338 lines
34 KiB
XML
338 lines
34 KiB
XML
<document id="E30890CC944D90736AF9A29BE44A0049" ID-CLB-Dataset="28593" ID-DOI="http://dx.doi.org/10.3897/zookeys.820.29632" ID-GBIF-Dataset="602cd784-a4d4-4417-86e3-b3417bf40916" ID-PMC="PMC6367594" ID-Pensoft-Pub="1313-2970-820-25" ID-PubMed="30745797" ID-ZooBank="7110E86F2AEC41A89A0156117B6A1556" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-820-25" ModsDocOrigin="ZooKeys 820" ModsDocTitle="The identity of the Sri Lankan Amblypharyngodon (Teleostei, Cyprinidae)" checkinTime="1548695297380" checkinUser="pensoft" docAuthor="Sudasinghe, Hiranya, Pethiyagoda, Rohan, Maduwage, Kalana & Madhava Meegaskumbura," docDate="2019" docId="84EC1C6056C96802DB6B3C6D1BF7C8EF" docLanguage="en" docName="ZooKeys 820: 25-49" docOrigin="ZooKeys 820" docSource="http://dx.doi.org/10.3897/zookeys.820.29632" docTitle="Amblypharyngodon grandisquamis Jordan & Starks 1917" docType="treatment" docVersion="6" lastPageNumber="35" masterDocId="3C177B16FFA2FFC4BF45E35483798907" masterDocTitle="The identity of the Sri Lankan Amblypharyngodon (Teleostei, Cyprinidae)" masterLastPageNumber="49" masterPageNumber="25" pageNumber="31" updateTime="1701374850711" updateUser="ExternalLinkService">
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<mods:title id="981A87E9FACDA64D559D4BE44D912422">The identity of the Sri Lankan Amblypharyngodon (Teleostei, Cyprinidae)</mods:title>
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<mods:namePart id="98EB541FB588E6A67AFE3CB6573FF629">Sudasinghe, Hiranya</mods:namePart>
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<mods:namePart id="6BCBF60062EE996ECB2B1B88361C6AEF">Pethiyagoda, Rohan</mods:namePart>
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<mods:namePart id="87BDD0A659B68D2CA5DD4D8C2A4EC1EE">Maduwage, Kalana</mods:namePart>
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<mods:namePart id="537C7742D4A476BBF83F991D6F290CCA">Madhava Meegaskumbura,</mods:namePart>
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<mods:title id="A990482325313FB0A8A71FEBAE78C1C9">ZooKeys</mods:title>
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<mods:date id="70DE1EEAF45301081AB7D406B534AA81">2019</mods:date>
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<mods:identifier id="431C8352A9E18B2F5A946DDDB405D5BA" type="DOI">http://dx.doi.org/10.3897/zookeys.820.29632</mods:identifier>
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<treatment id="84EC1C6056C96802DB6B3C6D1BF7C8EF" ID-GBIF-Taxon="154125471" LSID="urn:lsid:plazi:treatment:84EC1C6056C96802DB6B3C6D1BF7C8EF" httpUri="http://treatment.plazi.org/id/84EC1C6056C96802DB6B3C6D1BF7C8EF" lastPageId="10" lastPageNumber="35" pageId="6" pageNumber="31">
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<subSubSection id="E648A5CF40090DC110B0B1D5E8942C17" pageId="6" pageNumber="31" type="nomenclature">
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<paragraph id="25ACEAC03C4F38C1232BC11888C4DE83" pageId="6" pageNumber="31">
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<taxonomicName id="F8C411473EEF416E9C2A6341D5E7E7F2" ID-CoL="CMXW" authority="Jordan & Starks, 1917" authorityName="Jordan & Starks" authorityYear="1917" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="6" pageNumber="31" phylum="Chordata" rank="species" species="grandisquamis">
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<pageBreakToken id="96CC4D31FB1BC4FDE9009129E9B116A0" pageId="6" pageNumber="31" start="start">Amblypharyngodon</pageBreakToken>
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grandisquamis Jordan & Starks, 1917
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</taxonomicName>
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Figure 4
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</paragraph>
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</subSubSection>
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<subSubSection id="561654741C4F7BE43E4EB1B6B861526D" pageId="6" pageNumber="31" type="reference_group">
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<paragraph id="5467692BB72C568CCF60DB5557403A03" pageId="6" pageNumber="31">
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<taxonomicName id="7671FEA2C235B587E54BE8BF5A65EED5" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon melettinus" order="Cypriniformes" pageId="6" pageNumber="31" phylum="Chordata" rank="species" species="melettinus">Amblypharyngodon melettinus</taxonomicName>
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, Non-Valenciennes, 1844 (from Sri Lanka):
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<bibRefCitation id="B9CA4FD99DD1274ADD091610E16F09A7" author="Guenther, A" journalOrPublisher="British Museum, London" pageId="16" pageNumber="41" title="Catalogue of the fishes in the British Museum. Vol. 7. Catalogue of the Physostomi containing the families Heteropygii, Cyprinidae, Gonorhynchidae, Hyodontidae, Osteoglossidae, Clupeidae, Chirocentridae, Alepocephalidae, Notopteridae, Halosauridae, in the collection of the British Museum." year="1868">
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<normalizedToken id="F0A4D46A9B36E2F1A7D4515E8D33A05D" originalValue="Günther">Guenther</normalizedToken>
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1868
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</bibRefCitation>
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: 202;
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<bibRefCitation id="4952758C82B3B1259960A64CB31B5567" pageId="6" pageNumber="31">Day 1878</bibRefCitation>
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: 555;
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<bibRefCitation id="CD411E78646ED6D1576A026D5EDEEBA0" author="Day, F" journalOrPublisher="Taylor & Francis, London" pageId="16" pageNumber="41" title="The Fauna of British India, including Ceylon and Burma. Fishes Vol. 1." year="1889">Day 1889</bibRefCitation>
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: 292;
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<bibRefCitation id="19CA3D13E70F760F7582694C39719FF3" author="Duncker, G" journalOrPublisher="Mitteilungen aus dem Naturhistorischen (Zoologischen) Museum in Hamburg" pageId="16" pageNumber="41" pagination="241 - 272" title="Die Suesswasserfische Ceylons." volume="29" year="1912">Duncker 1912</bibRefCitation>
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: 265;
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<bibRefCitation id="5F6C3692A39BAD9EE2A25C32AF8B36E8" author="Deraniyagala, PEP" journalOrPublisher="National Museums of Ceylon, Colombo" pageId="16" pageNumber="41" title="A coloured atlas of some vertebrates from Ceylon, Vol. 1. Fishes." year="1952">Deraniyagala 1952</bibRefCitation>
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: 45;
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<bibRefCitation id="99A57FA0A7F99AB26DD7ABDCB8250E4D" pageId="6" pageNumber="31">Senanayake 1980</bibRefCitation>
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: 167;
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<bibRefCitation id="5C6518EC0CD09F7D16E50BD0A73ADD29" author="Pethiyagoda, R" journalOrPublisher="Wildlife Heritage Trust, Colombo, Sri Lanka" pageId="17" pageNumber="42" title="Freshwater fishes of Sri Lanka." year="1991">Pethiyagoda 1991</bibRefCitation>
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: 25;
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<bibRefCitation id="1528888906649FCD1521690965F29849" pageId="6" pageNumber="31">Pethiyagoda 2006</bibRefCitation>
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;
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<bibRefCitation id="14CD2D251559E0BC898380F35549C856" author="Menon, AGK" journalOrPublisher="Ministry of Environment, Colombo" pageId="17" pageNumber="42" title="The National Red List 2012 of Sri Lanka; conservation status of the fauna and flora." year="2012">MOE 2012</bibRefCitation>
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;
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<bibRefCitation id="F71F080F3C73787810E475C26022DE85" author="De Silva, MA" journalOrPublisher="Wildlife Conservation Society, Galle" pageId="16" pageNumber="41" title="Sri Lankan Freshwater Fishes." year="2015">De Silva et al. 2015</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3D105B1360467B1FAB785D1801D954E0" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31" type="materials_examined">
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<paragraph id="2EABB73135DD51A8AE7405FC463BA718" pageId="6" pageNumber="31">Material examined (all from Sri Lanka).</paragraph>
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<paragraph id="59B5591B6551E7E5B4F1732D07FD70F6" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31">
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SU 22868, 8 paratypes, 56.4-65.4 mm SL, river at Colombo (presumably the Kelani, which is the only river in the vicinity of the city); WHT 30275, 6, 65.0-76.4 mm SL, Bolgoda River basin, Bellanwila-Attidiya; WHT 64, 3, 45.4-47.5 mm SL, Kalu River basin, Ingiriya, Dombagaskanda; WHT 22, 2, 37.2-38.7 mm SL, Kalu River basin, Walandure, Kuruwita; WHT 225, 2, 30.2-34.3 mm SL, Kalu River basin, Ekneligoda, Kuruwita; WHT 30790, 3, 49.3-52.8 mm SL, Kelani River basin, Deraniyagala; WHT 32, 3, 49.6-60.1 mm SL, Attanagalu Oya basin, Weliweriya; WHT 30260, 58.2 mm SL, Nilwala River basin, Godapitiya, Akuressa; WHT 30167, 3, 54.2-74.7 mm SL, Malwathu Oya basin, Anuradhapura; DZ 3193, 7, 57.2-74.0 mm SL, Mahaweli River basin, Polonnaruwa; DZ 3564, 2, 57.2-62.3 mm SL, Mahaweli River basin, Badulu Oya, Kandeketiya; WHT 1694, 3,
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<pageBreakToken id="4209F090C55A9E9EC4FBC6C4F2A33F55" pageId="7" pageNumber="32" start="start">42.7</pageBreakToken>
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-45.1 mm SL, Mahaweli River basin, Wasgamuwa; WHT 45, 4, 54.6-71.9 mm SL, Mi Oya basin, Puttlam; WHT 1810, 4, 40.8-52.4 mm SL, Kirindi Oya basin, Weerawila; WHT 1851, 41.2 mm SL, Gal Oya basin, Rathugala near Bibile. Cleared and stained: WHT 11075, 64.2 mm SL, Bolgoda River basin, Bellanwila-Attidiya; WHT 11046, 54.2 mm SL, Mi Oya basin, Puttlam; WHT 11081, 45.1 mm SL, Mi Oya basin, Puttlam; WHT 11052, 45.9 mm SL, Kelani River basin, Deraniyagala; WHT 11070, 40.0 mm SL, Kelani River basin, Deraniyagala. Other material (identified for distribution data, but not measured): WHT 30789, Mahaweli River basin, Wasgamuwa; DZ 3434, Mahaweli River basin, Naula; DZ 1504, Mahaweli River basin, Ulhitiya; WHT 30263, Deduru Oya basin, Wewagama, Kuliyapitiya; WHT 30679, Deduru Oya basin; WHT 2158, Kirindi Oya basin, Tissamaharama; DZ 4423, Kirindi Oya basin, Lunugamwehera; WHT 30723, Heda Oya, Lahugala; WHT 30310, Kala Oya basin, Eluwankulam; WHT 30167, Malwathu Oya basin, Anuradhapura; DZ 4042, Attanagalu Oya basin, Yakkala; DZ 3577, Mi Oya basin, Galgamuwa; DZ 3292, Kalu River basin, Remuna.
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</paragraph>
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</subSubSection>
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<subSubSection id="18A32F8BD4CC2EB71DD99C270CDBE0BC" pageId="7" pageNumber="32" type="diagnosis">
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<paragraph id="8E60BE40D70FEA6C64860D5BD04F4280" pageId="7" pageNumber="32">Diagnosis.</paragraph>
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<paragraph id="177170F5388709A92362E09D13E12784" pageId="7" pageNumber="32">
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<taxonomicName id="3486DCA71FF3EE68D20F9B554D6A4588" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="7" pageNumber="32" phylum="Chordata" rank="species" species="grandisquamis">Amblypharyngodon grandisquamis</taxonomicName>
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is distinguished from
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<taxonomicName id="9A17776FB3FADB244FBB191E9151FDB1" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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by having a deeper body (26.9-31.2% SL in
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<taxonomicName id="5A30F2B72B94A2DED2F73C079340E293" lsidName="A. grandisquamis" pageId="7" pageNumber="32" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
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, vs. 22.9-26.3 in
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<taxonomicName id="E6760B72088606A91F9439EECB9F8267" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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); fewer pored lateral-line scales (8-16 (25), 19 (1), vs. 15(1), 17-21 (13) in
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<taxonomicName id="449D208B0D78193F7B7360AA34541D68" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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: Figure 3A); more caudal vertebrae (18-19 (5), vs. 17 (2) in
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<taxonomicName id="F44711815FA4BB7F99EBC7DE7AD82B70" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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); 2-4 minute foramina (absent in
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<taxonomicName id="AE7522AC075545DC7F488AFF17A73902" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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) in addition to a large foramen on the base of the lateral arm of the fifth ceratobranchial; and absence of a minute foramen at the base of the medial arm of the fifth ceratobranchial (present in
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<taxonomicName id="6588263DA6388DC2A35FAA9302032ACF" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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). It differs from
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<taxonomicName id="2CD968F17A82B5EDDA32F39CF70436AF" lsidName="A. microlepis" pageId="7" pageNumber="32" rank="species" species="microlepis">A. microlepis</taxonomicName>
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by having fewer scales in the lateral series (42-56 vs. 55-65 in
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<taxonomicName id="EEEA217205EF78BAC9A88C3E8B022105" lsidName="A. microlepis" pageId="7" pageNumber="32" rank="species" species="microlepis">A. microlepis</taxonomicName>
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) and a greater body depth (26.9-31.2% SL vs. 24.3-26.3 in
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<taxonomicName id="191E6172EF8DDB61F36004987C578DA5" lsidName="A. microlepis" pageId="7" pageNumber="32" rank="species" species="microlepis">A. microlepis</taxonomicName>
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); from
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<taxonomicName id="A413C77BF761792E99910ACCFE68731E" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
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by a lesser dorsal-fin height (21.1-27.6% SL vs. 27.8-29.2 in
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<taxonomicName id="1F87706D12DED5A905FB213FF413FF8A" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
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), shorter eye diameter (22.7-30.5% HL vs. 31.3-36.6 in
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<taxonomicName id="A8D8E7B7092E64B0F9F9F60F505D7DBF" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
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), fewer scales in the lateral series (42-56 vs. 69-73 in
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<taxonomicName id="F30C153C2655378A4E9A3ABA148A0A05" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
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: Figure 3B), fewer circumpeduncular scales (20-24 vs. 27-31 in
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<taxonomicName id="C77D67F5DC1E7DEA0BD21AF4A2772535" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
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: Figure 3C), and fewer scale rows between the origins of the dorsal and pelvic fins (14-17 vs. 23-25 in
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<taxonomicName id="A60EBC3F67E649E97DC5B1BF5A51754E" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
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: Figure 3D); from
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<taxonomicName id="6BF4BCE2D43845EF0F1B1BA8BA9D4283" lsidName="A. chulabhornae" pageId="7" pageNumber="32" rank="species" species="chulabhornae">A. chulabhornae</taxonomicName>
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by having more pored lateral-line scales (8-16 (25) or 19 (1) vs. 6-7 in
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<taxonomicName id="281CBDC9ED384B98EF72054179967FB6" lsidName="A. chulabhornae" pageId="7" pageNumber="32" rank="species" species="chulabhornae">A. chulabhornae</taxonomicName>
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) and more vertebrae (33-34 (5) vs. 31-32 in
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<taxonomicName id="84184F75FEEA0A85CCE0D4CD4EE06D6F" lsidName="A. chulabhornae" pageId="7" pageNumber="32" rank="species" species="chulabhornae">A. chulabhornae</taxonomicName>
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); and from
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<taxonomicName id="68AE0A1474442E272786CE1C6486263C" lsidName="A. atkinsonii" pageId="7" pageNumber="32" rank="species" species="atkinsonii">A. atkinsonii</taxonomicName>
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by having fewer scales in the lateral series (42-56 vs. 55-61 in
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<taxonomicName id="C4EE4F60AE1FB177063BCFA4A65CA390" lsidName="A. atkinsonii" pageId="7" pageNumber="32" rank="species" species="atkinsonii">A. atkinsonii</taxonomicName>
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), a lesser body depth (26.9-31.2% SL, vs. 40.5 in
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<taxonomicName id="9EE40225A8F14FAD134C87C266A60A13" lsidName="A. atkinsonii" pageId="7" pageNumber="32" rank="species" species="atkinsonii">A. atkinsonii</taxonomicName>
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), and fewer scale rows between the origins of the dorsal and pelvic fins (14-17 vs. 21 in
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<taxonomicName id="D4E0727E486240E0684C413C0F184C5F" lsidName="A. atkinsonii" pageId="7" pageNumber="32" rank="species" species="atkinsonii">A. atkinsonii</taxonomicName>
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).
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</paragraph>
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<caption id="3645FDC65FB2B19D8FFAA5ECD531733A" pageId="7" pageNumber="32">
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<paragraph id="7FFDDEFE94B245CB4C012F2845F5FDA0" pageId="7" pageNumber="32">
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Figure 3. Boxplots representing the distribution of A pored lateral-line scales B lateral series scales C circumpeduncular scales and D total transverse scales in recent material of
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<taxonomicName id="13EE1E8D8A1E508F45BE67659E1C8CBC" lsidName="A. grandisquamis" pageId="7" pageNumber="32" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
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,
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<taxonomicName id="4C68EE4B0D3EBFD25A3E1D1A7BD6A8DF" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
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, and
|
||
<taxonomicName id="2F4CBBA91BD90A4FDA1883B5CECFA654" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="0BA00F09AAE96C754D7E296F5023C54F" lastPageId="9" lastPageNumber="34" pageId="7" pageNumber="32" type="description">
|
||
<paragraph id="4C0C0F1E82B821076A605519B247119C" pageId="7" pageNumber="32">Description.</paragraph>
|
||
<paragraph id="3DF976E56E8BA638713651C43DF5A6CA" pageId="7" pageNumber="32">For general appearance, see Figure 4; morphometric data are provided in Table 5. Head and body oblong, slightly compressed. Head wider than body. Body depth greatest at dorsal-fin origin. Snout short, subequal to eye diameter, dorsally rounded, laterally subtriangular. Mouth terminal; symphysial knob present, small, elongated, fitting into shallow groove on inner margin of upper jaw with mouth closed.</paragraph>
|
||
<caption id="07AB45E7AE925E15AA73DD0C1BCCB6C5" pageId="7" pageNumber="32">
|
||
<paragraph id="F68C1D6CA20FD301F93E653B78F0BB15" pageId="7" pageNumber="32">
|
||
Table 5. Morphometric data for
|
||
<taxonomicName id="FB04693C843FC529D3511258DF83705D" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="7" pageNumber="32" phylum="Chordata" rank="species" species="grandisquamis">Amblypharyngodon grandisquamis</taxonomicName>
|
||
(SU 22868, WHT 30275, WHT 64, WHT 30790, WHT 32, WHT 30260, WHT 30167, WHT 45, DZ 3193, DZ 3564),
|
||
<taxonomicName id="1361EE0B45A38372F0ECA58FB1C84361" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
|
||
(MNHN 3812, WHT 7419, WHT 7420, WHT 30329, WHT 30787), and
|
||
<taxonomicName id="800C3D4EE1DB92AAB92CB86C4DFE7284" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
|
||
(WHT 30316).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5D0FA9CB621633B11A1DEE6E12050EEA" pageId="7" pageNumber="32">
|
||
<table id="E6EF965E46552F3D7070459685026958" pageId="7" pageNumber="32">
|
||
<tr id="8C668194CD0467D6BEB066C741C48BEF" pageId="7" pageNumber="32">
|
||
<th id="525A50C3F27EE985BECE0B71E59BD106" colspan="8" pageId="7" pageNumber="32" rowspan="1">
|
||
<taxonomicName id="83812D390A2217F435BF22D98EE3A366" lsidName="A. grandisquamis" pageId="7" pageNumber="32" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
</th>
|
||
<th id="6A4E8CB91BD82080892E2CD67DD84942" colspan="8" pageId="7" pageNumber="32" rowspan="1">
|
||
<taxonomicName id="DB51E0029C3984A8C108695F4162885D" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
|
||
</th>
|
||
<th id="54E7DE8AA69545DD5997230D96183ED0" colspan="4" pageId="7" pageNumber="32" rowspan="2">
|
||
<taxonomicName id="5BA06BCCC940549B5348B75673D8DA45" lsidName="A. mola" pageId="7" pageNumber="32" rank="species" species="mola">A. mola</taxonomicName>
|
||
(n = 5)
|
||
</th>
|
||
</tr>
|
||
<tr id="16F65E68BBA2EAD666CC9E1C776841FB" pageId="7" pageNumber="32">
|
||
<th id="58DD8C59ED5A8757FBC872EFA6A39CAC" colspan="4" pageId="7" pageNumber="32" rowspan="1">
|
||
paratypes of
|
||
<taxonomicName id="4E55F5D3816993D7E7960F642BE942E3" lsidName="A. grandisquamis" pageId="7" pageNumber="32" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
(n = 8)
|
||
</th>
|
||
<th id="D7FE42E7552DA73747D66167FC36C233" colspan="4" pageId="7" pageNumber="32" rowspan="1">
|
||
recently collected material of
|
||
<taxonomicName id="4956C8DB8E3DEDFAB766F293E7B4C074" lsidName="A. grandisquamis" pageId="7" pageNumber="32" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
(n = 23)
|
||
</th>
|
||
<th id="26A7B833ECAF277A03F197EAC239F9B7" colspan="4" pageId="7" pageNumber="32" rowspan="1">
|
||
syntypes of
|
||
<taxonomicName id="AB37ED473E0B59255BDCE66345A11430" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
|
||
(n = 5)
|
||
</th>
|
||
<th id="05E1D408A3BD3F807E333B6D2558CFC6" colspan="4" pageId="7" pageNumber="32" rowspan="1">
|
||
recently collected material of
|
||
<taxonomicName id="D67E2275B7DBE8BAABB8476AA11E2F0C" lsidName="A. melettinus" pageId="7" pageNumber="32" rank="species" species="melettinus">A. melettinus</taxonomicName>
|
||
(n = 14)
|
||
</th>
|
||
</tr>
|
||
<tr id="68F7F8FB6F3D11AE340424CF28D88AE5" pageId="7" pageNumber="32">
|
||
<th id="46154FEE2F96DBF18BA6E28987A3007E" colspan="1" pageId="7" pageNumber="32" rowspan="1">min</th>
|
||
<th id="3E0FC5B148DCA166F7D8E7245149E954" colspan="1" pageId="7" pageNumber="32" rowspan="1">max</th>
|
||
<th id="9180852F608BD6AB72869C67FA9BE843" colspan="1" pageId="7" pageNumber="32" rowspan="1">mean</th>
|
||
<th id="F1B37267139C779CF703C1E15191F095" colspan="1" pageId="7" pageNumber="32" rowspan="1">s.d.</th>
|
||
<th id="50A07DA4A4375BD613C7796D0E5B5E6A" colspan="1" pageId="7" pageNumber="32" rowspan="1">min</th>
|
||
<th id="F1A18899BF60E6479F55CC1A06E138B3" colspan="1" pageId="7" pageNumber="32" rowspan="1">max</th>
|
||
<th id="4FDCFA453F90FCD3C567645389BE29C3" colspan="1" pageId="7" pageNumber="32" rowspan="1">mean</th>
|
||
<th id="AF5E5EB111477510E13D7A70A5A8FB71" colspan="1" pageId="7" pageNumber="32" rowspan="1">s.d.</th>
|
||
<th id="6DE8AF829E946312DA2B30896250A3D4" colspan="1" pageId="7" pageNumber="32" rowspan="1">min</th>
|
||
<th id="687C225F668FF74FF755DAAA2FB82F87" colspan="1" pageId="7" pageNumber="32" rowspan="1">max</th>
|
||
<th id="0A8D9A23A5C0D8905FA4F05310F81693" colspan="1" pageId="7" pageNumber="32" rowspan="1">mean</th>
|
||
<th id="DBED17F534872701BAEBA44F4777C8D9" colspan="1" pageId="7" pageNumber="32" rowspan="1">s.d.</th>
|
||
<th id="A271C91D006F954AA9405FE283A07F21" colspan="1" pageId="7" pageNumber="32" rowspan="1">min</th>
|
||
<th id="6C428EC2C57644C5058E9E41E0029DB3" colspan="1" pageId="7" pageNumber="32" rowspan="1">max</th>
|
||
<th id="17E61C6DEC4B51BE2852E56C2BE63584" colspan="1" pageId="7" pageNumber="32" rowspan="1">mean</th>
|
||
<th id="2EFD0A69B2823CD49F95D9534A5D7101" colspan="1" pageId="7" pageNumber="32" rowspan="1">s.d.</th>
|
||
<th id="28B178F301B59E601117A64CAABDF960" colspan="1" pageId="7" pageNumber="32" rowspan="1">min</th>
|
||
<th id="68DA213E9AA0D0FD3BAA3FE91F97A18E" colspan="1" pageId="7" pageNumber="32" rowspan="1">max</th>
|
||
<th id="90D5F14D792D80174CC73115991CE3AF" colspan="1" pageId="7" pageNumber="32" rowspan="1">mean</th>
|
||
<th id="F7148F9D9677479CFF54FA8E87FAFE35" colspan="1" pageId="7" pageNumber="32" rowspan="1">s.d.</th>
|
||
</tr>
|
||
<tr id="5A5B63AC6DBED1B5389C3CB0EDAABD81" pageId="7" pageNumber="32">
|
||
<td id="25B60047E9511F98188129915F323F25" colspan="21" pageId="7" pageNumber="32" rowspan="1">As percentage of standard length</td>
|
||
</tr>
|
||
<tr id="DA1F0CB69C6AEA71B1DAFC757F603BB0" pageId="7" pageNumber="32">
|
||
<td id="72DD09B96F004959E00F7399B575A414" colspan="21" pageId="7" pageNumber="32" rowspan="1">As percentage of head length</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="D9968F59208F0964EEDC8946253E9F9C" pageId="7" pageNumber="32">
|
||
<paragraph id="BF0CE08946CB64C40958E9472DB1377B" pageId="7" pageNumber="32">
|
||
Figure 4.
|
||
<taxonomicName id="6D9DF1A7AD5405FFA51EDFA6F2EC3CE7" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="7" pageNumber="32" phylum="Chordata" rank="species" species="grandisquamis">Amblypharyngodon grandisquamis</taxonomicName>
|
||
, A in aquarium (~70 mm SL, not collected), Sri Lanka, Ulhitiya, Mahaweli River basin B in preservation (DZ 3193A, 74.0 mm SL), Sri Lanka, Polonnaruwa, Mahaweli River basin C illustration of the holotype of
|
||
<taxonomicName id="DF1C002B6789F47D8B26547A2D561FC8" lsidName="A. grandisquamis" pageId="7" pageNumber="32" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
by
|
||
<bibRefCitation id="6B2FEC5CCD6C8B79B3E0F3FFC35FC756" author="Jordan, DS" journalOrPublisher="Annals of the Carnegie Museum" pageId="16" pageNumber="41" pagination="430 - 460" title="Notes on a collection of fishes from Ceylon with descriptions of new species." volume="11" year="1917">Jordan and Starks (1917</bibRefCitation>
|
||
: pl. XLIV, FMNH 58964).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="376A620DEC81F1CC8BD1EB3E4907B8B8" lastPageId="8" lastPageNumber="33" pageId="7" pageNumber="32">
|
||
Lateral line incomplete, with 8 (1), 9 (2), 10 (4), 11 (4), 12 (6), 13 (5), 15 (1), 16 (2) or 19 (1) pored scales, 42 (1), 44 (1), 45 (4), 46 (3), 47 (4), 48 (4), 49 (2), 50 (1), 51 (1), 52 (2), 53 (2) or 56 (1) scales in lateral series plus 2-4 scales on base of caudal fin.
|
||
<pageBreakToken id="3B9BF2C1896E88D95FCA6C14557C5BDE" pageId="8" pageNumber="33" start="start">Predorsal</pageBreakToken>
|
||
scales 27 (2), 28 (3), 29 (2), 30 (2) or 32 (1). Prepelvic scales 25 (1), 26 (1), 27 (1), 28 (1), 29 (1), 30 (2), 31 (1), 33 (1) or 34 (1). Lateral scale rows between origins of dorsal and pelvic fins
|
||
<normalizedToken id="702AE6D13F6013EE787412733BDA3D4F" originalValue="½8+1+5½">1/28+1+51/2</normalizedToken>
|
||
(1),
|
||
<normalizedToken id="D42F1165BE60C6CDFBA3F34EF7E393A2" originalValue="½8">1/28</normalizedToken>
|
||
+1+6 (1),
|
||
<normalizedToken id="5C7F676A725F704CE647B9C2B274C156" originalValue="½9+1+4½">1/29+1+41/2</normalizedToken>
|
||
(2),
|
||
<normalizedToken id="F03EE7CF66C41889C7FB35DADED61581" originalValue="½9+1+5½">1/29+1+51/2</normalizedToken>
|
||
(8),
|
||
<normalizedToken id="1753BFB516C4DBE2D74478AA6017B6A3" originalValue="½9">1/29</normalizedToken>
|
||
+1+6 (1),
|
||
<normalizedToken id="B502EFB19371352CA563F654C1FB2C43" originalValue="½9+1+6½">1/29+1+61/2</normalizedToken>
|
||
(2),
|
||
<normalizedToken id="E23BF9D490BF89FEB4667763C353D930" originalValue="½10+1+4½">1/210+1+41/2</normalizedToken>
|
||
(1),
|
||
<normalizedToken id="3CA0759AFE82A618E45E567A76DAB343" originalValue="½10">1/210</normalizedToken>
|
||
+1+5 (1),
|
||
<normalizedToken id="3F3FA69B125612D5ECD9C7577F75C5C6" originalValue="½10+1+5½">1/210+1+51/2</normalizedToken>
|
||
(4), or
|
||
<normalizedToken id="79F9731CE78377B75784242A2CEE4B67" originalValue="½10+1+6½">1/210+1+61/2</normalizedToken>
|
||
(5). Circumpeduncular scales 20 (2), 21 (7), 22 (10), 23 (5) or 24 (2).
|
||
</paragraph>
|
||
<paragraph id="F3A8C84B2EBD8A6CE491C33F8B72222E" pageId="8" pageNumber="33">
|
||
Dorsal fin with two unbranched and seven branched rays, its origin just posterior to vertical through pelvic-fin origin, its distal margin straight. Anal fin with three unbranched and five branched rays, its origin slightly posterior to vertical through origin of dorsal fin, its distal margin slightly concave. Pectoral fin with a single unbranched and 11 (1), 12 (10), 13 (2), or 14 (1) branched rays, its origin anterior to posteriormost point of opercular opening, not reaching pelvic-fin origin when adpressed. Pectoral-fin axillary lobe rudimentary. Pelvic fin with one unbranched and 7 (3) or 8 (13) branched rays, its origin slightly closer to anal-fin origin than to origin of pectoral fin, its tip not reaching anal-fin origin when adpressed. Pelvic
|
||
<normalizedToken id="BCA8576E19D27102B1057E3E7E2B6CBA" originalValue="‘axillary’">'axillary'</normalizedToken>
|
||
scale present. Caudal fin with 9 + 8 (14) branched rays, forked, lobes rounded distally, upper and lower lobes subequal.
|
||
</paragraph>
|
||
<paragraph id="1F725624BA18850BC4996E6938FAC41D" pageId="9" pageNumber="34">
|
||
<pageBreakToken id="3A06490670673E060B6D7B82298A85AD" pageId="9" pageNumber="34" start="start">Vertebrae</pageBreakToken>
|
||
15 + 18 = 33 (1), 15 + 19 = 34 (4). Pharyngeal teeth 5 + 2 + 1 (4), 5 + 3 + 1 (1) (Figure 5
|
||
<normalizedToken id="8AF0332300578971E50BEEFB4DC9F05D" originalValue="A–C">A-C</normalizedToken>
|
||
). Fifth ceratobranchial with 2-4 minute foramina in addition to a large foramen at base of lateral arm; no foramen at base of medial arm (Figure 5
|
||
<normalizedToken id="F7B82B52172162AAA554F2D9BA43B2D7" originalValue="A–C">A-C</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<caption id="6684C554AEF87B9E8958BD783F74D17A" pageId="9" pageNumber="34">
|
||
<paragraph id="A7C051535E87E440A434D33A99D0F75F" pageId="9" pageNumber="34">
|
||
Figure 5. Comparison of the fifth ceratobranchial of
|
||
<normalizedToken id="90C51BAB126CD78E6179C2DF5F13C017" originalValue="A–C">A-C</normalizedToken>
|
||
<taxonomicName id="778C083E8D4A5FD04A7A3E134BDB1BAF" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="9" pageNumber="34" phylum="Chordata" rank="species" species="grandisquamis">Amblypharyngodon grandisquamis</taxonomicName>
|
||
and D
|
||
<taxonomicName id="0EF174C82E04F2AC353A244466346119" lsidName="A. melettinus" pageId="9" pageNumber="34" rank="species" species="melettinus">A. melettinus</taxonomicName>
|
||
. AWHT 11075, 64.2 mm SL BWHT 11046, 54.2 mm SL CWHT 11070, 40.0 mm SL DWHT 11072, 57.6 mm SL. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="A30647F1C33E46E2B7798FB442A37706" pageId="9" pageNumber="34" type="coloration">
|
||
<paragraph id="5B1404A976331D817AA8E53B8915133A" pageId="9" pageNumber="34">Coloration.</paragraph>
|
||
<paragraph id="702D41CF3FDCFD6D6EF7643202A5AA1D" pageId="9" pageNumber="34">
|
||
In 70% alcohol (Figure 4B), head and body silvery brown, darker dorsally, becoming lighter laterally, off-white ventrally. Head darker than body. Dusky-brown stripe 1
|
||
<normalizedToken id="5F1C8A5BFD320CBB0E71579FE4A117BE" originalValue="–1½">-11/2</normalizedToken>
|
||
scales wide on side of body, from immediately behind operculum, extending to caudal fin base, broader at middle, scales above it with prominent melanophores throughout, scales below it with scattered melanophores on margins, disappearing ventrally. Fins hyaline. Dorsal and caudal fins with scattered melanophores, more prominent on caudal fin.
|
||
</paragraph>
|
||
<paragraph id="2E5294D6B8D7905ACFB90546BE681763" pageId="9" pageNumber="34">In life (Figure 4A), head and body silvery grey to iridescent gold, lighter laterally. Scattered melanophores on side of body. A faint yellowish stripe extending from behind operculum to caudal fin base. Caudal fin yellowish, other fins mostly hyaline.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D55785B8DD8ECCB846EFA1A353FBF33" pageId="10" pageNumber="35" type="habitat">
|
||
<paragraph id="1EE69A50502D283BB60FA63E0B306DB8" pageId="10" pageNumber="35">
|
||
<pageBreakToken id="E8514F00A936075B828403781E37834B" pageId="10" pageNumber="35" start="start">Habitat</pageBreakToken>
|
||
, distribution, and natural history.
|
||
</paragraph>
|
||
<paragraph id="ACF305E195C97966C357EE63278ABA2D" pageId="10" pageNumber="35">
|
||
<taxonomicName id="0E62DD3D8E93CB841C901864744E12D6" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="10" pageNumber="35" phylum="Chordata" rank="species" species="grandisquamis">Amblypharyngodon grandisquamis</taxonomicName>
|
||
occurs in lotic habitats such as rivers and canals as well as in lentic habitats such as reservoirs and marshes. The species is recorded primarily from the lowland floodplain of Sri Lanka, in both the dry and the wet zones of the island (annual precipitation less than, and greater than, 2,000 mm, respectively), though much more frequently encountered in the dry zone (Figure 6). The highest elevation from which we recorded
|
||
<taxonomicName id="2BCF639A68DC7453D2D238B04979B845" lsidName="A. grandisquamis" pageId="10" pageNumber="35" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
was at ca 460 m a.s.l., in the Samanala reservoir on the Walawe River basin. It is a slow-swimming fish, usually encountered in large groups close to the surface. With the onset of the rains, adults are observed in rice paddies, probably migrating there to spawn. The relative abundance of
|
||
<taxonomicName id="B7E9B539511EF9DA6E9ECA7DD92DCC1C" lsidName="A. grandisquamis" pageId="10" pageNumber="35" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
, at least in the wet zone, appears to be seasonal, with more adults usually observed during the rainy season.
|
||
</paragraph>
|
||
<caption id="B3B725DEFBBB0A06607181D8365A8C68" pageId="10" pageNumber="35">
|
||
<paragraph id="11867A31B16A0BAF5C96E0B56CED06B1" pageId="10" pageNumber="35">
|
||
Figure 6. A Distribution of
|
||
<taxonomicName id="9961FF947537C66E5E0CCB6693CD9B60" family="Cyprinidae" genus="Amblypharyngodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblypharyngodon grandisquamis" order="Cypriniformes" pageId="10" pageNumber="35" phylum="Chordata" rank="species" species="grandisquamis">Amblypharyngodon grandisquamis</taxonomicName>
|
||
in Sri Lanka. Present records (green) and the type locality (red); habitat of
|
||
<taxonomicName id="B4A6F178464CE7699C6681FDDE8F7158" lsidName="A. grandisquamis" pageId="10" pageNumber="35" rank="species" species="grandisquamis">A. grandisquamis</taxonomicName>
|
||
in B Badulu Oya, Kandeketiya and C a reservoir at Galgamuwa.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |