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<document id="13763030DEDB18C2D0688F54E05E12EC" ID-CLB-Dataset="5017" ID-DOI="10.1649/0010-065X-72.4.717" ID-GBIF-Dataset="5d03d026-6f4b-4ec9-a01f-14b0c30e4ac8" ID-ISSN="1938-4394" ID-Zenodo-Dep="4789739" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621946783060" checkinUser="carolina" docAuthor="Ratcliffe, Brett C., Jameson, Mary Liz &amp; Zorn, Carsten" docDate="2018" docId="840D87F0FFC2FFC4FD7BFDACFEB7FE07" docLanguage="en" docName="TheColeopBull.72.4.717-735.pdf" docOrigin="The Coleopterists Bulletin 72 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-72.4.717" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Ganganomala SALTINI RATCLIFFE, JAMESON, AND ZORN, A 2018, new genus" docType="treatment" docUuid="13EA55C1-33DD-427C-957C-53BC75E115CE" docUuidSource="ZooBank" docVersion="7" lastPageNumber="731" masterDocId="7834FF88FFC7FFCAFFC5FF8AFFF7FFC1" masterDocTitle="Ganganomala Saltini Ratcliffe, Jameson, and Zorn, a New Genus and Species of Anomalini (Coleoptera: Scarabaeidae: Rutelinae) from Bangladesh and Nepal, with a Revised Circumscription of the Tribe" masterLastPageNumber="735" masterPageNumber="717" pageNumber="722" updateTime="1698989705229" updateUser="ExternalLinkService">
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<mods:title id="1A57FA665B9EC1FC883B36F22F881991">Ganganomala Saltini Ratcliffe, Jameson, and Zorn, a New Genus and Species of Anomalini (Coleoptera: Scarabaeidae: Rutelinae) from Bangladesh and Nepal, with a Revised Circumscription of the Tribe</mods:title>
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<mods:namePart id="AA35F6AF42570E04035BBD480FDD5728">Ratcliffe, Brett C.</mods:namePart>
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<mods:namePart id="6DC2F76A24F2CEDCC55F743237FB40A7">Jameson, Mary Liz</mods:namePart>
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<mods:namePart id="E9D19F09C99AA29D5BB98272110319E7">Zorn, Carsten</mods:namePart>
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<mods:title id="2C0C15E1E75A78C2608BB4899A600F28">The Coleopterists Bulletin</mods:title>
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<treatment id="840D87F0FFC2FFC4FD7BFDACFEB7FE07" ID-DOI="http://doi.org/10.5281/zenodo.5190684" ID-GBIF-Taxon="184090566" ID-Zenodo-Dep="5190684" LSID="urn:lsid:zoobank.org:act:13EA55C1-33DD-427C-957C-53BC75E115CE" httpUri="http://treatment.plazi.org/id/840D87F0FFC2FFC4FD7BFDACFEB7FE07" lastPageId="14" lastPageNumber="731" pageId="5" pageNumber="722">
<subSubSection id="44BE656DFFC2FFCFFD7BFDACFC21FD94" pageId="5" pageNumber="722" type="nomenclature">
<paragraph id="0C1B36E6FFC2FFCFFD7BFDACFC21FD94" blockId="5.[702,1172,550,677]" pageId="5" pageNumber="722">
<heading id="5753818AFFC2FFCFFD7BFDACFC21FD94" centered="true" fontSize="9" level="2" pageId="5" pageNumber="722" reason="2">
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<taxonomicName id="CBA44D65FFC2FFCFFD7BFDACFB88FDFA" authority="Ratcliffe, Jameson, and Zorn" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[702,1151,550,571]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="722" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3ED0EAF4FFC2FFCFFD7BFDACFCBCFDFA" bold="true" box="[702,843,550,571]" italics="true" pageId="5" pageNumber="722">Ganganomala</emphasis>
Ratcliffe, Jameson, and Zorn
</taxonomicName>
,
<taxonomicNameLabel id="25E3578FFFC2FFCFFCAEFDCAFC21FD94" box="[875,982,576,597]" pageId="5" pageNumber="722" rank="genus">new genus</taxonomicNameLabel>
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<paragraph id="0C1B36E6FFC2FFCFFD00FDD1FB88FD64" blockId="5.[702,1172,550,677]" pageId="5" pageNumber="722">
(
<figureCitation id="949F2A63FFC2FFCFFD09FDD1FCD1FDB1" box="[716,806,602,624]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="5" pageNumber="722">Figs. 1–4</figureCitation>
,
<figureCitation id="949F2A63FFC2FFCFFCEAFDD1FC6EFDB1" box="[815,921,602,624]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="5" pageNumber="722">7–9, 11–12</figureCitation>
,
<figureCitation id="949F2A63FFC2FFCFFC64FDD1FC28FDB1" box="[929,991,602,624]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="5" pageNumber="722">14–21</figureCitation>
,
<figureCitation id="949F2A63FFC2FFCFFC2DFDD1FBD2FDB1" box="[1000,1061,602,624]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="5.[175,206,521,540]" captionStartId-1="6.[197,236,1580,1599]" captionTargetBox-0="[155,633,196,503]" captionTargetBox-1="[208,1158,186,1562]" captionTargetId-0="figure-697@5.[153,635,194,505]" captionTargetId-1="figure-0@6.[206,1160,184,1563]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 23. Distribution of Ganganomala saltini in relation to the Ganges River Basin and locations in Nepal and Bangladesh. Map courtesy of A. Smith." captionText-1="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi-0="http://doi.org/10.5281/zenodo.4789844" figureDoi-1="http://doi.org/10.5281/zenodo.4789862" httpUri-0="https://zenodo.org/record/4789844/files/figure.png" httpUri-1="https://zenodo.org/record/4789862/files/figure.png" pageId="5" pageNumber="722">23–27</figureCitation>
,
<figureCitation id="949F2A63FFC2FFCFFBEBFDD1FB9BFDB1" box="[1070,1132,602,624]" captionStart="Figs" captionStartId="7.[175,214,677,696]" captionTargetBox="[156,1165,196,659]" captionTargetId="figure-532@7.[154,1169,194,661]" captionTargetPageId="7" captionText="Figs. 31–35. Protibiae of male and female of Ganganomala saltini, Anomala dorsalis, and Hoplopus marcens Peyerimhoff, dorsal views. 31) G. saltini, male, subapical spur lacking, three external teeth; 32) G. saltini, female, subapical spur present, three external teeth; 33) A. dorsalis (syntype of A. fraterna, MLUH), male, subapical spur present, three external teeth; 34) A. dorsalis, female, subapical spur present, three external teeth; 35) H. marcens, male, subapical spur lacking, two external teeth, third tooth nearly obsolete." figureDoi="http://doi.org/10.5281/zenodo.4789880" httpUri="https://zenodo.org/record/4789880/files/figure.png" pageId="5" pageNumber="722">31–32</figureCitation>
,
<figureCitation id="949F2A63FFC2FFCFFBB1FDD1FB7AFDB1" box="[1140,1165,603,624]" captionStart="Figs" captionStartId="12.[197,236,1602,1621]" captionTargetBox="[220,1151,198,1583]" captionTargetId="figure-0@12.[211,1154,194,1586]" captionTargetPageId="12" captionText="Figs. 36–46. Comparison of male Anomalina protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws. 36) Ganganomala saltini; 37) Anomala dorsalis; 38) Anomala keithi Zorn; 39) Anomala n. sp.; 40) Anomala zornella Prokofiev; 41) Singhala tenella Blanchard; 42) Mimela rugatipennis; 43) Anomala vittata; 44) Megapertha massageta Kirsch; 45) Cyriopertha arcuata; 46) Hoplopus marcens." figureDoi="http://doi.org/10.5281/zenodo.4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="5" pageNumber="722">36</figureCitation>
) Zoobank.org/
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urn:lsid:zoobank.org:act:
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<paragraph id="0C1B36E6FFC2FFC1FD00FD30FED2FE9D" blockId="5.[685,1173,698,1729]" lastBlockId="11.[151,639,194,1730]" lastPageId="11" lastPageNumber="728" pageId="5" pageNumber="722">
<emphasis id="3ED0EAF4FFC2FFCFFD00FD30FC7FFD0E" bold="true" box="[709,904,698,719]" pageId="5" pageNumber="722">Description. Form:</emphasis>
Elongate oval, robust, apices of elytra broadly rounded, pygidium exposed beyond elytra; dorsum convex, glabrous. Color dark reddish brown. Sexual dimorphism present on clypeus, protibia, and protarsomeres. Length 15.2– 16.2 mm; width 8.1–8.4 mm.
<emphasis id="3ED0EAF4FFC2FFCFFC23FCB5FBEBFC95" bold="true" box="[998,1052,831,852]" pageId="5" pageNumber="722">Head</emphasis>
(
<figureCitation id="949F2A63FFC2FFCFFBEAFCB5FB67FC95" box="[1071,1168,831,853]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="5" pageNumber="722">Figs. 1–4</figureCitation>
,
<figureCitation id="949F2A63FFC2FFCFFD68FCD0FD1DFCAE" box="[685,746,857,879]" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="5" pageNumber="722">24–27</figureCitation>
): Frons flat or weakly concave, lacking tubercles, surface punctate, punctures minute or large. Frontoclypeal suture poorly indicated, obsolete medially, weakly arcuate, reaching each lateral margin near ocular canthus. Clypeus with surface flat or weakly concave, punctate, punctures minute or large; apex of male quadrate, strongly or moderately reflexed with anterolateral angle square; apex of female weakly rounded, moderately reflexed with anterolateral margin rounded; margins of male bowed inward, base slightly wider than apex; margins of female parallel, slightly narrower at apex than at base. Eye canthus planar, not cariniform. Interocular width equals 3.0–5.2 transverse eye diameters. Antenna with 9 antennomeres, club subequal in length to antennomeres 2–6. Labrum quadrate at middle, sides rounded, not visible or weakly visible in dorsal view. Mandible (
<figureCitation id="949F2A63FFC2FFCFFB80FA97FB7BFAF3" box="[1093,1164,1309,1330]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="5" pageNumber="722">Fig. 15</figureCitation>
) rounded externally, dorsal surface flat, inner apex bifid (teeth short, acute), molar region with 12–16 lamellae, scissorial region brush-like (lacking teeth). Maxilla (
<figureCitation id="949F2A63FFC2FFCFFC8FFA02FC67FA5C" box="[842,912,1416,1437]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="5" pageNumber="722">Fig. 14</figureCitation>
) with 6 acute teeth, apical tooth and 1 subapical tooth fused with base with 3 equally separated teeth; palpus 4-segmented, terminal segment subequal in length to segments 2–3 combined.
<collectingRegion id="CE60F804FFC2FFCFFCDEFA78FC9CF9C6" box="[795,875,1522,1543]" country="Myanmar" name="Chin" pageId="5" pageNumber="722">Mentum</collectingRegion>
(
<figureCitation id="949F2A63FFC2FFCFFCB9FA78FC33F9C6" box="[892,964,1522,1543]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="5" pageNumber="722">Fig. 16</figureCitation>
) with surface flat or weakly convex, apex bisinuate; length subequal to 1.2 times greatest width; terminal segment of palpus subequal in length to segments 1–2 combined.
<emphasis id="3ED0EAF4FFC2FFCFFD68F9D6FCE4F9B0" bold="true" box="[685,787,1628,1649]" pageId="5" pageNumber="722">Pronotum</emphasis>
(
<figureCitation id="949F2A63FFC2FFCFFCE7F9D6FC60F9B0" box="[802,919,1628,1650]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="5" pageNumber="722">Figs. 1, 3–4</figureCitation>
): Widest at middle, width at base broad (subequal in width to elytral humeri), apical margin membranous, basal margin weakly protuberant posteriorly at middle. Disc evenly convex. Marginal bead present apically, laterally, and at base. Surface variably punctate, punctures with or without setae laterally.
<emphasis id="3ED0EAF4FFC0FFCDFE10FCFFFDB6FC4B" bold="true" box="[469,577,885,906]" pageId="7" pageNumber="724">Scutellum:</emphasis>
Parabolic, slightly wider than long, base declivous at elytral base. Surface variably punctate.
<emphasis id="3ED0EAF4FFC0FFCDFDEBFC20FF17FC1B" bold="true" pageId="7" pageNumber="724">Mesepimeron:</emphasis>
Base not exposed beyond elytral base in dorsal view. Basal width broader than base of elytron.
<emphasis id="3ED0EAF4FFC0FFCDFF1DFC70FED7FBCE" bold="true" box="[216,288,1018,1039]" pageId="7" pageNumber="724">Elytra:</emphasis>
Surface with longitudinal, punctate striae, punctures variable. Epipleuron from base to metacoxa rounded, beaded from base to near apex, with transparent membrane from metacoxa to apex. Apex subquadrately rounded.
<emphasis id="3ED0EAF4FFC0FFCDFE34FBEEFD89FBB8" bold="true" box="[497,638,1124,1145]" pageId="7" pageNumber="724">Propygidium:</emphasis>
Concealed by elytra.
<emphasis id="3ED0EAF4FFC0FFCDFEA4FBF5FE3EFB55" bold="true" box="[353,457,1151,1172]" pageId="7" pageNumber="724">Pygidium:</emphasis>
Width about twice length at middle. Shape subtriangular in caudal view, weakly convex in lateral view. Surface punctate, some punctures setose; setae moderately long, tawny or reddish. Apex and lateral margins beaded.
<emphasis id="3ED0EAF4FFC0FFCDFF52FA8EFF15FAD8" bold="true" box="[151,226,1284,1305]" pageId="7" pageNumber="724">Venter:</emphasis>
Prosternal process obsolete, not produced ventrally, not produced to trochanter, glabrous (
<figureCitation id="949F2A63FFC0FFCDFF5BFAB3FF20FA8F" box="[158,215,1337,1358]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="7" pageNumber="724">Fig. 2</figureCitation>
). Mesometasternal region lacking projection; mesocoxae nearly contiguous. Abdominal sternites lacking carinate ridge laterally (compare with
<figureCitation id="949F2A63FFC0FFCDFD9DFAE4FF58FA5F" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 13</figureCitation>
of
<taxonomicName id="CBA44D65FFC0FFCDFF14FA03FECEFA5F" box="[209,313,1417,1438]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFC0FFCDFF14FA03FECEFA5F" box="[209,313,1417,1438]" italics="true" pageId="7" pageNumber="724">A. dorsalis</emphasis>
</taxonomicName>
with carinate ridge). Apex of last abdominal sternite quadrate (male) or weakly rounded (female).
<emphasis id="3ED0EAF4FFC0FFCDFE8EFA34FE8CFA12" bold="true" box="[331,379,1470,1491]" pageId="7" pageNumber="724">Legs</emphasis>
(
<figureCitation id="949F2A63FFC0FFCDFE4FFA34FDC7FA12" box="[394,560,1469,1491]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Figs. 7–9, 11–12</figureCitation>
,
<figureCitation id="949F2A63FFC0FFCDFDF9FA34FD8DFA12" box="[572,634,1469,1491]" captionStart="Figs" captionStartId="7.[175,214,677,696]" captionTargetBox="[156,1165,196,659]" captionTargetId="figure-532@7.[154,1169,194,661]" captionTargetPageId="7" captionText="Figs. 31–35. Protibiae of male and female of Ganganomala saltini, Anomala dorsalis, and Hoplopus marcens Peyerimhoff, dorsal views. 31) G. saltini, male, subapical spur lacking, three external teeth; 32) G. saltini, female, subapical spur present, three external teeth; 33) A. dorsalis (syntype of A. fraterna, MLUH), male, subapical spur present, three external teeth; 34) A. dorsalis, female, subapical spur present, three external teeth; 35) H. marcens, male, subapical spur lacking, two external teeth, third tooth nearly obsolete." figureDoi="http://doi.org/10.5281/zenodo.4789880" httpUri="https://zenodo.org/record/4789880/files/figure.png" pageId="7" pageNumber="724">31–32</figureCitation>
,
<figureCitation id="949F2A63FFC0FFCDFF52FA52FF47FA2C" box="[151,176,1496,1517]" captionStart="Figs" captionStartId="12.[197,236,1602,1621]" captionTargetBox="[220,1151,198,1583]" captionTargetId="figure-0@12.[211,1154,194,1586]" captionTargetPageId="12" captionText="Figs. 36–46. Comparison of male Anomalina protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws. 36) Ganganomala saltini; 37) Anomala dorsalis; 38) Anomala keithi Zorn; 39) Anomala n. sp.; 40) Anomala zornella Prokofiev; 41) Singhala tenella Blanchard; 42) Mimela rugatipennis; 43) Anomala vittata; 44) Megapertha massageta Kirsch; 45) Cyriopertha arcuata; 46) Hoplopus marcens." figureDoi="http://doi.org/10.5281/zenodo.4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="7" pageNumber="724">36</figureCitation>
): Profemur lacking rounded, dilated apex. Protibia with 3 external teeth, teeth subequally spaced (
<figureCitation id="949F2A63FFC0FFCDFF5BF987FF0DF9E3" box="[158,250,1549,1571]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Figs. 8–9</figureCitation>
); subapical spur on inner margin lacking (male,
<figureCitation id="949F2A63FFC0FFCDFF19F9A2FED8F9FC" box="[220,303,1576,1598]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 7–8</figureCitation>
) or present (female,
<figureCitation id="949F2A63FFC0FFCDFDC7F9A2FDC8F9FC" box="[514,575,1576,1597]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 9</figureCitation>
); spur subequal in length to protarsomere 2; base without protibial notch. Male protarsomere 5 robust (subequal in length to protarsomeres 1–4, width about 1/2 length) with well-developed internomedial protuberance (
<figureCitation id="949F2A63FFC0FFCDFEDBF927FEA1F903" box="[286,342,1709,1730]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 7</figureCitation>
); female protarsomere 5 normal (subequal in length to protarsomeres 2–4, width about 1/3 length) with poorly developed internomedial protuberance (
<figureCitation id="949F2A63FFC0FFCDFC55FCFFFC3FFC4B" box="[912,968,885,906]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 9</figureCitation>
). Male inner protarsal claw entire (not split), robust (twice thickness of outer claw), and with inner, minute, apical tooth; female inner protarsal claw split, ventral ramus subequal to dorsal ramus. Unguitractor plate of protarsus produced beyond apex of protarsomere 5, laterally flattened, bisetose. Mesotibia widest at middle, weakly expanded at apex; external margin with 1 weak, oblique carina on basal 1/4 (with 5–7 spines) and 1 moderately developed, oblique carina at middle (with 6–8 spines); apex with 6–8 spines and 2 inner spurs, longer spur produced to near apex of mesotarsomere 2. Meso- and metatarsomeres 5 with moderately developed internomedial protuberance (male) or weakly developed protuberance (female). Male mesotarsus with claws simple, split in female with ventral ramus subequal in width to dorsal ramus. Unguitractor plate of meso- and metatarsi produced beyond apex of tarsomere 5, laterally flattened, bisetose. Metatrochanter with apex not or only weakly produced beyond posterior border of femur. Metatibia (
<figureCitation id="949F2A63FFC0FFCDFC10FA29FBD6FA79" box="[981,1057,1443,1464]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 11</figureCitation>
) widest at middle, weakly expanded at apex, external margin carinate with spines, spines on female more elongate than those on male; apex with 15–22 spines and 2 inner spurs, longer spur produced to near apex of metatarsomere 2. Metatarsus (
<figureCitation id="949F2A63FFC0FFCDFC16F9A2FBEDF9FC" box="[979,1050,1576,1597]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="7" pageNumber="724">Fig. 12</figureCitation>
) with claws simple in male and female.
<emphasis id="3ED0EAF4FFC0FFCDFC01F9C9FBCEF999" bold="true" box="[964,1081,1603,1624]" pageId="7" pageNumber="724">Hind wing:</emphasis>
Apex of AA
<subScript id="902034A3FFC0FFCDFD15F9EDFD06F9B4" attach="left" box="[720,753,1639,1653]" fontSize="6" pageId="7" pageNumber="724">1+2</subScript>
short, not united with CuA (
<figureCitation id="949F2A63FFC0FFCDFC3BF9D7FBB4F9B3" box="[1022,1091,1629,1650]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="7" pageNumber="724">Fig. 18</figureCitation>
). AP
<subScript id="902034A3FFC0FFCDFBB6F9EDFB63F9B4" attach="right" box="[1139,1172,1639,1653]" fontSize="6" pageId="7" pageNumber="724">3+4</subScript>
at base bulbous, with short, dense setae. ScA with reduced membrane and single row of poorly developed pegs. Region anterior to RA
<subScript id="902034A3FFC0FFCDFBE7F93CFBB4F905" attach="left" box="[1058,1091,1718,1732]" fontSize="6" pageId="7" pageNumber="724">3+4</subScript>
lacking setae (except sparse setae near fold).
<emphasis id="3ED0EAF4FFCCFFC1FDCCFF48FD8AFF16" bold="true" box="[521,637,194,215]" pageId="11" pageNumber="728">Parameres:</emphasis>
Parallel, simple, free (not fused), slightly overlapping at base (
<figureCitation id="949F2A63FFCCFFC1FEF2FF7DFE5BFECD" box="[311,428,246,268]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="11" pageNumber="728">Figs. 19–21</figureCitation>
).
<emphasis id="3ED0EAF4FFCCFFC1FE7AFF7DFD8AFECD" bold="true" box="[447,637,247,268]" pageId="11" pageNumber="728">Spiculum gastrale:</emphasis>
Weakly Y-shaped and with moderately developed sclerites (
<figureCitation id="949F2A63FFCCFFC1FF35FEA6FEC2FE80" box="[240,309,300,321]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="11" pageNumber="728">Fig. 17</figureCitation>
).
<emphasis id="3ED0EAF4FFCCFFC1FE83FEA6FE3EFE80" bold="true" box="[326,457,300,321]" pageId="11" pageNumber="728">Gonocoxites:</emphasis>
Poorly sclerotized, setose at apex.
</paragraph>
</subSubSection>
<caption id="58DB666EFFC1FFCCFF00F9A6FCA4F977" ID-DOI="http://doi.org/10.5281/zenodo.4789862" ID-Zenodo-Dep="4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="6" pageNumber="723" startId="6.[197,236,1580,1599]" targetBox="[208,1158,186,1562]" targetPageId="6">
<paragraph id="0C1B36E6FFC1FFCCFF00F9A6FCA4F977" blockId="6.[173,1193,1580,1718]" pageId="6" pageNumber="723">
<emphasis id="3ED0EAF4FFC1FFCCFF00F9A6FEC3F9FE" bold="true" box="[197,308,1580,1599]" pageId="6" pageNumber="723">Figs. 24–30.</emphasis>
Head comparisons of male and female
<taxonomicName id="CBA44D65FFC1FFCCFD75F9A6FC90F9FE" box="[688,871,1580,1599]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFC1FFCCFD75F9A6FC90F9FE" box="[688,871,1580,1599]" italics="true" pageId="6" pageNumber="723">Ganganomala saltini</emphasis>
</taxonomicName>
and similar
<taxonomicName id="CBA44D65FFC1FFCCFC1AF9A6FBC8F9FE" authorityName="Streubel" authorityYear="1839" box="[991,1087,1580,1599]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="6" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
, dorsal and dorso-oblique views.
<taxonomicName id="CBA44D65FFC1FFCCFEACF9CEFE4DF996" box="[361,442,1604,1623]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFC1FFCCFEACF9CEFE4DF996" box="[361,442,1604,1623]" italics="true" pageId="6" pageNumber="723">G. saltini</emphasis>
</taxonomicName>
:
<emphasis id="3ED0EAF4FFC1FFCCFE0DF9CEFDF3F996" bold="true" box="[456,516,1604,1623]" pageId="6" pageNumber="723">24–25)</emphasis>
Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base;
<emphasis id="3ED0EAF4FFC1FFCCFD07F9D1FD09F9AF" bold="true" box="[706,766,1627,1646]" pageId="6" pageNumber="723">26–27)</emphasis>
Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base.
<emphasis id="3ED0EAF4FFC1FFCCFCBBF9F9FC6DF947" bold="true" box="[894,922,1651,1670]" pageId="6" pageNumber="723">28)</emphasis>
<emphasis id="3ED0EAF4FFC1FFCCFC66F9F9FBD9F947" box="[931,1070,1651,1670]" italics="true" pageId="6" pageNumber="723">
<taxonomicName id="CBA44D65FFC1FFCCFC66F9F9FBDEF947" authorityName="Heller" authorityYear="1891" box="[931,1065,1651,1670]" class="Insecta" family="Rutelidae" genus="Singhala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="hindu">Singhala hindu</taxonomicName>
,
</emphasis>
male, clypeal apex strongly reflexed, emarginate at middle;
<emphasis id="3ED0EAF4FFC1FFCCFD8AF901FD7CF95F" bold="true" box="[591,651,1675,1694]" pageId="6" pageNumber="723">29–30)</emphasis>
<taxonomicName id="CBA44D65FFC1FFCCFD50F901FD01F95F" box="[661,758,1675,1694]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFC1FFCCFD50F901FD01F95F" box="[661,758,1675,1694]" italics="true" pageId="6" pageNumber="723">A. dorsalis</emphasis>
</taxonomicName>
, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex.
</paragraph>
</caption>
<caption id="58DB666EFFC0FFCDFF6AFD2FFCD2FCD6" ID-DOI="http://doi.org/10.5281/zenodo.4789880" ID-Zenodo-Dep="4789880" httpUri="https://zenodo.org/record/4789880/files/figure.png" pageId="7" pageNumber="724" startId="7.[175,214,677,696]" targetBox="[156,1165,196,659]" targetPageId="7">
<paragraph id="0C1B36E6FFC0FFCDFF6AFD2FFCD2FCD6" blockId="7.[151,1173,677,791]" pageId="7" pageNumber="724">
<emphasis id="3ED0EAF4FFC0FFCDFF6AFD2FFEE9FD79" bold="true" box="[175,286,677,696]" pageId="7" pageNumber="724">Figs. 31–35.</emphasis>
Protibiae of male and female of
<taxonomicName id="CBA44D65FFC0FFCDFD9EFD2FFCE6FD79" box="[603,785,677,696]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFC0FFCDFD9EFD2FFCE6FD79" box="[603,785,677,696]" italics="true" pageId="7" pageNumber="724">Ganganomala saltini</emphasis>
</taxonomicName>
,
<taxonomicName id="CBA44D65FFC0FFCDFCE4FD2FFC4DFD79" box="[801,954,677,696]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFC0FFCDFCE4FD2FFC4DFD79" box="[801,954,677,696]" italics="true" pageId="7" pageNumber="724">Anomala dorsalis</emphasis>
</taxonomicName>
, and
<taxonomicName id="CBA44D65FFC0FFCDFC34FD2FFEF5FD11" authority="Peyerimhoff" authorityName="Peyerimhoff" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="marcens">
<emphasis id="3ED0EAF4FFC0FFCDFC34FD2FFB63FD79" box="[1009,1172,677,696]" italics="true" pageId="7" pageNumber="724">Hoplopus marcens</emphasis>
Peyerimhoff
</taxonomicName>
, dorsal views.
<emphasis id="3ED0EAF4FFC0FFCDFE4FFD37FE51FD11" bold="true" box="[394,422,701,720]" pageId="7" pageNumber="724">31)</emphasis>
<taxonomicName id="CBA44D65FFC0FFCDFE6AFD37FDF6FD11" box="[431,513,701,720]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFC0FFCDFE6AFD37FDF6FD11" box="[431,513,701,720]" italics="true" pageId="7" pageNumber="724">G. saltini</emphasis>
</taxonomicName>
, male, subapical spur lacking, three external teeth;
<emphasis id="3ED0EAF4FFC0FFCDFC15FD37FC1BFD11" bold="true" box="[976,1004,701,720]" pageId="7" pageNumber="724">32)</emphasis>
<taxonomicName id="CBA44D65FFC0FFCDFC30FD37FBB0FD11" box="[1013,1095,701,720]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFC0FFCDFC30FD37FBB0FD11" box="[1013,1095,701,720]" italics="true" pageId="7" pageNumber="724">G. saltini</emphasis>
</taxonomicName>
, female, subapical spur present, three external teeth;
<emphasis id="3ED0EAF4FFC0FFCDFDE4FD5EFDCAFD26" bold="true" box="[545,573,724,743]" pageId="7" pageNumber="724">33)</emphasis>
<taxonomicName id="CBA44D65FFC0FFCDFD82FD5EFD51FD26" box="[583,678,724,743]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFC0FFCDFD82FD5EFD51FD26" box="[583,678,724,743]" italics="true" pageId="7" pageNumber="724">A. dorsalis</emphasis>
</taxonomicName>
(syntype of
<taxonomicName id="CBA44D65FFC0FFCDFCD9FD5EFC3AFD26" authority=", MLUH" authorityName="MLUH" box="[796,973,724,743]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFC0FFCDFCD9FD5EFC8AFD26" box="[796,893,724,743]" italics="true" pageId="7" pageNumber="724">A. fraterna</emphasis>
, MLUH
</taxonomicName>
), male, subapical spur present, three external teeth;
<emphasis id="3ED0EAF4FFC0FFCDFE59FD66FE4FFD3E" bold="true" box="[412,440,748,767]" pageId="7" pageNumber="724">34)</emphasis>
<taxonomicName id="CBA44D65FFC0FFCDFE07FD66FDD6FD3E" box="[450,545,748,767]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFC0FFCDFE07FD66FDD6FD3E" box="[450,545,748,767]" italics="true" pageId="7" pageNumber="724">A. dorsalis</emphasis>
</taxonomicName>
, female, subapical spur present, three external teeth;
<emphasis id="3ED0EAF4FFC0FFCDFBC6FD66FBD7FD3E" bold="true" box="[1027,1056,748,767]" pageId="7" pageNumber="724">35)</emphasis>
<taxonomicName id="CBA44D65FFC0FFCDFBECFD66FB78FD3E" box="[1065,1167,748,767]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="marcens">
<emphasis id="3ED0EAF4FFC0FFCDFBECFD66FB78FD3E" box="[1065,1167,748,767]" italics="true" pageId="7" pageNumber="724">H. marcens</emphasis>
</taxonomicName>
, male, subapical spur lacking, two external teeth, third tooth nearly obsolete.
</paragraph>
</caption>
<caption id="58DB666EFFCFFFC2FF6AFE42FA49FE32" ID-Table-UUID="58DB666EFFCFFFC2FF6AFE42FA49FE32" httpUri="http://table.plazi.org/id/58DB666EFFCFFFC2FF6AFE42FA49FE32" pageId="8" pageNumber="725" startId="8.[175,225,456,475]" targetBox="[151,1612,531,1424]" targetIsTable="true" targetPageId="8">
<paragraph id="0C1B36E6FFCFFFC2FF6AFE42FA49FE32" blockId="8.[151,1682,456,499]" pageId="8" pageNumber="725">
<emphasis id="3ED0EAF4FFCFFFC2FF6AFE42FF02FE1A" bold="true" box="[175,245,456,475]" pageId="8" pageNumber="725">Table 1.</emphasis>
Fifty genera and subgenera of world
<taxonomicName id="CBA44D65FFCFFFC2FD84FE42FD68FE1A" authorityName="Streubel" authorityYear="1839" box="[577,671,456,475]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="8" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
(tribe
<taxonomicName id="CBA44D65FFCFFFC2FD13FE42FCC6FE1A" box="[726,817,456,475]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
) from the Afrotropical Region [AFR], Australian Region [AUR], Nearctic Region [NAR], Neotropical Region [NTR], Oriental Region [ORR], Palaearctic Region [PAL], and Pacific Region [PAC]. Biogeographic regions are based on
<bibRefCitation id="68354B17FFCFFFC2FB38FE6AFA4DFE32" author="Lobl, I." box="[1277,1466,480,499]" pageId="8" pageNumber="725" refId="ref11285" refString="Lobl, I., and D. Lobl (editors). 2016. Catalogue of the Palaearctic Coleoptera. Volume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Koninklijke Brill, Leiden, Netherlands." type="journal volume" year="2016">Löbl and Löbl (2016)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="0C1B36E6FFCFFFC2FE81FD99F9BBFA51" pageId="8" pageNumber="725">
<table id="7EA4C446FFCF0035FF52FD99F9BBFA51" box="[151,1612,531,1424]" colsContinueIn="9.[151,1682,602,1040]" gridcols="3" gridrows="8" pageId="8" pageNumber="725">
<tr id="B29434A4FFCF0035FF52FD99F9BBFDE7" box="[151,1612,531,550]" gridrow="0" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FD99FDA2FDE7" box="[151,597,531,550]" gridcol="0" gridrow="0" pageId="8" pageNumber="725">
<emphasis id="3ED0EAF4FFCFFFC2FE81FD99FE1FFDE7" bold="true" box="[324,488,531,550]" pageId="8" pageNumber="725">Genus (Subgenus)</emphasis>
</th>
<th id="F1455DD8FFCF0035FD1AFD99FBBDFDE7" box="[735,1098,531,550]" gridcol="1" gridrow="0" pageId="8" pageNumber="725">
<emphasis id="3ED0EAF4FFCFFFC2FCF1FD99FC02FDE7" bold="true" box="[820,1013,531,550]" pageId="8" pageNumber="725">Biogeographic region</emphasis>
</th>
<th id="F1455DD8FFCF0035FB51FD99F9BBFDE7" box="[1172,1612,531,550]" gridcol="2" gridrow="0" pageId="8" pageNumber="725">
<emphasis id="3ED0EAF4FFCFFFC2FAA7FD99FA34FDE7" bold="true" box="[1378,1475,531,550]" pageId="8" pageNumber="725">References</emphasis>
</th>
</tr>
<tr id="B29434A4FFCF0035FF52FDB2F9BBFD1B" box="[151,1612,568,730]" gridrow="1" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FDB2FDA2FD1B" box="[151,597,568,730]" gridcol="0" gridrow="1" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FDB2FE6CFD8A" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[151,411,568,587]" class="Insecta" family="Rutelidae" genus="Adoretosoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FDB2FEF0FD8A" box="[151,263,568,587]" italics="true" pageId="8" pageNumber="725">Adoretosoma</emphasis>
Blanchard, 1851
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FDDAFE8FFDA2" authority="Samouelle, 1819" authorityName="Samouelle" authorityYear="1819" box="[151,376,592,611]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FDDAFF14FDA2" box="[151,227,592,611]" italics="true" pageId="8" pageNumber="725">Anomala</emphasis>
Samouelle, 1819
</taxonomicName>
1
<taxonomicName id="CBA44D65FFCFFFC2FF52FDE2FE1DFDBA" authority="Jameson Paucar-Cabrera" authorityName="Jameson Paucar-Cabrera" box="[151,490,616,635]" class="Insecta" family="Rutelidae" genus="Anomalorhina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FDE2FEE6FDBA" box="[151,273,616,635]" italics="true" pageId="8" pageNumber="725">Anomalorhina</emphasis>
Jameson Paucar-Cabrera
</taxonomicName>
, Sol´ıs, 2003
<taxonomicName id="CBA44D65FFCFFFC2FF52FD0AFE71FD52" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" box="[151,390,640,659]" class="Insecta" family="Rutelidae" genus="Anoplanomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FD0AFEE4FD52" box="[151,275,640,659]" italics="true" pageId="8" pageNumber="725">Anoplanomala</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FEDFFD0AFE71FD52" author="Arrow, G. J." box="[282,390,640,659]" pageId="8" pageNumber="725" refId="ref10166" refString="Arrow, G. J. 1917. The Fauna of British India, including Ceylon and Burma. Coleoptera, Lamellicornia, Part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor and Francis, London, UK." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FD1DFE7EFD6B" authority="Medvedev, 1949" authorityName="Medvedev" authorityYear="1949" box="[151,393,663,682]" class="Insecta" family="Scarabaeidae" genus="Asiopertha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FD1DFF04FD6B" box="[151,243,663,682]" italics="true" pageId="8" pageNumber="725">Asiopertha</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FF3FFD1DFE7EFD6B" author="Medvedev S. I." box="[250,393,663,682]" pageId="8" pageNumber="725" pagination="3" refId="ref11439" refString="Medvedev S. I. 1949. Fauna SSSR. Zhestkokrytye. Tom 10, vyo. 3. Plastinchatousye (Scarabaeidae) podsem. Rutelinae (khlebnye zhuki i blizkiegruppy). Akademii Nauk SSSR, Moscow, Union of Soviet Socialist Republics." type="journal article" year="1949">Medvedev, 1949</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FD25FE8DFD03" authority="Paucar, 2003" authorityName="Paucar" authorityYear="2003" box="[151,378,687,706]" class="Insecta" family="Rutelidae" genus="Balanogonia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FD25FEF3FD03" box="[151,260,687,706]" italics="true" pageId="8" pageNumber="725">Balanogonia</emphasis>
Paucar, 2003
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FD4DFE9BFD1B" authority="Reitter, 1903" authorityName="Reitter" authorityYear="1903" box="[151,364,711,730]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FD4DFF00FD1B" box="[151,247,711,730]" italics="true" pageId="8" pageNumber="725">Blitopertha</emphasis>
Reitter, 1903
</taxonomicName>
</th>
<td id="F1455DD8FFCF0035FD1AFDB2FBBDFD1B" box="[735,1098,568,730]" gridcol="1" gridrow="1" pageId="8" pageNumber="725">PAL, ORR AFR, AUS, NAR, NTR, ORR, PAC, PAL NTR ORR PAL NTR PAL</td>
<td id="F1455DD8FFCF0035FB51FDB2F9BBFD1B" box="[1172,1612,568,730]" gridcol="2" gridrow="1" pageId="8" pageNumber="725">
<bibRefCitation id="68354B17FFCFFFC2FB51FDB2FAC3FD8A" author="Machatschke, J. W." box="[1172,1332,568,587]" pageId="8" pageNumber="725" pagination="349 - 396" refId="ref11367" refString="Machatschke, J. W. 1955. Versuch einer Neugliederung der Arten des Genus Adoretosoma Blanchard. Beitrage zur Entomologie 5: 349 - 396." type="journal article" year="1955">Machatschke 1955</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FDDAFAC3FDA2" author="Machatschke, J. W." box="[1172,1332,592,611]" pageId="8" pageNumber="725" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FDE2FAB7FDBA" author="Jameson, M. L." box="[1172,1344,616,635]" pageId="8" pageNumber="725" pagination="415 - 432" refId="ref10890" refString="Jameson, M. L., A. Paucar-Cabrera, and A. Sol'is. 2003. Synopsis of the New World genera of Anomalini (Coleoptera: Scarabaeidae: Rutelinae) and description of a new genus from Costa Rica and Nicaragua. Annals of the Entomological Society of America 96: 415 - 432." type="journal article" year="2003">
Jameson
<emphasis id="3ED0EAF4FFCFFFC2FB26FDE2FAE7FDBA" box="[1251,1296,616,635]" italics="true" pageId="8" pageNumber="725">et al.</emphasis>
2003
</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FD0AFAC3FD52" author="Machatschke, J. W." box="[1172,1332,640,659]" pageId="8" pageNumber="725" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FD1DFAE9FD6B" author="Medvedev S. I." box="[1172,1310,663,682]" pageId="8" pageNumber="725" pagination="3" refId="ref11439" refString="Medvedev S. I. 1949. Fauna SSSR. Zhestkokrytye. Tom 10, vyo. 3. Plastinchatousye (Scarabaeidae) podsem. Rutelinae (khlebnye zhuki i blizkiegruppy). Akademii Nauk SSSR, Moscow, Union of Soviet Socialist Republics." type="journal article" year="1949">Medvedev 1949</bibRefCitation>
Paucar 2003
<bibRefCitation id="68354B17FFCFFFC2FB51FD4DFAF5FD1B" author="Baraud, J." box="[1172,1282,711,730]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCF0035FF52FD55F9BBFC73" box="[151,1612,735,946]" gridrow="2" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FD55FDA2FC73" box="[151,597,735,946]" gridcol="0" gridrow="2" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FD55FE78FD33" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[151,399,735,754]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FD55FF0CFD33" box="[151,251,735,754]" italics="true" pageId="8" pageNumber="725">Callistethus</emphasis>
Blanchard, 1851
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FD7DFE17FCCB" authority="Moron and Nogueira, 1998" authorityName="Moron and Nogueira" authorityYear="1998" box="[151,480,759,778]" class="Insecta" family="Rutelidae" genus="Chelilabia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FD7DFF07FCCB" box="[151,240,759,778]" italics="true" pageId="8" pageNumber="725">Chelilabia</emphasis>
Morón and Nogueira, 1998
</taxonomicName>
2
<taxonomicName id="CBA44D65FFCFFFC2FF52FC85FE70FCE3" authority="Kobayashi, 2008" authorityName="Kobayashi" authorityYear="2008" box="[151,391,783,802]" class="Insecta" family="Rutelidae" genus="Choumala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC85FF18FCE3" box="[151,239,783,802]" italics="true" pageId="8" pageNumber="725">Choumala</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FF33FC85FE70FCE3" author="Kobayashi, H." box="[246,391,783,802]" pageId="8" pageNumber="725" pagination="69 - 76" refId="ref11078" refString="Kobayashi, H., and W. - I. Chou. 2008. Description of a new genus of Anomalini and two new species of Hoplini and Melolonthini from Taiwan, with two new records of scarabaeid beetles (Coleoptera, Scarabaeidae). Kogane 9: 69 - 76." type="journal article" year="2008">Kobayashi, 2008</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FCADFF09FCFB" authorityName="Reitter" authorityYear="1903" box="[151,254,807,826]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FCADFF09FCFB" box="[151,254,807,826]" italics="true" pageId="8" pageNumber="725">Cyriopertha</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCFFFC2FECEFCADFE1FFCFB" authority="Reitter, 1903" authorityName="Reitter" authorityYear="1903" box="[267,488,807,826]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FECEFCADFE85FCFB" box="[267,370,807,826]" italics="true" pageId="8" pageNumber="725">Cyriopertha</emphasis>
Reitter, 1903
</taxonomicName>
)
<taxonomicName id="CBA44D65FFCFFFC2FF52FCB5FF09FC93" authorityName="Reitter" authorityYear="1903" box="[151,254,831,850]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FCB5FF09FC93" box="[151,254,831,850]" italics="true" pageId="8" pageNumber="725">Cyriopertha</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCFFFC2FECAFCB5FDF1FC93" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[271,518,831,850]" class="Insecta" family="Rutelidae" genus="Apleopertha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FECAFCB5FE80FC93" box="[271,375,831,850]" italics="true" pageId="8" pageNumber="725">Apleopertha</emphasis>
Semenov, 1903
</taxonomicName>
)
<taxonomicName id="CBA44D65FFCFFFC2FF52FCDDFF09FCAB" authorityName="Reitter" authorityYear="1903" box="[151,254,855,874]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FCDDFF09FCAB" box="[151,254,855,874]" italics="true" pageId="8" pageNumber="725">Cyriopertha</emphasis>
</taxonomicName>
(
<emphasis id="3ED0EAF4FFCFFFC2FECAFCDDFE9BFCAB" box="[271,364,855,874]" italics="true" pageId="8" pageNumber="725">Pleopertha</emphasis>
Reitter, 1903)
<taxonomicName id="CBA44D65FFCFFFC2FF52FCE5FE7CFC43" authority="Prokofiev, 2013" authorityName="Prokofiev" authorityYear="2013" box="[151,395,879,898]" class="Insecta" family="Rutelidae" genus="Dalatamala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FCE5FF0BFC43" box="[151,252,879,898]" italics="true" pageId="8" pageNumber="725">Dalatamala</emphasis>
Prokofiev, 2013
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FC0DFE85FC5B" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[151,370,903,922]" class="Insecta" family="Rutelidae" genus="Dilophochila" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC0DFEF1FC5B" box="[151,262,903,922]" italics="true" pageId="8" pageNumber="725">Dilophochila</emphasis>
Bates, 1888
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FC15FE61FC73" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[151,406,927,946]" class="Insecta" family="Rutelidae" genus="Epectinaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC15FEF5FC73" box="[151,258,927,946]" italics="true" pageId="8" pageNumber="725">Epectinaspis</emphasis>
Blanchard, 1851
</taxonomicName>
</th>
<td id="F1455DD8FFCF0035FD1AFD55FBBDFC73" box="[735,1098,735,946]" gridcol="1" gridrow="2" pageId="8" pageNumber="725">NTR, ORR, PAL NTR PAL PAL PAL PAL ORR NTR NTR</td>
<td id="F1455DD8FFCF0035FB51FD55F9BBFC73" box="[1172,1612,735,946]" gridcol="2" gridrow="2" pageId="8" pageNumber="725">
<bibRefCitation id="68354B17FFCFFFC2FB51FD55FAC3FD33" author="Machatschke, J. W." box="[1172,1332,735,754]" pageId="8" pageNumber="725" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
Morón and Nogueira 1998 Kobayashi and Chou 2008 Reitter 1903 Reitter 1903 Reitter 1903 Prokofiev 2013 Morón and Howden 2001 Paucar 2003; Ram´ırez-Ponce and Curoe 2014
</td>
</tr>
<tr id="B29434A4FFCF0035FF52FC3CF9BBFBD0" box="[151,1612,950,1041]" gridrow="3" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FC3CFDA2FBD0" box="[151,597,950,1041]" gridcol="0" gridrow="3" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FC3CFF15FC08" authorityName="Reitter" authorityYear="1903" box="[151,226,950,969]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC3CFF15FC08" box="[151,226,950,969]" italics="true" pageId="8" pageNumber="725">Exomala</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCFFFC2FF2AFC3CFE47FC08" authority="Reitter, 1903" authorityName="Reitter" authorityYear="1903" box="[239,432,950,969]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF2AFC3CFECDFC08" box="[239,314,950,969]" italics="true" pageId="8" pageNumber="725">Exomala</emphasis>
Reitter, 1903
</taxonomicName>
)
<taxonomicName id="CBA44D65FFCFFFC2FF52FC44FF15FC20" authorityName="Reitter" authorityYear="1903" box="[151,226,974,993]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC44FF15FC20" box="[151,226,974,993]" italics="true" pageId="8" pageNumber="725">Exomala</emphasis>
</taxonomicName>
(
<emphasis id="3ED0EAF4FFCFFFC2FF2AFC44FE99FC20" box="[239,366,974,993]" italics="true" pageId="8" pageNumber="725">Neoblitopertha</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FEB0FC44FE1EFC20" author="Baraud, J." box="[373,489,974,993]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud, 1991</bibRefCitation>
)
<taxonomicName id="CBA44D65FFCFFFC2FF52FC6CFF15FC38" authorityName="Reitter" authorityYear="1903" box="[151,226,998,1017]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC6CFF15FC38" box="[151,226,998,1017]" italics="true" pageId="8" pageNumber="725">Exomala</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCFFFC2FF2AFC6CFE33FC38" authority="Baraud, 1991" authorityName="Baraud" authorityYear="1991" box="[239,452,998,1017]" class="Insecta" family="Scarabaeidae" genus="Taxipertha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF2AFC6CFEBEFC38" box="[239,329,998,1017]" italics="true" pageId="8" pageNumber="725">Taxipertha</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FE94FC6CFE33FC38" author="Baraud, J." box="[337,452,998,1017]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud, 1991</bibRefCitation>
</taxonomicName>
)
<taxonomicName id="CBA44D65FFCFFFC2FF52FC74FF15FBD0" authorityName="Reitter" authorityYear="1903" box="[151,226,1022,1041]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FC74FF15FBD0" box="[151,226,1022,1041]" italics="true" pageId="8" pageNumber="725">Exomala</emphasis>
</taxonomicName>
(
<emphasis id="3ED0EAF4FFCFFFC2FF2AFC74FEAAFBD0" box="[239,349,1022,1041]" italics="true" pageId="8" pageNumber="725">Trichopertha</emphasis>
Reitter, 1903)
</th>
<td id="F1455DD8FFCF0035FD1AFC3CFBBDFBD0" box="[735,1098,950,1041]" gridcol="1" gridrow="3" pageId="8" pageNumber="725">NAR, PAC, PAL PAL PAL PAL</td>
<td id="F1455DD8FFCF0035FB51FC3CF9BBFBD0" box="[1172,1612,950,1041]" gridcol="2" gridrow="3" pageId="8" pageNumber="725">
Reitter 1903;
<bibRefCitation id="68354B17FFCFFFC2FACFFC3CFA8CFC08" author="Baraud, J." box="[1290,1403,950,969]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
; Shokhin 2017
<bibRefCitation id="68354B17FFCFFFC2FB51FC44FAF3FC20" author="Baraud, J." box="[1172,1284,974,993]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
; Rössner
<emphasis id="3ED0EAF4FFCFFFC2FA9DFC44FA72FC20" box="[1368,1413,974,993]" italics="true" pageId="8" pageNumber="725">et al.</emphasis>
2009
<bibRefCitation id="68354B17FFCFFFC2FB51FC6CFAF3FC38" author="Baraud, J." box="[1172,1284,998,1017]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
; Shokhin 2017
<bibRefCitation id="68354B17FFCFFFC2FB51FC74FAF3FBD0" author="Baraud, J." box="[1172,1284,1022,1041]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
; Shokhin 2017
</td>
</tr>
<tr id="B29434A4FFCF0035FF52FB9CF9BBFB48" box="[151,1612,1046,1161]" gridrow="4" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FB9CFDA2FB48" box="[151,597,1046,1161]" gridcol="0" gridrow="4" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FB9CFDB2FBE8" authority="Ratcliffe, Jameson, and Zorn, 2018" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[151,581,1046,1065]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FB9CFEE7FBE8" box="[151,272,1046,1065]" italics="true" pageId="8" pageNumber="725">Ganganomala</emphasis>
Ratcliffe, Jameson, and Zorn, 2018
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FBA4FE8AFB80" authority="Escalera, 1914" authorityName="Escalera" authorityYear="1914" box="[151,381,1070,1089]" class="Insecta" family="Rutelidae" genus="Hontoriella" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FBA4FF0EFB80" box="[151,249,1070,1089]" italics="true" pageId="8" pageNumber="725">Hontoriella</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FF3AFBA4FE8AFB80" author="Escalera, M. M. de la" box="[255,381,1070,1089]" pageId="8" pageNumber="725" pagination="1 - 553" refId="ref10653" refString="Escalera, M. M. de la 1914. Los Coleopteros de Marruecos. Trabajos del Museo de Ciencias Naturales (Serie Zoologica) 11: 1 - 553." type="journal article" year="1914">Escalera, 1914</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FBCCFE9FFB98" authority="Laporte, 1832" authorityName="Laporte" authorityYear="1832" box="[151,360,1094,1113]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FBCCFF1EFB98" box="[151,233,1094,1113]" italics="true" pageId="8" pageNumber="725">Hoplopus</emphasis>
Laporte, 1832
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FBD4FDFAFBB0" authority="Moron and Nogueira, 1998" authorityName="Moron and Nogueira" authorityYear="1998" box="[151,525,1118,1137]" class="Insecta" family="Rutelidae" genus="Mazahuapertha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FBD4FEEAFBB0" box="[151,285,1118,1137]" italics="true" pageId="8" pageNumber="725">Mazahuapertha</emphasis>
Morón and Nogueira, 1998
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FBFCFE83FB48" authority="Reitter, 1903" authorityName="Reitter" authorityYear="1903" box="[151,372,1142,1161]" class="Insecta" family="Scarabaeidae" genus="Megapertha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FBFCFF09FB48" box="[151,254,1142,1161]" italics="true" pageId="8" pageNumber="725">Megapertha</emphasis>
Reitter, 1903
</taxonomicName>
</th>
<td id="F1455DD8FFCF0035FD1AFB9CFBBDFB48" box="[735,1098,1046,1161]" gridcol="1" gridrow="4" pageId="8" pageNumber="725">ORR, PAL PAL PAL NTR PAL</td>
<td id="F1455DD8FFCF0035FB51FB9CF9BBFB48" box="[1172,1612,1046,1161]" gridcol="2" gridrow="4" pageId="8" pageNumber="725">
This publication
<bibRefCitation id="68354B17FFCFFFC2FB51FBA4FAFBFB80" author="Escalera, M. M. de la" box="[1172,1292,1070,1089]" pageId="8" pageNumber="725" pagination="1 - 553" refId="ref10653" refString="Escalera, M. M. de la 1914. Los Coleopteros de Marruecos. Trabajos del Museo de Ciencias Naturales (Serie Zoologica) 11: 1 - 553." type="journal article" year="1914">Escalera 1914</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FBCCFAF5FB98" author="Baraud, J." box="[1172,1282,1094,1113]" pageId="8" pageNumber="725" refId="ref10322" refString="Baraud, J. 1985. Coleopteres Scarabaeoidea. Faune du Nord de l' Afrique du Maroc au Sina ¨ i. Encyclopedie Entomologique - XLVI. Editions Lechevalier, Paris, France." type="book" year="1985">Baraud 1985</bibRefCitation>
Morón and Nogueira 1998 Reitter 1903;
<bibRefCitation id="68354B17FFCFFFC2FACFFBFCFA8EFB48" author="Baraud, J." box="[1290,1401,1142,1161]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCF0035FF52FB04F9BBFB29" box="[151,1612,1166,1256]" gridrow="5" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FB04FDA2FB29" box="[151,597,1166,1256]" gridcol="0" gridrow="5" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FB04FE84FB60" authority="Lin, 1980" authorityName="Lin" authorityYear="1980" box="[151,371,1166,1185]" class="Insecta" family="Rutelidae" genus="Melanopopillia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FB04FEEEFB60" box="[151,281,1166,1185]" italics="true" pageId="8" pageNumber="725">Melanopopillia</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FEDAFB04FE84FB60" author="Lin, P." box="[287,371,1166,1185]" pageId="8" pageNumber="725" pagination="297 - 301" refId="ref11220" refString="Lin, P. 1980. A new genus, Melanopopillia, from China (Coleoptera, Rutelidae). Entomotaxia 2: 297 - 301." type="journal article" year="1980">Lin, 1980</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FB2CFE8BFB78" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[151,380,1190,1209]" class="Insecta" family="Rutelidae" genus="Micranomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FB2CFEFDFB78" box="[151,266,1190,1209]" italics="true" pageId="8" pageNumber="725">Micranomala</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FED5FB2CFE8BFB78" author="Arrow, G. J." box="[272,380,1190,1209]" pageId="8" pageNumber="725" pagination="268 - 271" refId="ref10124" refString="Arrow, G. J. 1911. A new genus of Ruteline Coleoptera from the Indian region. The Annals and Magazine of Natural History, including Zoology, Botany and Geology (8) 8: 268 - 271." type="journal article" year="1911">Arrow, 1911</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FB37FE7EFB11" authority="Petrovitz, 1967" authorityName="Petrovitz" authorityYear="1967" box="[151,393,1213,1232]" class="Insecta" family="Rutelidae" genus="Microlontha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FB37FF08FB11" box="[151,255,1213,1232]" italics="true" pageId="8" pageNumber="725">Microlontha</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FEC3FB37FE7EFB11" author="Petrovitz, R." box="[262,393,1213,1232]" pageId="8" pageNumber="725" pagination="479 - 490" refId="ref11857" refString="Petrovitz, R. 1967. Osterreichische entomologische Expeditionen nach Persien und Afghanistan. Beitrage zur Coleopterologie. Teil XI. Lamellicornia. Annalen des Naturhistorischen Museums in Wien 70: 479 - 490." type="journal article" year="1967">Petrovitz, 1967</bibRefCitation>
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FB5FFE8CFB29" authority="Baraud, 1991" authorityName="Baraud" authorityYear="1991" box="[151,379,1237,1256]" class="Insecta" family="Rutelidae" genus="Micropertha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FB5FFEF7FB29" box="[151,256,1237,1256]" italics="true" pageId="8" pageNumber="725">Micropertha</emphasis>
<bibRefCitation id="68354B17FFCFFFC2FEC2FB5FFE8CFB29" author="Baraud, J." box="[263,379,1237,1256]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud, 1991</bibRefCitation>
</taxonomicName>
</th>
<td id="F1455DD8FFCF0035FD1AFB04FBBDFB29" box="[735,1098,1166,1256]" gridcol="1" gridrow="5" pageId="8" pageNumber="725">ORR, PAL ORR PAL PAL</td>
<td id="F1455DD8FFCF0035FB51FB04F9BBFB29" box="[1172,1612,1166,1256]" gridcol="2" gridrow="5" pageId="8" pageNumber="725">
<bibRefCitation id="68354B17FFCFFFC2FB51FB04FB14FB60" author="Lin, P." box="[1172,1251,1166,1185]" pageId="8" pageNumber="725" pagination="297 - 301" refId="ref11220" refString="Lin, P. 1980. A new genus, Melanopopillia, from China (Coleoptera, Rutelidae). Entomotaxia 2: 297 - 301." type="journal article" year="1980">Lin 1980</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FB2CFB0CFB78" author="Arrow, G. J." box="[1172,1275,1190,1209]" pageId="8" pageNumber="725" pagination="268 - 271" refId="ref10124" refString="Arrow, G. J. 1911. A new genus of Ruteline Coleoptera from the Indian region. The Annals and Magazine of Natural History, including Zoology, Botany and Geology (8) 8: 268 - 271." type="journal article" year="1911">Arrow 1911</bibRefCitation>
<bibRefCitation id="68354B17FFCFFFC2FB51FB37FAE4FB11" author="Petrovitz, R." box="[1172,1299,1213,1232]" pageId="8" pageNumber="725" pagination="479 - 490" refId="ref11857" refString="Petrovitz, R. 1967. Osterreichische entomologische Expeditionen nach Persien und Afghanistan. Beitrage zur Coleopterologie. Teil XI. Lamellicornia. Annalen des Naturhistorischen Museums in Wien 70: 479 - 490." type="journal article" year="1967">Petrovitz 1967</bibRefCitation>
; Sabatinelli and Pontuale 1999
<bibRefCitation id="68354B17FFCFFFC2FB51FB5FFAF3FB29" author="Baraud, J." box="[1172,1284,1237,1256]" pageId="8" pageNumber="725" pagination="46 - 61" refId="ref10356" refString="Baraud, J. 1991. Nouvelle classification proposee pour les especes du genre Blitopertha Reitter (1903). (Coleoptera Rutelidae). Lambillionea 91: 46 - 61." type="journal article" year="1991">Baraud 1991</bibRefCitation>
; Shokhin 2017
</td>
</tr>
<tr id="B29434A4FFCF0035FF52FB67F9BBFA89" box="[151,1612,1261,1352]" gridrow="6" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FB67FDA2FA89" box="[151,597,1261,1352]" gridcol="0" gridrow="6" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FB67FEB3FAC1" authority="Kirby, 1823" authorityName="Kirby" authorityYear="1823" box="[151,324,1261,1280]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FB67FF20FAC1" box="[151,215,1261,1280]" italics="true" pageId="8" pageNumber="725">Mimela</emphasis>
Kirby, 1823
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FA8FFF02FAD9" authorityName="Ramirez-Ponce" authorityYear="2015" box="[151,245,1285,1304]" class="Insecta" family="Rutelidae" genus="Moroniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FA8FFF02FAD9" box="[151,245,1285,1304]" italics="true" pageId="8" pageNumber="725">Moroniella</emphasis>
</taxonomicName>
<bibRefCitation id="68354B17FFCFFFC2FF3EFA8FFE45FAD9" author="Ram'irez-Ponce, A." box="[251,434,1285,1304]" pageId="8" pageNumber="725" pagination="261 - 266" refId="ref11921" refString="Ram'irez-Ponce, A. 2015. Moroniella, un nuevo genero americano de Anomalini (Coleoptera: Melolonthidae). Dugesiana 22: 261 - 266." type="journal article" year="2015">Ram´ırez-Ponce, 2015</bibRefCitation>
<taxonomicName id="CBA44D65FFCFFFC2FF52FA97FE25FAF1" authority="Moron and Nogueira, 1998" authorityName="Moron and Nogueira" authorityYear="1998" box="[151,466,1309,1328]" class="Insecta" family="Rutelidae" genus="Nayarita" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FA97FF15FAF1" box="[151,226,1309,1328]" italics="true" pageId="8" pageNumber="725">Nayarita</emphasis>
Morón and Nogueira, 1998
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FABFFE8EFA89" authority="Heller, 1896" authorityName="Heller" authorityYear="1896" box="[151,377,1333,1352]" class="Insecta" family="Rutelidae" genus="Nothophanus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FABFFEFFFA89" box="[151,264,1333,1352]" italics="true" pageId="8" pageNumber="725">Nothophanus</emphasis>
Heller, 1896
</taxonomicName>
</th>
<td id="F1455DD8FFCF0035FD1AFB67FBBDFA89" box="[735,1098,1261,1352]" gridcol="1" gridrow="6" pageId="8" pageNumber="725">AFR, ORR, PAL NTR NTR ORR</td>
<td id="F1455DD8FFCF0035FB51FB67F9BBFA89" box="[1172,1612,1261,1352]" gridcol="2" gridrow="6" pageId="8" pageNumber="725">
<bibRefCitation id="68354B17FFCFFFC2FB51FB67FB0AFAC1" author="Ohaus, F." box="[1172,1277,1261,1280]" pageId="8" pageNumber="725" pagination="65 - 88" refId="ref11760" refString="Ohaus, F. 1944. Revision der Gattung Mimela Kirby (Col. Scarab. Rutelin.). Deutsche Entomologische Zeitschrift 1943: 65 - 88." type="journal article" year="1944">Ohaus 1944</bibRefCitation>
;
<bibRefCitation id="68354B17FFCFFFC2FAC2FB67FA5DFAC1" author="Machatschke, J. W." box="[1287,1450,1261,1280]" pageId="8" pageNumber="725" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
;
<bibRefCitation id="68354B17FFCFFFC2FA71FB67F9F4FAC1" author="Lin P." box="[1460,1539,1261,1280]" pageId="8" pageNumber="725" refId="ref11247" refString="Lin P. 1993. A Systematic Revision of the China Mimela: (Coleoptera: Rutelidae). The Publishing Company of Zhong Shan University, Zhongshan, China. [in Chinese and English]." type="book" year="1993">Lin 1993</bibRefCitation>
Ramirez-Ponce 2015 Morón and Nogueira 1998
<bibRefCitation id="68354B17FFCFFFC2FB51FABFFAC3FA89" author="Machatschke, J. W." box="[1172,1332,1333,1352]" pageId="8" pageNumber="725" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCF0035FF52FAC6F9BBFA51" box="[151,1612,1356,1424]" gridrow="7" pageId="8" pageNumber="725">
<th id="F1455DD8FFCF0035FF52FAC6FDA2FA51" box="[151,597,1356,1424]" gridcol="0" gridrow="7" pageId="8" pageNumber="725">
<taxonomicName id="CBA44D65FFCFFFC2FF52FAC7FE6DFAA1" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[151,410,1357,1376]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FAC7FEF1FAA1" box="[151,262,1357,1376]" italics="true" pageId="8" pageNumber="725">Pachystethus</emphasis>
Blanchard, 1851
</taxonomicName>
<taxonomicName id="CBA44D65FFCFFFC2FF52FAEFFEF7FAB9" authorityName="Casey" authorityYear="1915" box="[151,256,1381,1400]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FF52FAEFFEF7FAB9" box="[151,256,1381,1400]" italics="true" pageId="8" pageNumber="725">Paranomala</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCFFFC2FECBFAEFFE10FAB9" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[270,487,1381,1400]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="725" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCFFFC2FECBFAEFFE8FFAB9" box="[270,376,1381,1400]" italics="true" pageId="8" pageNumber="725">Paranomala</emphasis>
Casey, 1915
</taxonomicName>
)3, 4
</th>
<td id="F1455DD8FFCF0035FD1AFAC6FBBDFA51" box="[735,1098,1356,1424]" gridcol="1" gridrow="7" pageId="8" pageNumber="725">NTR NAR, NTR</td>
<td id="F1455DD8FFCF0035FB51FAC6F9BBFA51" box="[1172,1612,1356,1424]" gridcol="2" gridrow="7" pageId="8" pageNumber="725">
Ram´ırez-Ponce and Morón 2009, 2012 Morón and Ram´ırez-Ponce 2012;
<bibRefCitation id="68354B17FFCFFFC2FB6CFAF6FA67FA51" author="Ram'irez-Ponce, A." box="[1193,1424,1404,1424]" pageId="8" pageNumber="725" pagination="1054 - 1060" refId="ref11948" refString="Ram'irez-Ponce, A., A. Bitar, and D. Curoe. 2014. Two new species of Paranomala (Coleoptera: Melolonthidae: Rutelinae: Anomalini) from Mexico and Costa Rica with remarkable modifications on the pronotum. Revista Mexicana de Biodiversidad 85: 1054 - 1060." type="journal article" year="2014">
Ram´ırez-Ponce
<emphasis id="3ED0EAF4FFCFFFC2FAF4FAF7FAA9FA51" box="[1329,1374,1405,1424]" italics="true" pageId="8" pageNumber="725">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="68354B17FFCFFFC2FA5FFAF6F9BBFA51" author="Ram'irez-Ponce, A." box="[1434,1612,1404,1424]" pageId="8" pageNumber="725" pagination="261 - 266" refId="ref11921" refString="Ram'irez-Ponce, A. 2015. Moroniella, un nuevo genero americano de Anomalini (Coleoptera: Melolonthidae). Dugesiana 22: 261 - 266." type="journal article" year="2015">Ram´ırez-Ponce 2015</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<paragraph id="0C1B36E6FFCFFFC2FA01FA29F965FA77" blockId="8.[1476,1682,1443,1462]" box="[1476,1682,1443,1462]" pageId="8" pageNumber="725">
<tableNote id="F1423768FFCFFFC2FA01FA29F965FA77" box="[1476,1682,1443,1462]" pageId="8" pageNumber="725" targetBox="[151,1612,531,1424]" targetPageId="8">Continued on next page</tableNote>
</paragraph>
<caption id="58DB666EFFCEFFC3FF6AFDADFE9CFDFB" box="[175,363,551,570]" isContinuationCaption="true" pageId="9" pageNumber="726" startId="9.[175,225,551,570]" targetBox="[151,1682,602,1040]" targetIsTable="true" targetPageId="9">
<paragraph id="0C1B36E6FFCEFFC3FF6AFDADFE9CFDFB" blockId="9.[175,363,551,570]" box="[175,363,551,570]" pageId="9" pageNumber="726">
<emphasis id="3ED0EAF4FFCEFFC3FF6AFDADFF00FDFB" bold="true" box="[175,247,551,570]" pageId="9" pageNumber="726">Table 1.</emphasis>
Continued.
</paragraph>
</caption>
<paragraph id="0C1B36E6FFCEFFC3FE81FDD0F9E1FBD1" pageId="9" pageNumber="726">
<table id="7EA4C446FFCE0035FF52FDD0F965FBD1" box="[151,1682,602,1040]" colsContinueFrom="8.[151,1612,531,1424]" gridcols="3" gridrows="16" pageId="9" pageNumber="726">
<tr id="B29434A4FFCE0035FF52FDD0F965FDAC" box="[151,1682,602,621]" gridrow="0" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FDD0FD77FDAC" box="[151,640,602,621]" gridcol="0" gridrow="0" pageId="9" pageNumber="726">
<emphasis id="3ED0EAF4FFCEFFC3FE81FDD0FE1FFDAC" bold="true" box="[324,488,602,621]" pageId="9" pageNumber="726">Genus (Subgenus)</emphasis>
</th>
<th id="F1455DD8FFCE0035FD1AFDD0FC02FDAC" box="[735,1013,602,621]" gridcol="1" gridrow="0" pageId="9" pageNumber="726">
<emphasis id="3ED0EAF4FFCEFFC3FCF1FDD0FC02FDAC" bold="true" box="[820,1013,602,621]" pageId="9" pageNumber="726">Biogeographic region</emphasis>
</th>
<th id="F1455DD8FFCE0035FB51FDD0F965FDAC" box="[1172,1682,602,621]" gridcol="2" gridrow="0" pageId="9" pageNumber="726">
<emphasis id="3ED0EAF4FFCEFFC3FAA7FDD0FA34FDAC" bold="true" box="[1378,1475,602,621]" pageId="9" pageNumber="726">References</emphasis>
</th>
</tr>
<tr id="B29434A4FFCE0035FF52FDF4F965FD53" box="[151,1682,638,658]" gridrow="1" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FDF4FD77FD53" box="[151,640,638,658]" gridcol="0" gridrow="1" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FDF5FEF7FD53" authorityName="Casey" authorityYear="1915" box="[151,256,639,658]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FDF5FEF7FD53" box="[151,256,639,658]" italics="true" pageId="9" pageNumber="726">Paranomala</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCEFFC3FECBFDF5FE7AFD53" authorityName="Ramirez-Ponce &amp; Moron" authorityYear="2012" box="[270,397,639,658]" class="Insecta" family="Rutelidae" genus="Bucaphallanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FECBFDF5FE7AFD53" box="[270,397,639,658]" italics="true" pageId="9" pageNumber="726">Bucaphallanus</emphasis>
</taxonomicName>
Ram´ırez-Ponce and Morón,
</th>
<td id="F1455DD8FFCE0035FD1AFDF4FC02FD53" box="[735,1013,638,658]" gridcol="1" gridrow="1" pageId="9" pageNumber="726">NAR, NTR</td>
<td id="F1455DD8FFCE0035FB51FDF4F965FD53" box="[1172,1682,638,658]" gridcol="2" gridrow="1" pageId="9" pageNumber="726">
Ram´ırez-Ponce and
<bibRefCitation id="68354B17FFCEFFC3FA84FDF5FA5CFD53" author="Moron, M. A." box="[1345,1451,639,658]" pageId="9" pageNumber="726" pagination="97 - 114" refId="ref11688" refString="Moron, M. A., and A. Ram'irez-Ponce. 2012. Mesoamerican genera of Anomalini (Coleoptera: Melolonthidae: Rutelinae): A brief review. Trends in Entomology 8: 97 - 114." type="journal article" year="2012">Morón 2012</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FD1DF965FD6B" box="[151,1682,663,682]" gridrow="2" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FD1DF965FD6B" box="[151,1682,663,682]" colspan="3" colspanRight="2" gridcol="0" gridrow="2" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FF69FD1DFF2FFD6B" author="Moron, M. A." box="[172,216,663,682]" pageId="9" pageNumber="726" pagination="97 - 114" refId="ref11688" refString="Moron, M. A., and A. Ram'irez-Ponce. 2012. Mesoamerican genera of Anomalini (Coleoptera: Melolonthidae: Rutelinae): A brief review. Trends in Entomology 8: 97 - 114." type="journal article" year="2012">2012</bibRefCitation>
)
</th>
</tr>
<tr id="B29434A4FFCE0035FF52FD25F965FD1B" box="[151,1682,687,730]" gridrow="3" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FD25FD77FD1B" box="[151,640,687,730]" gridcol="0" gridrow="3" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FD25FE8CFD03" authority="Ohaus, 1919" authorityName="Ohaus" authorityYear="1919" box="[151,379,687,706]" class="Insecta" family="Rutelidae" genus="Pentanomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FD25FEF0FD03" box="[151,263,687,706]" italics="true" pageId="9" pageNumber="726">Pentanomala</emphasis>
Ohaus, 1919
</taxonomicName>
<taxonomicName id="CBA44D65FFCEFFC3FF52FD4DFE7AFD1B" authority="Stephens, 1830" authorityName="Stephens" authorityYear="1830" box="[151,397,711,730]" class="Insecta" family="Rutelidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FD4DFEF2FD1B" box="[151,261,711,730]" italics="true" pageId="9" pageNumber="726">Phyllopertha</emphasis>
Stephens, 1830
</taxonomicName>
5
</th>
<td id="F1455DD8FFCE0035FD1AFD25FC02FD1B" box="[735,1013,687,730]" gridcol="1" gridrow="3" pageId="9" pageNumber="726">AFR PAL, ORR</td>
<td id="F1455DD8FFCE0035FB51FD25F965FD1B" box="[1172,1682,687,730]" gridcol="2" gridrow="3" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FD25FAC3FD03" author="Machatschke, J. W." box="[1172,1332,687,706]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
<bibRefCitation id="68354B17FFCEFFC3FB51FD4DFAC3FD1B" author="Machatschke, J. W." box="[1172,1332,711,730]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FD54F965FD30" box="[151,1682,734,753]" gridrow="4" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FD54FD77FD30" box="[151,640,734,753]" gridcol="0" gridrow="4" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FD54FE75FD30" authority="Reitter, 1903" authorityName="Reitter. The" authorityYear="1903" box="[151,386,734,753]" class="Insecta" family="Rutelidae" genus="Proagopertha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FD54FEFAFD30" box="[151,269,734,753]" italics="true" pageId="9" pageNumber="726">Proagopertha</emphasis>
Reitter, 1903
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFD54FC02FD30" box="[735,1013,734,753]" gridcol="1" gridrow="4" pageId="9" pageNumber="726">PAL</td>
<td id="F1455DD8FFCE0035FB51FD54F965FD30" box="[1172,1682,734,753]" gridcol="2" gridrow="4" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FD54FAC3FD30" author="Machatschke, J. W." box="[1172,1332,734,753]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FD7CF965FCC8" box="[151,1682,758,777]" gridrow="5" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FD7CFD77FCC8" box="[151,640,758,777]" gridcol="0" gridrow="5" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FD7CFE78FCC8" authority="Heller, 1891" authorityName="Heller" authorityYear="1891" box="[151,399,758,777]" class="Insecta" family="Rutelidae" genus="Pseudosinghala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FD7CFEE9FCC8" box="[151,286,758,777]" italics="true" pageId="9" pageNumber="726">Pseudosinghala</emphasis>
Heller, 1891
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFD7CFC02FCC8" box="[735,1013,758,777]" gridcol="1" gridrow="5" pageId="9" pageNumber="726">ORR, PAL</td>
<td id="F1455DD8FFCE0035FB51FD7CF965FCC8" box="[1172,1682,758,777]" gridcol="2" gridrow="5" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FD7CFAC1FCC8" author="Machatschke, J. W." box="[1172,1334,758,777]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
; Kobayashi and Fujioka 2013
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FC84F965FCE0" box="[151,1682,782,801]" gridrow="6" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FC84FD77FCE0" box="[151,640,782,801]" gridcol="0" gridrow="6" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FC84FE4DFCE0" authority="Keith, 2007" authorityName="Keith" authorityYear="2007" box="[151,442,782,801]" class="Insecta" family="Rutelidae" genus="Pseudotrigonocnemis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FC84FEBAFCE0" box="[151,333,782,801]" italics="true" pageId="9" pageNumber="726">Pseudotrigonocnemis</emphasis>
<bibRefCitation id="68354B17FFCEFFC3FE91FC84FE4DFCE0" author="Keith, D." box="[340,442,782,801]" pageId="9" pageNumber="726" pagination="466 - 470" refId="ref11034" refString="Keith, D. 2007. Un nouveau genre de Rutelinae du Vietnam. Lambillionea 107: 466 - 470." type="journal article" year="2007">Keith, 2007</bibRefCitation>
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFC84FC02FCE0" box="[735,1013,782,801]" gridcol="1" gridrow="6" pageId="9" pageNumber="726">ORR</td>
<td id="F1455DD8FFCE0035FB51FC84F965FCE0" box="[1172,1682,782,801]" gridcol="2" gridrow="6" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FC84FB03FCE0" author="Keith, D." box="[1172,1268,782,801]" pageId="9" pageNumber="726" pagination="466 - 470" refId="ref11034" refString="Keith, D. 2007. Un nouveau genre de Rutelinae du Vietnam. Lambillionea 107: 466 - 470." type="journal article" year="2007">Keith 2007</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FCACF965FCF8" box="[151,1682,806,825]" gridrow="7" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FCACFD77FCF8" box="[151,640,806,825]" gridcol="0" gridrow="7" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FCACFE50FCF8" authority="Machatschke, 1957" authorityName="Machatschke" authorityYear="1957" box="[151,423,806,825]" class="Insecta" family="Rutelidae" genus="Rugopertha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FCACFF0CFCF8" box="[151,251,806,825]" italics="true" pageId="9" pageNumber="726">Rugopertha</emphasis>
<bibRefCitation id="68354B17FFCEFFC3FEC4FCACFE50FCF8" author="Machatschke, J. W." box="[257,423,806,825]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke, 1957</bibRefCitation>
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFCACFC02FCF8" box="[735,1013,806,825]" gridcol="1" gridrow="7" pageId="9" pageNumber="726">NTR</td>
<td id="F1455DD8FFCE0035FB51FCACF965FCF8" box="[1172,1682,806,825]" gridcol="2" gridrow="7" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FCACFAB5FCF8" author="Jameson, M. L." box="[1172,1346,806,825]" pageId="9" pageNumber="726" pagination="415 - 432" refId="ref10890" refString="Jameson, M. L., A. Paucar-Cabrera, and A. Sol'is. 2003. Synopsis of the New World genera of Anomalini (Coleoptera: Scarabaeidae: Rutelinae) and description of a new genus from Costa Rica and Nicaragua. Annals of the Entomological Society of America 96: 415 - 432." type="journal article" year="2003">
Jameson
<emphasis id="3ED0EAF4FFCEFFC3FB26FCACFAE7FCF8" box="[1251,1296,806,825]" italics="true" pageId="9" pageNumber="726">et al.</emphasis>
2003
</bibRefCitation>
; Ram´ırez-Ponce and
<bibRefCitation id="68354B17FFCEFFC3FA3CFCACF993FCF8" author="Moron, M. A." box="[1529,1636,806,825]" pageId="9" pageNumber="726" pagination="97 - 114" refId="ref11688" refString="Moron, M. A., and A. Ram'irez-Ponce. 2012. Mesoamerican genera of Anomalini (Coleoptera: Melolonthidae: Rutelinae): A brief review. Trends in Entomology 8: 97 - 114." type="journal article" year="2012">Morón 2012</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FCB4F965FC90" box="[151,1682,830,849]" gridrow="8" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FCB4FD77FC90" box="[151,640,830,849]" gridcol="0" gridrow="8" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FCB4FE81FC90" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[151,374,830,849]" class="Insecta" family="Rutelidae" genus="Singhala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FCB4FF15FC90" box="[151,226,830,849]" italics="true" pageId="9" pageNumber="726">Singhala</emphasis>
Blanchard, 1851
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFCB4FC02FC90" box="[735,1013,830,849]" gridcol="1" gridrow="8" pageId="9" pageNumber="726">ORR</td>
<td id="F1455DD8FFCE0035FB51FCB4F965FC90" box="[1172,1682,830,849]" gridcol="2" gridrow="8" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FCB4FAC3FC90" author="Machatschke, J. W." box="[1172,1332,830,849]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FCDCF965FCA8" box="[151,1682,854,873]" gridrow="9" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FCDCFD77FCA8" box="[151,640,854,873]" gridcol="0" gridrow="9" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FCDCFE8DFCA8" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" box="[151,378,854,873]" class="Insecta" family="Rutelidae" genus="Spinanomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FCDCFEF0FCA8" box="[151,263,854,873]" italics="true" pageId="9" pageNumber="726">Spinanomala</emphasis>
Ohaus, 1910
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFCDCFC02FCA8" box="[735,1013,854,873]" gridcol="1" gridrow="9" pageId="9" pageNumber="726">ORR, PAL</td>
<td id="F1455DD8FFCE0035FB51FCDCF965FCA8" box="[1172,1682,854,873]" gridcol="2" gridrow="9" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FCDCFB12FCA8" author="Lin, P." box="[1172,1253,854,873]" pageId="9" pageNumber="726" pagination="184 - 187" refId="ref11193" refString="Lin, P. 1979. Two new species of Rutelidae from south China (Coleoptera). Acta Entomologica Sinica 22: 184 - 187." type="journal article" year="1979">Lin 1979</bibRefCitation>
;
<bibRefCitation id="68354B17FFCEFFC3FB2AFCDCFA8CFCA8" author="Kobayashi, H." box="[1263,1403,854,873]" pageId="9" pageNumber="726" pagination="37 - 48" refId="ref11055" refString="Kobayashi, H. 2012. Notes on the genus Spinanomala from the Oriental region. Kogane 13: 37 - 48." type="journal article" year="2012">Kobayashi 2012</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FCE4F965FC40" box="[151,1682,878,897]" gridrow="10" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FCE4FD77FC40" box="[151,640,878,897]" gridcol="0" gridrow="10" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FCE4FE6DFC40" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[151,410,878,897]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FCE4FEF7FC40" box="[151,256,878,897]" italics="true" pageId="9" pageNumber="726">Strigoderma</emphasis>
<bibRefCitation id="68354B17FFCEFFC3FEC2FCE4FE6DFC40" author="Burmeister, H." box="[263,410,878,897]" pageId="9" pageNumber="726" refId="ref10413" refString="Burmeister, H. 1844. Handbuch der Entomologie, Volume 4, Part 1. T. C. F. Enslin, Berlin, Germany." type="book" year="1844">Burmeister, 1844</bibRefCitation>
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFCE4FC02FC40" box="[735,1013,878,897]" gridcol="1" gridrow="10" pageId="9" pageNumber="726">NAR, NTR</td>
<td id="F1455DD8FFCE0035FB51FCE4F965FC40" box="[1172,1682,878,897]" gridcol="2" gridrow="10" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FCE4FB00FC40" author="Bader, A. M." box="[1172,1271,878,897]" pageId="9" pageNumber="726" pagination="269 - 330" refId="ref10243" refString="Bader, A. M. 1992. A review of the North and Central American Strigoderma (Coleoptera: Scarabaeidae). Transactions of the American Entomological Society 118 (2): 269 - 330." type="journal article" year="1992">Bader 1992</bibRefCitation>
;
<bibRefCitation id="68354B17FFCEFFC3FB3AFCE4FA50FC40" author="Katbeh-Bader, A." box="[1279,1447,878,897]" pageId="9" pageNumber="726" pagination="293 - 296" refId="ref11004" refString="Katbeh-Bader, A. 2000. Description of two new species of Strigoderma from Central America (Coleoptera, Rutelidae). Fragmenta Entomologica 32: 293 - 296." type="journal article" year="2000">Katbeh-Bader 2000</bibRefCitation>
; Delgado and Mora-Aguilar
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FC0FF965FC58" box="[151,1682,901,921]" gridrow="11" pageId="9" pageNumber="726" rowspan-0="1" rowspan-1="1">
<td id="F1455DD8FFCE0035FB51FC0FF965FC58" box="[1172,1682,901,921]" gridcol="2" gridrow="11" pageId="9" pageNumber="726">2012; Ram´ırez-Ponce and Curoe 2014</td>
</tr>
<tr id="B29434A4FFCE0035FF52FC14F965FC70" box="[151,1682,926,945]" gridrow="12" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FC14FD77FC70" box="[151,640,926,945]" gridcol="0" gridrow="12" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FC14FE8BFC70" authority="Kolbe, 1897" authorityName="Kolbe" authorityYear="1897" box="[151,380,926,945]" class="Insecta" family="Rutelidae" genus="Stomanomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FC14FEFCFC70" box="[151,267,926,945]" italics="true" pageId="9" pageNumber="726">Stomanomala</emphasis>
Kolbe, 1897
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFC14FC02FC70" box="[735,1013,926,945]" gridcol="1" gridrow="12" pageId="9" pageNumber="726">AFR, PAL</td>
<td id="F1455DD8FFCE0035FB51FC14F965FC70" box="[1172,1682,926,945]" gridcol="2" gridrow="12" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FC14FAC3FC70" author="Machatschke, J. W." box="[1172,1332,926,945]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FC3CF965FC08" box="[151,1682,950,969]" gridrow="13" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FC3CFD77FC08" box="[151,640,950,969]" gridcol="0" gridrow="13" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FC3CFE77FC08" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" box="[151,384,950,969]" class="Insecta" family="Rutelidae" genus="Trichanomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FC3CFEFAFC08" box="[151,269,950,969]" italics="true" pageId="9" pageNumber="726">Trichanomala</emphasis>
<bibRefCitation id="68354B17FFCEFFC3FED1FC3CFE77FC08" author="Arrow, G. J." box="[276,384,950,969]" pageId="9" pageNumber="726" refId="ref10166" refString="Arrow, G. J. 1917. The Fauna of British India, including Ceylon and Burma. Coleoptera, Lamellicornia, Part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor and Francis, London, UK." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFC3CFC02FC08" box="[735,1013,950,969]" gridcol="1" gridrow="13" pageId="9" pageNumber="726">ORR, PAL</td>
<td id="F1455DD8FFCE0035FB51FC3CF965FC08" box="[1172,1682,950,969]" gridcol="2" gridrow="13" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FC3CFAC1FC08" author="Machatschke, J. W." box="[1172,1334,950,969]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
;
<bibRefCitation id="68354B17FFCEFFC3FA85FC3CFA78FC08" author="Lin, P." box="[1344,1423,950,969]" pageId="9" pageNumber="726" pagination="184 - 187" refId="ref11193" refString="Lin, P. 1979. Two new species of Rutelidae from south China (Coleoptera). Acta Entomologica Sinica 22: 184 - 187." type="journal article" year="1979">Lin 1979</bibRefCitation>
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FC44F965FC39" box="[151,1682,974,1016]" gridrow="14" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FC44FD77FC39" box="[151,640,974,1016]" gridcol="0" gridrow="14" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FC44FECFFC39" authority="Kraatz, 1894 Xochicotlia Moron" authorityName="Kraatz" authorityYear="1894" class="Insecta" family="Rutelidae" genus="Trigonocnemis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FC44FEE3FC20" box="[151,276,974,993]" italics="true" pageId="9" pageNumber="726">Trigonocnemis</emphasis>
Kraatz, 1894
<emphasis id="3ED0EAF4FFCEFFC3FF52FC6FFF0FFC39" box="[151,248,997,1016]" italics="true" pageId="9" pageNumber="726">Xochicotlia</emphasis>
Morón
</taxonomicName>
and Ram´ırez-Ponce, 2012 6
</th>
<td id="F1455DD8FFCE0035FD1AFC44FC02FC39" box="[735,1013,974,1016]" gridcol="1" gridrow="14" pageId="9" pageNumber="726">PAL NTR</td>
<td id="F1455DD8FFCE0035FB51FC44F965FC39" box="[1172,1682,974,1016]" gridcol="2" gridrow="14" pageId="9" pageNumber="726">
<bibRefCitation id="68354B17FFCEFFC3FB51FC44FAC3FC20" author="Machatschke, J. W." box="[1172,1332,974,993]" pageId="9" pageNumber="726" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
Morón and Ram´ırez-Ponce 2012; Filippini
<emphasis id="3ED0EAF4FFCEFFC3F9C2FC6FF9C3FC39" box="[1543,1588,997,1016]" italics="true" pageId="9" pageNumber="726">et al.</emphasis>
2015
</td>
</tr>
<tr id="B29434A4FFCE0035FF52FC77F965FBD1" box="[151,1682,1021,1040]" gridrow="15" pageId="9" pageNumber="726">
<th id="F1455DD8FFCE0035FF52FC77FD77FBD1" box="[151,640,1021,1040]" gridcol="0" gridrow="15" pageId="9" pageNumber="726">
<taxonomicName id="CBA44D65FFCEFFC3FF52FC77FE00FBD1" authority="Moron and Nogueira, 1998" authorityName="Moron and Nogueira" authorityYear="1998" box="[151,503,1021,1040]" class="Insecta" family="Rutelidae" genus="Yaaxkumukia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FF52FC77FEF0FBD1" box="[151,263,1021,1040]" italics="true" pageId="9" pageNumber="726">Yaaxkumukia</emphasis>
Morón and Nogueira, 1998
</taxonomicName>
</th>
<td id="F1455DD8FFCE0035FD1AFC77FC02FBD1" box="[735,1013,1021,1040]" gridcol="1" gridrow="15" pageId="9" pageNumber="726">NTR</td>
<td id="F1455DD8FFCE0035FB51FC77F965FBD1" box="[1172,1682,1021,1040]" gridcol="2" gridrow="15" pageId="9" pageNumber="726">
Morón and Nogueira 1998;
<bibRefCitation id="68354B17FFCEFFC3FA40FC77F9E1FBD1" author="Mico, E." box="[1413,1558,1021,1040]" pageId="9" pageNumber="726" pagination="1 - 6" refId="ref11484" refString="Mico, E., B. Gomez, and E. Galante. 2006. The Mesoamerican genus Yaaxkumukia: Biogeography and descriptions of new species (Coleoptera: Rutelidae). Annals of the Entomological Society of America 99: 1 - 6." type="journal article" year="2006">
Micó
<emphasis id="3ED0EAF4FFCEFFC3FA7DFC77FA12FBD1" box="[1464,1509,1021,1040]" italics="true" pageId="9" pageNumber="726">et al.</emphasis>
2006
</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<tableNote id="F1423768FFCEFFC3FF52FBA2FD73FA96" pageId="9" pageNumber="726" targetBox="[151,1682,602,1040]" targetPageId="9">
<paragraph id="0C1B36E6FFCEFFC3FF52FBA2FEB2FB9A" blockId="9.[151,1683,1064,1367]" pageId="9" pageNumber="726">
<superScript id="FBD19BAEFFCEFFC3FF52FBA2FF68FBF5" attach="left" box="[151,159,1064,1076]" fontSize="5" pageId="9" pageNumber="726">1</superScript>
Ram´ırez-Ponce and Morón (2009) considered
<taxonomicName id="CBA44D65FFCEFFC3FDE8FBA7FCF2FB81" authority="Saylor, 1935" authorityName="Saylor" authorityYear="1935" box="[557,773,1069,1088]" class="Insecta" family="Rutelidae" genus="Leptohoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FDE8FBA7FD64FB81" box="[557,659,1069,1088]" italics="true" pageId="9" pageNumber="726">Leptohoplia</emphasis>
Saylor, 1935
</taxonomicName>
as a member of
<taxonomicName id="CBA44D65FFCEFFC3FC57FBA7FC29FB81" authorityName="Samouelle" authorityYear="1819" box="[914,990,1069,1088]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FC57FBA7FC29FB81" box="[914,990,1069,1088]" italics="true" pageId="9" pageNumber="726">Anomala</emphasis>
</taxonomicName>
. This action also resulted in the synonymy of the subtribe
<taxonomicName id="CBA44D65FFCEFFC3FA14FBA7F9B0FB81" authorityName="Potts" authorityYear="1974" box="[1489,1607,1069,1088]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="subTribe" subTribe="Leptohopliina">Leptohopliina</taxonomicName>
with the subtribe
<taxonomicName id="CBA44D65FFCEFFC3FF24FBC2FEB6FB9A" authorityName="Streubel" authorityYear="1839" box="[225,321,1096,1115]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="9" pageNumber="726" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
.
</paragraph>
<paragraph id="0C1B36E6FFCEFFC3FF52FBECFA01FB59" blockId="9.[151,1683,1064,1367]" pageId="9" pageNumber="726">
<superScript id="FBD19BAEFFCEFFC3FF52FBECFF68FBB3" attach="left" box="[151,159,1126,1138]" fontSize="5" pageId="9" pageNumber="726">2</superScript>
Morón and Ram´ırez-Ponce (2012) stated that
<taxonomicName id="CBA44D65FFCEFFC3FDE4FBE0FD8DFBBC" authorityName="Moron and Nogueira" authorityYear="1998" box="[545,634,1130,1149]" class="Insecta" family="Rutelidae" genus="Chelilabia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FDE4FBE0FD8DFBBC" box="[545,634,1130,1149]" italics="true" pageId="9" pageNumber="726">Chelilabia</emphasis>
</taxonomicName>
was a subgenus of
<taxonomicName id="CBA44D65FFCEFFC3FCDFFBE0FC73FBBC" authorityName="Casey" authorityYear="1915" box="[794,900,1130,1149]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FCDFFBE0FC73FBBC" box="[794,900,1130,1149]" italics="true" pageId="9" pageNumber="726">Paranomala</emphasis>
</taxonomicName>
(also referencing Ram´ırez-Ponce and
<bibRefCitation id="68354B17FFCEFFC3FB07FBE0FADAFBBC" author="Moron, M. A." box="[1218,1325,1130,1149]" pageId="9" pageNumber="726" pagination="97 - 114" refId="ref11688" refString="Moron, M. A., and A. Ram'irez-Ponce. 2012. Mesoamerican genera of Anomalini (Coleoptera: Melolonthidae: Rutelinae): A brief review. Trends in Entomology 8: 97 - 114." type="journal article" year="2012">Morón 2012</bibRefCitation>
), but no taxonomic change has been made. They also stated that
<taxonomicName id="CBA44D65FFCEFFC3FE8AFB0FFE42FB59" authorityName="Saylor" authorityYear="1935" box="[335,437,1157,1176]" class="Insecta" family="Rutelidae" genus="Leptohoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FE8AFB0FFE42FB59" box="[335,437,1157,1176]" italics="true" pageId="9" pageNumber="726">Leptohoplia</emphasis>
</taxonomicName>
was a subgenus of
<taxonomicName id="CBA44D65FFCEFFC3FDA4FB0FFD3CFB59" authorityName="Casey" authorityYear="1915" box="[609,715,1157,1176]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FDA4FB0FFD3CFB59" box="[609,715,1157,1176]" italics="true" pageId="9" pageNumber="726">Paranomala</emphasis>
</taxonomicName>
(also referencing Ram´ırez-Ponce and
<bibRefCitation id="68354B17FFCEFFC3FBD1FB0FFB76FB59" author="Moron, M. A." box="[1044,1153,1157,1176]" pageId="9" pageNumber="726" pagination="97 - 114" refId="ref11688" refString="Moron, M. A., and A. Ram'irez-Ponce. 2012. Mesoamerican genera of Anomalini (Coleoptera: Melolonthidae: Rutelinae): A brief review. Trends in Entomology 8: 97 - 114." type="journal article" year="2012">Morón 2012</bibRefCitation>
), but no taxonomic change has been made.
</paragraph>
<paragraph id="0C1B36E6FFCEFFC3FF52FB29FB80FB14" blockId="9.[151,1683,1064,1367]" pageId="9" pageNumber="726">
<superScript id="FBD19BAEFFCEFFC3FF52FB29FF68FB6E" attach="left" box="[151,159,1187,1199]" fontSize="5" pageId="9" pageNumber="726">3</superScript>
According to Morón and Ram´ırez-Ponce (2012),
<taxonomicName id="CBA44D65FFCEFFC3FD87FB2DFD5BFB7B" authorityName="Casey" authorityYear="1915" box="[578,684,1191,1210]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FD87FB2DFD5BFB7B" box="[578,684,1191,1210]" italics="true" pageId="9" pageNumber="726">Paranomala</emphasis>
</taxonomicName>
is widely distributed in the Americas, which likely includes many species currently classified as
<taxonomicName id="CBA44D65FFCEFFC3FA1DFB2DF9D3FB7B" authorityName="Samouelle" authorityYear="1819" box="[1496,1572,1191,1210]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FA1DFB2DF9D3FB7B" box="[1496,1572,1191,1210]" italics="true" pageId="9" pageNumber="726">Anomala</emphasis>
</taxonomicName>
. The authors also include Africa, India, and Southeast Asia in the distributional data of
<taxonomicName id="CBA44D65FFCEFFC3FCD2FB48FC76FB14" authorityName="Casey" authorityYear="1915" box="[791,897,1218,1237]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FCD2FB48FC76FB14" box="[791,897,1218,1237]" italics="true" pageId="9" pageNumber="726">Paranomala</emphasis>
</taxonomicName>
without further explanation.
</paragraph>
<paragraph id="0C1B36E6FFCEFFC3FF52FB6AFC37FB39" blockId="9.[151,1683,1064,1367]" box="[151,960,1248,1272]" pageId="9" pageNumber="726">
<superScript id="FBD19BAEFFCEFFC3FF52FB6AFF68FB2D" attach="left" box="[151,159,1248,1260]" fontSize="5" pageId="9" pageNumber="726">4</superScript>
Ram´ırez-Ponce and Morón (2009) synonymized
<taxonomicName id="CBA44D65FFCEFFC3FD82FB6FFCD7FB39" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[583,800,1253,1272]" class="Insecta" family="Rutelidae" genus="Anomalacra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FD82FB6FFD58FB39" box="[583,687,1253,1272]" italics="true" pageId="9" pageNumber="726">Anomalacra</emphasis>
Casey, 1915
</taxonomicName>
with
<taxonomicName id="CBA44D65FFCEFFC3FC97FB6FFC4CFB39" authorityName="Casey" authorityYear="1915" box="[850,955,1253,1272]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCEFFC3FC97FB6FFC4CFB39" box="[850,955,1253,1272]" italics="true" pageId="9" pageNumber="726">Paranomala</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0C1B36E6FFCEFFC3FF52FA88FA5BFADB" blockId="9.[151,1683,1064,1367]" box="[151,1452,1282,1306]" pageId="9" pageNumber="726">
<superScript id="FBD19BAEFFCEFFC3FF52FA88FF68FACF" attach="left" box="[151,159,1282,1294]" fontSize="5" pageId="9" pageNumber="726">5</superScript>
<taxonomicName id="CBA44D65FFCEFFC3FF61FA8DFE42FADB" authority="Ohaus" authorityName="Ohaus" authorityYear="1905" box="[164,437,1287,1306]" class="Insecta" family="Rutelidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="726" phylum="Arthropoda" rank="species" species="carinicollis">
<emphasis id="3ED0EAF4FFCEFFC3FF61FA8DFE80FADB" box="[164,375,1287,1306]" italics="true" pageId="9" pageNumber="726">Phyllopertha carinicollis</emphasis>
Ohaus
</taxonomicName>
from Vietnam is the only species of the genus in the Oriental Region and may be incorrectly classified in that genus.
</paragraph>
<paragraph id="0C1B36E6FFCEFFC3FF52FAAFFD73FA96" blockId="9.[151,1683,1064,1367]" pageId="9" pageNumber="726">
<superScript id="FBD19BAEFFCEFFC3FF52FAAFFF68FAF0" attach="left" box="[151,159,1317,1329]" fontSize="5" pageId="9" pageNumber="726">6</superScript>
Although Morón and Ram´ırez-Ponce (2012) referred to
<emphasis id="3ED0EAF4FFCEFFC3FDB3FAA0FD20FAFC" box="[630,727,1322,1341]" italics="true" pageId="9" pageNumber="726">Xochicotlia</emphasis>
as “unpublished”, they provided all requisite information required by ICZN Article 13.1 (type species designation, description, images), thus making this an available name.
</paragraph>
</tableNote>
<caption id="58DB666EFFCDFFC0FF6AFE69FB3CFE37" ID-Table-UUID="58DB666EFFCDFFC0FF6AFE69FB3CFE37" box="[175,1227,483,502]" httpUri="http://table.plazi.org/id/58DB666EFFCDFFC0FF6AFE69FB3CFE37" pageId="10" pageNumber="727" startId="10.[175,225,483,502]" targetBox="[151,1682,530,1091]" targetIsTable="true" targetPageId="10">
<paragraph id="0C1B36E6FFCDFFC0FF6AFE69FB3CFE37" blockId="10.[175,1227,483,502]" box="[175,1227,483,502]" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FF6AFE69FF00FE37" bold="true" box="[175,247,483,502]" pageId="10" pageNumber="727">Table 2.</emphasis>
Comparative characters and character states for Old World
<taxonomicName id="CBA44D65FFCDFFC0FCCCFE69FC91FE37" authorityName="Streubel" authorityYear="1839" box="[777,870,483,502]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="10" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
taxa that possess similar character states.
</paragraph>
</caption>
<paragraph id="0C1B36E6FFCDFFC0FCD7FD98F9BDFB82" pageId="10" pageNumber="727">
<table id="7EA4C446FFCD0035FF52FD98F965FB82" box="[151,1682,530,1091]" gridcols="7" gridrows="25" pageId="10" pageNumber="727">
<tr id="B29434A4FFCD0035FF52FD98F965FDE3" box="[151,1682,530,546]" gridrow="0" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="F1455DD8FFCD0035FCCBFD98FC58FDE3" box="[782,943,530,546]" gridcol="3" gridrow="0" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FCD7FD98FC5CFDE3" bold="true" box="[786,939,530,546]" pageId="10" pageNumber="727">Male protarsomere</emphasis>
</th>
</tr>
<tr id="B29434A4FFCD0035FF52FDA2F965FDF9" box="[151,1682,552,568]" gridrow="1" pageId="10" pageNumber="727" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="F1455DD8FFCD0035FEB0FDA2FDE1FDF9" box="[373,534,552,568]" gridcol="1" gridrow="1" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FE48FDA2FE09FDF9" bold="true" box="[397,510,552,568]" pageId="10" pageNumber="727">Reflexed male</emphasis>
</td>
<td id="F1455DD8FFCD0035FCCBFDA2FC58FDF9" box="[782,943,552,568]" gridcol="3" gridrow="1" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FC83FDA2FC8FFDF9" bold="true" box="[838,888,552,568]" pageId="10" pageNumber="727">5 with</emphasis>
</td>
</tr>
<tr id="B29434A4FFCD0035FF52FDB7F965FD8C" box="[151,1682,573,589]" gridrow="2" pageId="10" pageNumber="727" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="F1455DD8FFCD0035FEB0FDB7FDE1FD8C" box="[373,534,573,589]" gridcol="1" gridrow="2" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FE45FDB7FDFCFD8C" bold="true" box="[384,523,573,589]" pageId="10" pageNumber="727">clypeal apex with</emphasis>
</td>
<td id="F1455DD8FFCD0035FCCBFDB7FC58FD8C" box="[782,943,573,589]" gridcol="3" gridrow="2" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FCE0FDB7FC6EFD8C" bold="true" box="[805,921,573,589]" pageId="10" pageNumber="727">well-developed</emphasis>
</td>
</tr>
<tr id="B29434A4FFCD0035FF52FDD8F965FDA3" box="[151,1682,594,610]" gridrow="3" pageId="10" pageNumber="727" rowspan-0="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<td id="F1455DD8FFCD0035FEB0FDD8FDE1FDA3" box="[373,534,594,610]" gridcol="1" gridrow="3" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FEB0FDD8FDE1FDA3" bold="true" box="[373,534,594,610]" pageId="10" pageNumber="727">square anterolateral</emphasis>
</td>
<td id="F1455DD8FFCD0035FCCBFDD8FC58FDA3" box="[782,943,594,610]" gridcol="3" gridrow="3" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FCEDFDD8FC61FDA3" bold="true" box="[808,918,594,610]" pageId="10" pageNumber="727">internomedial</emphasis>
</td>
<td id="F1455DD8FFCD0035FC14FDD8FB70FDA3" box="[977,1159,594,610]" gridcol="4" gridrow="3" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FC34FDD8FB9FFDA3" bold="true" box="[1009,1128,594,610]" pageId="10" pageNumber="727">Male protarsal</emphasis>
</td>
</tr>
<tr id="B29434A4FFCD0035FF52FDEDF965FDB9" box="[151,1682,615,632]" gridrow="4" pageId="10" pageNumber="727" rowspan-0="1">
<td id="F1455DD8FFCD0035FEB0FDEDFDE1FDB9" box="[373,534,615,632]" gridcol="1" gridrow="4" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FE48FDE2FE09FDB9" bold="true" box="[397,510,616,632]" pageId="10" pageNumber="727">angles present</emphasis>
</td>
<td id="F1455DD8FFCD0035FDF2FDEDFD1AFDB9" box="[567,749,615,632]" gridcol="2" gridrow="4" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FD8DFDE2FD2CFDB9" bold="true" box="[584,731,616,632]" pageId="10" pageNumber="727">Protibia tridentate</emphasis>
</td>
<td id="F1455DD8FFCD0035FCCBFDEDFC58FDB9" box="[782,943,615,632]" gridcol="3" gridrow="4" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FCEFFDEDFC63FDB6" bold="true" box="[810,916,615,631]" pageId="10" pageNumber="727">protuberance</emphasis>
</td>
<td id="F1455DD8FFCD0035FC14FDEDFB70FDB9" box="[977,1159,615,632]" gridcol="4" gridrow="4" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FC38FDEDFBACFDB6" bold="true" box="[1021,1115,615,631]" pageId="10" pageNumber="727">claws entire</emphasis>
</td>
<td id="F1455DD8FFCD0035FB6DFDEDFAA7FDB9" box="[1192,1360,615,632]" gridcol="5" gridrow="4" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FB6BFDEDFABDFDB6" bold="true" box="[1198,1354,615,631]" pageId="10" pageNumber="727">Male protibial spur</emphasis>
</td>
<td id="F1455DD8FFCD0035FAB4FDEDF965FDB9" box="[1393,1682,615,632]" gridcol="6" gridrow="4" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FA56FDEDF986FDB6" bold="true" box="[1427,1649,615,631]" pageId="10" pageNumber="727">Male protarsal claw robust,</emphasis>
</td>
</tr>
<tr id="B29434A4FFCD0035FF52FDF7F965FD4C" box="[151,1682,637,653]" gridrow="5" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FDF7FEA4FD4C" box="[151,339,637,653]" gridcol="0" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FF18FDF7FEF9FD4C" bold="true" box="[221,270,637,653]" pageId="10" pageNumber="727">Taxon</emphasis>
</th>
<td id="F1455DD8FFCD0035FEB0FDF7FDE1FD4C" box="[373,534,637,653]" gridcol="1" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FE65FDF7FE1DFD4C" bold="true" box="[416,490,637,653]" pageId="10" pageNumber="727">or absent</emphasis>
</td>
<td id="F1455DD8FFCD0035FDF2FDF7FD1AFD4C" box="[567,749,637,653]" gridcol="2" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FD95FDF7FD23FD4C" bold="true" box="[592,724,637,653]" pageId="10" pageNumber="727">or not tridentate</emphasis>
</td>
<td id="F1455DD8FFCD0035FCCBFDF7FC58FD4C" box="[782,943,637,653]" gridcol="3" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FCDFFDF7FC53FD4C" bold="true" box="[794,932,637,653]" pageId="10" pageNumber="727">present or absent</emphasis>
</td>
<td id="F1455DD8FFCD0035FC14FDF7FB70FD4C" box="[977,1159,637,653]" gridcol="4" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FC23FDF7FB85FD4C" bold="true" box="[998,1138,637,653]" pageId="10" pageNumber="727">(not split) or split</emphasis>
</td>
<td id="F1455DD8FFCD0035FB6DFDF7FAA7FD4C" box="[1192,1360,637,653]" gridcol="5" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FB72FDF7FAB6FD4C" bold="true" box="[1207,1345,637,653]" pageId="10" pageNumber="727">present or absent</emphasis>
</td>
<td id="F1455DD8FFCD0035FAB4FDF7F965FD4C" box="[1393,1682,637,653]" gridcol="6" gridrow="5" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FA77FDF7F9A5FD4C" bold="true" box="[1458,1618,637,653]" pageId="10" pageNumber="727">strongly bent or not</emphasis>
</td>
</tr>
<tr id="B29434A4FFCD0035FF52FD15F965FD71" box="[151,1682,671,688]" gridrow="6" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FD15FEA4FD71" box="[151,339,671,688]" gridcol="0" gridrow="6" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FD15FEF3FD6E" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[151,260,671,687]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FD15FEF3FD6E" bold="true" box="[151,260,671,687]" italics="true" pageId="10" pageNumber="727">Ganganomala</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FD15FDE1FD71" box="[373,534,671,688]" gridcol="1" gridrow="6" pageId="10" pageNumber="727">Present (Figs. 24–25)</td>
<td id="F1455DD8FFCD0035FDF2FD15FD1AFD71" box="[567,749,671,688]" gridcol="2" gridrow="6" pageId="10" pageNumber="727">Tridentate (Figs. 31–32)</td>
<td id="F1455DD8FFCD0035FCCBFD15FC58FD71" box="[782,943,671,688]" gridcol="3" gridrow="6" pageId="10" pageNumber="727">Present (Fig. 36)</td>
<td id="F1455DD8FFCD0035FC14FD15FB70FD71" box="[977,1159,671,688]" gridcol="4" gridrow="6" pageId="10" pageNumber="727">Entire (Fig. 36)</td>
<td id="F1455DD8FFCD0035FB6DFD15FAA7FD71" box="[1192,1360,671,688]" gridcol="5" gridrow="6" pageId="10" pageNumber="727">Absent (Figs. 7, 31)</td>
<td id="F1455DD8FFCD0035FAB4FD15F965FD71" box="[1393,1682,671,688]" gridcol="6" gridrow="6" pageId="10" pageNumber="727">Robust, strongly bent (Figs. 7, 36)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FD3EF965FD04" box="[151,1682,692,709]" gridrow="7" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FD3EFEA4FD04" box="[151,339,692,709]" gridcol="0" gridrow="7" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FD3EFEE8FD05" box="[151,287,692,708]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCDFFC0FF52FD3EFEE8FD05" bold="true" box="[151,287,692,708]" italics="true" pageId="10" pageNumber="727">Anomala dorsalis</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FD3EFDE1FD04" box="[373,534,692,709]" gridcol="1" gridrow="7" pageId="10" pageNumber="727">Present (Figs. 29–30)</td>
<td id="F1455DD8FFCD0035FDF2FD3EFD1AFD04" box="[567,749,692,709]" gridcol="2" gridrow="7" pageId="10" pageNumber="727">Tridentate (Figs. 33–34)</td>
<td id="F1455DD8FFCD0035FCCBFD3EFC58FD04" box="[782,943,692,709]" gridcol="3" gridrow="7" pageId="10" pageNumber="727">Present (Fig. 37)</td>
<td id="F1455DD8FFCD0035FC14FD3EFB70FD04" box="[977,1159,692,709]" gridcol="4" gridrow="7" pageId="10" pageNumber="727">Split (Fig. 37)</td>
<td id="F1455DD8FFCD0035FB6DFD3EFAA7FD04" box="[1192,1360,692,709]" gridcol="5" gridrow="7" pageId="10" pageNumber="727">Present (Fig. 33)</td>
<td id="F1455DD8FFCD0035FAB4FD3EF965FD04" box="[1393,1682,692,709]" gridcol="6" gridrow="7" pageId="10" pageNumber="727">Robust, strongly bent (Fig. 37)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FD4CF965FD1B" box="[151,1682,710,730]" gridrow="8" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FD4CFEA4FD1B" box="[151,339,710,730]" gridcol="0" gridrow="8" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FD40FF16FD1B" authorityName="Laporte" authorityYear="1832" box="[151,225,714,730]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FD40FF16FD1B" bold="true" box="[151,225,714,730]" italics="true" pageId="10" pageNumber="727">Hoplopus</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FD4CFDE1FD1B" box="[373,534,710,730]" gridcol="1" gridrow="8" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FD4CFD1AFD1B" box="[567,749,710,730]" gridcol="2" gridrow="8" pageId="10" pageNumber="727">Tridentate1 (Fig. 35)</td>
<td id="F1455DD8FFCD0035FCCBFD4CFC58FD1B" box="[782,943,710,730]" gridcol="3" gridrow="8" pageId="10" pageNumber="727">Absent (Fig. 46)</td>
<td id="F1455DD8FFCD0035FC14FD4CFB70FD1B" box="[977,1159,710,730]" gridcol="4" gridrow="8" pageId="10" pageNumber="727">Entire (Fig. 46)</td>
<td id="F1455DD8FFCD0035FB6DFD4CFAA7FD1B" box="[1192,1360,710,730]" gridcol="5" gridrow="8" pageId="10" pageNumber="727">Absent2 (Figs. 35, 46)</td>
<td id="F1455DD8FFCD0035FAB4FD4CF965FD1B" box="[1393,1682,710,730]" gridcol="6" gridrow="8" pageId="10" pageNumber="727">Not robust and strongly</td>
</tr>
<tr id="B29434A4FFCD0035FF52FD55F965FD2E" box="[151,1682,735,751]" gridrow="9" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="F1455DD8FFCD0035FAB4FD55F965FD2E" box="[1393,1682,735,751]" gridcol="6" gridrow="9" pageId="10" pageNumber="727">bent (Fig. 46)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FD7EF965FCDB" box="[151,1682,756,794]" gridrow="10" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FD7EFEA4FCDB" box="[151,339,756,794]" gridcol="0" gridrow="10" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FD7EFF2AFCC5" authorityName="Samouelle" authorityYear="1819" box="[151,221,756,772]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FD7EFF2AFCC5" bold="true" box="[151,221,756,772]" italics="true" pageId="10" pageNumber="727">Anomala</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FD7EFDE1FCDB" box="[373,534,756,794]" gridcol="1" gridrow="10" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FD7EFD1AFCDB" box="[567,749,756,794]" gridcol="2" gridrow="10" pageId="10" pageNumber="727">Tridentate or not tridentate</td>
<td id="F1455DD8FFCD0035FCCBFD7EFC58FCDB" box="[782,943,756,794]" gridcol="3" gridrow="10" pageId="10" pageNumber="727">Present (Figs. 37–40) or absent (Fig. 43)</td>
<td id="F1455DD8FFCD0035FC14FD7EFB70FCDB" box="[977,1159,756,794]" gridcol="4" gridrow="10" pageId="10" pageNumber="727">Entire (Fig. 43) or split (Figs. 38–40)3</td>
<td id="F1455DD8FFCD0035FB6DFD7EFAA7FCDB" box="[1192,1360,756,794]" gridcol="5" gridrow="10" pageId="10" pageNumber="727">Present (Fig. 38)</td>
<td id="F1455DD8FFCD0035FAB4FD7EF965FCDB" box="[1393,1682,756,794]" gridcol="6" gridrow="10" pageId="10" pageNumber="727">Robust, strongly bent (Fig. 38) or not robust and strongly</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC95F965FCEE" box="[151,1682,799,815]" gridrow="11" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="F1455DD8FFCD0035FAB4FC95F965FCEE" box="[1393,1682,799,815]" gridcol="6" gridrow="11" pageId="10" pageNumber="727">bent (Fig. 40)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FCBEF965FC85" box="[151,1682,820,836]" gridrow="12" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FCBEFEA4FC85" box="[151,339,820,836]" gridcol="0" gridrow="12" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FCBEFEA4FC85" authority="" box="[151,339,820,836]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="subGenus" subGenus="Trichopertha">
<emphasis id="3ED0EAF4FFCDFFC0FF52FCBEFEA4FC85" bold="true" box="[151,339,820,836]" pageId="10" pageNumber="727">
<emphasis id="3ED0EAF4FFCDFFC0FF52FCBEFF2BFC85" bold="true" box="[151,220,820,836]" italics="true" pageId="10" pageNumber="727">Exomala</emphasis>
(
<emphasis id="3ED0EAF4FFCDFFC0FF2DFCBEFEBAFC85" bold="true" box="[232,333,820,836]" italics="true" pageId="10" pageNumber="727">Trichopertha</emphasis>
)
</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FCBEFDE1FC85" box="[373,534,820,836]" gridcol="1" gridrow="12" pageId="10" pageNumber="727">Present</td>
<td id="F1455DD8FFCD0035FDF2FCBEFD1AFC85" box="[567,749,820,836]" gridcol="2" gridrow="12" pageId="10" pageNumber="727">Not tridentate</td>
<td id="F1455DD8FFCD0035FCCBFCBEFC58FC85" box="[782,943,820,836]" gridcol="3" gridrow="12" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FC14FCBEFB70FC85" box="[977,1159,820,836]" gridcol="4" gridrow="12" pageId="10" pageNumber="727">Split</td>
<td id="F1455DD8FFCD0035FB6DFCBEFAA7FC85" box="[1192,1360,820,836]" gridcol="5" gridrow="12" pageId="10" pageNumber="727">Present</td>
<td id="F1455DD8FFCD0035FAB4FCBEF965FC85" box="[1393,1682,820,836]" gridcol="6" gridrow="12" pageId="10" pageNumber="727">Not robust and strongly bent</td>
</tr>
<tr id="B29434A4FFCD0035FF52FCCFF965FC98" box="[151,1682,837,857]" gridrow="13" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FCCFFEA4FC98" box="[151,339,837,857]" gridcol="0" gridrow="13" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FCC3FF2BFC98" authorityName="Blanchard" authorityYear="1851" box="[151,220,841,857]" class="Insecta" family="Rutelidae" genus="Singhala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FCC3FF2BFC98" bold="true" box="[151,220,841,857]" italics="true" pageId="10" pageNumber="727">Singhala</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FCCFFDE1FC98" box="[373,534,837,857]" gridcol="1" gridrow="13" pageId="10" pageNumber="727">Present (Fig. 28)</td>
<td id="F1455DD8FFCD0035FDF2FCCFFD1AFC98" box="[567,749,837,857]" gridcol="2" gridrow="13" pageId="10" pageNumber="727">Not tridentate</td>
<td id="F1455DD8FFCD0035FCCBFCCFFC58FC98" box="[782,943,837,857]" gridcol="3" gridrow="13" pageId="10" pageNumber="727">Present4 (Fig. 41)</td>
<td id="F1455DD8FFCD0035FC14FCCFFB70FC98" box="[977,1159,837,857]" gridcol="4" gridrow="13" pageId="10" pageNumber="727">Split (Fig. 41)</td>
<td id="F1455DD8FFCD0035FB6DFCCFFAA7FC98" box="[1192,1360,837,857]" gridcol="5" gridrow="13" pageId="10" pageNumber="727">Present or absent</td>
<td id="F1455DD8FFCD0035FAB4FCCFF965FC98" box="[1393,1682,837,857]" gridcol="6" gridrow="13" pageId="10" pageNumber="727">Not robust and strongly bent (Fig. 41)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FCD5F965FCAE" box="[151,1682,863,879]" gridrow="14" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="F1455DD8FFCD0035FB6DFCD5FAA7FCAE" box="[1192,1360,863,879]" gridcol="5" gridrow="14" pageId="10" pageNumber="727">(Fig. 41)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FCFAF965FC45" box="[151,1682,880,900]" gridrow="15" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FCFAFEA4FC45" box="[151,339,880,900]" gridcol="0" gridrow="15" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FCFEFF04FC45" authorityName="Reitter" authorityYear="1903" box="[151,243,884,900]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FCFEFF04FC45" bold="true" box="[151,243,884,900]" italics="true" pageId="10" pageNumber="727">Cyriopertha</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FCFAFDE1FC45" box="[373,534,880,900]" gridcol="1" gridrow="15" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FCFAFD1AFC45" box="[567,749,880,900]" gridcol="2" gridrow="15" pageId="10" pageNumber="727">Not tridentate</td>
<td id="F1455DD8FFCD0035FCCBFCFAFC58FC45" box="[782,943,880,900]" gridcol="3" gridrow="15" pageId="10" pageNumber="727">Absent (Fig. 45)</td>
<td id="F1455DD8FFCD0035FC14FCFAFB70FC45" box="[977,1159,880,900]" gridcol="4" gridrow="15" pageId="10" pageNumber="727">Entire or split (Fig. 45)5</td>
<td id="F1455DD8FFCD0035FB6DFCFAFAA7FC45" box="[1192,1360,880,900]" gridcol="5" gridrow="15" pageId="10" pageNumber="727">Absent (Fig. 45)</td>
<td id="F1455DD8FFCD0035FAB4FCFAF965FC45" box="[1393,1682,880,900]" gridcol="6" gridrow="15" pageId="10" pageNumber="727">Not robust and strongly</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC03F965FC58" box="[151,1682,905,921]" gridrow="16" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="F1455DD8FFCD0035FAB4FC03F965FC58" box="[1393,1682,905,921]" gridcol="6" gridrow="16" pageId="10" pageNumber="727">bent (Fig. 45)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC10F965FC6F" box="[151,1682,922,942]" gridrow="17" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FC10FEA4FC6F" box="[151,339,922,942]" gridcol="0" gridrow="17" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FC14FF03FC6F" authorityName="Reitter" authorityYear="1903" box="[151,244,926,942]" class="Insecta" family="Scarabaeidae" genus="Megapertha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FC14FF03FC6F" bold="true" box="[151,244,926,942]" italics="true" pageId="10" pageNumber="727">Megapertha</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FC10FDE1FC6F" box="[373,534,922,942]" gridcol="1" gridrow="17" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FC10FD1AFC6F" box="[567,749,922,942]" gridcol="2" gridrow="17" pageId="10" pageNumber="727">Not tridentate</td>
<td id="F1455DD8FFCD0035FCCBFC10FC58FC6F" box="[782,943,922,942]" gridcol="3" gridrow="17" pageId="10" pageNumber="727">Absent (Fig. 44)</td>
<td id="F1455DD8FFCD0035FC14FC10FB70FC6F" box="[977,1159,922,942]" gridcol="4" gridrow="17" pageId="10" pageNumber="727">Entire or split (Fig. 44)6</td>
<td id="F1455DD8FFCD0035FB6DFC10FAA7FC6F" box="[1192,1360,922,942]" gridcol="5" gridrow="17" pageId="10" pageNumber="727">Present (Fig. 44)</td>
<td id="F1455DD8FFCD0035FAB4FC10F965FC6F" box="[1393,1682,922,942]" gridcol="6" gridrow="17" pageId="10" pageNumber="727">Not robust and strongly</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC39F965FC02" box="[151,1682,947,963]" gridrow="18" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="F1455DD8FFCD0035FAB4FC39F965FC02" box="[1393,1682,947,963]" gridcol="6" gridrow="18" pageId="10" pageNumber="727">bent (Fig. 44)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC4FF965FC18" box="[151,1682,965,985]" gridrow="19" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FC4FFEA4FC18" box="[151,339,965,985]" gridcol="0" gridrow="19" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FC43FEF7FC18" authorityName="Reitter. The" authorityYear="1903" box="[151,256,969,985]" class="Insecta" family="Rutelidae" genus="Proagopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FC43FEF7FC18" bold="true" box="[151,256,969,985]" italics="true" pageId="10" pageNumber="727">Proagopertha</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FC4FFDE1FC18" box="[373,534,965,985]" gridcol="1" gridrow="19" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FC4FFD1AFC18" box="[567,749,965,985]" gridcol="2" gridrow="19" pageId="10" pageNumber="727">Not tridentate</td>
<td id="F1455DD8FFCD0035FCCBFC4FFC58FC18" box="[782,943,965,985]" gridcol="3" gridrow="19" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FC14FC4FFB70FC18" box="[977,1159,965,985]" gridcol="4" gridrow="19" pageId="10" pageNumber="727">Split</td>
<td id="F1455DD8FFCD0035FB6DFC4FFAA7FC18" box="[1192,1360,965,985]" gridcol="5" gridrow="19" pageId="10" pageNumber="727">Present or absent7</td>
<td id="F1455DD8FFCD0035FAB4FC4FF965FC18" box="[1393,1682,965,985]" gridcol="6" gridrow="19" pageId="10" pageNumber="727">Not robust and strongly bent</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC54F965FC2F" box="[151,1682,990,1006]" gridrow="20" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FC54FEA4FC2F" box="[151,339,990,1006]" gridcol="0" gridrow="20" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FC54FF25FC2F" authorityName="Kirby" authorityYear="1823" box="[151,210,990,1006]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FC54FF25FC2F" bold="true" box="[151,210,990,1006]" italics="true" pageId="10" pageNumber="727">Mimela</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FC54FDE1FC2F" box="[373,534,990,1006]" gridcol="1" gridrow="20" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FC54FD1AFC2F" box="[567,749,990,1006]" gridcol="2" gridrow="20" pageId="10" pageNumber="727">Tridentate or</td>
<td id="F1455DD8FFCD0035FCCBFC54FC58FC2F" box="[782,943,990,1006]" gridcol="3" gridrow="20" pageId="10" pageNumber="727">Absent (Fig. 42)</td>
<td id="F1455DD8FFCD0035FC14FC54FB70FC2F" box="[977,1159,990,1006]" gridcol="4" gridrow="20" pageId="10" pageNumber="727">Entire (Fig. 42) or split</td>
<td id="F1455DD8FFCD0035FB6DFC54FAA7FC2F" box="[1192,1360,990,1006]" gridcol="5" gridrow="20" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FAB4FC54F965FC2F" box="[1393,1682,990,1006]" gridcol="6" gridrow="20" pageId="10" pageNumber="727">Not robust and strongly</td>
</tr>
<tr id="B29434A4FFCD0035FF52FC79F965FBC2" box="[151,1682,1011,1027]" gridrow="21" pageId="10" pageNumber="727" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="F1455DD8FFCD0035FDF2FC79FD1AFBC2" box="[567,749,1011,1027]" gridcol="2" gridrow="21" pageId="10" pageNumber="727">not tridentate</td>
<td id="F1455DD8FFCD0035FAB4FC79F965FBC2" box="[1393,1682,1011,1027]" gridcol="6" gridrow="21" pageId="10" pageNumber="727">bent (Fig. 42)</td>
</tr>
<tr id="B29434A4FFCD0035FF52FB82F965FBD9" box="[151,1682,1032,1048]" gridrow="22" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FB82FEA4FBD9" box="[151,339,1032,1048]" gridcol="0" gridrow="22" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FB82FF08FBD9" authorityName="Arrow" authorityYear="1911" box="[151,255,1032,1048]" class="Insecta" family="Rutelidae" genus="Micranomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FB82FF08FBD9" bold="true" box="[151,255,1032,1048]" italics="true" pageId="10" pageNumber="727">Micranomala</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FB82FDE1FBD9" box="[373,534,1032,1048]" gridcol="1" gridrow="22" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FB82FD1AFBD9" box="[567,749,1032,1048]" gridcol="2" gridrow="22" pageId="10" pageNumber="727">Tridentate</td>
<td id="F1455DD8FFCD0035FCCBFB82FC58FBD9" box="[782,943,1032,1048]" gridcol="3" gridrow="22" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FC14FB82FB70FBD9" box="[977,1159,1032,1048]" gridcol="4" gridrow="22" pageId="10" pageNumber="727">Split</td>
<td id="F1455DD8FFCD0035FB6DFB82FAA7FBD9" box="[1192,1360,1032,1048]" gridcol="5" gridrow="22" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FAB4FB82F965FBD9" box="[1393,1682,1032,1048]" gridcol="6" gridrow="22" pageId="10" pageNumber="727">Not robust and strongly bent</td>
</tr>
<tr id="B29434A4FFCD0035FF52FB90F965FBEF" box="[151,1682,1050,1070]" gridrow="23" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FB90FEA4FBEF" box="[151,339,1050,1070]" gridcol="0" gridrow="23" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FB94FF00FBEF" authorityName="Petrovitz" authorityYear="1967" box="[151,247,1054,1070]" class="Insecta" family="Rutelidae" genus="Microlontha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FB94FF00FBEF" bold="true" box="[151,247,1054,1070]" italics="true" pageId="10" pageNumber="727">Microlontha</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FB90FDE1FBEF" box="[373,534,1050,1070]" gridcol="1" gridrow="23" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FB90FD1AFBEF" box="[567,749,1050,1070]" gridcol="2" gridrow="23" pageId="10" pageNumber="727">Tridentate</td>
<td id="F1455DD8FFCD0035FCCBFB90FC58FBEF" box="[782,943,1050,1070]" gridcol="3" gridrow="23" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FC14FB90FB70FBEF" box="[977,1159,1050,1070]" gridcol="4" gridrow="23" pageId="10" pageNumber="727">Entire8</td>
<td id="F1455DD8FFCD0035FB6DFB90FAA7FBEF" box="[1192,1360,1050,1070]" gridcol="5" gridrow="23" pageId="10" pageNumber="727">Absent8</td>
<td id="F1455DD8FFCD0035FAB4FB90F965FBEF" box="[1393,1682,1050,1070]" gridcol="6" gridrow="23" pageId="10" pageNumber="727">Not robust and strongly bent</td>
</tr>
<tr id="B29434A4FFCD0035FF52FBB9F965FB82" box="[151,1682,1075,1091]" gridrow="24" pageId="10" pageNumber="727">
<th id="F1455DD8FFCD0035FF52FBB9FEA4FB82" box="[151,339,1075,1091]" gridcol="0" gridrow="24" pageId="10" pageNumber="727">
<taxonomicName id="CBA44D65FFCDFFC0FF52FBB9FF0AFB82" box="[151,253,1075,1091]" class="Insecta" family="Rutelidae" genus="Pentanomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FBB9FF0AFB82" bold="true" box="[151,253,1075,1091]" italics="true" pageId="10" pageNumber="727">Pentanomala</emphasis>
</taxonomicName>
</th>
<td id="F1455DD8FFCD0035FEB0FBB9FDE1FB82" box="[373,534,1075,1091]" gridcol="1" gridrow="24" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FDF2FBB9FD1AFB82" box="[567,749,1075,1091]" gridcol="2" gridrow="24" pageId="10" pageNumber="727">Tridentate</td>
<td id="F1455DD8FFCD0035FCCBFBB9FC58FB82" box="[782,943,1075,1091]" gridcol="3" gridrow="24" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FC14FBB9FB70FB82" box="[977,1159,1075,1091]" gridcol="4" gridrow="24" pageId="10" pageNumber="727">Split</td>
<td id="F1455DD8FFCD0035FB6DFBB9FAA7FB82" box="[1192,1360,1075,1091]" gridcol="5" gridrow="24" pageId="10" pageNumber="727">Absent</td>
<td id="F1455DD8FFCD0035FAB4FBB9F965FB82" box="[1393,1682,1075,1091]" gridcol="6" gridrow="24" pageId="10" pageNumber="727">Not robust and strongly bent</td>
</tr>
</table>
</paragraph>
<tableNote id="F1423768FFCDFFC0FF52FBD6FCD4FA5A" pageId="10" pageNumber="727" targetBox="[151,1682,530,1091]" targetPageId="10">
<paragraph id="0C1B36E6FFCDFFC0FF52FBD6FD78FBB5" blockId="10.[151,1683,1116,1435]" box="[151,655,1116,1140]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FBD6FF68FBA9" attach="left" box="[151,159,1116,1128]" fontSize="5" pageId="10" pageNumber="727">1</superScript>
The third (basal) tooth is weekly developed in
<taxonomicName id="CBA44D65FFCDFFC0FDFDFBEBFD7DFBB5" authorityName="Laporte" authorityYear="1832" box="[568,650,1121,1140]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FDFDFBEBFD7DFBB5" box="[568,650,1121,1140]" italics="true" pageId="10" pageNumber="727">Hoplopus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FBF5FAC1FB56" blockId="10.[151,1683,1116,1435]" box="[151,1334,1151,1175]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FBF5FF68FB4A" attach="left" box="[151,159,1151,1163]" fontSize="5" pageId="10" pageNumber="727">2</superScript>
According to
<bibRefCitation id="68354B17FFCDFFC0FEDEFB0EFE6FFB56" author="Baraud, J." box="[283,408,1156,1175]" pageId="10" pageNumber="727" refId="ref10322" refString="Baraud, J. 1985. Coleopteres Scarabaeoidea. Faune du Nord de l' Afrique du Maroc au Sina ¨ i. Encyclopedie Entomologique - XLVI. Editions Lechevalier, Paris, France." type="book" year="1985">Baraud (1985)</bibRefCitation>
, the rudimentary spur in
<taxonomicName id="CBA44D65FFCDFFC0FDB7FB0EFD33FB56" authorityName="Laporte" authorityYear="1832" box="[626,708,1156,1175]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FDB7FB0EFD33FB56" box="[626,708,1156,1175]" italics="true" pageId="10" pageNumber="727">Hoplopus</emphasis>
</taxonomicName>
is sometimes missing in specimens of
<taxonomicName id="CBA44D65FFCDFFC0FBD3FB0EFAC5FB56" authority="Chobaut, 1896" authorityName="Chobaut" authorityYear="1896" box="[1046,1330,1156,1175]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="species" species="bleusei">
<emphasis id="3ED0EAF4FFCDFFC0FBD3FB0EFB5DFB56" box="[1046,1194,1156,1175]" italics="true" pageId="10" pageNumber="727">Hoplopus bleusei</emphasis>
Chobaut, 1896
</taxonomicName>
.
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FB2BFC2AFB2F" blockId="10.[151,1683,1116,1435]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FB2BFF68FB6C" attach="left" box="[151,159,1185,1197]" fontSize="5" pageId="10" pageNumber="727">3</superScript>
Entire (not split) claws in males combined with split pro- and mesotarsal claws in females are present only in the species of the Afrotropical
<taxonomicName id="CBA44D65FFCDFFC0FA8DFB2CFA03FB78" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[1352,1524,1190,1209]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="species" species="emortualis">
<emphasis id="3ED0EAF4FFCDFFC0FA8DFB2CFA03FB78" box="[1352,1524,1190,1209]" italics="true" pageId="10" pageNumber="727">Anomala emortualis</emphasis>
</taxonomicName>
species-group and
<taxonomicName id="CBA44D65FFCDFFC0FF52FB4BFEECFB15" baseAuthorityName="Wiedemann" baseAuthorityYear="1821" box="[151,283,1217,1236]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="species" species="vetula">
<emphasis id="3ED0EAF4FFCDFFC0FF52FB4BFEECFB15" box="[151,283,1217,1236]" italics="true" pageId="10" pageNumber="727">Anomala vetula</emphasis>
</taxonomicName>
species-group (
<bibRefCitation id="68354B17FFCDFFC0FE5AFB4BFDC8FB15" author="Machatschke, J. W." box="[415,575,1217,1236]" pageId="10" pageNumber="727" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
) as well as several
<taxonomicName id="CBA44D65FFCDFFC0FD25FB4BFC99FB15" box="[736,878,1217,1236]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3ED0EAF4FFCDFFC0FD25FB4BFCDBFB15" box="[736,812,1217,1236]" italics="true" pageId="10" pageNumber="727">Anomala</emphasis>
species
</taxonomicName>
from Central Asia and the Himalayas, of which many were previously included in the subgenus
<taxonomicName id="CBA44D65FFCDFFC0FF52FB51FEDAFB2F" box="[151,301,1243,1262]" class="Insecta" family="Scarabaeidae" genus="Psammoscapheus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF52FB51FEDAFB2F" box="[151,301,1243,1262]" italics="true" pageId="10" pageNumber="727">Psammoscapheus</emphasis>
</taxonomicName>
. In these species, the protarsal claws are always simple, not strongly thickened.
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FB73FBB2FAD0" blockId="10.[151,1683,1116,1435]" box="[151,1093,1273,1297]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FB73FF68FAC4" attach="left" box="[151,159,1273,1285]" fontSize="5" pageId="10" pageNumber="727">4</superScript>
Males of
<taxonomicName id="CBA44D65FFCDFFC0FF33FB74FEB6FAD0" authorityName="Blanchard" authorityYear="1851" box="[246,321,1278,1297]" class="Insecta" family="Rutelidae" genus="Singhala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FF33FB74FEB6FAD0" box="[246,321,1278,1297]" italics="true" pageId="10" pageNumber="727">Singhala</emphasis>
</taxonomicName>
have thickened protarsomeres 5 and a moderately developed internomedial protuberance.
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FA96FB86FAF2" blockId="10.[151,1683,1116,1435]" box="[151,1137,1308,1331]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FA96FF68FAE9" attach="left" box="[151,159,1308,1320]" fontSize="5" pageId="10" pageNumber="727">5</superScript>
The protarsal inner claw of
<taxonomicName id="CBA44D65FFCDFFC0FE51FAAAFE0CFAF2" authorityName="Reitter" authorityYear="1903" box="[404,507,1312,1331]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FE51FAAAFE0CFAF2" box="[404,507,1312,1331]" italics="true" pageId="10" pageNumber="727">Cyriopertha</emphasis>
</taxonomicName>
is entire (not split) or with a rudimentary split (scored as entire
<emphasis id="3ED0EAF4FFCDFFC0FBE3FAAAFBCEFAF2" box="[1062,1081,1312,1331]" italics="true" pageId="10" pageNumber="727">or</emphasis>
split).
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FAB4FB4CFA97" blockId="10.[151,1683,1116,1435]" box="[151,1211,1342,1366]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FAB4FF68FA8B" attach="left" box="[151,159,1342,1354]" fontSize="5" pageId="10" pageNumber="727">6</superScript>
The protarsal inner claw of some male specimens of
<taxonomicName id="CBA44D65FFCDFFC0FDA8FAC9FD23FA97" authorityName="Reitter" authorityYear="1903" box="[621,724,1347,1366]" class="Insecta" family="Scarabaeidae" genus="Megapertha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCDFFC0FDA8FAC9FD23FA97" box="[621,724,1347,1366]" italics="true" pageId="10" pageNumber="727">Megapertha</emphasis>
</taxonomicName>
have a very rudimentary split (scored as entire
<emphasis id="3ED0EAF4FFCDFFC0FBB5FAC9FB74FA97" box="[1136,1155,1347,1366]" italics="true" pageId="10" pageNumber="727">or</emphasis>
split).
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FAEBFA08FAB9" blockId="10.[151,1683,1116,1435]" box="[151,1535,1377,1400]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FAEBFF68FAAC" attach="left" box="[151,159,1377,1389]" fontSize="5" pageId="10" pageNumber="727">7</superScript>
<taxonomicName id="CBA44D65FFCDFFC0FF61FAEFFDF7FAB9" authority="Nomura, 1965" authorityName="Nomura" authorityYear="1965" box="[164,512,1381,1400]" class="Insecta" family="Rutelidae" genus="Proagopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="727" phylum="Arthropoda" rank="species" species="ohbayashii">
<emphasis id="3ED0EAF4FFCDFFC0FF61FAEFFE8AFAB9" box="[164,381,1381,1400]" italics="true" pageId="10" pageNumber="727">Proagopertha ohbayashii</emphasis>
Nomura, 1965
</taxonomicName>
is unique in the genus due to the presence of a protibial spur, and its classification in this genus requires examination.
</paragraph>
<paragraph id="0C1B36E6FFCDFFC0FF52FA09FCD4FA5A" blockId="10.[151,1683,1116,1435]" box="[151,803,1411,1435]" pageId="10" pageNumber="727">
<superScript id="FBD19BAEFFCDFFC0FF52FA09FF68FA4E" attach="left" box="[151,159,1411,1423]" fontSize="5" pageId="10" pageNumber="727">8</superScript>
Character state from
<bibRefCitation id="68354B17FFCDFFC0FE9DFA02FE14FA5A" author="Petrovitz, R." box="[344,483,1416,1435]" pageId="10" pageNumber="727" pagination="479 - 490" refId="ref11857" refString="Petrovitz, R. 1967. Osterreichische entomologische Expeditionen nach Persien und Afghanistan. Beitrage zur Coleopterologie. Teil XI. Lamellicornia. Annalen des Naturhistorischen Museums in Wien 70: 479 - 490." type="journal article" year="1967">Petrovitz (1967)</bibRefCitation>
and Sabatinelli and Pontuale (1999).
</paragraph>
</tableNote>
<subSubSection id="44BE656DFFCCFFC1FF6AFEEBFBD2FBB8" pageId="11" pageNumber="728" type="type_taxon">
<paragraph id="0C1B36E6FFCCFFC1FF6AFEEBFE2BFE50" blockId="11.[151,639,194,1730]" pageId="11" pageNumber="728">
<emphasis id="3ED0EAF4FFCCFFC1FF6AFEEBFECAFEB7" bold="true" box="[175,317,353,374]" pageId="11" pageNumber="728">Type Species.</emphasis>
<taxonomicName id="CBA44D65FFCCFFC1FE8CFEEBFEB8FE50" ID-CoL="6JWXY" authority="Ratcliffe, Jameson, and Zorn" authorityName="Ratcliffe, Jameson, and Zorn" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCCFFC1FE8CFEEBFDEEFEB7" box="[329,537,353,374]" italics="true" pageId="11" pageNumber="728">Ganganomala saltini</emphasis>
Ratcliffe, Jameson, and Zorn
</taxonomicName>
by monotypy.
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FF6AFE1CFEEFFD11" blockId="11.[151,639,194,1730]" pageId="11" pageNumber="728">
<emphasis id="3ED0EAF4FFCCFFC1FF6AFE1CFE39FE6A" bold="true" box="[175,462,406,427]" pageId="11" pageNumber="728">Higher-Level Classification.</emphasis>
We place
<taxonomicName id="CBA44D65FFCCFFC1FD84FE1CFF1DFE07" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FD84FE1CFF1DFE07" italics="true" pageId="11" pageNumber="728">Ganganomala</emphasis>
</taxonomicName>
in the subtribe
<taxonomicName id="CBA44D65FFCCFFC1FE48FE3BFE01FE07" authorityName="Streubel" authorityYear="1839" box="[397,502,433,454]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="11" pageNumber="728" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
based on the following characters: 1) pronotal base anterior to scutellum evenly rounded, never emarginate anteriad (emarginate anteriad in
<taxonomicName id="CBA44D65FFCCFFC1FE69FD8BFDFBFDD7" ID-CoL="LFC" authorityName="Ohaus" authorityYear="1918" box="[428,524,513,534]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="subTribe" subTribe="Popilliina">Popilliina</taxonomicName>
); 2) labrum more or less hidden beneath anterior margin of clypeus in dorsal view (projecting anteriorly beyond margin of clypeus in
<taxonomicName id="CBA44D65FFCCFFC1FE69FDDAFDF4FDA4" authorityName="Peringuey" authorityYear="1902" box="[428,515,592,613]" class="Insecta" family="Rutelidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="subTribe" subTribe="Isopliina">Isopliina</taxonomicName>
); 3) anterior coxae transverse (longitudinal in
<taxonomicName id="CBA44D65FFCCFFC1FE0AFDE1FDD1FD41" authorityName="Peringuey" authorityYear="1902" box="[463,550,619,640]" class="Insecta" family="Rutelidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="subTribe" subTribe="Isopliina">Isopliina</taxonomicName>
); 4) clypeus not strongly elongated, constricted, and reflexed (strongly elongated, constricted, and reflexed in
<taxonomicName id="CBA44D65FFCCFFC1FF52FD31FEFAFD11" ID-CoL="LBQ" authorityName="Burmeister" authorityYear="1844" box="[151,269,699,720]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="subTribe" subTribe="Anisopliina">Anisopliina</taxonomicName>
).
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FF6AFD5FFE5CFCC4" blockId="11.[151,639,194,1730]" pageId="11" pageNumber="728">
<emphasis id="3ED0EAF4FFCCFFC1FF6AFD5FFE83FD2B" bold="true" box="[175,372,725,746]" pageId="11" pageNumber="728">Generic Diagnosis.</emphasis>
The following characters serve to diagnose the genus:
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FF6AFC80FF2BFBB8" blockId="11.[151,639,194,1730]" pageId="11" pageNumber="728">
1. Protibia with three well-developed teeth on the external margin (
<figureCitation id="949F2A63FFCCFFC1FE83FCAFFE52FCFB" box="[326,421,804,826]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Figs. 8–9</figureCitation>
) (in most species of
<taxonomicName id="CBA44D65FFCCFFC1FF52FCCAFF08FC94" ID-CoL="KLB" box="[151,255,832,853]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
, the external margin of the protibia has two teeth, although
<taxonomicName id="CBA44D65FFCCFFC1FE96FCD0FE04FCAE" box="[339,499,858,879]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3ED0EAF4FFCCFFC1FE96FCD0FE5EFCAE" box="[339,425,858,879]" italics="true" pageId="11" pageNumber="728">Anomala</emphasis>
species
</taxonomicName>
may have 1–3 teeth, according to
<bibRefCitation id="68354B17FFCCFFC1FE91FCFFFE2BFC4B" author="Arrow, G. J." box="[340,476,885,906]" pageId="11" pageNumber="728" refId="ref10166" refString="Arrow, G. J. 1917. The Fauna of British India, including Ceylon and Burma. Coleoptera, Lamellicornia, Part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor and Francis, London, UK." type="book" year="1917">Arrow [1917]</bibRefCitation>
). Several exceptions to this character state illustrate problems with the current classification (or homoplasy of the character state) and include some species of
<taxonomicName id="CBA44D65FFCCFFC1FDE6FC4EFD80FC18" authorityName="Samouelle" authorityYear="1819" box="[547,631,964,985]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FDE6FC4EFD80FC18" box="[547,631,964,985]" italics="true" pageId="11" pageNumber="728">Anomala</emphasis>
</taxonomicName>
, a few Asian
<taxonomicName id="CBA44D65FFCCFFC1FEECFC55FE60FC35" authorityName="Blanchard" authorityYear="1851" box="[297,407,991,1012]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FEECFC55FE60FC35" box="[297,407,991,1012]" italics="true" pageId="11" pageNumber="728">Callistethus</emphasis>
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCCFFC1FE69FC55FD97FC35" authority="Laporte" authorityName="Laporte" authorityYear="1832" box="[428,608,991,1012]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FE69FC55FDF0FC35" box="[428,519,991,1012]" italics="true" pageId="11" pageNumber="728">Hoplopus</emphasis>
Laporte
</taxonomicName>
, a few species of
<taxonomicName id="CBA44D65FFCCFFC1FEF7FC70FE4DFBCE" authority="Kirby" authorityName="Kirby" authorityYear="1823" box="[306,442,1018,1039]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FEF7FC70FE8EFBCE" box="[306,377,1018,1039]" italics="true" pageId="11" pageNumber="728">Mimela</emphasis>
Kirby
</taxonomicName>
from Sulawesi and Africa,
<taxonomicName id="CBA44D65FFCCFFC1FF1BFB9EFE55FBE8" authority="Arrow" authorityName="Arrow" authorityYear="1911" box="[222,418,1044,1065]" class="Insecta" family="Rutelidae" genus="Micranomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FF1BFB9EFEAAFBE8" box="[222,349,1044,1065]" italics="true" pageId="11" pageNumber="728">Micranomala</emphasis>
Arrow
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCCFFC1FE6EFB9EFD8CFBE8" authority="Petrovitz" authorityName="Petrovitz" authorityYear="1967" box="[427,635,1044,1065]" class="Insecta" family="Rutelidae" genus="Microlontha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FE6EFB9EFDE9FBE8" box="[427,542,1044,1065]" italics="true" pageId="11" pageNumber="728">Microlontha</emphasis>
Petrovitz
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCCFFC1FF52FBA5FEA6FB85" authority="Reitter" authorityName="Reitter" authorityYear="1903" box="[151,337,1071,1092]" class="Insecta" family="Rutelidae" genus="Tribopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FF52FBA5FEF1FB85" box="[151,262,1071,1092]" italics="true" pageId="11" pageNumber="728">Tribopertha</emphasis>
Reitter
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCCFFC1FE98FBA5FDD8FB85" authority="Burmeister" authorityName="Burmeister" authorityYear="1844" box="[349,559,1071,1092]" class="Insecta" family="Rutelidae" genus="Rhinyptia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FE98FBA5FE4CFB85" box="[349,443,1071,1092]" italics="true" pageId="11" pageNumber="728">Rhinyptia</emphasis>
Burmeister
</taxonomicName>
(except
<taxonomicName id="CBA44D65FFCCFFC1FF52FBC3FE46FB9F" authority="Arrow" authorityName="Arrow" authorityYear="1917" box="[151,433,1097,1118]" class="Insecta" family="Rutelidae" genus="Rhinyptia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="laeviceps">
<emphasis id="3ED0EAF4FFCCFFC1FF52FBC3FEA8FB9F" box="[151,351,1097,1118]" italics="true" pageId="11" pageNumber="728">Rhinyptia laeviceps</emphasis>
Arrow
</taxonomicName>
), and
<taxonomicName id="CBA44D65FFCCFFC1FE39FBC3FF2BFBB8" authority="Ohaus." authorityName="Ohaus." class="Insecta" family="Rutelidae" genus="Pentanomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FE39FBC3FD8AFB9F" box="[508,637,1097,1118]" italics="true" pageId="11" pageNumber="728">Pentanomala</emphasis>
Ohaus.
</taxonomicName>
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FF6AFBF5FDA7FA8F" blockId="11.[151,639,194,1730]" pageId="11" pageNumber="728">
2. Autapomorphic clypeal form in the male. The male clypeal apex is strongly reflexed, quadrate, with square anterolateral angles and the margins bowed inward (broader at apex than base) (
<figureCitation id="949F2A63FFCCFFC1FD8BFB44FF21FB3F" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="11" pageNumber="728">Figs. 24–25</figureCitation>
). The female clypeal apex is broadly rounded, moderately reflexed, with the margins subequal or lightly narrower at the apex than at base (
<figureCitation id="949F2A63FFCCFFC1FF5BFAB3FEE5FA8F" box="[158,274,1336,1358]" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="11" pageNumber="728">Figs. 26–27</figureCitation>
) (shared with many
<taxonomicName id="CBA44D65FFCCFFC1FE19FAB3FDB1FA8F" ID-CoL="KLB" box="[476,582,1337,1358]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
).
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FF6AFAD9FDA0F9C9" blockId="11.[151,639,194,1730]" pageId="11" pageNumber="728">
3. Male protarsus 5 greatly enlarged (
<figureCitation id="949F2A63FFCCFFC1FDD4FAD9FDA3FAA9" box="[529,596,1363,1384]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Figs. 7</figureCitation>
,
<figureCitation id="949F2A63FFCCFFC1FD9BFAD9FD80FAA9" box="[606,631,1363,1384]" captionStart="Figs" captionStartId="12.[197,236,1602,1621]" captionTargetBox="[220,1151,198,1583]" captionTargetId="figure-0@12.[211,1154,194,1586]" captionTargetPageId="12" captionText="Figs. 36–46. Comparison of male Anomalina protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws. 36) Ganganomala saltini; 37) Anomala dorsalis; 38) Anomala keithi Zorn; 39) Anomala n. sp.; 40) Anomala zornella Prokofiev; 41) Singhala tenella Blanchard; 42) Mimela rugatipennis; 43) Anomala vittata; 44) Megapertha massageta Kirsch; 45) Cyriopertha arcuata; 46) Hoplopus marcens." figureDoi="http://doi.org/10.5281/zenodo.4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="11" pageNumber="728">36</figureCitation>
) (subequal in length to tarsomeres 1–4, widest width half length) and with well-developed internomedial protuberance (autapomorphic for the genus). This compares with male
<taxonomicName id="CBA44D65FFCCFFC1FE9EFA34FE36FA12" box="[347,449,1470,1491]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCCFFC1FE9EFA34FE36FA12" box="[347,449,1470,1491]" italics="true" pageId="11" pageNumber="728">A. dorsalis</emphasis>
</taxonomicName>
(which is similar to
<taxonomicName id="CBA44D65FFCCFFC1FF52FA52FEE8FA2C" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[151,287,1496,1517]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FF52FA52FEE8FA2C" box="[151,287,1496,1517]" italics="true" pageId="11" pageNumber="728">Ganganomala</emphasis>
</taxonomicName>
) in which the internomedial protuberance is less strongly developed (
<figureCitation id="949F2A63FFCCFFC1FDC6FA79FDBBF9C9" box="[515,588,1523,1544]" captionStart="Figs" captionStartId="12.[197,236,1602,1621]" captionTargetBox="[220,1151,198,1583]" captionTargetId="figure-0@12.[211,1154,194,1586]" captionTargetPageId="12" captionText="Figs. 36–46. Comparison of male Anomalina protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws. 36) Ganganomala saltini; 37) Anomala dorsalis; 38) Anomala keithi Zorn; 39) Anomala n. sp.; 40) Anomala zornella Prokofiev; 41) Singhala tenella Blanchard; 42) Mimela rugatipennis; 43) Anomala vittata; 44) Megapertha massageta Kirsch; 45) Cyriopertha arcuata; 46) Hoplopus marcens." figureDoi="http://doi.org/10.5281/zenodo.4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="11" pageNumber="728">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FF6AF987FCE2FD2B" blockId="11.[151,639,194,1730]" lastBlockId="11.[685,1173,194,1730]" pageId="11" pageNumber="728">
4. All claws entire (simple or not split) in the male (shared with some species of
<taxonomicName id="CBA44D65FFCCFFC1FE6DF9A2FE09F9FC" authorityName="Samouelle" authorityYear="1819" box="[424,510,1576,1597]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FE6DF9A2FE09F9FC" box="[424,510,1576,1597]" italics="true" pageId="11" pageNumber="728">Anomala</emphasis>
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCCFFC1FDCCF9A2FDA6F9FC" authorityName="Kirby" authorityYear="1823" box="[521,593,1576,1597]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FDCCF9A2FDA6F9FC" box="[521,593,1576,1597]" italics="true" pageId="11" pageNumber="728">Mimela</emphasis>
</taxonomicName>
, and
<taxonomicName id="CBA44D65FFCCFFC1FF52F9C9FEA4F999" authority="Reitter" authorityName="Reitter" authorityYear="1903" box="[151,339,1603,1624]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FF52F9C9FEFCF999" box="[151,267,1603,1624]" italics="true" pageId="11" pageNumber="728">Cyriopertha</emphasis>
Reitter
</taxonomicName>
and all species of
<taxonomicName id="CBA44D65FFCCFFC1FDCDF9C9FD89F999" authorityName="Petrovitz" authorityYear="1967" box="[520,638,1603,1624]" class="Insecta" family="Rutelidae" genus="Microlontha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FDCDF9C9FD89F999" box="[520,638,1603,1624]" italics="true" pageId="11" pageNumber="728">Microlontha</emphasis>
</taxonomicName>
and
<taxonomicName id="CBA44D65FFCCFFC1FF0EF9D7FEDFF9B3" authorityName="Laporte" authorityYear="1832" box="[203,296,1629,1650]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FF0EF9D7FEDFF9B3" box="[203,296,1629,1650]" italics="true" pageId="11" pageNumber="728">Hoplopus</emphasis>
</taxonomicName>
) with the protarsal inner claw thickened (twice thickness of outer claw), strongly curved, and with inner, minute, apical tooth (
<figureCitation id="949F2A63FFCCFFC1FDFDF918FD86F966" box="[568,625,1682,1703]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Fig. 7</figureCitation>
); inner claws of pro- and mesotarsus split in female, metatarsal inner claw simple (
<figureCitation id="949F2A63FFCCFFC1FC0EFF48FBE7FF16" box="[971,1040,194,215]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Fig. 12</figureCitation>
) (shared with many
<taxonomicName id="CBA44D65FFCCFFC1FD2CFF56FCA4FF30" ID-CoL="KLB" box="[745,851,220,241]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
). Old World
<taxonomicName id="CBA44D65FFCCFFC1FC15FF56FBCEFF30" authorityName="Streubel" authorityYear="1839" box="[976,1081,220,241]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="11" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
in which all claws are entire (not split) include
<taxonomicName id="CBA44D65FFCCFFC1FBF2FF7DFB63FECD" authorityName="Laporte" authorityYear="1832" box="[1079,1172,247,268]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FBF2FF7DFB63FECD" box="[1079,1172,247,268]" italics="true" pageId="11" pageNumber="728">Hoplopus</emphasis>
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FC84FE9BFBA6FEE7" authority="Arrow" authorityName="Arrow" authorityYear="1899" box="[833,1105,273,294]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="calcarata">
<emphasis id="3ED0EAF4FFCCFFC1FC84FE9BFBF4FEE7" box="[833,1027,273,294]" italics="true" pageId="11" pageNumber="728">Anomala calcarata</emphasis>
Arrow
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FD3FFEA6FC14FE80" authority="Gebler" authorityName="Gebler" authorityYear="1841" box="[762,995,300,321]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="3ED0EAF4FFCCFFC1FD3FFEA6FC60FE80" box="[762,919,300,321]" italics="true" pageId="11" pageNumber="728">Anomala vittata</emphasis>
Gebler
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FBFBFEA6FCACFE9A" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="errans">
<emphasis id="3ED0EAF4FFCCFFC1FBFBFEA6FD1CFE9A" italics="true" pageId="11" pageNumber="728">Anomala errans</emphasis>
(Fabricius)
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FC75FECCFD26FEB7" authority="Semenov" authorityName="Semenov" authorityYear="1891" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="oxiana">
<emphasis id="3ED0EAF4FFCCFFC1FC75FECCFBB8FE9A" box="[944,1103,326,347]" italics="true" pageId="11" pageNumber="728">Anomala oxiana</emphasis>
Semenov
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FCDAFEEBFBBCFEB7" authority="(Medvedev)" baseAuthorityName="Medvedev" baseAuthorityYear="1949" box="[799,1099,353,374]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="semenovi">
<emphasis id="3ED0EAF4FFCCFFC1FCDAFEEBFC25FEB7" box="[799,978,353,374]" italics="true" pageId="11" pageNumber="728">Anomala semenovi</emphasis>
(Medvedev)
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FD68FEF6FBC9FE50" authority="(Semenov and Medvedev)" baseAuthorityName="Semenov and Medvedev" box="[685,1086,380,401]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="3ED0EAF4FFCCFFC1FD68FEF6FCEFFE50" box="[685,792,380,401]" italics="true" pageId="11" pageNumber="728">A. pallens</emphasis>
(Semenov and Medvedev)
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FD68FE1CFC72FE6A" authority="Arrow" authorityName="Arrow" authorityYear="1917" box="[685,901,406,427]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="euops">
<emphasis id="3ED0EAF4FFCCFFC1FD68FE1CFCB6FE6A" box="[685,833,406,427]" italics="true" pageId="11" pageNumber="728">Anomala euops</emphasis>
Arrow
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FC11FE1CFD1CFE07" authority="Arrow" authorityName="Arrow" authorityYear="1917" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="praenitens">
<emphasis id="3ED0EAF4FFCCFFC1FC11FE1CFB63FE6A" box="[980,1172,406,427]" italics="true" pageId="11" pageNumber="728">Anomala praenitens</emphasis>
Arrow
</taxonomicName>
(male),
<taxonomicName id="CBA44D65FFCCFFC1FC92FE3BFC0BFE07" baseAuthorityName="Wiedemann" baseAuthorityYear="1821" box="[855,1020,433,454]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="vetula">
<emphasis id="3ED0EAF4FFCCFFC1FC92FE3BFC0BFE07" box="[855,1020,433,454]" italics="true" pageId="11" pageNumber="728">Anomala vetula</emphasis>
</taxonomicName>
species-group (male),
<taxonomicName id="CBA44D65FFCCFFC1FD3CFE41FC4BFE21" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[761,956,459,480]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="emortualis">
<emphasis id="3ED0EAF4FFCCFFC1FD3CFE41FC4BFE21" box="[761,956,459,480]" italics="true" pageId="11" pageNumber="728">Anomala emortualis</emphasis>
</taxonomicName>
species-group (male),
<taxonomicName id="CBA44D65FFCCFFC1FD68FE6CFC43FE3A" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[685,948,486,507]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="3ED0EAF4FFCCFFC1FD68FE6CFCB6FE3A" box="[685,833,486,507]" italics="true" pageId="11" pageNumber="728">Mimela aurata</emphasis>
(Fabricius)
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FB8EFE6CFCB9FDD7" authority="Duftschmid" authorityName="Duftschmid" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="junii">
<emphasis id="3ED0EAF4FFCCFFC1FB8EFE6CFD2FFDD7" italics="true" pageId="11" pageNumber="728">Mimela junii</emphasis>
Duftschmid
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FC0AFD8BFD06FDF1" authority="Graells" authorityName="Graells" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="rugatipennis">
<emphasis id="3ED0EAF4FFCCFFC1FC0AFD8BFB63FDD7" box="[975,1172,513,534]" italics="true" pageId="11" pageNumber="728">Mimela rugatipennis</emphasis>
Graëlls
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FC4CFD91FB63FDF1" authority="(Ohaus)" baseAuthorityName="Ohaus" box="[905,1172,539,560]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="3ED0EAF4FFCCFFC1FC4CFD91FBCCFDF1" box="[905,1083,539,560]" italics="true" pageId="11" pageNumber="728">Mimela lusitanica</emphasis>
(Ohaus)
</taxonomicName>
(probably both sexes, although
<bibRefCitation id="68354B17FFCCFFC1FBC5FDBCFB63FD8A" author="Branco, T." box="[1024,1172,566,587]" pageId="11" pageNumber="728" pagination="49 - 61" refId="ref10388" refString="Branco, T. 2005. Contributions towards a catalogue of Scarabaeoidea of Portugal (Coleoptera). Elytron 19: 49 - 61." type="journal article" year="2005">Branco (2005)</bibRefCitation>
noted a slightly cleft protarsal claw on the left side in the only known male),
<taxonomicName id="CBA44D65FFCCFFC1FC02FDE1FCC7FD5A" authority="Matchatschke" authorityName="Matchatschke" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="holosericea">
<emphasis id="3ED0EAF4FFCCFFC1FC02FDE1FB64FD41" box="[967,1171,619,640]" italics="true" pageId="11" pageNumber="728">Mimela holosericea</emphasis>
Matchatschke
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FC78FD0CFC8AFD74" authority="Gebler" authorityName="Gebler" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="arcuata" subGenus="Pleopertha">
<emphasis id="3ED0EAF4FFCCFFC1FC78FD0CFBC6FD5A" box="[957,1073,646,667]" italics="true" pageId="11" pageNumber="728">Cyriopertha</emphasis>
(
<emphasis id="3ED0EAF4FFCCFFC1FB84FD0CFD3CFD74" italics="true" pageId="11" pageNumber="728">Pleopertha</emphasis>
)
<emphasis id="3ED0EAF4FFCCFFC1FD26FD2AFCDBFD74" box="[739,812,672,693]" italics="true" pageId="11" pageNumber="728">arcuata</emphasis>
Gebler
</taxonomicName>
(both sexes),
<taxonomicName id="CBA44D65FFCCFFC1FBE5FD2AFD1AFD11" authority="Reitter" authorityName="Reitter" authorityYear="1903" class="Insecta" family="Scarabaeidae" genus="Megapertha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FBE5FD2AFB63FD74" box="[1056,1172,672,693]" italics="true" pageId="11" pageNumber="728">Megapertha</emphasis>
Reitter
</taxonomicName>
(males), and
<taxonomicName id="CBA44D65FFCCFFC1FCBEFD31FC06FD11" authorityName="Petrovitz" authorityYear="1967" box="[891,1009,699,720]" class="Insecta" family="Rutelidae" genus="Microlontha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FCBEFD31FC06FD11" box="[891,1009,699,720]" italics="true" pageId="11" pageNumber="728">Microlontha</emphasis>
</taxonomicName>
(males; females unknown).
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FD00FD7AFB7FFC65" blockId="11.[685,1173,194,1730]" pageId="11" pageNumber="728">
5. Protibial spur lacking in male (
<figureCitation id="949F2A63FFCCFFC1FBCAFD7AFB9CFCC4" box="[1039,1131,751,773]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Figs. 7–8</figureCitation>
). In
<taxonomicName id="CBA44D65FFCCFFC1FD68FC80FCEEFCDE" authorityName="Streubel" authorityYear="1839" box="[685,793,778,799]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="11" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
, this character state is shared with species of
<taxonomicName id="CBA44D65FFCCFFC1FCE4FCAFFC62FCFB" authorityName="Reitter" authorityYear="1903" box="[801,917,805,826]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FCE4FCAFFC62FCFB" box="[801,917,805,826]" italics="true" pageId="11" pageNumber="728">Cyriopertha</emphasis>
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCCFFC1FC6DFCAFFBE9FCFB" authorityName="Petrovitz" authorityYear="1967" box="[936,1054,805,826]" class="Insecta" family="Rutelidae" genus="Microlontha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FC6DFCAFFBE9FCFB" box="[936,1054,805,826]" italics="true" pageId="11" pageNumber="728">Microlontha</emphasis>
</taxonomicName>
, and some species of
<taxonomicName id="CBA44D65FFCCFFC1FCDCFCCAFC2FFC94" authority="Blanchard" authorityName="Blanchard" authorityYear="1851" box="[793,984,832,853]" class="Insecta" family="Rutelidae" genus="Singhala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FCDCFCCAFC99FC94" box="[793,878,832,853]" italics="true" pageId="11" pageNumber="728">Singhala</emphasis>
Blanchard
</taxonomicName>
and
<taxonomicName id="CBA44D65FFCCFFC1FBCAFCCAFCD7FCAE" authority="Reitter. The" authorityName="Reitter. The" authorityYear="1903" class="Insecta" family="Rutelidae" genus="Proagopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FBCAFCCAFB63FC94" box="[1039,1172,832,853]" italics="true" pageId="11" pageNumber="728">Proagopertha</emphasis>
Reitter. The
</taxonomicName>
spur is minute but usually present in
<taxonomicName id="CBA44D65FFCCFFC1FD68FCFFFCFDFC4B" authorityName="Laporte" authorityYear="1832" box="[685,778,885,906]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FD68FCFFFCFDFC4B" box="[685,778,885,906]" italics="true" pageId="11" pageNumber="728">Hoplopus</emphasis>
</taxonomicName>
; present and subapical in female
<taxonomicName id="CBA44D65FFCCFFC1FB92FCFFFCF7FC65" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FB92FCFFFCF7FC65" italics="true" pageId="11" pageNumber="728">Ganganomala</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCCFFC1FCCAFC05FCBDFC65" box="[783,842,911,932]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Fig. 9</figureCitation>
) (shared with many
<taxonomicName id="CBA44D65FFCCFFC1FBD1FC05FB89FC65" ID-CoL="KLB" box="[1044,1150,911,932]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
).
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FD00FC20FBC6FB85" blockId="11.[685,1173,194,1730]" pageId="11" pageNumber="728">
6. Meso- and metatarsomeres 5 with moderately developed, internomedial protuberance (male) (
<figureCitation id="949F2A63FFCCFFC1FBA8FC4FFD32FC35" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">Fig. 12</figureCitation>
) or weakly developed protuberance (female) (weakly developed protuberance shared with many
<taxonomicName id="CBA44D65FFCCFFC1FD68FB9EFCE0FBE8" ID-CoL="KLB" box="[685,791,1044,1065]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
). In male
<taxonomicName id="CBA44D65FFCCFFC1FCB2FB9EFC28FBE8" box="[887,991,1044,1065]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCCFFC1FCB2FB9EFC28FBE8" box="[887,991,1044,1065]" italics="true" pageId="11" pageNumber="728">A. dorsalis</emphasis>
</taxonomicName>
, the internomedial protuberance is less strongly developed.
</paragraph>
<paragraph id="0C1B36E6FFCCFFC1FD00FBC3FBD2FBB8" blockId="11.[685,1173,194,1730]" pageId="11" pageNumber="728">
7. Sexual dimorphism present (
<figureCitation id="949F2A63FFCCFFC1FC38FBC3FBACFB9F" box="[1021,1115,1097,1119]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="11" pageNumber="728">Figs. 1–3</figureCitation>
,
<figureCitation id="949F2A63FFCCFFC1FBAFFBC3FB67FB9F" box="[1130,1168,1097,1119]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="11" pageNumber="728">8–9</figureCitation>
,
<figureCitation id="949F2A63FFCCFFC1FD68FBEEFD1DFBB8" box="[685,746,1123,1145]" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="11" pageNumber="728">24–27</figureCitation>
) (shared with some
<taxonomicName id="CBA44D65FFCCFFC1FC74FBEEFBECFBB8" ID-CoL="KLB" box="[945,1051,1124,1145]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="44BE656DFFCCFFC7FD00FBF5FBC8F9C9" lastPageId="13" lastPageNumber="730" pageId="11" pageNumber="728" type="discussion">
<paragraph id="0C1B36E6FFCCFFC7FD00FBF5FE90FE80" blockId="11.[685,1173,194,1730]" lastBlockId="13.[151,639,194,1730]" lastPageId="13" lastPageNumber="730" pageId="11" pageNumber="728">
<emphasis id="3ED0EAF4FFCCFFC1FD00FBF5FCD0FB55" bold="true" box="[709,807,1151,1172]" pageId="11" pageNumber="728">Remarks.</emphasis>
With approximately 1,125 species of
<taxonomicName id="CBA44D65FFCCFFC1FD68FB13FCF4FB6F" authorityName="Samouelle" authorityYear="1819" box="[685,771,1177,1198]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FD68FB13FCF4FB6F" box="[685,771,1177,1198]" italics="true" pageId="11" pageNumber="728">Anomala</emphasis>
</taxonomicName>
in the Old World (P. Limbourg, personal communication,
<date id="781A1026FFCCFFC1FC97FB3EFC3AFB08" box="[850,973,1204,1225]" pageId="11" pageNumber="728" value="2018-03">March 2018</date>
), very few regional Old World treatises for genera or species, and the lack of a useful dichotomous key to Old World genera (Machatschke’ s 1957 work is badly out of date), the starting point for an understanding of
<taxonomicName id="CBA44D65FFCCFFC1FD68FAB3FCE1FA8F" authorityName="Streubel" authorityYear="1839" box="[685,790,1337,1358]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="11" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
biodiversity is a list of taxa.
<bibRefCitation id="68354B17FFCCFFC1FBF9FAB3FCE8FAA9" author="Lobl, I." pageId="11" pageNumber="728" refId="ref11285" refString="Lobl, I., and D. Lobl (editors). 2016. Catalogue of the Palaearctic Coleoptera. Volume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Koninklijke Brill, Leiden, Netherlands." type="journal volume" year="2016">Löbl and Löbl (2016)</bibRefCitation>
listed 19 genera of
<taxonomicName id="CBA44D65FFCCFFC1FC1CFAD9FBB5FAA9" authorityName="Streubel" authorityYear="1839" box="[985,1090,1363,1384]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="11" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
and 294 species and subspecies of
<taxonomicName id="CBA44D65FFCCFFC1FC67FAE4FC0FFA42" authorityName="Samouelle" authorityYear="1819" box="[930,1016,1390,1411]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FC67FAE4FC0FFA42" box="[930,1016,1390,1411]" italics="true" pageId="11" pageNumber="728">Anomala</emphasis>
</taxonomicName>
that occur in the Asian portion of the Palearctic region (including
<collectingCountry id="74B37676FFCCFFC1FD68FA29FD12FA79" box="[685,741,1443,1464]" name="Nepal" pageId="11" pageNumber="728">Nepal</collectingCountry>
but not including
<collectingCountry id="74B37676FFCCFFC1FC4AFA29FBF5FA79" box="[911,1026,1443,1464]" name="Bangladesh" pageId="11" pageNumber="728">Bangladesh</collectingCountry>
). For the Indian subcontinent, the only work that synthesized ruteline biodiversity was Arrow’ s (1917)
<emphasis id="3ED0EAF4FFCCFFC1FBF3FA52FCDFF9C9" italics="true" pageId="11" pageNumber="728">
Fauna of
<collectingCountry id="74B37676FFCCFFC1FD68FA79FCDFF9C9" box="[685,808,1523,1544]" name="India" pageId="11" pageNumber="728">British India</collectingCountry>
</emphasis>
. Even at the time of publication, this work was incomplete because so much of the area had not been adequately explored for scarab beetles, and our knowledge of the fauna there was still rudimentary. More recently, scarab research in the Ganges River basin did not discover
<taxonomicName id="CBA44D65FFCCFFC1FBC0F9F2FB7AF94C" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[1029,1165,1656,1677]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FBC0F9F2FB7AF94C" box="[1029,1165,1656,1677]" italics="true" pageId="11" pageNumber="728">Ganganomala</emphasis>
</taxonomicName>
. A survey of
<taxonomicName id="CBA44D65FFCCFFC1FCF6F918FC7EF966" authorityName="Samouelle" authorityYear="1819" box="[819,905,1682,1703]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCCFFC1FCF6F918FC7EF966" box="[819,905,1682,1703]" italics="true" pageId="11" pageNumber="728">Anomala</emphasis>
</taxonomicName>
from Buxa Tiger Reserve (
<collectingRegion id="CE60F804FFCCFFC1FD71F927FCC3F903" box="[692,820,1709,1730]" country="India" name="West Bengal" pageId="11" pageNumber="728">West Bengal</collectingRegion>
,
<collectingCountry id="74B37676FFCCFFC1FC84F927FC84F903" box="[833,883,1709,1730]" name="India" pageId="11" pageNumber="728">India</collectingCountry>
) (Sarkar
<emphasis id="3ED0EAF4FFCCFFC1FC17F927FBF1F903" box="[978,1030,1709,1730]" italics="true" pageId="11" pageNumber="728">et al.</emphasis>
2017), use of pheromone and light traps in Saharanpur for detection of white grubs associated with sugarcane (Sushil
<emphasis id="3ED0EAF4FFCAFFC7FF27FF7DFEE3FECD" box="[226,276,247,268]" italics="true" pageId="13" pageNumber="730">et al.</emphasis>
2017), and agricultural pest management reports (Sharma
<emphasis id="3ED0EAF4FFCAFFC7FEA3FE9BFE62FEE7" box="[358,405,273,294]" italics="true" pageId="13" pageNumber="730">et al.</emphasis>
2002) did not, of course, reveal
<taxonomicName id="CBA44D65FFCAFFC7FF1CFEA6FE96FE80" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[217,353,300,321]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FF1CFEA6FE96FE80" box="[217,353,300,321]" italics="true" pageId="13" pageNumber="730">Ganganomala</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="58DB666EFFCBFFC6FF00F9C8FB7CF974" ID-DOI="http://doi.org/10.5281/zenodo.4789890" ID-Zenodo-Dep="4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="12" pageNumber="729" startId="12.[197,236,1602,1621]" targetBox="[220,1151,198,1583]" targetPageId="12">
<paragraph id="0C1B36E6FFCBFFC6FF00F9C8FB7CF974" blockId="12.[173,1194,1602,1717]" pageId="12" pageNumber="729">
<emphasis id="3ED0EAF4FFCBFFC6FF00F9C8FEC3F994" bold="true" box="[197,308,1602,1621]" pageId="12" pageNumber="729">Figs. 36–46.</emphasis>
Comparison of male
<taxonomicName id="CBA44D65FFCBFFC6FDCCF9C8FD90F994" authorityName="Streubel" authorityYear="1839" box="[521,615,1602,1621]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="12" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws.
<emphasis id="3ED0EAF4FFCBFFC6FD4EF9F8FD5FF944" bold="true" box="[651,680,1650,1669]" pageId="12" pageNumber="729">36)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FD6AF9F8FC93F944" box="[687,868,1650,1669]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCBFFC6FD6AF9F8FC93F944" box="[687,868,1650,1669]" italics="true" pageId="12" pageNumber="729">Ganganomala saltini</emphasis>
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FCB7F9F8FC79F944" bold="true" box="[882,910,1650,1669]" pageId="12" pageNumber="729">37)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FC53F9F8FBD9F944" box="[918,1070,1650,1669]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCBFFC6FC53F9F8FBD9F944" box="[918,1070,1650,1669]" italics="true" pageId="12" pageNumber="729">Anomala dorsalis</emphasis>
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FBFEF9F8FBA0F944" bold="true" box="[1083,1111,1650,1669]" pageId="12" pageNumber="729">38)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FB9AF9F8FEE6F95C" authority="Zorn" authorityName="Zorn" authorityYear="2011" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="keithi">
<emphasis id="3ED0EAF4FFCBFFC6FB9AF9F8FF2EF95C" italics="true" pageId="12" pageNumber="729">Anomala keithi</emphasis>
Zorn
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FEE4F900FECBF95C" bold="true" box="[289,316,1674,1693]" pageId="12" pageNumber="729">39)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FE8FF900FE63F95C" authority="Ratcliffe &amp; Jameson &amp; Zorn, 2018" authorityName="Ratcliffe &amp; Jameson &amp; Zorn" authorityYear="2018" box="[330,404,1674,1693]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="3ED0EAF4FFCBFFC6FE8FF900FE63F95C" box="[330,404,1674,1693]" italics="true" pageId="12" pageNumber="729">Anomala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25E3578FFFCBFFC6FE64F900FE2FF95C" box="[417,472,1674,1693]" pageId="12" pageNumber="729" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="3ED0EAF4FFCBFFC6FE2DF900FDF4F95C" bold="true" box="[488,515,1674,1693]" pageId="12" pageNumber="729">40)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FDD4F900FCFDF95C" authority="Prokofiev" authorityName="Prokofiev" authorityYear="2015" box="[529,778,1674,1693]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="zornella">
<emphasis id="3ED0EAF4FFCBFFC6FDD4F900FD5DF95C" box="[529,682,1674,1693]" italics="true" pageId="12" pageNumber="729">Anomala zornella</emphasis>
Prokofiev
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FCDFF900FCC2F95C" bold="true" box="[794,821,1674,1693]" pageId="12" pageNumber="729">41)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FC86F900FBC6F95C" authority="Blanchard" authorityName="Blanchard" authorityYear="1851" box="[835,1073,1674,1693]" class="Insecta" family="Rutelidae" genus="Singhala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="tenella">
<emphasis id="3ED0EAF4FFCBFFC6FC86F900FC39F95C" box="[835,974,1674,1693]" italics="true" pageId="12" pageNumber="729">Singhala tenella</emphasis>
Blanchard
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FB87F900FBAAF95C" bold="true" box="[1090,1117,1674,1693]" pageId="12" pageNumber="729">42)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FBAEF900FEE5F974" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="rugatipennis">
<emphasis id="3ED0EAF4FFCBFFC6FBAEF900FEE5F974" italics="true" pageId="12" pageNumber="729">Mimela rugatipennis</emphasis>
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FEDBF928FECEF974" bold="true" box="[286,313,1698,1717]" pageId="12" pageNumber="729">43)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FE85F928FE35F974" authorityName="Gebler" authorityYear="1841" box="[320,450,1698,1717]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="3ED0EAF4FFCBFFC6FE85F928FE35F974" box="[320,450,1698,1717]" italics="true" pageId="12" pageNumber="729">Anomala vittata</emphasis>
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FE0BF928FE15F974" bold="true" box="[462,482,1698,1717]" pageId="12" pageNumber="729">44</emphasis>
)
<taxonomicName id="CBA44D65FFCBFFC6FE35F928FD1AF974" authority="Kirsch" authorityName="Kirsch" box="[496,749,1698,1717]" class="Insecta" family="Scarabaeidae" genus="Megapertha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="massageta">
<emphasis id="3ED0EAF4FFCBFFC6FE35F928FD58F974" box="[496,687,1698,1717]" italics="true" pageId="12" pageNumber="729">Megapertha massageta</emphasis>
Kirsch
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FD32F928FCE5F974" bold="true" box="[759,786,1698,1717]" pageId="12" pageNumber="729">45)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FCDDF928FC48F974" box="[792,959,1698,1717]" class="Insecta" family="Rutelidae" genus="Cyriopertha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="3ED0EAF4FFCBFFC6FCDDF928FC48F974" box="[792,959,1698,1717]" italics="true" pageId="12" pageNumber="729">Cyriopertha arcuata</emphasis>
</taxonomicName>
;
<emphasis id="3ED0EAF4FFCBFFC6FC0EF928FC10F974" bold="true" box="[971,999,1698,1717]" pageId="12" pageNumber="729">46)</emphasis>
<taxonomicName id="CBA44D65FFCBFFC6FC28F928FB71F974" box="[1005,1158,1698,1717]" class="Insecta" family="Scarabaeidae" genus="Hoplopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="729" phylum="Arthropoda" rank="species" species="marcens">
<emphasis id="3ED0EAF4FFCBFFC6FC28F928FB71F974" box="[1005,1158,1698,1717]" italics="true" pageId="12" pageNumber="729">Hoplopus marcens</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="0C1B36E6FFCAFFC7FF6AFECCFE74FBE8" blockId="13.[151,639,194,1730]" pageId="13" pageNumber="730">
Ascertaining the classification status of this new taxon was at first perplexing. The taxon strongly resembles some
<taxonomicName id="CBA44D65FFCAFFC7FEF8FEF6FE50FE50" authorityName="MacLeay" authorityYear="1819" box="[317,423,380,401]" class="Insecta" family="Dynastinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="family">Dynastinae</taxonomicName>
based on male characters (the protarsal claws are unequal in size and entire, and the meso- and metatarsal claws are subequal in size), but it is indicative of
<taxonomicName id="CBA44D65FFCAFFC7FDDDFE41FD8AFE21" box="[536,637,459,480]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
based on male and female characters (all tarsal claws are independently moveable, and the elytral margins possess a slender membrane) (
<bibRefCitation id="68354B17FFCAFFC7FE05FD91FD8EFDF1" author="Machatschke, J. W." box="[448,633,539,560]" pageId="13" pageNumber="730" pagination="199" refId="ref11394" refString="Machatschke, J. W. 1957. Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae 2. In: Genera Insectorum 199 (B) (P. A. G. Wytsman, editor). Editions Mercurius, Antwerp, Belgium." type="book chapter" year="1957">Machatschke 1957</bibRefCitation>
;
<bibRefCitation id="68354B17FFCAFFC7FF52FDBCFE86FD8A" author="Jameson, M. L." box="[151,369,566,587]" pageId="13" pageNumber="730" pagination="415 - 432" refId="ref10890" refString="Jameson, M. L., A. Paucar-Cabrera, and A. Sol'is. 2003. Synopsis of the New World genera of Anomalini (Coleoptera: Scarabaeidae: Rutelinae) and description of a new genus from Costa Rica and Nicaragua. Annals of the Entomological Society of America 96: 415 - 432." type="journal article" year="2003">
Jameson
<emphasis id="3ED0EAF4FFCAFFC7FF32FDBCFED8FD8A" box="[247,303,566,587]" italics="true" pageId="13" pageNumber="730">et al.</emphasis>
2003
</bibRefCitation>
). Employing the subtribal classification of
<taxonomicName id="CBA44D65FFCAFFC7FEF7FDDAFE6DFDA4" box="[306,410,592,613]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
, we place the new taxon in the subtribe
<taxonomicName id="CBA44D65FFCAFFC7FEF4FDE1FE6DFD41" authorityName="Streubel" authorityYear="1839" box="[305,410,619,640]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="13" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
(see “Classification of World
<taxonomicName id="CBA44D65FFCAFFC7FF18FD0FFEB5FD5B" box="[221,322,645,666]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
” above). Genera in
<taxonomicName id="CBA44D65FFCAFFC7FDD1FD0FFD8AFD5B" authorityName="Streubel" authorityYear="1839" box="[532,637,645,666]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="13" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
share the following character states: the pro- and mesosternal processes are lacking; the base of the pronotum is as broad as the elytra; the mesepimera are hidden at the base of the elytra; and the elytra are longer than broad. Sufficient differences clearly separate the new genus from other genera of
<taxonomicName id="CBA44D65FFCAFFC7FF52FCCAFEF7FC94" authorityName="Streubel" authorityYear="1839" box="[151,256,832,853]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="13" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
(see “Diagnosis”): the absence of a protibial spur in the male; the unsplit claws in the male, with the protarsal inner claw strongly thickened and curved (autapomorphic in
<taxonomicName id="CBA44D65FFCAFFC7FE2DFC05FDA6FC65" box="[488,593,911,932]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
); the greatly enlarged male protarsus 5 with a welldeveloped internomedial protuberance; and the autapomorphic form of the male clypeus (apex strongly reflexed with square anterolateral angles, margins bowed inward).
</paragraph>
<paragraph id="0C1B36E6FFCAFFC7FF6AFBA5FD0DFC65" blockId="13.[151,639,194,1730]" lastBlockId="13.[685,1173,194,1730]" pageId="13" pageNumber="730">
We considered that
<taxonomicName id="CBA44D65FFCAFFC7FEB6FBA5FE0AFB85" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[371,509,1071,1092]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FEB6FBA5FE0AFB85" box="[371,509,1071,1092]" italics="true" pageId="13" pageNumber="730">Ganganomala</emphasis>
</taxonomicName>
might represent a currently named taxon within the
<taxonomicName id="CBA44D65FFCAFFC7FDC8FBC3FD8DFB9F" authorityName="Streubel" authorityYear="1839" box="[525,634,1097,1118]" class="Globothalamea" family="Anomalinidae" kingdom="Chromista" order="Rotaliida" pageId="13" pageNumber="730" phylum="Foraminifera" rank="subTribe" subTribe="Anomalina">Anomalina</taxonomicName>
. All
<taxonomicName id="CBA44D65FFCAFFC7FF78FBEEFE96FBB8" box="[189,353,1124,1145]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3ED0EAF4FFCAFFC7FF78FBEEFEE3FBB8" box="[189,276,1124,1145]" italics="true" pageId="13" pageNumber="730">Anomala</emphasis>
species
</taxonomicName>
have a protibial spur present in both sexes, tarsal claws of mostly unequal size on at least the front and middle legs (subequal claws on the middle legs present in several species of the otherwise very different former subgenus
<taxonomicName id="CBA44D65FFCAFFC7FD84FB44FE61FB3F" authority="Motschulsky" authorityName="Motschulsky" class="Insecta" family="Scarabaeidae" genus="Psammoscapheus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FD84FB44FEFBFB3F" italics="true" pageId="13" pageNumber="730">Psammoscapheus</emphasis>
Motschulsky
</taxonomicName>
), and at least one claw on the pro- and/or mesotarsus is split at its apex (exceptions are listed in “Generic Diagnosis”). Contrastingly, all the meso- and metatarsal claws on the male of
<taxonomicName id="CBA44D65FFCAFFC7FEE0FAD9FE58FAA9" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[293,431,1363,1384]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FEE0FAD9FE58FAA9" box="[293,431,1363,1384]" italics="true" pageId="13" pageNumber="730">Ganganomala</emphasis>
</taxonomicName>
are of equal size, and all claws are entire at their apices. We carefully considered similarities of the new taxon with
<taxonomicName id="CBA44D65FFCAFFC7FDACFA02FF13FA79" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FDACFA02FF13FA79" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FF37FA29FEB8FA79" box="[242,335,1443,1465]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 5–6</figureCitation>
,
<figureCitation id="949F2A63FFCAFFC7FE9CFA29FE6FFA79" box="[345,408,1443,1464]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">10, 13</figureCitation>
,
<figureCitation id="949F2A63FFCAFFC7FE67FA29FE4BFA79" box="[418,444,1443,1464]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="13" pageNumber="730">22</figureCitation>
,
<figureCitation id="949F2A63FFCAFFC7FE03FA29FDF2FA79" box="[454,517,1443,1465]" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="13" pageNumber="730">29–30</figureCitation>
,
<figureCitation id="949F2A63FFCAFFC7FDCAFA29FDBAFA79" box="[527,589,1443,1465]" captionStart="Figs" captionStartId="7.[175,214,677,696]" captionTargetBox="[156,1165,196,659]" captionTargetId="figure-532@7.[154,1169,194,661]" captionTargetPageId="7" captionText="Figs. 31–35. Protibiae of male and female of Ganganomala saltini, Anomala dorsalis, and Hoplopus marcens Peyerimhoff, dorsal views. 31) G. saltini, male, subapical spur lacking, three external teeth; 32) G. saltini, female, subapical spur present, three external teeth; 33) A. dorsalis (syntype of A. fraterna, MLUH), male, subapical spur present, three external teeth; 34) A. dorsalis, female, subapical spur present, three external teeth; 35) H. marcens, male, subapical spur lacking, two external teeth, third tooth nearly obsolete." figureDoi="http://doi.org/10.5281/zenodo.4789880" httpUri="https://zenodo.org/record/4789880/files/figure.png" pageId="13" pageNumber="730">33–34</figureCitation>
,
<figureCitation id="949F2A63FFCAFFC7FD9DFA29FD85FA79" box="[600,626,1443,1464]" captionStart="Figs" captionStartId="12.[197,236,1602,1621]" captionTargetBox="[220,1151,198,1583]" captionTargetId="figure-0@12.[211,1154,194,1586]" captionTargetPageId="12" captionText="Figs. 36–46. Comparison of male Anomalina protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws. 36) Ganganomala saltini; 37) Anomala dorsalis; 38) Anomala keithi Zorn; 39) Anomala n. sp.; 40) Anomala zornella Prokofiev; 41) Singhala tenella Blanchard; 42) Mimela rugatipennis; 43) Anomala vittata; 44) Megapertha massageta Kirsch; 45) Cyriopertha arcuata; 46) Hoplopus marcens." figureDoi="http://doi.org/10.5281/zenodo.4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="13" pageNumber="730">37</figureCitation>
), formerly in the subgenus
<taxonomicName id="CBA44D65FFCAFFC7FE4FFA37FE1FFA13" authorityName="Burmeister" authorityYear="1844" box="[394,488,1469,1490]" class="Insecta" family="Scarabaeidae" genus="Rhinoplia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FE4FFA37FE1FFA13" box="[394,488,1469,1490]" italics="true" pageId="13" pageNumber="730">Rhinoplia</emphasis>
</taxonomicName>
. This subgenus was proposed by
<bibRefCitation id="68354B17FFCAFFC7FE83FA52FE0FFA2C" author="Burmeister, H." box="[326,504,1496,1517]" pageId="13" pageNumber="730" refId="ref10413" refString="Burmeister, H. 1844. Handbuch der Entomologie, Volume 4, Part 1. T. C. F. Enslin, Berlin, Germany." type="book" year="1844">Burmeister (1844)</bibRefCitation>
and included East Indian species with a strongly reflexed clypeal apex and “sharp” lateral corners as well as tridentate protibia.
<taxonomicName id="CBA44D65FFCAFFC7FF35F9A2FE6BF9FC" box="[240,412,1576,1597]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FF35F9A2FE6BF9FC" box="[240,412,1576,1597]" italics="true" pageId="13" pageNumber="730">Anomala dorsalis</emphasis>
</taxonomicName>
is the
<typeStatus id="D31F8844FFCAFFC7FE27F9A2FDFCF9FC" box="[482,523,1576,1597]" pageId="13" pageNumber="730">type</typeStatus>
of the subgenus, which includes three names that are synonyms of
<taxonomicName id="CBA44D65FFCAFFC7FF2AF9D7FEAFF9B3" box="[239,344,1629,1650]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FF2AF9D7FEAFF9B3" box="[239,344,1629,1650]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<taxonomicName id="CBA44D65FFCAFFC7FEADF9D7FD8DF9B3" authority="Nonfried" authorityName="Nonfried" authorityYear="1893" box="[360,634,1629,1650]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="3ED0EAF4FFCAFFC7FEADF9D7FDEDF9B3" box="[360,538,1629,1650]" italics="true" pageId="13" pageNumber="730">Anomala centralis</emphasis>
Nonfried
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCAFFC7FF52F9F2FE40F94C" authority="Nonfried" authorityName="Nonfried" authorityYear="1894" box="[151,439,1656,1677]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="3ED0EAF4FFCAFFC7FF52F9F2FEB8F94C" box="[151,335,1656,1677]" italics="true" pageId="13" pageNumber="730">Anomala imitatrix</emphasis>
Nonfried
</taxonomicName>
,
<taxonomicName id="CBA44D65FFCAFFC7FE0CF9F2FEF2F966" authority="Burmeister" authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FE0CF9F2FD8AF94C" box="[457,637,1656,1677]" italics="true" pageId="13" pageNumber="730">Anomala fraterna</emphasis>
Burmeister
</taxonomicName>
).
<taxonomicName id="CBA44D65FFCAFFC7FED0F918FE48F966" box="[277,447,1682,1703]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FED0F918FE48F966" box="[277,447,1682,1703]" italics="true" pageId="13" pageNumber="730">Anomala dorsalis</emphasis>
</taxonomicName>
is similar in overall appearance, size, and reflexed clypeal apex in the male. Females of
<taxonomicName id="CBA44D65FFCAFFC7FC9AFF48FC4CFF16" box="[863,955,194,215]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FC9AFF48FC4CFF16" box="[863,955,194,215]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
and
<taxonomicName id="CBA44D65FFCAFFC7FC2BFF48FBAFFF16" box="[1006,1112,194,215]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FC2BFF48FBAFFF16" box="[1006,1112,194,215]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
share split pro- and mesotarsal claws (
<figureCitation id="949F2A63FFCAFFC7FC2CFF56FBA6FF30" box="[1001,1105,220,242]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Figs. 9–10</figureCitation>
) and a protibial spur (spur absent in the male of
<taxonomicName id="CBA44D65FFCAFFC7FBEBFF7DFB71FECD" box="[1070,1158,247,268]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FBEBFF7DFB71FECD" box="[1070,1158,247,268]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
). However, the male of
<taxonomicName id="CBA44D65FFCAFFC7FC59FE9BFC0BFEE7" box="[924,1020,273,294]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FC59FE9BFC0BFEE7" box="[924,1020,273,294]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
has an entire, strongly thickened and curved protarsal claw (
<figureCitation id="949F2A63FFCAFFC7FBA8FEA6FD4CFE9A" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Fig. 7</figureCitation>
), whereas it is split in
<taxonomicName id="CBA44D65FFCAFFC7FC7CFECCFBD0FE9A" box="[953,1063,326,347]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FC7CFECCFBD0FE9A" box="[953,1063,326,347]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FBFFFECCFB7EFE9A" box="[1082,1161,326,347]" captionStart="Figs" captionStartId="12.[197,236,1602,1621]" captionTargetBox="[220,1151,198,1583]" captionTargetId="figure-0@12.[211,1154,194,1586]" captionTargetPageId="12" captionText="Figs. 36–46. Comparison of male Anomalina protarsomeres showing form of claws, protarsomere 5, apex of protarsomere 4, and relative lengths of protarsomeres 1–5 (see Table 2). Figs. 37–41 show taxa with split claws; Figs. 42–46 show taxa with entire (not split, simple) claws. 36) Ganganomala saltini; 37) Anomala dorsalis; 38) Anomala keithi Zorn; 39) Anomala n. sp.; 40) Anomala zornella Prokofiev; 41) Singhala tenella Blanchard; 42) Mimela rugatipennis; 43) Anomala vittata; 44) Megapertha massageta Kirsch; 45) Cyriopertha arcuata; 46) Hoplopus marcens." figureDoi="http://doi.org/10.5281/zenodo.4789890" httpUri="https://zenodo.org/record/4789890/files/figure.png" pageId="13" pageNumber="730">Fig. 37</figureCitation>
). Additional characters that separate these taxa are: the lateral margins of the abdominal sternites (simple in
<taxonomicName id="CBA44D65FFCAFFC7FCDDFE1CFC83FE6A" box="[792,884,406,427]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FCDDFE1CFC83FE6A" box="[792,884,406,427]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
but strongly carinate in both sexes of
<taxonomicName id="CBA44D65FFCAFFC7FD3BFE3BFC93FE07" box="[766,868,433,454]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FD3BFE3BFC93FE07" box="[766,868,433,454]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FCB5FE3BFC44FE07" box="[880,947,433,454]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Fig. 13</figureCitation>
)); the lateral margins of the male clypeus (concave in
<taxonomicName id="CBA44D65FFCAFFC7FC07FE41FBEDFE21" box="[962,1050,459,480]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FC07FE41FBEDFE21" box="[962,1050,459,480]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FBE0FE41FB9DFE21" box="[1061,1130,459,480]" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="13" pageNumber="730">Fig. 25</figureCitation>
) but slightly convex in
<taxonomicName id="CBA44D65FFCAFFC7FCA5FE6CFC30FE3A" box="[864,967,486,507]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FCA5FE6CFC30FE3A" box="[864,967,486,507]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FC10FE6CFBEBFE3A" box="[981,1052,486,507]" captionStart="Figs" captionStartId="6.[197,236,1580,1599]" captionTargetBox="[208,1158,186,1562]" captionTargetId="figure-0@6.[206,1160,184,1563]" captionTargetPageId="6" captionText="Figs. 24–30. Head comparisons of male and female Ganganomala saltini and similar Anomalina, dorsal and dorso-oblique views. G. saltini: 24–25) Male, clypeus with apex strongly reflexed and quadrate, anterolateral angles square, margins bowed inward with apex broader than base; 26–27) Female, clypeus with apex broadly rounded and moderately reflexed, margins subequal or slightly narrower at apex than at base. 28) Singhala hindu, male, clypeal apex strongly reflexed, emarginate at middle; 29–30) A. dorsalis, male, clypeus with apex strongly reflexed and subquadrate, anterolateral angles acute, margins converging from base to apex." figureDoi="http://doi.org/10.5281/zenodo.4789862" httpUri="https://zenodo.org/record/4789862/files/figure.png" pageId="13" pageNumber="730">Fig. 30</figureCitation>
)); the dorsal coloration (
<taxonomicName id="CBA44D65FFCAFFC7FCDAFD8BFC8DFDD7" box="[799,890,513,534]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FCDAFD8BFC8DFDD7" box="[799,890,513,534]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
is a bright, chestnut reddish brown (
<figureCitation id="949F2A63FFCAFFC7FD3CFD91FCA2FDF1" box="[761,853,539,561]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 1–3</figureCitation>
) but usually testaceous to bright reddish brown and with or without black dorsal markings in
<taxonomicName id="CBA44D65FFCAFFC7FCE2FDDAFC78FDA4" box="[807,911,592,613]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FCE2FDDAFC78FDA4" box="[807,911,592,613]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FC5BFDDAFC0DFDA4" box="[926,1018,592,614]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 5–6</figureCitation>
)); and the body shape (compact and suboval in
<taxonomicName id="CBA44D65FFCAFFC7FC36FDE1FBA5FD41" box="[1011,1106,619,640]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FC36FDE1FBA5FD41" box="[1011,1106,619,640]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FBA1FDE1FD26FD5A" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 1–3</figureCitation>
) but elongate in
<taxonomicName id="CBA44D65FFCAFFC7FCB9FD0CFC12FD5A" box="[892,997,646,667]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FCB9FD0CFC12FD5A" box="[892,997,646,667]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FC30FD0CFBA5FD5A" box="[1013,1106,645,667]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 5–6</figureCitation>
)). The peculiar shape of the male protarsus in
<taxonomicName id="CBA44D65FFCAFFC7FBFDFD2AFB63FD74" box="[1080,1172,672,693]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FBFDFD2AFB63FD74" box="[1080,1172,672,693]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FD70FD31FD19FD11" box="[693,750,699,720]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Fig. 7</figureCitation>
), the form of the claws on all legs (
<figureCitation id="949F2A63FFCAFFC7FB84FD31FB7FFD11" box="[1089,1160,699,720]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Fig. 12</figureCitation>
), the absence of a protibial spur in the male (
<figureCitation id="949F2A63FFCAFFC7FB88FD5FFB7FFD2B" box="[1101,1160,725,746]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Fig. 7</figureCitation>
), the simple lateral margins of abdominal sternites (not carinate, compare with
<figureCitation id="949F2A63FFCAFFC7FC7BFC80FBF2FCDE" box="[958,1029,778,799]" captionStart="Figs" captionStartId="3.[175,214,1518,1537]" captionTargetBox="[156,1167,196,1500]" captionTargetId="figure-1@3.[154,1169,191,1502]" captionTargetPageId="3" captionText="Figs. 7–13. Comparative characters of Ganganomala saltini and Anomala dorsalis. 7) G. saltini, male, greatly enlarged protarsus 5, well-developed internomedial protuberance, simple claws, and protibial spur lacking; 8) G. saltini, male, external margin of protibia with three well-developed teeth; 9) G. saltini, female, protibial spur, claw split, and tibia with three external teeth; 10) A. dorsalis, female, protarsus similar to that of female G. saltini; 11) G. saltini, male, metatibial apex and metatarsomere 1; 12) G. saltini, male, moderately developed, internomedial protuberance on metatarsomere 5 and simple claws; 13) A. dorsalis, lateral ventrites with well-defined carina." figureDoi="http://doi.org/10.5281/zenodo.4789770" httpUri="https://zenodo.org/record/4789770/files/figure.png" pageId="13" pageNumber="730">Fig. 13</figureCitation>
), and the form of the parameres (ventrally setose in
<taxonomicName id="CBA44D65FFCAFFC7FBF1FCAFFB64FCFB" box="[1076,1171,805,826]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="saltini">
<emphasis id="3ED0EAF4FFCAFFC7FBF1FCAFFB64FCFB" box="[1076,1171,805,826]" italics="true" pageId="13" pageNumber="730">G. saltini</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FD70FCCAFCC5FC94" box="[693,818,831,853]" captionStart="Figs" captionStartId="4.[197,236,1469,1488]" captionTargetBox="[177,1184,199,1446]" captionTargetId="figure-58@4.[175,1190,194,1453]" captionTargetPageId="4" captionText="Figs. 14–22. Morphological characters for Ganganomala saltini and Anomala dorsalis. G. saltini: 14) Left maxilla, ventral view; 15) Left mandible, dorsal view; 16) Mentum, ventral view; 17) Spiculum gastrale and associated sclerites; 18) Hind wing, ventral view; 19) Parameres, dorsal view; 20) Parameres, lateral view; 21) Parameres, ventral view. 22) A. dorsalis (syntype of A. fraterna, MLUH), parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789824" httpUri="https://zenodo.org/record/4789824/files/figure.png" pageId="13" pageNumber="730">Figs. 20–21</figureCitation>
)
<emphasis id="3ED0EAF4FFCAFFC7FC8DFCCAFC71FC94" box="[840,902,832,853]" italics="true" pageId="13" pageNumber="730">versus</emphasis>
glabrous ventrally in
<taxonomicName id="CBA44D65FFCAFFC7FBBAFCCAFD0CFCAE" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FBBAFCCAFD0CFCAE" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
) demonstrate that this similarity is only superficial and not the result of close taxonomic affinity.
</paragraph>
<paragraph id="0C1B36E6FFCAFFC7FD00FC20FBC8F9C9" blockId="13.[685,1173,194,1730]" pageId="13" pageNumber="730">
The overall resemblance of
<taxonomicName id="CBA44D65FFCAFFC7FC0BFC20FBA1FC7E" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[974,1110,938,959]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FC0BFC20FBA1FC7E" box="[974,1110,938,959]" italics="true" pageId="13" pageNumber="730">Ganganomala</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FBA1FC20FD26FC1B" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 1–3</figureCitation>
) and
<taxonomicName id="CBA44D65FFCAFFC7FCD7FC4FFC8AFC1B" box="[786,893,965,986]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FCD7FC4FFC8AFC1B" box="[786,893,965,986]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
(
<figureCitation id="949F2A63FFCAFFC7FC55FC4FFC19FC1B" box="[912,1006,964,986]" captionStart="Figs" captionStartId="2.[197,236,1576,1595]" captionTargetBox="[178,1187,196,1553]" captionTargetId="figure-1@2.[175,1190,194,1561]" captionTargetPageId="2" captionText="Figs. 1–6. Habitus of Ganganomala saltini in comparison with Anomala dorsalis. 1) G. saltini, male holotype, dorsal view; 2) G. saltini, male holotype, ventral view; 3) G. saltini, female allotype, dorsal view; 4) G. saltini, male paratype, lateral view; 5) A. dorsalis, male, dorsal view (syntype of Anomala fraterna, MLUH); 6) A. dorsalis, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4789761" httpUri="https://zenodo.org/record/4789761/files/figure.png" pageId="13" pageNumber="730">Figs. 5–6</figureCitation>
) has resulted in taxonomic confusion.
<bibRefCitation id="68354B17FFCAFFC7FC47FC55FBEFFC35" author="Chandra, K." box="[898,1048,991,1012]" pageId="13" pageNumber="730" pagination="292 - 294" refId="ref10535" refString="Chandra, K. 1991. New genitalic characters for distinguishing Anomala dorsalis Fabricius and Anomala fraterna Burmeister (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 45: 292 - 294." type="journal article" year="1991">Chandra (1991)</bibRefCitation>
attempted to separate
<taxonomicName id="CBA44D65FFCAFFC7FCCFFC70FC8DFBCE" box="[778,890,1018,1039]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FCCFFC70FC8DFBCE" box="[778,890,1018,1039]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
from a similar anomaline species, both of which are distributed in Dehradun,
<collectingCountry id="74B37676FFCAFFC7FD68FBA5FD29FB85" box="[685,734,1071,1092]" name="India" pageId="13" pageNumber="730">India</collectingCountry>
(
<bibRefCitation id="68354B17FFCAFFC7FD2FFBA5FC83FB85" author="Chandra, K." box="[746,884,1071,1092]" pageId="13" pageNumber="730" pagination="292 - 294" refId="ref10535" refString="Chandra, K. 1991. New genitalic characters for distinguishing Anomala dorsalis Fabricius and Anomala fraterna Burmeister (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 45: 292 - 294." type="journal article" year="1991">Chandra 1991</bibRefCitation>
). Chandra’ s diagnosis of these sympatric species was based on male genitalia, eye size, punctation of the head and pronotum, and color. Description of the form of the claws, protibial teeth, and clypeal form were lacking, and the diagnosis was not based on comparison of type specimens. Chandra proposed that one morphotype was conspecific with
<taxonomicName id="CBA44D65FFCAFFC7FC4BFB63FC0CFB3F" box="[910,1019,1257,1278]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FC4BFB63FC0CFB3F" box="[910,1019,1257,1278]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
and the other morphotype was conspecific with “
<taxonomicName id="CBA44D65FFCAFFC7FBDEFA8EFCB5FAF2" authority="&quot; (Chandra 1991)" authorityName="&quot; (Chandra" authorityYear="1991" baseAuthorityName="Chandra" baseAuthorityYear="1991" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FBDEFA8EFB7EFAD8" box="[1051,1161,1284,1305]" italics="true" pageId="13" pageNumber="730">A. fraterna</emphasis>
” (
<bibRefCitation id="68354B17FFCAFFC7FD71FA94FCCCFAF2" author="Chandra, K." box="[692,827,1310,1331]" pageId="13" pageNumber="730" pagination="292 - 294" refId="ref10535" refString="Chandra, K. 1991. New genitalic characters for distinguishing Anomala dorsalis Fabricius and Anomala fraterna Burmeister (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 45: 292 - 294." type="journal article" year="1991">Chandra 1991</bibRefCitation>
)
</taxonomicName>
. Chandra proposed renewed species status for “
<taxonomicName id="CBA44D65FFCAFFC7FCD0FAB3FC8AFA8F" authorityName="&quot; (Chandra" authorityYear="1991" box="[789,893,1337,1358]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FCD0FAB3FC8AFA8F" box="[789,893,1337,1358]" italics="true" pageId="13" pageNumber="730">A. fraterna</emphasis>
</taxonomicName>
”, which was then considered a synonym of
<taxonomicName id="CBA44D65FFCAFFC7FCF5FAD9FC63FAA9" box="[816,916,1363,1384]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="3ED0EAF4FFCAFFC7FCF5FAD9FC63FAA9" box="[816,916,1363,1384]" italics="true" pageId="13" pageNumber="730">A. dorsalis</emphasis>
</taxonomicName>
. However, examination (by CZ) of Burmeister’ s
<typeStatus id="D31F8844FFCAFFC7FCA9FAE4FC49FA42" box="[876,958,1390,1411]" pageId="13" pageNumber="730" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="CBA44D65FFCAFFC7FC1EFAE4FBB7FA42" authorityName="&quot; (Chandra" authorityYear="1991" box="[987,1088,1390,1411]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FC1EFAE4FBB7FA42" box="[987,1088,1390,1411]" italics="true" pageId="13" pageNumber="730">A. fraterna</emphasis>
</taxonomicName>
revealed that the pro- and mesotarsi have split claws (also concurring with Burmeister’ s description of
<taxonomicName id="CBA44D65FFCAFFC7FB90FA29FD29FA12" authorityName="&quot; (Chandra" authorityYear="1991" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FB90FA29FD29FA12" italics="true" pageId="13" pageNumber="730">A. fraterna</emphasis>
</taxonomicName>
). Based on the images in
<bibRefCitation id="68354B17FFCAFFC7FC12FA34FB9AFA12" author="Chandra, K." box="[983,1133,1470,1491]" pageId="13" pageNumber="730" pagination="292 - 294" refId="ref10535" refString="Chandra, K. 1991. New genitalic characters for distinguishing Anomala dorsalis Fabricius and Anomala fraterna Burmeister (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 45: 292 - 294." type="journal article" year="1991">Chandra (1991)</bibRefCitation>
, we suspect that Chandra’ s “
<taxonomicName id="CBA44D65FFCAFFC7FC58FA52FBFFFA2C" authorityName="&quot; (Chandra" authorityYear="1991" box="[925,1032,1496,1517]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FC58FA52FBFFFA2C" box="[925,1032,1496,1517]" italics="true" pageId="13" pageNumber="730">A. fraterna</emphasis>
</taxonomicName>
” morphotype may be conspecific with
<emphasis id="3ED0EAF4FFCAFFC7FC6EFA79FBC8F9C9" box="[939,1087,1523,1544]" italics="true" pageId="13" pageNumber="730">
<taxonomicName id="CBA44D65FFCAFFC7FC6EFA79FBCBF9C9" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[939,1084,1523,1544]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">Ganganomala</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="44BE656DFFCAFFC4FD00F987FE5EFF30" lastPageId="14" lastPageNumber="731" pageId="13" pageNumber="730" type="distribution">
<paragraph id="0C1B36E6FFCAFFC4FD00F987FE5EFF30" blockId="13.[685,1173,194,1730]" lastBlockId="14.[173,659,193,454]" lastPageId="14" lastPageNumber="731" pageId="13" pageNumber="730">
<emphasis id="3ED0EAF4FFCAFFC7FD00F987FCB1F9E3" bold="true" box="[709,838,1549,1570]" pageId="13" pageNumber="730">Distribution.</emphasis>
<taxonomicName id="CBA44D65FFCAFFC7FC9DF987FC17F9E3" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[856,992,1549,1570]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FC9DF987FC17F9E3" box="[856,992,1549,1570]" italics="true" pageId="13" pageNumber="730">Ganganomala</emphasis>
</taxonomicName>
is known from
<collectingCountry id="74B37676FFCAFFC7FD68F9A2FD12F9FC" box="[685,741,1576,1597]" name="Nepal" pageId="13" pageNumber="730">Nepal</collectingCountry>
and
<collectingCountry id="74B37676FFCAFFC7FCD3F9A2FC72F9FC" box="[790,901,1576,1597]" name="Bangladesh" pageId="13" pageNumber="730">Bangladesh</collectingCountry>
in the greater Ganges River drainage and the Indo-Gangetic Plain (floodplain of the Indus and Ganga-Brahmaputra River systems) (
<figureCitation id="949F2A63FFCAFFC7FD34F9F2FCCDF94C" box="[753,826,1656,1677]" captionStart="Fig" captionStartId="5.[175,206,521,540]" captionTargetBox="[155,633,196,503]" captionTargetId="figure-697@5.[153,635,194,505]" captionTargetPageId="5" captionText="Fig. 23. Distribution of Ganganomala saltini in relation to the Ganges River Basin and locations in Nepal and Bangladesh. Map courtesy of A. Smith." figureDoi="http://doi.org/10.5281/zenodo.4789844" httpUri="https://zenodo.org/record/4789844/files/figure.png" pageId="13" pageNumber="730">Fig. 23</figureCitation>
).
<bibRefCitation id="68354B17FFCAFFC7FC88F9F2FC12F94C" author="Chandra, K." box="[845,997,1656,1677]" pageId="13" pageNumber="730" pagination="292 - 294" refId="ref10535" refString="Chandra, K. 1991. New genitalic characters for distinguishing Anomala dorsalis Fabricius and Anomala fraterna Burmeister (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 45: 292 - 294." type="journal article" year="1991">Chandra (1991)</bibRefCitation>
recorded putative
<taxonomicName id="CBA44D65FFCAFFC7FD68F918FCC2F966" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[685,821,1682,1703]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFCAFFC7FD68F918FCC2F966" box="[685,821,1682,1703]" italics="true" pageId="13" pageNumber="730">Ganganomala</emphasis>
</taxonomicName>
(as “
<taxonomicName id="CBA44D65FFCAFFC7FCB5F918FC2BF966" authorityName="&quot; (Chandra" authorityYear="1991" box="[880,988,1682,1703]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="730" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="3ED0EAF4FFCAFFC7FCB5F918FC2BF966" box="[880,988,1682,1703]" italics="true" pageId="13" pageNumber="730">A. fraterna</emphasis>
</taxonomicName>
”) from Dehradun,
<collectingCountry id="74B37676FFCAFFC7FD68F927FD29F903" box="[685,734,1709,1730]" name="India" pageId="13" pageNumber="730">India</collectingCountry>
, also in the Indo-Gangetic Plain, but examination of Chandra’ s specimens would be needed to corroborate an Indian record.
</paragraph>
</subSubSection>
<subSubSection id="44BE656DFFC9FFC4FF00FF7DFEC7FE80" pageId="14" pageNumber="731" type="biology_ecology">
<paragraph id="0C1B36E6FFC9FFC4FF00FF7DFEC7FE80" blockId="14.[173,659,193,454]" pageId="14" pageNumber="731">
<emphasis id="3ED0EAF4FFC9FFC4FF00FF7DFEE1FECD" bold="true" box="[197,278,247,268]" pageId="14" pageNumber="731">Biology.</emphasis>
Adults are attracted to lights at night, but otherwise we know nothing of this seemingly rare scarab beetle.
</paragraph>
</subSubSection>
<subSubSection id="44BE656DFFC9FFC4FF00FECCFEB7FE07" pageId="14" pageNumber="731" type="etymology">
<paragraph id="0C1B36E6FFC9FFC4FF00FECCFEB7FE07" blockId="14.[173,659,193,454]" pageId="14" pageNumber="731">
<emphasis id="3ED0EAF4FFC9FFC4FF00FECCFEC0FE9A" bold="true" box="[197,311,326,347]" pageId="14" pageNumber="731">Etymology.</emphasis>
<taxonomicName id="CBA44D65FFC9FFC4FE83FECCFE39FE9A" authorityName="SALTINI RATCLIFFE, JAMESON, AND ZORN, A" authorityYear="2018" box="[326,462,326,347]" class="Insecta" family="Rutelidae" genus="Ganganomala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="731" phylum="Arthropoda" rank="genus">
<emphasis id="3ED0EAF4FFC9FFC4FE83FECCFE39FE9A" box="[326,462,326,347]" italics="true" pageId="14" pageNumber="731">Ganganomala</emphasis>
</taxonomicName>
is named for the Ganga River (Ganges River) that flows from the Himalayan Mountains, through the Gangetic Plains in
<collectingCountry id="74B37676FFC9FFC4FF03FE1CFF0EFE6A" box="[198,249,406,427]" name="India" pageId="14" pageNumber="731">India</collectingCountry>
, through
<collectingCountry id="74B37676FFC9FFC4FE90FE1CFE31FE6A" box="[341,454,406,427]" name="Bangladesh" pageId="14" pageNumber="731">Bangladesh</collectingCountry>
, and empties into the Bay of Bengal.
</paragraph>
</subSubSection>
</treatment>
</document>
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