473 lines
72 KiB
XML
473 lines
72 KiB
XML
<document id="9D2E2F51E65C6D176FB215E3328EF6A8" ID="10.11646/zootaxa.4250.4.9SLASH10733" ID-CLB-Dataset="33283" ID-DOI="10.11646/zootaxa.4250.4.9" ID-GBIF-Dataset="758720c0-3a9e-4ca4-9ecd-a3461bacc7aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="495328" ID-ZooBank="16DD8BB1-55CA-44AA-BCD4-170B277F875A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491465532197" checkinUser="plazi" docAuthor="Giraldes, Bruno Welter, Al-Maslamani, Ibrahim & Smyth, David" docDate="2017" docId="843E5B23A82AF068FF79FF6AFD1E7052" docLanguage="en" docName="zootaxa.4250.4.9SLASH10733.pdf" docOrigin="Zootaxa 4250 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleusia janani Giraldes, Al-Maslamani & Smyth, 2017, sp. nov." docType="treatment" docUuid="B72209CC-1992-4A98-A42F-9D8BA7EF6F2E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="393" masterDocId="7807235BA82BF06CFFEEFFFDFF94777E" masterDocTitle="A new species of leucosiid crab (Decapoda: Brachyura: Leucosiidae) from the Arabian Gulf" masterLastPageNumber="395" masterPageNumber="389" pageNumber="390" updateTime="1698438750671" updateUser="ExternalLinkService">
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<mods:mods id="0820F3F98CB0503A87DDCADAF0792CA5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="75DAC1ECE1180EC83F19CD25AFB9CC44">
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<mods:title id="79119D38B9CBA96E542AE41D11989352">A new species of leucosiid crab (Decapoda: Brachyura: Leucosiidae) from the Arabian Gulf</mods:title>
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</mods:titleInfo>
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<mods:name id="F3B8D5D23C9B46AA68D03A1609E4A9AF" type="personal">
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<mods:role id="D9C6B0C6362444CBF9DDF99784F36F7F">
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<mods:roleTerm id="A761195CEDAF5DE822F345706068FBF7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6CDD30C6CA784076855800D355A6FF0C">Giraldes, Bruno Welter</mods:namePart>
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</mods:name>
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<mods:name id="6F4B1A19356AF84C49CB97D2AF020577" type="personal">
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<mods:roleTerm id="DD8DA5E0C38EC58339CB13E97C2CAABB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58059FA14B770BD8816F90120293CB3A">Al-Maslamani, Ibrahim</mods:namePart>
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</mods:name>
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<mods:name id="C81DDE22711051B678657211FDCB7FF5" type="personal">
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<mods:roleTerm id="AC96F262E3BD72806492426A154C640C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="31360EAF2A930729AC50F3E77A0A8509">Smyth, David</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D080B6EBAC090D256828DA5AC41F363D">text</mods:typeOfResource>
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<mods:titleInfo id="47218CB4ADFC333662EEF28EF99A7347">
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<mods:title id="E9D179E5B9647DD7D4968273C762105C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="45CC4F7AE646D2714878FA67DB3DC73E">
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<mods:date id="107DA28F0E004626E365C07C7F31823C">2017</mods:date>
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<mods:number id="5BA1D5111F4E620F0A6FA953EFC55241">4250</mods:number>
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<mods:number id="141017839EB19E76662187C9EDEA5AA6">4</mods:number>
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<mods:start id="F78759A7F487188C47680E09FB4DA06F">389</mods:start>
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<mods:end id="105EC774941F63B3EEF486C420FD61CA">395</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:classification id="55B1C2D34A401B93335FFB74E1E2D6A8">journal article</mods:classification>
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<mods:identifier id="D43DF762B8E9914E4D36779C8BBD1CB4" type="CLB-Dataset">33283</mods:identifier>
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<mods:identifier id="B52F81858617397767402B6B372C75BE" type="DOI">10.11646/zootaxa.4250.4.9</mods:identifier>
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<mods:identifier id="1E071C77B83765560BC7EC8FFD6A6901" type="GBIF-Dataset">758720c0-3a9e-4ca4-9ecd-a3461bacc7aa</mods:identifier>
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<mods:identifier id="09A3F1B1E970FB3D79973212887093E8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AFAB7BD5775DF223570C9826B9099A58" type="Zenodo-Dep">495328</mods:identifier>
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<mods:identifier id="C3C22EC242DBC640F1BCEC0730C8EBA4" type="ZooBank">16DD8BB1-55CA-44AA-BCD4-170B277F875A</mods:identifier>
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</mods:mods>
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<treatment id="843E5B23A82AF068FF79FF6AFD1E7052" ID-DOI="http://doi.org/10.5281/zenodo.6025708" ID-GBIF-Taxon="128344066" ID-Zenodo-Dep="6025708" LSID="urn:lsid:zoobank.org:act:B72209CC-1992-4A98-A42F-9D8BA7EF6F2E" httpUri="http://treatment.plazi.org/id/843E5B23A82AF068FF79FF6AFD1E7052" lastPageId="4" lastPageNumber="393" pageId="1" pageNumber="390">
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<subSubSection id="448DB9BEA82AF06DFF79FF6AFE8077AD" pageId="1" pageNumber="390" type="nomenclature">
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<paragraph id="0C28EA35A82AF06DFF79FF6AFE2F77CC" blockId="1.[151,443,151,211]" box="[151,443,151,178]" pageId="1" pageNumber="390">
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<heading id="57605D59A82AF06DFF79FF6AFE2F77CC" bold="true" box="[151,443,151,178]" fontSize="11" level="1" pageId="1" pageNumber="390" reason="1">
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<emphasis id="3EE33627A82AF06DFF79FF6AFE2F77CC" bold="true" box="[151,443,151,178]" pageId="1" pageNumber="390">
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<taxonomicName id="CB9791B6A82AF06DFF79FF6AFEC277CF" ID-CoL="X2C7" box="[151,342,151,177]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
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||
<emphasis id="3EE33627A82AF06DFF79FF6AFEC277CF" bold="true" box="[151,342,151,177]" italics="true" pageId="1" pageNumber="390">Coleusia janani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="25D08B5CA82AF06DFEB3FF65FE2F77CC" box="[349,443,152,178]" pageId="1" pageNumber="390" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="0C28EA35A82AF06DFF79FF47FE8077AD" blockId="1.[151,443,151,211]" box="[151,276,186,211]" pageId="1" pageNumber="390">
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(
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||
<figureCitation id="94ACF6B0A82AF06DFF71FF47FF7E77AD" box="[159,234,186,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
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||
¯3)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="448DB9BEA82AF06DFF79FEFFFEC47537" pageId="1" pageNumber="390" type="materials_examined">
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<paragraph id="0C28EA35A82AF06DFF79FEFFFEC47537" blockId="1.[151,1437,258,2025]" pageId="1" pageNumber="390">
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||
<emphasis id="3EE33627A82AF06DFF79FEFFFE007665" bold="true" box="[151,404,258,283]" pageId="1" pageNumber="390">
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||
Type material.
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||
<emphasis id="3EE33627A82AF06DFEBDFEFEFE007665" bold="true" box="[339,404,259,283]" italics="true" pageId="1" pageNumber="390">Qatar</emphasis>
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||
</emphasis>
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||
.
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<materialsCitation id="BCFFE068A82AF06DFE4BFEFFFB55763C" ID-GBIF-Occurrence="1455705574" collectingDate="2014-11-12" collectionCode="ESCMC" country="Qatar" latitude="25.75017" location="Arabian Gulf" longLatPrecision="1" longitude="51.873497" pageId="1" pageNumber="390" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="3EE33627A82AF06DFE4BFEFFFD807665" bold="true" box="[421,532,258,283]" italics="true" pageId="1" pageNumber="390">
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||
<typeStatus id="D32C5497A82AF06DFE4BFEFFFD847665" box="[421,528,258,283]" pageId="1" pageNumber="390" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
male (cl
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||
<quantity id="CB6F47D0A82AF06DFD66FEFFFD677665" box="[648,755,258,283]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="390" unit="mm" value="13.6">13.6 mm</quantity>
|
||
),
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||
<collectionCode id="6A8672F0A82AF06DFCE4FEFEFCFD7665" box="[778,873,259,283]" pageId="1" pageNumber="390">ESCMC</collectionCode>
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||
0 0 161, 22 m, near offshore oyster bed in
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<collectingCountry id="7480AAA5A82AF06DFAB0FEFEFA087665" box="[1374,1436,259,283]" name="Qatar" pageId="1" pageNumber="390">Qatar</collectingCountry>
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||
Marine Zone within the
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||
<location id="0948BCEEA82AF06DFE5EFED4FDDE763C" LSID="urn:lsid:plazi:treatment:843E5B23A82AF068FF79FF6AFD1E7052:0948BCEEA82AF06DFE5EFED4FDDE763C" box="[432,586,297,322]" country="Qatar" latitude="25.75017" longLatPrecision="1" longitude="51.873497" name="Arabian Gulf" pageId="1" pageNumber="390">Arabian Gulf</location>
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||
,
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||
<geoCoordinate id="69A38CF2A82AF06DFDB6FED4FC94763C" box="[600,768,294,322]" direction="north" orientation="latitude" pageId="1" pageNumber="390" precision="1" value="25.75017">25o45’0.612”N</geoCoordinate>
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<geoCoordinate id="69A38CF2A82AF06DFCE4FED4FC54763C" box="[778,960,294,322]" direction="east" orientation="longitude" pageId="1" pageNumber="390" precision="1" value="51.873497">51o52’24.599”E</geoCoordinate>
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||
, grab survey,
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||
<collectingDate id="686D351DA82AF06DFB8CFED4FB55763C" box="[1122,1217,297,322]" pageId="1" pageNumber="390" value="2014-11-12">
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<date id="7829CCF5A82AF06DFB8CFED4FB55763C" box="[1122,1217,297,322]" pageId="1" pageNumber="390" value="2014-11-12">12.xi.14</date>
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</collectingDate>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="BCFFE068A82AF06DFB20FED7FCAE76F3" ID-GBIF-Occurrence="1455705576" collectingDate="2014-11-12" collectionCode="ESCMC" country="Qatar" latitude="25.75017" location="Arabian Gulf" longLatPrecision="1" longitude="51.873497" pageId="1" pageNumber="390" specimenCount="1" typeStatus="paratype">
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<emphasis id="3EE33627A82AF06DFB20FED7FAD2763F" bold="true" box="[1230,1350,298,321]" italics="true" pageId="1" pageNumber="390">
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<typeStatus id="D32C5497A82AF06DFB20FED7FAD6763F" box="[1230,1346,298,321]" pageId="1" pageNumber="390" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
female juvenile (cl
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||
<quantity id="CB6F47D0A82AF06DFECCFEB0FEEA7618" box="[290,382,333,358]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="390" unit="mm" value="9.5">9.5 mm</quantity>
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||
),
|
||
<collectionCode id="6A8672F0A82AF06DFE7AFEB3FE677618" box="[404,499,334,358]" pageId="1" pageNumber="390">ESCMC</collectionCode>
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||
0 0 162, 22 m, near offshore oyster bed in
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||
<collectingCountry id="7480AAA5A82AF06DFC31FEB3FB897618" box="[991,1053,334,358]" name="Qatar" pageId="1" pageNumber="390">Qatar</collectingCountry>
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||
Marine Zone within the
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||
<location id="0948BCEEA82AF06DFAACFEB0FF5976F3" LSID="urn:lsid:plazi:treatment:843E5B23A82AF068FF79FF6AFD1E7052:0948BCEEA82AF06DFAACFEB0FF5976F3" country="Qatar" latitude="25.75017" longLatPrecision="1" longitude="51.873497" name="Arabian Gulf" pageId="1" pageNumber="390">Arabian Gulf</location>
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||
,
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||
<geoCoordinate id="69A38CF2A82AF06DFF37FE89FE1576F3" box="[217,385,369,397]" direction="north" orientation="latitude" pageId="1" pageNumber="390" precision="1" value="25.75017">25o45’0.612”N</geoCoordinate>
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||
<geoCoordinate id="69A38CF2A82AF06DFE67FE89FDAB76F3" box="[393,575,369,397]" direction="east" orientation="longitude" pageId="1" pageNumber="390" precision="1" value="51.873497">51o52’24.599”E</geoCoordinate>
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||
, grab survey,
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||
<collectingDate id="686D351DA82AF06DFD32FE89FCAE76F3" box="[732,826,372,397]" pageId="1" pageNumber="390" value="2014-11-12">
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||
<date id="7829CCF5A82AF06DFD32FE89FCAE76F3" box="[732,826,372,397]" pageId="1" pageNumber="390" value="2014-11-12">12.xi.14</date>
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||
</collectingDate>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="BCFFE068A82AF06DFCA9FE89FDB376A4" ID-GBIF-Occurrence="1455705577" collectingDate="2014-11-11" collectionCode="ESCMC" country="Qatar" latitude="25.558699" location="Arabian Gulf" longLatPrecision="1" longitude="52.0168" pageId="1" pageNumber="390" specimenCount="1" typeStatus="paratype">
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||
female (cl
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||
<quantity id="CB6F47D0A82AF06DFC2EFE88FBBD76F3" box="[960,1065,373,397]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="390" unit="mm" value="13.1">13.1 mm</quantity>
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||
),
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||
<collectionCode id="6A8672F0A82AF06DFBD2FE88FB0F76F3" box="[1084,1179,373,397]" pageId="1" pageNumber="390">ESCMC</collectionCode>
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||
0 0 163, 27 m, offshore soft substrate peripheral to oyster bed in
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||
<collectingCountry id="7480AAA5A82AF06DFD73FE66FD4F76CD" box="[669,731,411,435]" name="Qatar" pageId="1" pageNumber="390">Qatar</collectingCountry>
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||
Marine Zone within the
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||
<location id="0948BCEEA82AF06DFBF1FE66FB2B76CA" LSID="urn:lsid:plazi:treatment:843E5B23A82AF068FF79FF6AFD1E7052:0948BCEEA82AF06DFBF1FE66FB2B76CA" box="[1055,1215,411,436]" country="Qatar" latitude="25.558699" longLatPrecision="1" longitude="52.0168" name="Arabian Gulf" pageId="1" pageNumber="390">Arabian Gulf</location>
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||
,
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<geoCoordinate id="69A38CF2A82AF06DFB3AFE66FA0876CD" box="[1236,1436,407,436]" direction="north" orientation="latitude" pageId="1" pageNumber="390" precision="1" value="25.558699">25o 33’ 31.32”N</geoCoordinate>
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||
<geoCoordinate id="69A38CF2A82AF06DFF79FE3CFEA576A4" box="[151,305,446,474]" direction="east" orientation="longitude" pageId="1" pageNumber="390" precision="1" value="52.0168">52o01’0.48”E</geoCoordinate>
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||
, grab survey,
|
||
<collectingDate id="686D351DA82AF06DFE24FE3CFDB376A4" box="[458,551,449,474]" pageId="1" pageNumber="390" value="2014-11-11">
|
||
<date id="7829CCF5A82AF06DFE24FE3CFDB376A4" box="[458,551,449,474]" pageId="1" pageNumber="390" value="2014-11-11">11.xi.14</date>
|
||
</collectingDate>
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||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="BCFFE068A82AF06DFDDDFE3CFED87537" ID-GBIF-Occurrence="1455705573" collectingDate="2014-11-13" collectionCode="OUMNH" country="Qatar" latitude="26.03083" location="Arabian Gulf" longLatPrecision="1" longitude="51.76917" pageId="1" pageNumber="390" specimenCount="1" typeStatus="paratype">
|
||
female (cl
|
||
<quantity id="CB6F47D0A82AF06DFD47FE3FFC8576A4" box="[681,785,450,474]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="390" unit="mm" value="12.8">12.8 mm</quantity>
|
||
),
|
||
<collectionCode id="6A8672F0A82AF06DFCCDFE3CFC0176A4" box="[803,917,449,474]" pageId="1" pageNumber="390">OUMNH</collectionCode>
|
||
.ZC.2016-01-112, same data; male (cl
|
||
<quantity id="CB6F47D0A82AF06DFAD7FE3FFA0976A7" box="[1337,1437,450,474]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="390" unit="mm" value="14.2">14.2 mm</quantity>
|
||
[broken triangular rostrum and lacking red circles]),
|
||
<collectionCode id="6A8672F0A82AF06DFCFFFE18FC177680" box="[785,899,485,510]" pageId="1" pageNumber="390">OUMNH</collectionCode>
|
||
.ZC.2016-01-113, 26m, offshore soft substrate peripheral to oyster bed in
|
||
<collectingCountry id="7480AAA5A82AF06DFE39FDF0FD81755B" box="[471,533,525,549]" name="Qatar" pageId="1" pageNumber="390">Qatar</collectingCountry>
|
||
Marine Zone within the
|
||
<location id="0948BCEEA82AF06DFCD1FDF1FC4F755B" LSID="urn:lsid:plazi:treatment:843E5B23A82AF068FF79FF6AFD1E7052:0948BCEEA82AF06DFCD1FDF1FC4F755B" box="[831,987,524,549]" country="Qatar" latitude="26.03083" longLatPrecision="1" longitude="51.76917" name="Arabian Gulf" pageId="1" pageNumber="390">Arabian Gulf</location>
|
||
,
|
||
<geoCoordinate id="69A38CF2A82AF06DFC04FDF1FB34755B" box="[1002,1184,521,549]" direction="north" orientation="latitude" pageId="1" pageNumber="390" precision="1" value="26.03083">26o01’50.988”N</geoCoordinate>
|
||
<geoCoordinate id="69A38CF2A82AF06DFB45FDF1FAC7755B" box="[1195,1363,521,549]" direction="east" orientation="longitude" pageId="1" pageNumber="390" precision="1" value="51.76917">51o46’9.012”E</geoCoordinate>
|
||
, trawl survey,
|
||
<collectingDate id="686D351DA82AF06DFF00FDCDFED87537" box="[238,332,560,585]" pageId="1" pageNumber="390" value="2014-11-13">
|
||
<date id="7829CCF5A82AF06DFF00FDCDFED87537" box="[238,332,560,585]" pageId="1" pageNumber="390" value="2014-11-13">13.xi.14</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="448DB9BEA82AF06EFF29FDA9FEB2768B" lastPageId="2" lastPageNumber="391" pageId="1" pageNumber="390" type="description">
|
||
<paragraph id="0C28EA35A82AF06DFF29FDA9FDD774CF" blockId="1.[151,1437,258,2025]" pageId="1" pageNumber="390">
|
||
<emphasis id="3EE33627A82AF06DFF29FDA9FECD7513" bold="true" box="[199,345,596,621]" pageId="1" pageNumber="390">Description.</emphasis>
|
||
<typeStatus id="D32C5497A82AF06DFE8DFDA8FE5D7513" box="[355,457,597,621]" pageId="1" pageNumber="390" type="holotype">
|
||
<emphasis id="3EE33627A82AF06DFE8DFDA8FE5D7513" box="[355,457,597,621]" italics="true" pageId="1" pageNumber="390">Holotype</emphasis>
|
||
</typeStatus>
|
||
. Carapace globose to rhomboidal, (
|
||
<figureCitation id="94ACF6B0A82AF06DFC87FDA9FC2E7513" box="[873,954,596,621]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="94ACF6B0A82AF06DFC25FDA8FC4D7513" box="[971,985,597,621]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">2</figureCitation>
|
||
A) 1.19 times longer than wide; dorsal surface shiny, smooth, closely punctate only in anterior central area; posteriorly slightly punctate. Large, concave margin in hepatic region, about 0.6 times as large as maximum carapace width. Antero-lateral border sharply crenulated (
|
||
<figureCitation id="94ACF6B0A82AF06DFEF3FD3DFEF975A7" box="[285,365,704,729]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
A, G). Lateral margin of carapace with sequential granules, minutes anteriorly, increasing in size medially above the thoracic sinus, ending above third pereiopod (
|
||
<figureCitation id="94ACF6B0A82AF06DFC5AFD19FB977583" box="[948,1027,740,765]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
A, D). Posterobasal margin laterally expanded, forming small wings responsible for square shape of posterior carapace (
|
||
<figureCitation id="94ACF6B0A82AF06DFBBEFCF5FB0B745F" box="[1104,1183,776,801]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="94ACF6B0A82AF06DFB41FCF4FB2A745F" box="[1199,1214,777,801]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">2</figureCitation>
|
||
H); line of granules extending to basolateral margin in line with the granules at thoracic sinus, overlapped with anterolateral line of granules between chelipeds, third walking pereiopod (
|
||
<figureCitation id="94ACF6B0A82AF06DFCFBFCADFCC97417" box="[789,861,848,873]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
D). Pterygostomial region smooth (
|
||
<figureCitation id="94ACF6B0A82AF06DFB10FCADFAC47417" box="[1278,1360,848,873]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
A, D). Thoracic sinus plain, pterygostomial invagination with shallow, subtle Y-shaped cavity; with 4 perliform granules above cheliped basis (
|
||
<figureCitation id="94ACF6B0A82AF06DFE60FC65FE4F74CF" box="[398,475,920,945]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
A, B, D).
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A82AF06DFF29FC41FBE073AF" blockId="1.[151,1437,258,2025]" pageId="1" pageNumber="390">
|
||
Frontal region of carapace produced upward, up-curved, forming central longitudinal, triangular elevation; concave dorsolaterally (
|
||
<figureCitation id="94ACF6B0A82AF06DFE46FC1DFE637487" box="[424,503,992,1017]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
A, G). Frontal margin medially unidentate, small triangular rostrum with anterior margin rounded (
|
||
<figureCitation id="94ACF6B0A82AF06DFEB7FBF9FE0E7363" box="[345,410,1028,1053]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Fig. 1</figureCitation>
|
||
). Antennular fossa and orbit in line at the upward margin; antennular fossa sealed by short antennae (
|
||
<figureCitation id="94ACF6B0A82AF06DFEE3FBD5FEC0733F" box="[269,340,1064,1089]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
G). Orbit small, with a dorsolateral spine in the outer orbital margin, up the small fossa. Eyes retractable, small eyestalk. Width of upward frontal region (between orbits) about 0.21 times shorter than total carapace width. External maxillipeds flat, concealing trapezoid buccal opening; endopod with long triangular merus, passing ventral edge of antennular cavity in frontal margin of carapace (
|
||
<figureCitation id="94ACF6B0A82AF06DFBF4FB69FBCF73D3" box="[1050,1115,1172,1197]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Fig. 1</figureCitation>
|
||
G); merus 1.27 times longer than ischium. Exopod of external maxillipeds with merus 1.6 times longer than ischium.
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A82AF06DFF29FB21FDA572D7" blockId="1.[151,1437,258,2025]" pageId="1" pageNumber="390">
|
||
Thoracic sternites, abdomen punctate. Abdomen, smooth with narrow somites 1, 2; somites 3–6 fused with 2 convex (
|
||
<figureCitation id="94ACF6B0A82AF06DFF14FAFDFED57267" box="[250,321,1280,1305]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
F), rounded protuberances proximally (
|
||
<figureCitation id="94ACF6B0A82AF06DFCE8FAFDFCD97267" box="[774,845,1280,1305]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
H); somite 6 separated from proximal segments by suture, dorsally smooth with subparallel lateral margins converging distally only near the 4/5 of its length; telson triangular, slightly elongated, 1.1 times longer than wide (
|
||
<figureCitation id="94ACF6B0A82AF06DFCF2FAB5FCCB721F" box="[796,863,1352,1377]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
F). Abdominal cavity nearly reaching buccal cavity (
|
||
<figureCitation id="94ACF6B0A82AF06DFF71FA91FF7872FB" box="[159,236,1388,1413]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
F, G). Male first pleopod (G1) with shaft coiled three times on its axis; with apical process pointed, straight, upward, not curved (
|
||
<figureCitation id="94ACF6B0A82AF06DFE91FA6DFE5872D7" box="[383,460,1424,1449]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
|
||
A, C, E).
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A82AF06DFF29FA49FAF47193" blockId="1.[151,1437,258,2025]" pageId="1" pageNumber="390">
|
||
Chelipeds with merus, carpus, chela punctate. Cheliped merus trigonal; 0.6 times as long as carapace; anterior, posterior margins lined with perliform granules that slightly diminishing in size distally in approximately 5/6 of its length; upper surface with 6 small granules grouped proximally, followed by 2 large perliform granules, distal 3/4 smooth (
|
||
<figureCitation id="94ACF6B0A82AF06DFF17F9DDFED27147" box="[249,326,1568,1593]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="94ACF6B0A82AF06DFEBAF9DCFEF77147" box="[340,355,1569,1593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">2</figureCitation>
|
||
A); lower surface, with anterior margin dense granulated proximally, decreasing distally, smooth posteriorly (
|
||
<figureCitation id="94ACF6B0A82AF06DFECFF9B9FEF37123" box="[289,359,1604,1629]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
F). Carpus smooth, without granules (
|
||
<figureCitation id="94ACF6B0A82AF06DFCF9F9B9FCF37123" box="[791,871,1604,1629]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="94ACF6B0A82AF06DFC98F9B8FC117123" box="[886,901,1605,1629]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">2</figureCitation>
|
||
A). Palm elongated; propodus 1.3 times longer than wide, its upper margin rounded, lower margin with row of small granules; dactyls almost as long as propodi (
|
||
<figureCitation id="94ACF6B0A82AF06DFF71F971FF6471DB" box="[159,240,1676,1701]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="94ACF6B0A82AF06DFEECF970FE8471DB" box="[258,272,1677,1701]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">2</figureCitation>
|
||
A, F). Walking pereiopods unequal in size, decreasing from second to fifth (
|
||
<figureCitation id="94ACF6B0A82AF06DFB7FF971FB7671DB" box="[1169,1250,1676,1701]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="94ACF6B0A82AF06DFB1DF970FA9571DB" box="[1267,1281,1677,1701]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">2</figureCitation>
|
||
A, F). Larger second pereiopod reach the cheliped carpus, 0.8 times length of carapace. Pereiopodal meri with margin lined with minute granules along upper margin. Pereiopodal carpi rounded, smooth; propodi dorsally, ventrally carinate.
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A82AF06EFF29F904FF2D77AA" blockId="1.[151,1437,258,2025]" lastBlockId="2.[151,1437,151,719]" lastPageId="2" lastPageNumber="391" pageId="1" pageNumber="390">
|
||
<emphasis id="3EE33627A82AF06DFF29F904FE07706E" box="[199,403,1785,1809]" italics="true" pageId="1" pageNumber="390">
|
||
Female
|
||
<typeStatus id="D32C5497A82AF06DFECBF904FE07706E" box="[293,403,1785,1808]" pageId="1" pageNumber="390" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
(Figs. I-L): Shorter front 0.19 times width of carapace. Third maxilliped (
|
||
<figureCitation id="94ACF6B0A82AF06DFB1AF905FAAD706F" box="[1268,1337,1784,1809]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">Fig. 2</figureCitation>
|
||
J) setose along endopod, margin of exopod merus; keel-like ischium of endopod, longitudinal convex (not flattened) elevation. Chelipeds proportionally short (
|
||
<figureCitation id="94ACF6B0A82AF06DFD98F8BDFD2F7027" box="[630,699,1856,1881]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">
|
||
Fig.
|
||
<date id="7829CCF5A82AF06DFD45F8BCFD2F7027" box="[683,699,1857,1881]" pageId="1" pageNumber="390">2</date>
|
||
</figureCitation>
|
||
I), with merus 0.47 times the length of carapace, shorter propodi. Pereiopods shorter than male pereiopods, with longer second pereiopod 0.7 times length of carapace. Abdomen with somites 3¯6 fused, greatly enlarged; thick, convex, shield-like shape covering entire thorax; suture separating segment 6 absent; segment 1, 2, telson free (
|
||
<figureCitation id="94ACF6B0A82AF06DFD57F851FC9770BB" box="[697,771,1964,1989]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">
|
||
Fig.
|
||
<date id="7829CCF5A82AF06DFD1DF850FC9770BB" box="[755,771,1965,1989]" pageId="1" pageNumber="390">2</date>
|
||
</figureCitation>
|
||
I). Abdominal cavity reaching buccal cavity with telson overlapping part of endopod of third maxilliped (
|
||
<figureCitation id="94ACF6B0A82AF06DFD3FF82DFCB57097" box="[721,801,2000,2025]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="1" pageNumber="390">
|
||
Figs.
|
||
<date id="7829CCF5A82AF06DFCFDF82CFCB57097" box="[787,801,2001,2025]" pageId="1" pageNumber="390">2</date>
|
||
</figureCitation>
|
||
I, J, K). Gonopore (vagina) semicircular, in sternite 5; strong inner edge, rounded and with deeper groove; outer edge more straight with longitudinal fissure (
|
||
<figureCitation id="94ACF6B0A829F06EFADCFF6AFA1577CE" box="[1330,1409,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1834,1856]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@3.[151,1436,193,1813]" captionTargetPageId="3" captionText="FIGURE 2. Coleusia janani n. sp., holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E ̅ G, I), BWG (C, D, H, J, K, L)." httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="2" pageNumber="391">Figs. 2</figureCitation>
|
||
K, L).
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A829F06EFF29FF22FEB2768B" blockId="2.[151,1437,151,719]" pageId="2" pageNumber="391">
|
||
<emphasis id="3EE33627A829F06EFF29FF22FEFF7786" bold="true" box="[199,363,223,248]" pageId="2" pageNumber="391">Colour in life</emphasis>
|
||
(
|
||
<figureCitation id="94ACF6B0A829F06EFE90FF22FE567786" box="[382,450,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="2" pageNumber="391">Fig. 1</figureCitation>
|
||
). Carapace with bright orange/greyish background. Pair of large ocelli on the gastric region on each side of the carapace; with small white centres and lined by thin red outer rings overlapped with an inclined 8-shape. Intestinal region with reddish blotches on either side and without lined margins. Carapace with three bright orange markings, two anterolateral and one posterolateral, on lateral margins. Chelipeds and walking pereiopods with orange transversal bands. Meri of chelipeds with white granules and large proximal orange band covering 4/5 of its length; orange articulations in the carpi; propodus with light orange band proximally; dactyls with white tips and bright red band on the proximal region. Walking pereiopod white with one bright orange band per segment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="448DB9BEA829F06EFF29FE02FDEF7566" box="[199,635,511,536]" pageId="2" pageNumber="391" type="materials_examined">
|
||
<paragraph id="0C28EA35A829F06EFF29FE02FDEF7566" blockId="2.[151,1437,151,719]" box="[199,635,511,536]" pageId="2" pageNumber="391">
|
||
<materialsCitation id="BCFFE068A829F06EFF29FE02FDEF7566" ID-GBIF-Occurrence="1455705575" box="[199,635,511,536]" country="Qatar" location="Arabian Gulf" pageId="2" pageNumber="391" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="3EE33627A829F06EFF29FE02FEF17566" bold="true" box="[199,357,511,536]" pageId="2" pageNumber="391">
|
||
<typeStatus id="D32C5497A829F06EFF29FE02FE967566" box="[199,258,511,536]" pageId="2" pageNumber="391">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="0948BCEEA829F06EFE82FE02FD907566" LSID="urn:lsid:plazi:treatment:843E5B23A82AF068FF79FF6AFD1E7052:0948BCEEA829F06EFE82FE02FD907566" box="[364,516,511,536]" country="Qatar" name="Arabian Gulf" pageId="2" pageNumber="391">Arabian Gulf</location>
|
||
; off
|
||
<collectingCountry id="7480AAA5A829F06EFDD8FDFDFDE27566" box="[566,630,512,536]" name="Qatar" pageId="2" pageNumber="391">Qatar</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="448DB9BEA829F06EFF29FDD9FEC6751E" pageId="2" pageNumber="391" type="etymology">
|
||
<paragraph id="0C28EA35A829F06EFF29FDD9FEC6751E" blockId="2.[151,1437,151,719]" pageId="2" pageNumber="391">
|
||
<emphasis id="3EE33627A829F06EFF29FDD9FEDA7543" bold="true" box="[199,334,548,573]" pageId="2" pageNumber="391">Etymology.</emphasis>
|
||
In recognition of the
|
||
<collectingCountry id="7480AAA5A829F06EFDBBFDD9FD077542" box="[597,659,548,572]" name="Qatar" pageId="2" pageNumber="391">Qatar</collectingCountry>
|
||
University Oceanographic RV
|
||
<emphasis id="3EE33627A829F06EFBEDFDD8FBD37542" box="[1027,1095,549,572]" italics="true" pageId="2" pageNumber="391">Janan</emphasis>
|
||
used in the collection of the present material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="448DB9BEA829F06EFF29FD91FA9A75B1" pageId="2" pageNumber="391" type="distribution">
|
||
<paragraph id="0C28EA35A829F06EFF29FD91FCC975FB" blockId="2.[151,1437,151,719]" box="[199,861,620,645]" pageId="2" pageNumber="391">
|
||
<emphasis id="3EE33627A829F06EFF29FD91FEF475FB" bold="true" box="[199,352,620,645]" pageId="2" pageNumber="391">Distribution.</emphasis>
|
||
Currently only known from the
|
||
<typeStatus id="D32C5497A829F06EFD22FD90FD6875FB" box="[716,764,621,645]" pageId="2" pageNumber="391">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A829F06EFF29FD72FA9A75B1" blockId="2.[151,1437,151,719]" pageId="2" pageNumber="391">
|
||
<emphasis id="3EE33627A829F06EFF29FD72FEBF75D6" bold="true" box="[199,299,655,680]" pageId="2" pageNumber="391">Ecology.</emphasis>
|
||
Offshore zones with sandy mixed substrates of gravel and mud; depth between 22 and
|
||
<specimenCount id="1A9121BCA829F06EFB19FD6DFAA375D9" box="[1271,1335,656,680]" pageId="2" pageNumber="391" type="generic">
|
||
<quantity id="CB6F47D0A829F06EFB19FD6DFAA775D9" box="[1271,1331,656,680]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="391" unit="m" value="27.0">27 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Material collected with water temperature that varied between 27 and 28o C and salinity between 39 and 41 ppt.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="58E8BABDA829F06EFF79F84BFEBF7095" httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="2" pageNumber="391" targetBox="[151,1437,738,1950]" targetPageId="2">
|
||
<paragraph id="0C28EA35A829F06EFF79F84BFEBF7095" blockId="2.[151,1436,1974,2027]" pageId="2" pageNumber="391">
|
||
<emphasis id="3EE33627A829F06EFF79F84BFE8270B5" bold="true" box="[151,278,1974,1996]" pageId="2" pageNumber="391">FIGURE 1.</emphasis>
|
||
<taxonomicName id="CB9791B6A829F06EFEF1F84BFE5070B2" box="[287,452,1974,1996]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
|
||
<emphasis id="3EE33627A829F06EFEF1F84BFE5070B2" box="[287,452,1974,1996]" italics="true" pageId="2" pageNumber="391">Coleusia janani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="25D08B5CA829F06EFE23F84AFD9D70B2" box="[461,521,1975,1996]" pageId="2" pageNumber="391" rank="species">
|
||
<emphasis id="3EE33627A829F06EFE23F84AFD9D70B2" bold="true" box="[461,521,1975,1996]" pageId="2" pageNumber="391">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="58E8BABDA828F06FFF79F8D7FC2170C5" httpUri="https://zenodo.org/record/495330/files/figure.png" pageId="3" pageNumber="392" targetBox="[151,1436,193,1813]" targetPageId="3">
|
||
<paragraph id="0C28EA35A828F06FFF79F8D7FC2170C5" blockId="3.[151,1436,1834,1979]" pageId="3" pageNumber="392">
|
||
<emphasis id="3EE33627A828F06FFF79F8D7FE867041" bold="true" box="[151,274,1834,1856]" pageId="3" pageNumber="392">FIGURE 2.</emphasis>
|
||
<taxonomicName id="CB9791B6A828F06FFEF6F8D6FE2D703E" box="[280,441,1835,1856]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
|
||
<emphasis id="3EE33627A828F06FFEF6F8D6FE2D703E" box="[280,441,1835,1856]" italics="true" pageId="3" pageNumber="392">Coleusia janani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="25D08B5CA828F06FFE50F8D6FE63703E" box="[446,503,1835,1856]" pageId="3" pageNumber="392" rank="species">
|
||
<emphasis id="3EE33627A828F06FFE50F8D6FE63703E" bold="true" box="[446,503,1835,1856]" pageId="3" pageNumber="392">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (carapace length 13.6 mm; ESCMC 00161), A, lateral view; B, details of the thoracic sinus; C, apical detail of male first pleopod; D, thoraxic sinus and pterygostomian region; E, abdomen and both male first pleopods; F, ventral view; G, frontal region and external maxiliped; H, posterior region. Paratype female (carapace length 13.1 mm; ESCMC 00163), I, ventral view; J, external maxilliped; K, opened abdomen with sternites and gonopores; L, right gonopore. Photos: Najat H. Al-Omary (A, B, E̅G, I), BWG (C, D, H, J, K, L).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="58E8BABDA82FF068FF79FBB5FC3C7320" box="[151,936,1096,1118]" httpUri="https://zenodo.org/record/495331/files/figure.png" pageId="4" pageNumber="393" targetBox="[151,1436,185,1083]" targetPageId="4">
|
||
<paragraph id="0C28EA35A82FF068FF79FBB5FC3C7320" blockId="4.[151,936,1096,1118]" box="[151,936,1096,1118]" pageId="4" pageNumber="393">
|
||
<emphasis id="3EE33627A82FF068FF79FBB5FE867323" bold="true" box="[151,274,1096,1118]" pageId="4" pageNumber="393">FIGURE 3.</emphasis>
|
||
Geographical distribution of the species of
|
||
<taxonomicName id="CB9791B6A82FF068FD20FBB4FC307320" authority="Galil, 2006" authorityName="Galil" authorityYear="2006" box="[718,932,1096,1118]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EE33627A82FF068FD20FBB4FCBC7320" box="[718,808,1097,1118]" italics="true" pageId="4" pageNumber="393">Coleusia</emphasis>
|
||
Galil, 2006
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="448DB9BEA82FF068FF29FB76FD1E7052" pageId="4" pageNumber="393" type="discussion">
|
||
<paragraph id="0C28EA35A82FF068FF29FB76FC457299" blockId="4.[151,1436,1163,1836]" pageId="4" pageNumber="393">
|
||
<emphasis id="3EE33627A82FF068FF29FB76FEAF73DA" bold="true" box="[199,315,1163,1188]" pageId="4" pageNumber="393">Remarks.</emphasis>
|
||
The present study increases to seven the number of species within the genus
|
||
<taxonomicName id="CB9791B6A82FF068FB46FB76FA9873DD" box="[1192,1292,1163,1187]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EE33627A82FF068FB46FB76FA9873DD" box="[1192,1292,1163,1187]" italics="true" pageId="4" pageNumber="393">Coleusia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="680697C4A82FF068FAF5FB76FA0373DA" author="Galil" box="[1307,1431,1163,1188]" pageId="4" pageNumber="393" refString="Galil, B. S. (2006) Contributions to the knowledge of Leucosiidae V. Coleusia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 4, 55 - 69." type="journal article" year="2006">Galil 2006</bibRefCitation>
|
||
; Ng
|
||
<emphasis id="3EE33627A82FF068FF2EFB4DFF6C73B9" box="[192,248,1199,1223]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2008, 2014;
|
||
<bibRefCitation id="680697C4A82FF068FE64FB52FDED73B9" author="Promdam" box="[394,633,1199,1224]" pageId="4" pageNumber="393" refString="Promdam, R., Nabhitabhata, J. & Galil, B. S. (2014) A new species of Coleusia Galil, 2006 (Decapoda: Brachyura: Leucosiidae) from southern Asia. Zootaxa, 3786 (2), 135 - 140." type="journal article" year="2014">
|
||
Promdam
|
||
<emphasis id="3EE33627A82FF068FE10FB4DFDA273B9" box="[510,566,1199,1223]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="CB9791B6A82FF068FD65FB52FCA873B9" box="[651,828,1199,1223]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
|
||
<emphasis id="3EE33627A82FF068FD65FB52FCA873B9" box="[651,828,1199,1223]" italics="true" pageId="4" pageNumber="393">Coleusia janani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="25D08B5CA82FF068FCAAFB52FC1673B6" box="[836,898,1199,1224]" pageId="4" pageNumber="393" rank="species">
|
||
<emphasis id="3EE33627A82FF068FCAAFB52FC1673B6" bold="true" box="[836,898,1199,1224]" pageId="4" pageNumber="393">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from all other species in the genus in the apical shape of the male G1 (
|
||
<figureCitation id="94ACF6B0A82FF068FE0DFB2EFDB97392" box="[483,557,1235,1260]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1996]" captionTargetBox="[151,1437,738,1950]" captionTargetId="figure@2.[151,1437,738,1950]" captionTargetPageId="2" captionText="FIGURE 1. Coleusia janani n. sp., holotype male (carapace length 13.6 mm) (ESCMC 00161). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/495329/files/figure.png" pageId="4" pageNumber="393">Figs. 1</figureCitation>
|
||
F, 2E), which is pointed, straight, and not curved (
|
||
<bibRefCitation id="680697C4A82FF068FB87FB2EFB717392" author="Galil" box="[1129,1253,1235,1260]" pageId="4" pageNumber="393" refString="Galil, B. S. (2006) Contributions to the knowledge of Leucosiidae V. Coleusia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 4, 55 - 69." type="journal article" year="2006">Galil 2006</bibRefCitation>
|
||
; Ng
|
||
<emphasis id="3EE33627A82FF068FAF2FB29FAC07395" box="[1308,1364,1235,1259]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2014;
|
||
<bibRefCitation id="680697C4A82FF068FF79FB0AFE187271" author="Promdam" box="[151,396,1271,1296]" pageId="4" pageNumber="393" refString="Promdam, R., Nabhitabhata, J. & Galil, B. S. (2014) A new species of Coleusia Galil, 2006 (Decapoda: Brachyura: Leucosiidae) from southern Asia. Zootaxa, 3786 (2), 135 - 140." type="journal article" year="2014">
|
||
Promdam
|
||
<emphasis id="3EE33627A82FF068FEE2FB05FED37271" box="[268,327,1271,1295]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="CB9791B6A82FF068FE4EFB0AFDC07271" box="[416,596,1271,1295]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
|
||
<emphasis id="3EE33627A82FF068FE4EFB0AFDC07271" box="[416,596,1271,1295]" italics="true" pageId="4" pageNumber="393">Coleusia janani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="25D08B5CA82FF068FDB3FB0AFD0A726E" box="[605,670,1271,1296]" pageId="4" pageNumber="393" rank="species">
|
||
<emphasis id="3EE33627A82FF068FDB3FB0AFD0A726E" bold="true" box="[605,670,1271,1296]" pageId="4" pageNumber="393">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is otherwise similar to
|
||
<taxonomicName id="CB9791B6A82FF068FC43FB05FBDE7271" box="[941,1098,1271,1295]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FC43FB05FBDE7271" box="[941,1098,1271,1295]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
in some features, such as the thoracic sinus with a row of four perliform granules, and in the colour pattern, with a pair of large orange ocelli on the gastric region, with very broad red outer rings.
|
||
<taxonomicName id="CB9791B6A82FF068FD35FAC2FC197229" box="[731,909,1343,1367]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
|
||
<emphasis id="3EE33627A82FF068FD35FAC2FC197229" box="[731,909,1343,1367]" italics="true" pageId="4" pageNumber="393">Coleusia janani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3EE33627A82FF068FC78FAC2FC497226" bold="true" box="[918,989,1343,1368]" pageId="4" pageNumber="393">
|
||
<taxonomicNameLabel id="25D08B5CA82FF068FC78FAC2FC4C7226" box="[918,984,1343,1368]" pageId="4" pageNumber="393" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
however, differs from
|
||
<taxonomicName id="CB9791B6A82FF068FB0DFABDFAEA7229" box="[1251,1406,1343,1367]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FB0DFABDFAEA7229" box="[1251,1406,1343,1367]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
in the straight upward apical shape of the G1, being twisted and curved outwards in the latter; the palm of
|
||
<taxonomicName id="CB9791B6A82FF068FADDFA99FA087205" box="[1331,1436,1380,1403]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="janani">
|
||
<emphasis id="3EE33627A82FF068FADDFA99FA087205" box="[1331,1436,1380,1403]" italics="true" pageId="4" pageNumber="393">C. janani</emphasis>
|
||
</taxonomicName>
|
||
being less elongated (1.3 times as long as wide in contrast to
|
||
<quantity id="CB6F47D0A82FF068FCAEFA7AFC1472DE" box="[832,896,1415,1440]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="4" pageNumber="393" unit="in" value="1.4">1.4 in</quantity>
|
||
<taxonomicName id="CB9791B6A82FF068FC68FA75FB8B72E1" box="[902,1055,1415,1439]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FC68FA75FB8B72E1" box="[902,1055,1415,1439]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
); the carapace being only densely punctate in proximal regions of
|
||
<taxonomicName id="CB9791B6A82FF068FDEFFA51FDFD72BD" box="[513,617,1452,1475]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="janani" status="sp. nov.">
|
||
<emphasis id="3EE33627A82FF068FDEFFA51FDFD72BD" box="[513,617,1452,1475]" italics="true" pageId="4" pageNumber="393">C. janani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="25D08B5CA82FF068FD9CFA56FD2572BA" box="[626,689,1451,1476]" pageId="4" pageNumber="393" rank="species">
|
||
<emphasis id="3EE33627A82FF068FD9CFA56FD2572BA" bold="true" box="[626,689,1451,1476]" pageId="4" pageNumber="393">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and only being sparsely punctate in
|
||
<taxonomicName id="CB9791B6A82FF068FBBEFA51FB7F72BD" box="[1104,1259,1451,1475]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FBBEFA51FB7F72BD" box="[1104,1259,1451,1475]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
; and in thin six reddish spots, which are more anteriorly positioned than in C.
|
||
<taxonomicName id="CB9791B6A82FF068FCBFFA32FC5E7299" box="[849,970,1487,1511]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FCBFFA32FC5E7299" box="[849,970,1487,1511]" italics="true" pageId="4" pageNumber="393">biannulata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="0C28EA35A82FF068FF29FA0EFD1E7052" blockId="4.[151,1436,1163,1836]" pageId="4" pageNumber="393">
|
||
The physicochemical and biological parameters in the waters of the eastern coast of the Arabian Gulf are similar to those found in the Gulf of
|
||
<collectingCountry id="7480AAA5A82FF068FDD9F9EAFDE87151" box="[567,636,1559,1583]" name="Oman" pageId="4" pageNumber="393">Oman</collectingCountry>
|
||
and the Pakistani coast where
|
||
<taxonomicName id="CB9791B6A82FF068FC34F9E5FBE07151" box="[986,1140,1559,1583]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FC34F9E5FBE07151" box="[986,1140,1559,1583]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
occurs (
|
||
<figureCitation id="94ACF6B0A82FF068FB3BF9EAFA8F7151" box="[1237,1307,1559,1584]" captionStart="FIGURE 3" captionStartId="4.[151,250,1096,1118]" captionTargetBox="[151,1436,185,1083]" captionTargetId="figure@4.[151,1436,185,1083]" captionTargetPageId="4" captionText="FIGURE 3. Geographical distribution of the species of Coleusia Galil, 2006." httpUri="https://zenodo.org/record/495331/files/figure.png" pageId="4" pageNumber="393">Fig. 3</figureCitation>
|
||
). There are nevertheless considerable differences in the waters of the western Arabian coast (
|
||
<bibRefCitation id="680697C4A82FF068FBCFF9C6FB7D712A" author="Bosch" box="[1057,1257,1595,1620]" pageId="4" pageNumber="393" refString="Bosch, D. T., Dance, S. P., Moolenbeek, R. G. & Oliver, P. G. (2008) Seashells of Eastern Arabia. Motivate Publishing, London, 296 pp." type="book" year="2008">
|
||
Bosch
|
||
<emphasis id="3EE33627A82FF068FB9EF9C1FB3C712D" box="[1136,1192,1595,1619]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="680697C4A82FF068FB1BF9C6FF477109" author="Riegl" pageId="4" pageNumber="393" refString="Riegl, B. & Purkis, S. (2012) Coral reefs of the Gulf: adaptation to climatic extremes in the world's hottest sea. Lol. 3. Springer Netherlands. Available from: http: // link. springer. com / chapter / 10.1007 / 978 - 94 - 007 - 3008 - 3 _ 1 (accessed 30 March 2015)" type="journal article" year="2012">Riegl & Purkis 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="680697C4A82FF068FF31F9A2FE427106" author="Al-Ansari" box="[223,470,1631,1656]" pageId="4" pageNumber="393" refString="Al-Ansari, E. M. A. S., Rowe, G., Abdel-Moati, M. A. R., Yigiterhan, O., Al-Maslamani, I., Al-Yafei, M. A., Al-Shaikh, I. & Upstill-Goddard, R. (2015) Hypoxia in the central Arabian Gulf Exclusive Economic Zone (EEZ) of Qatar during summer season. Estuarine, Coastal and Shelf Science, 159, 60 - 68." type="journal article" year="2015">
|
||
Al-Ansari
|
||
<emphasis id="3EE33627A82FF068FEB6F99DFE067109" box="[344,402,1631,1655]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
; Al-Maslamani
|
||
<emphasis id="3EE33627A82FF068FD60F99DFD537109" box="[654,711,1631,1655]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
|
||
2015). It is quite possible that
|
||
<taxonomicName id="CB9791B6A82FF068FBC1F99DFB5F7109" box="[1071,1227,1631,1655]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FBC1F99DFB5F7109" box="[1071,1227,1631,1655]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CB9791B6A82FF068FAEBF99DFAE47109" box="[1285,1392,1632,1655]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="janani">
|
||
<emphasis id="3EE33627A82FF068FAEBF99DFAE47109" box="[1285,1392,1632,1655]" italics="true" pageId="4" pageNumber="393">C. janani</emphasis>
|
||
</taxonomicName>
|
||
cooccur in some regions of the Arabian Gulf where environmental conditions overlap. Conversely the dissimilarities in temperature and salinities between the east and west coast are certainly extreme enough to create a biogeographic barrier could be isolating the populations and thereby maintaining vicariance. For this reason, we do not dismiss the presence of
|
||
<taxonomicName id="CB9791B6A82FF068FE20F90DFDFC7079" box="[462,616,1775,1799]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="biannulata">
|
||
<emphasis id="3EE33627A82FF068FE20F90DFDFC7079" box="[462,616,1775,1799]" italics="true" pageId="4" pageNumber="393">C. biannulata</emphasis>
|
||
</taxonomicName>
|
||
in the Arabian Gulf as reported by Alcock (1896, as
|
||
<taxonomicName id="CB9791B6A82FF068FB53F90DFDEA7052" authority="A. Mine Edwards, 1874" authorityName="A. Mine Edwards" authorityYear="1874" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="variety" species="longifrons" variety="neocaledonica">
|
||
<emphasis id="3EE33627A82FF068FB53F90DFEF27055" italics="true" pageId="4" pageNumber="393">Leucosia longifrons var neocaledonica</emphasis>
|
||
A. Mine Edwards, 1874
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |