160 lines
12 KiB
XML
160 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZBK="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt & Liew, Thor-Seng" docDate="2017" docId="845A6A0AFC29D60B42CC9B353367C57F" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus virgogravida Foon & Liew, 2017, sp. n." docType="treatment" docUuid="5A58FAD8-6020-49FE-87CB-21D245D8EEC6" docUuidSource="ZooBank" docVersion="6" lastPageNumber="49" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="47" updateTime="1668164728433" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Foon, Junn Kitt</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>692</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>81</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.692.14706</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.692.14706</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-692-1</mods:identifier>
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<mods:identifier type="ZBK">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
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<mods:identifier type="ZooBank">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="132786253" LSID="urn:lsid:zoobank.org:act:5A58FAD8-6020-49FE-87CB-21D245D8EEC6" httpUri="http://treatment.plazi.org/id/845A6A0AFC29D60B42CC9B353367C57F" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
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<subSubSection pageId="46" pageNumber="47" type="nomenclature">
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<paragraph pageId="46" pageNumber="47">
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<taxonomicName LSID="http://zoobank.org/5A58FAD8-6020-49FE-87CB-21D245D8EEC6" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus virgogravida" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="virgogravida">Alycaeus virgogravida</taxonomicName>
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<taxonomicNameLabel pageId="46" pageNumber="47">sp. n.</taxonomicNameLabel>
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Figures 30, 31S
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</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="type locality">
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<paragraph pageId="46" pageNumber="47">Type locality.</paragraph>
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<paragraph pageId="46" pageNumber="47">
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Limestone hill at east side of Pulau Dayang Bunting, off Langkawi Island, Kedah (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="6.207222">6°12'26"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="99.78445">99°47'04"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="type material">
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<paragraph pageId="46" pageNumber="47">Type material.</paragraph>
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<paragraph pageId="46" pageNumber="47">Holotype. Limestone hill at east side of Pulau Dayang Bunting, off Langkawi Island, Kedah: BOR/MOL 12978. Paratypes. Limestone hill at east side of Pulau Dayang Bunting, off Langkawi Island, Kedah: BOR/MOL 12979/1, BOR/MOL 6248/6.</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="etymology">
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<paragraph pageId="46" pageNumber="47">Etymology.</paragraph>
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<paragraph pageId="46" pageNumber="47">Latin, meaning pregnant maiden. Named after the type locality, Pulau Dayang Bunting, meaning island of the pregnant maiden.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
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<paragraph pageId="46" pageNumber="47">Description.</paragraph>
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<paragraph pageId="46" pageNumber="47">Protoconch. Smooth.</paragraph>
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<paragraph pageId="46" pageNumber="47">Shell shape. Conical. Shell height: 4.71-5.14 mm. Shell width: 4.44-4.97 mm.</paragraph>
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<paragraph pageId="46" pageNumber="47">
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Spire. Spire height: 1.68-1.77 mm. Spire width: 2.19-2.50 mm. Number of whorls: up to 4
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<normalizedToken originalValue="½">1/2</normalizedToken>
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. Spire shape: oblong conical. Whorl periphery rounded. Umbilicus open.
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</paragraph>
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<paragraph pageId="46" pageNumber="47">
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Whorl constriction. At about 4
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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whorls posterior of protoconch.
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</paragraph>
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<paragraph pageId="46" pageNumber="47">Breathing tube. Length: 0.49-0.51 mm.</paragraph>
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<paragraph pageId="46" pageNumber="47">Aperture and peristome. Aperture circular, expanded. Aperture height: 2.10-2.36 mm. Aperture width: 2.10-2.29 mm. Peristome double, not thickened, notched at suture. Peristome orientation 29-33° oblique with respect to the coiling axis.</paragraph>
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<paragraph pageId="46" pageNumber="47">Spiral lines. Distinct. Regularly spaced. Approximately 18-34 lines per 1 mm.</paragraph>
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<paragraph pageId="46" pageNumber="47">Radial ribs running anterior of breathing tube. Radial ribs pronounced, evenly spaced. Approximately 11-15 ribs per 1 mm.</paragraph>
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<paragraph pageId="46" pageNumber="47">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced and thicker than those anterior of breathing tube, evenly spaced. Approximately 14-17 ribs per 1 mm.</paragraph>
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<paragraph pageId="46" pageNumber="47">Radial ribs running posterior of breathing tube. Radial ribs pronounced, unevenly spaced throughout.</paragraph>
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<paragraph pageId="46" pageNumber="47">Operculum. Concave, rounded. Exterior calcareous layer thick, smooth and polished. Exterior nucleus punctured through, with a cup-like projection. Four concentric ridges were distributed in the tunnel within the thick, cup-like projection. Interior calcareous, with dense concentric circles of multimellae, nucleus punctured through. Operculum edge teethed, etched with regularly spaced grooves.</paragraph>
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<paragraph pageId="46" pageNumber="47">Shell colour. Whorls yellow. Peristome white.</paragraph>
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<paragraph pageId="47" pageNumber="48">
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<pageBreakToken pageId="47" pageNumber="48" start="start">Living</pageBreakToken>
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animal. Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="47" pageNumber="48" type="habitat">
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<paragraph pageId="47" pageNumber="48">Habitat and ecology.</paragraph>
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<paragraph pageId="47" pageNumber="48">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="47" pageNumber="48" type="distribution">
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<paragraph pageId="47" pageNumber="48">Distribution range.</paragraph>
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<paragraph pageId="47" pageNumber="48">Known only from the type locality Pulau Dayang Bunting, Langkawi archipelago.</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="differential diagnsosis">
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<paragraph pageId="48" pageNumber="49">
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<pageBreakToken pageId="48" pageNumber="49" start="start">Differential</pageBreakToken>
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diagnsosis.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus virgogravida" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="virgogravida">Alycaeus virgogravida</taxonomicName>
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sp. n. is similar to
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<taxonomicName lsidName="A. kapayanensis" pageId="48" pageNumber="49" rank="species" species="kapayanensis">A. kapayanensis</taxonomicName>
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in having a conical shell but is distinguished by its more expanded whorls, spacing of radial and spiral ribs, and notably, the thick, calcareous operculum with cup-shaped nucleus.
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus virgogravida" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="virgogravida">Alycaeus virgogravida</taxonomicName>
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sp. n. is distinguished from the globose-shelled
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus liratulus" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="liratulus">Alycaeus liratulus</taxonomicName>
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, the only other species in Peninsular Malaysia with cup-shaped operculum nucleus, by its conical shell.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="discussion">
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<paragraph pageId="48" pageNumber="49">Discussion.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus virgogravida" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="virgogravida">Alycaeus virgogravida</taxonomicName>
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sp. n. is the only small, conical and yellow
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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species in Peninsular Malaysia with cup-shaped operculum.
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</paragraph>
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<caption pageId="48" pageNumber="49">
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<paragraph pageId="48" pageNumber="49">
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Figure 30.
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus virgogravida" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="virgogravida">Alycaeus virgogravida</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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Shell of holotype, BOR/MOL 12978
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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Close up of shell of holotype, BOR/MOL 12978
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<normalizedToken originalValue="I–L">I-L</normalizedToken>
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Operculum of holotype, BOR/MOL 12978. Scale bars:
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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, F, G 1 mm; H,
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<normalizedToken originalValue="I–L">I-L</normalizedToken>
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0.5 mm. All photographs by Junn Kitt Foon.
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</paragraph>
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</caption>
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<caption pageId="48" pageNumber="49">
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<paragraph pageId="48" pageNumber="49">
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Figure 31. Synoptic view of the 23
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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taxa in Peninsular Malaysia. Scale bar: 5 mm. Photograph of
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus carinata" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="carinata">Alycaeus carinata</taxonomicName>
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by Thor Seng Liew. Photograph of
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus altispirus" order="Architaenioglossa" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="altispirus">Alycaeus altispirus</taxonomicName>
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by
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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Barna, courtesy of R. Janssen, SMF. All other photographs by Junn Kitt Foon.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |