225 lines
26 KiB
XML
225 lines
26 KiB
XML
<document id="9080726702CF0CBC9C8496BB862AE461" ID-CLB-Dataset="35587" ID-DOI="http://dx.doi.org/10.3897/zookeys.311.5408" ID-GBIF-Dataset="24d5415a-77c5-4b27-a8fc-3508a0a8aa28" ID-PMC="PMC3698554" ID-Pensoft-Pub="1313-2970-311-1" ID-PubMed="23825440" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-311-1" ModsDocOrigin="ZooKeys 311" ModsDocTitle="On the Monacha species of Lebanon (Gastropoda, Hygromiidae)" checkinTime="1451247198070" checkinUser="pensoft" docAuthor="Neubert, Eike & Bariche, Michel" docDate="2013" docId="84D92537380016A6F50915F97DEC4C73" docLanguage="en" docName="ZooKeys 311: 1-18" docOrigin="ZooKeys 311" docSource="http://dx.doi.org/10.3897/zookeys.311.5408" docTitle="Monacha (Monacha) solitudinis Bourguignat 1852" docType="treatment" docVersion="5" lastPageNumber="13" masterDocId="FFDCFF8EEC3E5769BF5B283CFF86FF88" masterDocTitle="On the Monacha species of Lebanon (Gastropoda, Hygromiidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="11" updateTime="1732941228043" updateUser="ExternalLinkService">
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<mods:title id="C24E16C0BFBC8825A1A1758C3266A587">On the Monacha species of Lebanon (Gastropoda, Hygromiidae)</mods:title>
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<mods:namePart id="F32E2E11C4894CD141399DF0623A7DEA">Neubert, Eike</mods:namePart>
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<mods:namePart id="87FE71B9604E0DA9DDB25B0AA5BDAEE6">Bariche, Michel</mods:namePart>
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<mods:title id="8CBA19C44E34D35A771647E84BFD1D1A">ZooKeys</mods:title>
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<mods:date id="16A10EDDCB21F89B11D9168B1323EFCF">2013</mods:date>
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<mods:number id="35B0CDA2A5C6BD72319EA6D6DDD63305">311</mods:number>
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<mods:url id="159CF5EC78723A264A14E1CEC163151F">http://dx.doi.org/10.3897/zookeys.311.5408</mods:url>
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<mods:classification id="031BA1B4B7ED8417F936C67BE7578280">journal article</mods:classification>
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<mods:identifier id="71550013CC168A1374764CAE275B5E6D" type="DOI">http://dx.doi.org/10.3897/zookeys.311.5408</mods:identifier>
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<mods:identifier id="C575A317D493B7DAADED74956D1E00AE" type="Pensoft-Pub">1313-2970-311-1</mods:identifier>
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<treatment id="84D92537380016A6F50915F97DEC4C73" ID-GBIF-Taxon="152045443" LSID="urn:lsid:plazi:treatment:84D92537380016A6F50915F97DEC4C73" httpUri="http://treatment.plazi.org/id/84D92537380016A6F50915F97DEC4C73" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" scope_class="Gastropoda" scope_family="Hygromiidae">
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<subSubSection id="872357DBFADC1329D54B050764B84797" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="6622AD2B8257E40DAE2BF0B2AEAA5F64" pageId="10" pageNumber="11">
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<taxonomicName id="85D1B756E646911B6D903A5BB99E3016" ID-CoL="8QGRR" LSID="http://species-id.net/wiki/Monacha_solitudinis" authority="Bourguignat, 1852" authorityName="Bourguignat" authorityYear="1852" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha (Monacha) solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis" subGenus="Monacha">Monacha (Monacha) solitudinis (Bourguignat, 1852)</taxonomicName>
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Figs 12-14, 17
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</paragraph>
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</subSubSection>
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<subSubSection id="2C4A7504D5EE26B68BAC138DBF2CF533" pageId="10" pageNumber="11" type="reference_group">
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<paragraph id="B0C4ED68A09304070CB0526F03AD2080" pageId="10" pageNumber="11">
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<taxonomicName id="63784E54B21072B82EF5222214CB3A27" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis">Helix solitudinis</taxonomicName>
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Bourguignat,
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<taxonomicName id="81BEC32295EFC26A70F90BFB6D7DF0A4" genus="Testacea" lsidName="Testacea" pageId="10" pageNumber="11" rank="genus">Testacea</taxonomicName>
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novissimae
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<normalizedToken id="7172CE9C2015EC91D7C94C6676AB7660" originalValue="…">...</normalizedToken>
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: 15 [Circa Heliopolim Syriae (Baalbek) habitat]. 1852
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</paragraph>
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<paragraph id="5AFE4E24467D9F3D5832DB0C54173AF0" pageId="10" pageNumber="11">
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<taxonomicName id="C28CED81C1DCB96C07A01546E5B20EFD" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis">Helix solitudinis</taxonomicName>
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, - L. Pfeiffer, Monographia heliceorum viventium IV: 280. 1859
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</paragraph>
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<paragraph id="E831E7DD3CFFBDF5DF9442F00A08F7D0" pageId="10" pageNumber="11">
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<taxonomicName id="AC6FDDE259A60D54DBA8417554C29820" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis">Helix solitudinis</taxonomicName>
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, - L. Pfeiffer, Monographia heliceorum viventium V: 365. 1868
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</paragraph>
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<paragraph id="C5737D8D043FDD5BA3F01E7A193B6699" pageId="10" pageNumber="11">
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<taxonomicName id="49BCF458DECDB30F865C385D35B930A9" class="Gastropoda" family="Geomitridae" genus="Helicella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helicella (Monacha) solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis" subGenus="Monacha">Helicella (Monacha) solitudinis</taxonomicName>
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, - Gude, Journal of Malacology, 9: 128. 1902
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</paragraph>
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<paragraph id="757C7D3C5DB161ACCD4EA0EF5B1730E6" pageId="10" pageNumber="11">
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<taxonomicName id="BFA2A77DFB72FBA92862FF9BD5E6C600" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Monacha) solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis" subGenus="Monacha">Helix (Monacha) solitudinis</taxonomicName>
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- Germain, Mollusques terrestres et fluviatiles de Syrie I: 156. 1921
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</paragraph>
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<paragraph id="93A4334E287962E21C716FCB1DFC1755" pageId="10" pageNumber="11">
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<taxonomicName id="44F4A14FDBBC64C8AFBD18B1B180340A" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metafruticicola solitudinis" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="solitudinis">Metafruticicola solitudinis</taxonomicName>
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, - Pallary,
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<normalizedToken id="C64CA090C67AF7CB66DCB4E67E0267E8" originalValue="Mémoires">Memoires</normalizedToken>
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<normalizedToken id="8475B11B5D6C049A7C70D60C0C3A6DB3" originalValue="présentés">presentes</normalizedToken>
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a
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<normalizedToken id="A519E73C461935EE1ADCEA08CC2295AA" originalValue="l’Institut">l'Institut</normalizedToken>
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<normalizedToken id="C95065D5F98CF2E459FCAD15381C29E8" originalValue="d’Ègypte">d'Egypte</normalizedToken>
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39: 8. 1939
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</paragraph>
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<paragraph id="112ED1D6E95E8FAEC75E7B5B006679C3" pageId="10" pageNumber="11">
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<taxonomicName id="45F7510F84472E26EAD9B41D6D574FE7" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">Monacha</taxonomicName>
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sp., - Boessneck, Zoology in the Middle East, 54: figs 5a, 42, 48. 2011
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</paragraph>
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</subSubSection>
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<subSubSection id="DF932CF38FAB157262EE2C53A91E8481" pageId="10" pageNumber="11" type="type specimens">
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<paragraph id="35297211A589340BE89A7972ED78D7AB" pageId="10" pageNumber="11">Type specimens.</paragraph>
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<paragraph id="87C7AC2AF71D0D3A594E666EDC628294" pageId="10" pageNumber="11">
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Figured syntype NHMG 16032a; second syntype NHMG 16032b; Baalbek
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<geoCoordinate id="B4EC90528A6943ADAA784D96AC212066" direction="north" orientation="latitude" precision="5" value="34.0061">34.0061°N</geoCoordinate>
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,
