitude="11.26838" location="Nilambur" longLatPrecision="1" longitude="76.20617" municipality="Malappuram" pageId="0" pageNumber="441" specimenCode="ADSH2831" specimenCount="4" specimenCount-female="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="paratype">
<emphasis id="3F9BEAF6FF9CFFDC7855F9DAFA59AE16" bold="true" box="[1290,1413,1598,1621]" pageId="0" pageNumber="441">
<typeStatus id="D2548846FF9CFFDC7855F9DAFA5CAE16" box="[1290,1408,1598,1621]" pageId="0" pageNumber="441" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="1BE9FD6DFF9CFFDC78D1F9DAFF12AE3B" pageId="0" pageNumber="441" type="male">1 male</specimenCount>
,
<specimenCount id="1BE9FD6DFF9CFFDC7D87F985FE9EAE3B" box="[216,322,1633,1656]" pageId="0" pageNumber="441" type="female">2 females</specimenCount>
,
<specimenCount id="1BE9FD6DFF9CFFDC7C13F985FE6AAE3B" box="[332,438,1633,1656]" pageId="0" pageNumber="441" type="juvenile">1 juvenile</specimenCount>
(
<specimenCode id="5D499E9FFF9CFFDC7C9BF985FD94AE3B" box="[452,584,1633,1656]" collectionCode="ADSH" country="India" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="0" pageNumber="441">ADSH2831</specimenCode>
B), same data as for
<typeStatus id="D2548846FF9CFFDC7E41F985FCA3AE3B" box="[798,895,1633,1656]" pageId="0" pageNumber="441" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="45F5656FFF9CFFDC7D98F967FE21AEFE" pageId="0" pageNumber="441" type="etymology">
<paragraph id="0D5036E4FF9CFFDC7D98F967FE21AEFE" blockId="0.[151,1436,1388,2037]" pageId="0" pageNumber="441">
<emphasis id="3F9BEAF6FF9CFFDC7D98F967FE9BAED9" bold="true" box="[199,327,1667,1690]" pageId="0" pageNumber="441">Etymology.</emphasis>
The specific epithet is a neuter plural in apposition, derived from the leaf litter-dwelling habit of the new species. Latin: folium = leaf.
</paragraph>
</subSubSection>
<subSubSection id="45F5656FFF9CFFDD7D98F92DFCCCA8B7" lastPageId="1" lastPageNumber="442" pageId="0" pageNumber="441" type="diagnosis">
<paragraph id="0D5036E4FF9CFFDD7D98F92DFCCCA8B7" blockId="0.[151,1436,1388,2037]" lastBlockId="1.[151,1435,151,244]" lastPageId="1" lastPageNumber="442" pageId="0" pageNumber="441">
<emphasis id="3F9BEAF6FF9CFFDC7D98F92DFEE0AEA3" bold="true" box="[199,316,1737,1760]" pageId="0" pageNumber="441">Diagnosis.</emphasis>
<taxonomicName id="CAEF4D67FF9CFFDC7C17F92CFE3AAEA3" authority="Sankaran &amp; Sebastian, 2017" authorityName="Sankaran &amp; Sebastian" authorityYear="2017" box="[328,486,1736,1760]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="folia" status="sp. nov.">
<emphasis id="3F9BEAF6FF9CFFDC7C17F92CFE3AAEA3" box="[328,486,1736,1760]" italics="true" pageId="0" pageNumber="441">Anaxibia folia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24A8578DFF9CFFDC7CACF92DFDBAAEA3" box="[499,614,1737,1760]" pageId="0" pageNumber="441" rank="species">
<emphasis id="3F9BEAF6FF9CFFDC7CACF92DFDBAAEA3" bold="true" box="[499,614,1737,1760]" pageId="0" pageNumber="441">spec. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="CAEF4D67FF9CFFDC7E14F92CFACCAEA3" authority="(Kulczynski, 1908)" authorityName="Kulczynski" authorityYear="1908" baseAuthorityName="Kulczynski" baseAuthorityYear="1908" box="[843,1296,1736,1760]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="pictithorax">
<emphasis id="3F9BEAF6FF9CFFDC7E14F92CFBF1AEA3" box="[843,1069,1736,1760]" italics="true" pageId="0" pageNumber="441">Anaxibia pictithorax</emphasis>
(
<bibRefCitation id="697E4B15FF9CFFDC791DF92DFAD5AEA3" author="Kulczynski" box="[1090,1289,1737,1760]" pageId="0" pageNumber="441" refString="Kulczynski, W. (1908) Symbola ad faunam Aranearum Javae et Sumatrae cognoscendam. I. Mygalomorphae et Cribellatae. Bulletin International de l'Academie des Sciences de Cracovie, 1908, 527 - 581." type="journal article" year="1908">Kulczyński, 1908</bibRefCitation>
)
</taxonomicName>
, but can be separated from the latter by the following combination of characters: cymbial caudal lobe (CL;
<figureCitation id="95D42A61FF9CFFDC7997F90FFA9EAF41" box="[1224,1346,1771,1794]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="0" pageNumber="441">Fig. 2B, D</figureCitation>
) almost straight and remaining covered by tibia in ventral view (cymbial caudal lobe of
<taxonomicName id="CAEF4D67FF9CFFDC7EBAF8EAFBABAF66" baseAuthorityName="Kulczynski" baseAuthorityYear="1908" box="[997,1143,1805,1829]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="pictithorax">
<emphasis id="3F9BEAF6FF9CFFDC7EBAF8EAFBABAF66" box="[997,1143,1805,1829]" italics="true" pageId="0" pageNumber="441">A. pictithorax</emphasis>
</taxonomicName>
with strong baso-prolateral bend and visible in ventral view), tibia with ventral bulging (tibia of
<taxonomicName id="CAEF4D67FF9CFFDC7EDAF8D4FBC5AF0B" baseAuthorityName="Kulczynski" baseAuthorityYear="1908" box="[901,1049,1840,1864]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="pictithorax">
<emphasis id="3F9BEAF6FF9CFFDC7EDAF8D4FBC5AF0B" box="[901,1049,1840,1864]" italics="true" pageId="0" pageNumber="441">A. pictithorax</emphasis>
</taxonomicName>
lacks ventral bulging), dorsal tibial apophysis (DTA;
<figureCitation id="95D42A61FF9CFFDC7C06F8B7FE72AF29" box="[345,430,1875,1898]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="0" pageNumber="441">Fig. 2D</figureCitation>
) short and distant from cymbial caudal lobe (
<taxonomicName id="CAEF4D67FF9CFFDC7EF4F8B7FB9CAF29" baseAuthorityName="Kulczynski" baseAuthorityYear="1908" box="[939,1088,1874,1898]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="pictithorax">
<emphasis id="3F9BEAF6FF9CFFDC7EF4F8B7FB9CAF29" box="[939,1088,1874,1898]" italics="true" pageId="0" pageNumber="441">A. pictithorax</emphasis>
</taxonomicName>
with broad DTA lying close to cymbial caudal lobe) and epigyne with sclerotised horizontal plates (SHP;
<figureCitation id="95D42A61FF9CFFDC7EE1F892FBD2AFCE" box="[958,1038,1910,1933]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="0" pageNumber="441">Fig. 2E</figureCitation>
) and without lateral foveae (epigyne of
<taxonomicName id="CAEF4D67FF9CFFDC7DE8F87CFE90AFF3" baseAuthorityName="Kulczynski" baseAuthorityYear="1908" box="[183,332,1944,1968]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="pictithorax">
<emphasis id="3F9BEAF6FF9CFFDC7DE8F87CFE90AFF3" box="[183,332,1944,1968]" italics="true" pageId="0" pageNumber="441">A. pictithorax</emphasis>
</taxonomicName>
with lateral foveae and without sclerotised horizontal plates) (compare
<figureCitation id="95D42A61FF9CFFDC7935F87DFB03AFF3" box="[1130,1247,1945,1968]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="0" pageNumber="441">Figs 2BE</figureCitation>
with
<bibRefCitation id="697E4B15FF9CFFDC787DF87DFF0CAF91" author="Kulczynski" pageId="0" pageNumber="441" refString="Kulczynski, W. (1908) Symbola ad faunam Aranearum Javae et Sumatrae cognoscendam. I. Mygalomorphae et Cribellatae. Bulletin International de l'Academie des Sciences de Cracovie, 1908, 527 - 581." type="journal article" year="1908">Kulczyński 1908</bibRefCitation>
: plate 23, figs 2729). Females of
<taxonomicName id="CAEF4D67FF9CFFDC7F1EF85FFD53AF91" authority="Sankaran &amp; Sebastian, 2017" authorityName="Sankaran &amp; Sebastian" authorityYear="2017" box="[577,655,1978,2002]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="folia" status="sp. nov.">
<emphasis id="3F9BEAF6FF9CFFDC7F1EF85FFD53AF91" box="[577,655,1978,2002]" italics="true" pageId="0" pageNumber="441">A. folia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24A8578DFF9CFFDC7FC9F85FFCDFAF91" box="[662,771,1979,2002]" pageId="0" pageNumber="441" rank="species">
<emphasis id="3F9BEAF6FF9CFFDC7FC9F85FFCDFAF91" bold="true" box="[662,771,1979,2002]" pageId="0" pageNumber="441">spec. nov.</emphasis>
</taxonomicNameLabel>
can be separated from the females of
<taxonomicName id="CAEF4D67FF9CFFDC79C7F85FFB31AF91" baseAuthorityName="Tikader" baseAuthorityYear="1966" box="[1176,1261,1978,2002]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="rebai">
<emphasis id="3F9BEAF6FF9CFFDC79C7F85FFB31AF91" box="[1176,1261,1978,2002]" italics="true" pageId="0" pageNumber="441">A. rebai</emphasis>
</taxonomicName>
, the only Indian congener by lacking paired brown or black patches on the posterior half of mid-dorsal opisthosoma and epigyne with sclerotised horizontal plates and without lateral foveae, whereas the posterior half of mid-dorsal opisthosoma of
<taxonomicName id="CAEF4D67FF9DFFDD7818FF73FA47A8EC" baseAuthorityName="Tikader" baseAuthorityYear="1966" box="[1351,1435,151,175]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="rebai">
<emphasis id="3F9BEAF6FF9DFFDD7818FF73FA47A8EC" box="[1351,1435,151,175]" italics="true" pageId="1" pageNumber="442">A. rebai</emphasis>
</taxonomicName>
with paired conspicuous deep brown or black patches and epigyne with lateral foveae and lacks sclerotised horizontal plates (compare
<figureCitation id="95D42A61FF9DFFDD7C18FF39FE40A8B7" box="[327,412,221,244]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Figs 1B</figureCitation>
,
<figureCitation id="95D42A61FF9DFFDD7CF7FF39FE1AA8B7" box="[424,454,221,244]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="1" pageNumber="442">2E</figureCitation>
with
<bibRefCitation id="697E4B15FF9DFFDD7F5DFF39FD48A8B7" author="Tikader" box="[514,660,221,244]" pageId="1" pageNumber="442" refString="Tikader, B. K. (1966) Studies on some spiders of the genus Dictyna (family: Dictynidae) from India. Proceedings of the Linnean Society of London, 177, 45 - 53. https: // doi. org / 10.1111 / j. 1095 - 8312.1966. tb 00950. x" type="journal article" year="1966">Tikader 1966</bibRefCitation>
: figs 1ab).
</paragraph>
</subSubSection>
<caption id="5990666CFF9DFFDD7DC8F9C1FB91AE36" httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442" targetBox="[161,1428,307,1541]" targetPageId="1">
<paragraph id="0D5036E4FF9DFFDD7DC8F9C1FB91AE36" blockId="1.[151,1436,1573,1653]" pageId="1" pageNumber="442">
<emphasis id="3F9BEAF6FF9DFFDD7DC8F9C1FE9FAE78" bold="true" box="[151,323,1573,1595]" pageId="1" pageNumber="442">FIGURES 1AF.</emphasis>
<taxonomicName id="CAEF4D67FF9DFFDD7C15F9C1FE0EAE79" authority="Sankaran &amp; Sebastian, 2017" authorityName="Sankaran &amp; Sebastian" authorityYear="2017" box="[330,466,1573,1594]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="folia" status="sp. nov.">
<emphasis id="3F9BEAF6FF9DFFDD7C15F9C1FE0EAE79" box="[330,466,1573,1594]" italics="true" pageId="1" pageNumber="442">Anaxibia folia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24A8578DFF9DFFDD7C86F9C1FDE0AE78" box="[473,572,1573,1595]" pageId="1" pageNumber="442" rank="species">
<emphasis id="3F9BEAF6FF9DFFDD7C86F9C1FDE0AE78" bold="true" box="[473,572,1573,1595]" pageId="1" pageNumber="442">spec. nov.</emphasis>
</taxonomicNameLabel>
AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Female left metatarsus IV showing calamistrum, prolateral. Scale bars: AB 0.5 mm; CF 0.2 mm.
</paragraph>
</caption>
<subSubSection id="45F5656FFF9DFFDF7D98F97FFE23AE59" lastPageId="3" lastPageNumber="444" pageId="1" pageNumber="442" type="description">
<paragraph id="0D5036E4FF9DFFDD7D98F97FFA40AFA9" blockId="1.[151,1436,1691,2026]" pageId="1" pageNumber="442">
<emphasis id="3F9BEAF6FF9DFFDD7D98F97FFE8EAEF1" bold="true" box="[199,338,1691,1714]" pageId="1" pageNumber="442">Description.</emphasis>
Male (
<typeStatus id="D2548846FF9DFFDD7CC0F97FFDDCAEF1" box="[415,512,1691,1714]" pageId="1" pageNumber="442" type="holotype">holotype</typeStatus>
,
<figureCitation id="95D42A61FF9DFFDD7F53F97FFD78AEF1" box="[524,676,1691,1714]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Figs 1A, C, E</figureCitation>
,
<figureCitation id="95D42A61FF9DFFDD7FEFF97FFD08AEF1" box="[688,724,1691,1714]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="1" pageNumber="442">2A</figureCitation>
). Carapace brownish with dark brown striae, with cephalic region elevated more than thoracic region (
<figureCitation id="95D42A61FF9DFFDD7F6BF959FD4EAE97" box="[564,658,1725,1748]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Figs 1A</figureCitation>
,
<figureCitation id="95D42A61FF9DFFDD7FFBF959FD14AE97" box="[676,712,1725,1748]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="1" pageNumber="442">2A</figureCitation>
). Fovea absent. Ocular area, clypeus, chelicerae, fangs, labium, maxillae, sternum brownish. AER recurved, PER straight; AME black, all others pearly white (
<figureCitation id="95D42A61FF9DFFDD79D9F904FB06AEB4" box="[1158,1242,1760,1783]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Fig. 1A</figureCitation>
). Boss prominent. Cheliceral promargin with three teeth, retromargin with one. Maxillae with reduced scopulae. Sternum covered with black hairs, weak medially; truncate between coxae IV, with blunt tip. Opisthosoma widely oval, creamy white with inconspicuous broad greyish patches, covered with a uniform mat of fine greyish hairs; dorsum provided with two pairs of sigillae (
<figureCitation id="95D42A61FF9DFFDD7C4AF88FFEB1AFC1" box="[277,365,1899,1922]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Fig. 1A</figureCitation>
). Anal tubercle prominent (
<figureCitation id="95D42A61FF9DFFDD7FC0F88FFD2AAFC1" box="[671,758,1899,1922]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Fig. 1C</figureCitation>
). Cribellum reduced (
<figureCitation id="95D42A61FF9DFFDD7EB7F88FFBE2AFC1" box="[1000,1086,1899,1922]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Fig. 1C</figureCitation>
). Spinnerets creamy white with greenish shades only on posterior lateral spinneret; anterior lateral spinneret widest, posterior lateral spinneret longest (
<figureCitation id="95D42A61FF9DFFDD7DC0F854FF2EAF84" box="[159,242,1968,1991]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Fig. 1C</figureCitation>
). Legs slender, spineless, setaceous, brownish with green shades; metatarsus IV with reduced calamistrum (
<figureCitation id="95D42A61FF9DFFDD782FF854FF6AAFA9" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="1" pageNumber="442">Fig. 1E</figureCitation>
). Body length 1.35. Prosoma length 0.56, width 0.50. Opisthosoma length 0.79, width 0.58. Eye diameters: ALE 0.03.
</paragraph>
<paragraph id="0D5036E4FF9EFFDE7DC8FF7CFD0BAA14" blockId="2.[151,1436,152,599]" pageId="2" pageNumber="443">
AME 0.04. PLE 0.02. PME 0.02. Eye interdistances: AMEAME 0.02. AMEALE 0.02. AMEPME 0.02. PMEPME 0.05. PMEPLE 0.06. Clypeus height at AMEs 0.07, at ALEs 0.08. Chelicerae length 0.23. Measurements of pedipalp and legs. Pedipalp 0.60 [0.17, 0.11, 0.09, 0.23], I 1.73 [0.50, 0.18, 0.44, 0.36, 0.25], II 1.57 [0.46, 0.16, 0.37, 0.34, 0.24], III 1.35 [0.40, 0.14, 0.30, 0.30, 0.21], IV 1.52 [0.46, 0.15, 0.35, 0.36, 0.20]. Leg formula: 1243. Pedipalp (
<figureCitation id="95D42A61FF9EFFDE784BFEE4FA4CA954" box="[1300,1424,256,279]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Figs 2BD</figureCitation>
): Palpal segments uniformly brownish, spineless; femur disto-dorsally with a short bulging (
<figureCitation id="95D42A61FF9EFFDE7901FEC1FB0BA97F" box="[1118,1239,293,316]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Figs 2CD</figureCitation>
,
<figureCitation id="95D42A61FF9EFFDE79BDFEC1FB33A97F" box="[1250,1263,293,316]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="2" pageNumber="443">1</figureCitation>
<taxonomicName id="CAEF4D67FF9EFFDE79B0FEC7FA9FA97F" box="[1263,1347,291,316]" pageId="2" pageNumber="443" rank="subSpecies" subSpecies="arrow">st arrow</taxonomicName>
); patella retrolaterally with a short, triangular process (PP;
<figureCitation id="95D42A61FF9EFFDE7F9DFEAEFCC4A922" box="[706,792,330,353]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2B</figureCitation>
); tibia with ventral bulging (
<figureCitation id="95D42A61FF9EFFDE7906FEAEFB32A922" box="[1113,1262,330,353]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Figs 2BC, 2</figureCitation>
nd arrow), distoventrally with short, knob-like ventral tibial apophysis (VTA;
<figureCitation id="95D42A61FF9EFFDE7E76FE89FCA0A9C7" box="[809,892,365,388]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2C</figureCitation>
), with an obliquely oriented, plate-like dorsal tibial apophysis (DTA;
<figureCitation id="95D42A61FF9EFFDE7C00FE74FE66A9E4" box="[351,442,400,423]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2D</figureCitation>
). Cymbium proximo-prolaterally with a moderately long caudal lobe (CL;
<figureCitation id="95D42A61FF9EFFDE784FFE74FA53A9E4" box="[1296,1423,400,423]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2B, D</figureCitation>
). Tegulum moderately large. Dictynid terminal apophysis stout, medio-retrolaterally placed (dTA;
<figureCitation id="95D42A61FF9EFFDE79C4FE56FB31A98A" box="[1179,1261,434,457]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2C</figureCitation>
), disto-ventrally with a horizontal ridge (
<figureCitation id="95D42A61FF9EFFDE7CC9FE3CFE35A9AC" box="[406,489,472,495]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2B</figureCitation>
,
<figureCitation id="95D42A61FF9EFFDE7CABFE3CFDDDA9AC" box="[500,513,472,495]" captionStart="FIGURE 3" captionStartId="3.[151,244,994,1016]" captionTargetBox="[163,1424,189,976]" captionTargetId="figure@3.[151,1436,180,989]" captionTargetPageId="3" captionText="FIGURE 3. Anaxibiafolia spec. nov., type locality." httpUri="https://zenodo.org/record/1108260/files/figure.png" pageId="2" pageNumber="443">3</figureCitation>
rd arrow). Subtegulum small, globular (ST;
<figureCitation id="95D42A61FF9EFFDE7E96FE3CFB9CA9AC" box="[969,1088,472,495]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Figs 2BC</figureCitation>
). Embolus originating prolaterodistally from tegulum, moderately long, with wedge-shaped proximad tip resting near base of conductor (E;
<figureCitation id="95D42A61FF9EFFDE7865FE1EFA53AA52" box="[1338,1423,506,529]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2C</figureCitation>
). Conductor originating retrolatero-proximally from tegulum, unmodified, smoothly curved (C;
<figureCitation id="95D42A61FF9EFFDE79D4FDF9FADEAA77" box="[1163,1282,541,564]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Figs 2BC</figureCitation>
), with straight basal part and prolaterad apical part (uC, tC;
<figureCitation id="95D42A61FF9EFFDE7F28FDA4FD16AA14" box="[631,714,576,599]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443">Fig. 2B</figureCitation>
).
</paragraph>
<caption id="5990666CFF9EFFDE7DC8F8DEFB11AFA1" httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="2" pageNumber="443" targetBox="[212,1385,657,1829]" targetPageId="2">
<paragraph id="0D5036E4FF9EFFDE7DC8F8DEFB11AFA1" blockId="2.[151,1436,1850,2018]" pageId="2" pageNumber="443">
<emphasis id="3F9BEAF6FF9EFFDE7DC8F8DEFE9FAF13" bold="true" box="[151,323,1850,1872]" pageId="2" pageNumber="443">FIGURES 2AF.</emphasis>
<taxonomicName id="CAEF4D67FF9EFFDE7C15F8DEFE0EAF0C" authority="Sankaran &amp; Sebastian, 2017" authorityName="Sankaran &amp; Sebastian" authorityYear="2017" box="[330,466,1850,1871]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="folia" status="sp. nov.">
<emphasis id="3F9BEAF6FF9EFFDE7C15F8DEFE0EAF0C" box="[330,466,1850,1871]" italics="true" pageId="2" pageNumber="443">Anaxibia folia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24A8578DFF9EFFDE7C86F8DEFDE7AF13" box="[473,571,1850,1872]" pageId="2" pageNumber="443" rank="species">
<emphasis id="3F9BEAF6FF9EFFDE7C86F8DEFDE7AF13" bold="true" box="[473,571,1850,1872]" pageId="2" pageNumber="443">spec. nov.</emphasis>
</taxonomicNameLabel>
A Male prosoma, retrolateral. BD Male left pedipalp (B prolateral, C ventral, D dorsal). E Epigyne, ventral. F Vulva, dorsal. Abbreviations: C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynid terminal apophysis, DTA—dorsal tibial apophysis, E—embolus, FD—fertilization duct, LLMD—lateral lobe of median diverticulum, MD—median diverticulum, PP—patellar process, S—spermatheca, SHP—sclerotised horizontal plate, STsubtegulum, tC—apical part of conductor, T—tegulum, uC—basal part of conductor, VTA—ventral tibial apophysis. Scale bars: AF, 0.2 mm. Arrows: 1 indicates femoral bulging, 2 tibial bulging, 3 horizontal ridge of dictynid terminal apophysis.
</paragraph>
</caption>
<caption id="5990666CFF9FFFDF7DC8FC06FD57ABBB" box="[151,651,994,1016]" httpUri="https://zenodo.org/record/1108260/files/figure.png" pageId="3" pageNumber="444" targetBox="[163,1424,189,976]" targetPageId="3">
<paragraph id="0D5036E4FF9FFFDF7DC8FC06FD57ABBB" blockId="3.[151,651,994,1016]" box="[151,651,994,1016]" pageId="3" pageNumber="444">
<emphasis id="3F9BEAF6FF9FFFDF7DC8FC06FED0ABBB" bold="true" box="[151,268,994,1016]" pageId="3" pageNumber="444">FIGURE 3.</emphasis>
<taxonomicName id="CAEF4D67FF9FFFDF7C4DFC07FE46ABBB" authority="Sankaran &amp; Sebastian, 2017" authorityName="Sankaran &amp; Sebastian" authorityYear="2017" box="[274,410,995,1016]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="folia" status="sp. nov.">
<emphasis id="3F9BEAF6FF9FFFDF7C4DFC07FE46ABBB" box="[274,410,995,1016]" italics="true" pageId="3" pageNumber="444">Anaxibia folia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24A8578DFF9FFFDF7CFFFC06FDDEABBB" box="[416,514,994,1016]" pageId="3" pageNumber="444" rank="species">
<emphasis id="3F9BEAF6FF9FFFDF7CFFFC06FDDEABBB" bold="true" box="[416,514,994,1016]" pageId="3" pageNumber="444">spec. nov.</emphasis>
</taxonomicNameLabel>
, type locality.
</paragraph>
</caption>
<paragraph id="0D5036E4FF9FFFDF7D98FBFAFE23AE59" blockId="3.[151,1436,1054,1666]" pageId="3" pageNumber="444">
Female (
<typeStatus id="D2548846FF9FFFDF7C7CFBFAFE5FAC76" box="[291,387,1054,1077]" pageId="3" pageNumber="444" type="paratype">paratype</typeStatus>
,
<figureCitation id="95D42A61FF9FFFDF7CD1FBFAFDFEAC76" box="[398,546,1054,1077]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="3" pageNumber="444">Figs 1B, D, F</figureCitation>
). Like male except the following. Carapace, ocular area, clypeus, chelicerae, fangs, maxillae, labium, sternum, legs, spinnerets, palps straw coloured (
<figureCitation id="95D42A61FF9FFFDF7E3BFBA5FC64AC1B" box="[868,952,1089,1112]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="3" pageNumber="444">Fig. 1B</figureCitation>
). Opisthosoma lacking greyish patches (
<figureCitation id="95D42A61FF9FFFDF782FFBA5FF6BAC39" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="3" pageNumber="444">Fig. 1B</figureCitation>
). Cribellum well marked, unipartite (
<figureCitation id="95D42A61FF9FFFDF7F0EFB87FD7BAC39" box="[593,679,1123,1146]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="3" pageNumber="444">Fig. 1D</figureCitation>
). Calamistrum prominent, uniseriate (
<figureCitation id="95D42A61FF9FFFDF7919FB87FB44AC39" box="[1094,1176,1123,1146]" captionStart="FIGURES 1" captionStartId="1.[151,258,1573,1595]" captionTargetBox="[161,1428,307,1541]" captionTargetId="figure@1.[151,1436,295,1551]" captionTargetPageId="1" captionText="FIGURES 1AF. Anaxibia folia spec. nov. AB Habitus, dorsal (A male, B female). CD Posterior part showing reduced cribellum, anal tubercle and spinnerets, ventral (C male, D female). E Male left metatarsus IV showing reduced calamistrum, retrolateral. F Femaleleft metatarsus IV showing calamistrum, prolateral.Scale bars:AB 0.5 mm; CF0.2 mm." httpUri="https://zenodo.org/record/1108256/files/figure.png" pageId="3" pageNumber="444">Fig. 1F</figureCitation>
). Palpal tibiae and tarsi bear spine-like macrosetae. Body length 1.59. Prosoma length 0.60, width 0.51. Opisthosoma length 0.99, width 0.65. Eye diameters: ALE 0.03. AME 0.03. PLE 0.02. PME 0.02. Eye interdistances: AMEAME 0.03. AMEALE 0.02. AMEPME 0.02. PMEPME 0.05. PMEPLE 0.05. Clypeus height at AMEs 0.05, at ALEs 0.06. Chelicerae length 0.21. Measurements of palp and legs. Palp 0.53 [0.15, 0.08, 0.10, 0.20], I 1.88 [0.57, 0.19, 0.46, 0.41, 0.25], II 1.75 [0.54, 0.18, 0.41, 0.38, 0.24], III 1.54 [0.49, 0.17, 0.33, 0.34, 0.21], IV 1.82 [0.59, 0.19, 0.42, 0.40, 0.22]. Leg formula: 1423. Copulatory organ (
<figureCitation id="95D42A61FF9FFFDF7C3EFAD7FE0AAD09" box="[353,470,1331,1354]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Figs 2EF</figureCitation>
): Epigyne simple, hirsute, widely flat, laterally with distinct, sclerotised, smoothly curved horizontal plates (SHP;
<figureCitation id="95D42A61FF9FFFDF7CC8FAB2FE36AD2E" box="[407,490,1366,1389]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Fig. 2E</figureCitation>
). Copulatory openings crescent-like, widely separated, lying near to the posterior border of epigyne (CO;
<figureCitation id="95D42A61FF9FFFDF7C07FA9DFE72ADD3" box="[344,430,1401,1424]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Fig. 2E</figureCitation>
). Copulatory ducts short, sclerotised, basally wide, basal part remains contiguous with an enlarged sac-like median diverticulum (CD;
<figureCitation id="95D42A61FF9FFFDF7FDCFA7FFD0BADF1" box="[643,727,1435,1458]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Fig. 2F</figureCitation>
); median diverticulum roughly squarish with anterior concavity, postero-laterally with paired, broad lobes overhanging to the basal part of copulatory ducts (MD, LLMD;
<figureCitation id="95D42A61FF9FFFDF7864FA5AFA53AD96" box="[1339,1423,1470,1493]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Fig. 2F</figureCitation>
). Spermathecae small, sclerotised, circular (S;
<figureCitation id="95D42A61FF9FFFDF7FDCFA05FD09ADBB" box="[643,725,1505,1528]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Fig. 2F</figureCitation>
). Fertilization ducts narrow, angular, originating postero-laterally from spermathecae (FD;
<figureCitation id="95D42A61FF9FFFDF7CFDF9E7FE2EAE59" box="[418,498,1539,1562]" captionStart="FIGURES 2" captionStartId="2.[151,258,1850,1872]" captionTargetBox="[212,1385,657,1829]" captionTargetId="figure@2.[196,1401,637,1841]" captionTargetPageId="2" captionText="FIGURES 2AF. Anaxibia folia spec. nov. A Male prosoma, retrolateral.BD Male left pedipalp(B prolateral, C ventral, D dorsal). E Epigyne,ventral.F Vulva, dorsal.Abbreviations:C—conductor, CD—copulatory duct, CL—cymbial caudal lobe, CO—copulatory opening, dTA—dictynidterminalapophysis,DTA—dorsaltibial apophysis,E—embolus,FD—fertilization duct,LLMD—laterallobe of median diverticulum,MD—median diverticulum,PP—patellar process,S—spermatheca,SHP—sclerotised horizontal plate, STsubtegulum, tC—apicalpart of conductor,T—tegulum,uC—basal part of conductor, VTA—ventral tibial apophysis.Scale bars:AF, 0.2 mm.Arrows:1indicates femoralbulging,2 tibial bulging, 3 horizontal ridge of dictynid terminalapophysis." httpUri="https://zenodo.org/record/1108258/files/figure.png" pageId="3" pageNumber="444">Fig. 2F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="45F5656FFF9FFFDF7D98F9C2FF00AEC1" pageId="3" pageNumber="444" type="distribution">
<paragraph id="0D5036E4FF9FFFDF7D98F9C2FCF2AE7E" blockId="3.[151,1436,1054,1666]" box="[199,814,1574,1597]" pageId="3" pageNumber="444">
<emphasis id="3F9BEAF6FF9FFFDF7D98F9C2FE85AE7E" bold="true" box="[199,345,1574,1597]" pageId="3" pageNumber="444">Distribution.</emphasis>
Only known from the
<typeStatus id="D2548846FF9FFFDF7F13F9C2FDA6AE7E" box="[588,634,1574,1597]" pageId="3" pageNumber="444">type</typeStatus>
locality (
<figureCitation id="95D42A61FF9FFFDF7F80F9C2FCFCAE7E" box="[735,800,1574,1597]" captionStart="FIGURE 3" captionStartId="3.[151,244,994,1016]" captionTargetBox="[163,1424,189,976]" captionTargetId="figure@3.[151,1436,180,989]" captionTargetPageId="3" captionText="FIGURE 3. Anaxibiafolia spec. nov., type locality." httpUri="https://zenodo.org/record/1108260/files/figure.png" pageId="3" pageNumber="444">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="0D5036E4FF9FFFDF7D98F9ADFF00AEC1" blockId="3.[151,1436,1054,1666]" pageId="3" pageNumber="444">
<emphasis id="3F9BEAF6FF9FFFDF7D98F9ADFEDFAE23" bold="true" box="[199,259,1609,1632]" pageId="3" pageNumber="444">Note.</emphasis>
Males of
<taxonomicName id="CAEF4D67FF9FFFDF7C2BF9ACFE17AE23" baseAuthorityName="Tikader" baseAuthorityYear="1966" box="[372,459,1608,1632]" class="Arachnida" family="Dictynidae" genus="Anaxibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="rebai">
<emphasis id="3F9BEAF6FF9FFFDF7C2BF9ACFE17AE23" box="[372,459,1608,1632]" italics="true" pageId="3" pageNumber="444">A. rebai</emphasis>
</taxonomicName>
are misidentified and belonging to
<taxonomicName id="CAEF4D67FF9FFFDF7E0CF9ADFB43AE23" authority="(Tikader, 1966)" authorityName="Tikader" authorityYear="1966" baseAuthorityName="Tikader" baseAuthorityYear="1966" box="[851,1183,1608,1632]" class="Arachnida" family="Dictynidae" genus="Nigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="shiprai">
<emphasis id="3F9BEAF6FF9FFFDF7E0CF9ADFC32AE23" box="[851,1006,1608,1632]" italics="true" pageId="3" pageNumber="444">Nigma shiprai</emphasis>
(
<bibRefCitation id="697E4B15FF9FFFDF7EA1F9ADFB4BAE23" author="Tikader" box="[1022,1175,1609,1632]" pageId="3" pageNumber="444" refString="Tikader, B. K. (1966) Studies on some spiders of the genus Dictyna (family: Dictynidae) from India. Proceedings of the Linnean Society of London, 177, 45 - 53. https: // doi. org / 10.1111 / j. 1095 - 8312.1966. tb 00950. x" type="journal article" year="1966">Tikader, 1966</bibRefCitation>
)
</taxonomicName>
(World Spider Catalog 2017).
</paragraph>
</subSubSection>
</treatment>
</document>