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<geoCoordinate id="305FC78695C235325D8EC166579E2F91" direction="east" orientation="longitude" precision="5" value="36.2122">36.2122°E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="2E8AF0BFA6CEEDB76D91E0066E0CD4EC" pageId="10" pageNumber="11" type="specimens examined">
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<paragraph id="66EA4DD2CEE0772A0509373E5A04915B" pageId="10" pageNumber="11">Specimens examined.</paragraph>
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<paragraph id="85ACB1114D1E823ACC364EA990BF0E9E" pageId="10" pageNumber="11">
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Bcharre, Jabal el Mekmel ca. 7 km SE of Bcharre, loose calcareous vegetation,
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<geoCoordinate id="539B73F2CC0D92E159ADDC98867854BC" direction="north" orientation="latitude" precision="5" value="34.2138">34.2138°N</geoCoordinate>
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,
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<geoCoordinate id="0F9DB729BE988899A40852E8C5403F31" direction="east" orientation="longitude" precision="5" value="36.0683">36.0683°E</geoCoordinate>
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, 2600 m alt, leg. U. Boessneck et al.; NMBE 508041/1, NMBE 508044/12, AUBM-MOLL, Falougha above Soha water plant,
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<geoCoordinate id="C75FF5141369135D8D47E438AF659B68" direction="north" orientation="latitude" precision="55" value="33.835">33.835°N</geoCoordinate>
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,
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<geoCoordinate id="8CCB88EC77671B10573049D7DBBD5FAC" direction="east" orientation="longitude" precision="55" value="35.756">35.756°E</geoCoordinate>
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, 1610 m alt., 17.08.2008; NMBE 508043/3, Falougha next to Soha water plant,
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<geoCoordinate id="CC6DEA09B67EA53BEB963A17397DC049" direction="north" orientation="latitude" precision="55" value="33.835">33.835°N</geoCoordinate>
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,
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<geoCoordinate id="7CCDDD7E5EB68DED596DBF37E2A9D856" direction="east" orientation="longitude" precision="55" value="35.752">35.752°E</geoCoordinate>
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, 1474 m alt., 17.08.2008; NMBE 508042/8, Falougha,
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<geoCoordinate id="268C9191933673A90D1A5EA3A8FB5413" direction="north" orientation="latitude" precision="5" value="33.8352">33.8352°N</geoCoordinate>
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,
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<geoCoordinate id="5A22C003F44AC371F8F8CD84E235A9D6" direction="east" orientation="longitude" precision="5" value="35.7561">35.7561°E</geoCoordinate>
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, 1591 m alt., 29.10.2011; NMBE 515472/1, ditto (preserved).
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</paragraph>
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</subSubSection>
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<subSubSection id="DE277EED157DEC115C60F94C5934582E" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="9A9EFEF9FAC12CD7581C5969919B3391" pageId="10" pageNumber="11">Diagnosis.</paragraph>
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<paragraph id="32B9D64C375322174CBBAF2DB1A65E07" pageId="10" pageNumber="11">shell medium sized, with a white spiral band at the periphery of the last whorl, aperture subrectangular, umbilicus open, cylindrical, penial papilla large and stout, flagellum longer than epiphallus.</paragraph>
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</subSubSection>
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<subSubSection id="7FF50933A5B1FB29F4A0554F8534A853" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
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<paragraph id="B1794A845A12042521DB0D192F710017" pageId="10" pageNumber="11">Re-description.</paragraph>
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<paragraph id="9DF2D6CC9B2E8D8EB8411D6AD57FCDF6" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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shell medium sized, spire depressed; protoconch consists of 2 smooth whorls; shell colour light brown to yellowish brown with a white spiral band at the periphery of the last whorl; teleoconch with a malleate sculpture and fine axial riblets; suture deep, simple; teleoconch with up to seven densely coiled whorls; last whorl slightly bending towards the aperture; aperture broadly subrectangular, reinforced by a
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<pageBreakToken id="7317C65BDFFF641B87A86F057885523D" pageId="11" pageNumber="12" start="start">small</pageBreakToken>
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white lip; peristomial rim sharply bounded, simple; umbilicus open, cylindrical, a triangular columellar callus is indicated.
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</paragraph>
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<paragraph id="97E7331EF76C05917B5FA33DC442CD2F" pageId="11" pageNumber="12">
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Genital organs (Figs 12a, b): The only specimen that could be investigated was quite strongly contracted; it is a species of
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<taxonomicName id="D699CF55BEF6FFAFD3D961E8D7087FD6" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Monacha</taxonomicName>
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sensu str., because the genitalia show an appendicula, but the penial retractor muscle is missing. Penis very short (0.8 mm), penial papilla large, stout with a central perforation completely filling the atrial lumen (preservation artefact?), basis of the papilla with a strong collar (Fig. 12b); epiphallus a thick-walled tube, its lumen filled with several finely crenulated pilasters (1.9 mm); flagellum short (2.1 mm), but surmounting the length of the epiphallus; appendicula (4.9 mm) branching off the atrium, subdivided in a thicker basal part, and a shorter part with a narrower lumen; two glandulae mucosae poorly ramified, inserting in the middle of the vagina; vesicle of bursa copulatrix large, hammer-like, pedunculus quite long (4.7 mm); atrial lumen with two pilasters, one of them large, knob-like, the other narrow and elongate; right ommatophoran retractor passes left to the genital organs.
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</paragraph>
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<paragraph id="233FCA110D4A26315F8B5BB84831D272" pageId="11" pageNumber="12">Measurements. Syntype figured: H = 9.2; D = 14.6; PH = 4.81; PD = 7.92. Fig. 14: H = 7.28; D = 13.31; PH = 4.34; PD = 7.32.</paragraph>
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<caption id="94201E94B5D48CE8E7FDBDD4F2062D78" pageId="11" pageNumber="12">
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<paragraph id="60C5AB78CA3C72DF481313D4441A5226" pageId="11" pageNumber="12">
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Figure 12. Anatomical drawing
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<taxonomicName id="FCB6AEB37EE08B9BE085D50E23193FAB" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha (Monacha) solitudinis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="solitudinis" subGenus="Monacha">Monacha (Monacha) solitudinis</taxonomicName>
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a NMBE 515472, situs of genital organs, total length 10.9 mm b detail of atrium and penial papilla, not to scale. Abbreviations used: ap = appendicula; at = atrium; bc = bursa copulatrix; ep = epiphallus; fl = flagellum; ped = pedunculus; pp = penial papilla; vd = vas deferens.
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</paragraph>
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</caption>
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<caption id="A2F2836A573853715F8E1A2E9B0CF1E2" pageId="11" pageNumber="12">
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<paragraph id="E715BE9A7F375BE210CE382DE91086A3" pageId="11" pageNumber="12">
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Figures 13-17.
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<taxonomicName id="21360798769BBD212A613D9BA18196A6" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Monacha</taxonomicName>
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spp. 13
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<taxonomicName id="C975495E73321997CC2A8F31F79BCEFB" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha (Monacha) solitudinis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="solitudinis" subGenus="Monacha">Monacha (Monacha) solitudinis</taxonomicName>
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. 13 syntype NHMG 16032a, Baalbek, D = 14.6 mm 14 NMBE 508041, Falougha above Soha water plant, D = 13.31 mm 15
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<taxonomicName id="184119B01F16E088A64F20C86B8B1816" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha (Monacha) bari" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="bari" subGenus="Monacha">Monacha (Monacha) bari</taxonomicName>
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, paratype NHMB 11167a, Israel, Mt. Hermon, D = 10.13 mm 16
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<taxonomicName id="8051731897D6BCCB7BEC079A63F7D76E" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha (Monacha) bari" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="bari" subGenus="Monacha">Monacha (Monacha) bari</taxonomicName>
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, NMBE 508040, below Barouk Cedar Forest, D = 10.19 mm 17
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<taxonomicName id="D74D1EAED39A5C93F6FD62D477FB18B2" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha (Monacha)" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="subGenus" subGenus="Monacha">Monacha (Monacha)</taxonomicName>
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cf. compingtae, Aammiq, D = XX mm. -
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<normalizedToken id="07FCD3E56A14FF9325BB815C2303AB23" originalValue="×">x</normalizedToken>
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4, phot. E. Bochud.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="42346ABED139FF488272D7886E96AB90" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="distribution">
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<paragraph id="3A690B3A383F76811F821CE6388E8312" pageId="11" pageNumber="12">Distribution.</paragraph>
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<paragraph id="408341024DED87026B2B55D269C8E38C" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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This species has only been recorded from two localities in Lebanon so far. The type locality Baalbek is not too far away and they were probably collected
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<pageBreakToken id="11C2A502642CD6DC48EB34C8F7122C21" pageId="12" pageNumber="13" start="start">on</pageBreakToken>
|
||
the way towards Baalbek. Both new localities are in the central chain of Mt. Lebanon at high altitudes. The habitats are characterised by coarse limestone boulders with interspersed subalpine grassland vegetation and is covered by snow during winter. In October 2011, two living animals were found actively crawling over the rocks (ca. 10°C and heavy rainfall), while in August 2008, only dead shells were encountered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75463424276DD596E7A7271D3D6648DA" pageId="12" pageNumber="13" type="differential diagnosis">
|
||
<paragraph id="F5EE791470864B7215CA048597971A53" pageId="12" pageNumber="13">Differential diagnosis.</paragraph>
|
||
<paragraph id="1DF3F0D67F817CC2199BFFFE077F5FF9" pageId="12" pageNumber="13">
|
||
This species differs from all other species in the Levante area by the shape of its umbilicus, which is cylindrical like a borehole. All other
|
||
<taxonomicName id="09340737C4590E5182BED6FFB793F1E2" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Monacha</taxonomicName>
|
||
species with an open umbilicus deeply differ by either having a keeled (
|
||
<taxonomicName id="6336480B9AB017E1A9F17C0A59FB572D" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha nummus" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="nummus">Monacha nummus</taxonomicName>
|
||
) or a smaller and hairy shell (
|
||
<taxonomicName id="25DAA2459D7D11BFD3FBB987AA0E0607" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha crispulata" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="crispulata">Monacha crispulata</taxonomicName>
|
||
).
|
||
<taxonomicName id="ADC0D9DB77A48A652CCC89F9954DDA35" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha bari" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="bari">Monacha bari</taxonomicName>
|
||
and
|
||
<taxonomicName id="F96A85875688D1360FD72280D0CD8F37" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha compingtae" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="compingtae">Monacha compingtae</taxonomicName>
|
||
are conchologically similar to
|
||
<taxonomicName id="280835566393034C5298ECBAE0174B04" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metafruticicola solitudinis" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="solitudinis">Metafruticicola solitudinis</taxonomicName>
|
||
, but these species have a narrow and almost closed umbilicus (refer to the discussion of
|
||
<taxonomicName id="8D009F0DE9AF275A8655ECEDD27B8656" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha bari" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="bari">Monacha bari</taxonomicName>
|
||
). Information on the genital organs of the two latter species is scarce, only
|
||
<bibRefCitation id="ED20ED620BB045A1660F518F3BD8DF82" pageId="12" pageNumber="13">Hausdorf (2000: 84)</bibRefCitation>
|
||
reported on a subadult specimen of
|
||
<taxonomicName id="BA928F0830B9A8F9E5AA29162DD4A4F8" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha compingtae" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="compingtae">Monacha compingtae</taxonomicName>
|
||
from southern Turkey. In this species, the penial papilla is short (large but stout in
|
||
<taxonomicName id="647622928B8FB5F297457B293691243E" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metafruticicola solitudinis" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="solitudinis">Metafruticicola solitudinis</taxonomicName>
|
||
), the epiphallus is longer than the flagellum (vice versa in
|
||
<taxonomicName id="0F3A5C2661E305015BCA421AA9CC0A98" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metafruticicola solitudinis" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="solitudinis">Metafruticicola solitudinis</taxonomicName>
|
||
), and the appendicula is much shorter than in
|
||
<taxonomicName id="4AA10204F88FACCA93B101BEB975642D" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metafruticicola solitudinis" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="solitudinis">Metafruticicola solitudinis</taxonomicName>
|
||
. The specimen figured by
|
||
<bibRefCitation id="9A815CE849CAE97944F5F88E22BD06D2" author="Boessneck, U" journalOrPublisher="Zoology in the Middle East" pageId="15" pageNumber="16" pagination="35 - 52" title="New records of freshwater and land molluscs from Lebanon (Mollusca: Gastropoda & Bivalvia)." url="10.1080/09397140.2011.10648879" volume="54" year="2011">Boessneck (2011)</bibRefCitation>
|
||
was re-investigated here; it matches the type specimen from Baalbek quite well, it is somewhat larger than the specimens from Falougha.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E72FB936EA488E2EEB3C282674907560" pageId="12" pageNumber="13" type="remark">
|
||
<paragraph id="2AA8D601C259ED7FE73AB213348F889E" pageId="12" pageNumber="13">Remark.</paragraph>
|
||
<paragraph id="EF685EE47FF32612C24245F467706AC3" pageId="12" pageNumber="13">
|
||
The correct generic affiliation of this species remained unclear until now. In his original description, Bourguignat compared it to his
|
||
<taxonomicName id="75F4CDADE6B10040BB099DF341E2B289" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix camelina" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="camelina">Helix camelina</taxonomicName>
|
||
, which is a species of the
|
||
<taxonomicName id="961D50E062D0C9D750A4469006D2AAD3" class="Gastropoda" family="Oxychilidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="family">Oxychilidae</taxonomicName>
|
||
; this text was uncritically copied by
|
||
<bibRefCitation id="B0F3940B8411DCEFC0B62EBC583813EE" pageId="12" pageNumber="13">Pfeiffer (1859)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="487086EE4599BEC6ADCEBD507F48EB5A" author="Gude, GK" journalOrPublisher="The Journal of Malacology," pageId="15" pageNumber="16" pagination="112 - 129" title="A classified list of the Helicoid land shells of Asia, IV." volume="9" year="1902">Gude (1902)</bibRefCitation>
|
||
was the first to list it under the genus
|
||
<taxonomicName id="11CE807947C203C4630668D4711F0733" class="Gastropoda" family="Hygromiidae" genus="Monacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacha" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Monacha</taxonomicName>
|
||
, but without any justification for doing so.
|
||
<bibRefCitation id="979F5A9B60E2658469C20E9694E69227" pageId="12" pageNumber="13">Germain (1921)</bibRefCitation>
|
||
, who remarked that this affiliation needs to be corroborated by an investigation of the genital organs, shared this point of view. Later,
|
||
<bibRefCitation id="C08B63E92A02B7F566BA861C905DB501" pageId="12" pageNumber="13">Pallary (1939)</bibRefCitation>
|
||
studied the type specimen and concluded that it should be classified within the genus
|
||
<taxonomicName id="CFCC44186C8CAA9273232DE59FECFACA" genus="Metafruticiola" lsidName="Metafruticiola" pageId="12" pageNumber="13" rank="genus">Metafruticiola</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |