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<document id="5257E112F9120F1523B38E1C90B698A4" ID-CLB-Dataset="21486" ID-DOI="10.3161/150811013X678955" ID-GBIF-Dataset="69c434f8-762e-4e43-bc29-d0da90994625" ID-ISSN="1733-5329" ID-Zenodo-Dep="3943563" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594705346288" checkinUser="felipe" docAuthor="Bates, Paul J. J., Cameron, Kenneth, Pearch, Malcolm J. &amp; Hayes, Benjamin" docDate="2013" docId="837C87DBFF9EC071FC00FD6BFD83FDE1" docLanguage="en" docName="ActaChiropterologica.15.2.313-340.pdf.imf" docOrigin="Acta Chiropterologica 15 (2)" docStyle="DocumentStyle:A870862413AAF9DEC7D3C2B4FC7FB2AC.1:ActaChiropterol.2008-2015.journal_article" docStyleId="A870862413AAF9DEC7D3C2B4FC7FB2AC" docStyleName="ActaChiropterol.2008-2015.journal_article" docStyleVersion="1" docTitle="Nycteris hispida" docType="treatment" docVersion="9" lastPageNumber="323" masterDocId="7F45FFA3FF94C07AFF9CFFFFFFC9FFC4" masterDocTitle="A review of the bats (Chiroptera) of the Republic of Congo, including eight species new to the country" masterLastPageNumber="340" masterPageNumber="313" pageNumber="322" updateTime="1698843371229" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2AC856B9550C03131E9D00A9246E1B02">A review of the bats (Chiroptera) of the Republic of Congo, including eight species new to the country</mods:title>
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<mods:namePart id="D52DC75E18EEF696F2083DCB08E37917">Bates, Paul J. J.</mods:namePart>
<mods:affiliation id="DC6492FF706B1DD8F8E5457477C4F31B">Harrison Institute, Centre for Systematics and Biodiversity Research, Bowerwood House, St Botolphs Road, Sevenoaks, Kent, TN 13 3 AQ, United Kingdom &amp; Corresponding author: E-mail: pjjbates 2 @ hotmail. com</mods:affiliation>
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<mods:namePart id="898E93A7EB7201555C1E09E4A8ABEAAF">Cameron, Kenneth</mods:namePart>
<mods:affiliation id="6592A36D1D6B247564A25C27E53B17AB">Wildlife Conservation Society, 2300 Southern Boulevard, Bronx, New York 10460, USA</mods:affiliation>
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<mods:namePart id="AFDEE3494037A06AF186A1A7C0BEB849">Pearch, Malcolm J.</mods:namePart>
<mods:affiliation id="0B30EE72E70526239C7AE1371CB9FDBE">Harrison Institute, Centre for Systematics and Biodiversity Research, Bowerwood House, St Botolphs Road, Sevenoaks, Kent, TN 13 3 AQ, United Kingdom</mods:affiliation>
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<mods:namePart id="2D36F5B4E92257F8DFC886A8D14A153F">Hayes, Benjamin</mods:namePart>
<mods:affiliation id="DDA83E3A31DBBD59D6E9B1FF8E4A0FEB">Monadh, Inveruglas, Kingussie, Inverness-shire, United Kingdom</mods:affiliation>
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<mods:title id="78FB114AD5663ABB49D761EA2EEA064A">Acta Chiropterologica</mods:title>
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<mods:part id="85B1B22DD7CDC3022453112227B79A7E">
<mods:date id="8E69912A7EAC3E4B4B03957A6D791E0E">2013</mods:date>
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<mods:number id="F15CB0926F669C418C6B078C19B40685">15</mods:number>
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<treatment id="837C87DBFF9EC071FC00FD6BFD83FDE1" ID-DOI="http://doi.org/10.5281/zenodo.4332340" ID-GBIF-Taxon="165088893" ID-Zenodo-Dep="4332340" LSID="urn:lsid:plazi:treatment:837C87DBFF9EC071FC00FD6BFD83FDE1" httpUri="http://treatment.plazi.org/id/837C87DBFF9EC071FC00FD6BFD83FDE1" lastPageId="11" lastPageNumber="323" pageId="10" pageNumber="322">
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<paragraph id="0B6A36CDFF9EC070FC00FD6BFAE0FD6A" blockId="10.[816,1351,660,746]" box="[924,1321,660,686]" pageId="10" pageNumber="322">
<heading id="502281A1FF9EC070FC00FD6BFAE0FD6A" box="[924,1321,660,686]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="322" reason="2">
<taxonomicName id="CCD54D4EFF9EC070FC00FD6BFAE0FD6A" ID-CoL="485DK" authority="(Schreber, 1774)" baseAuthorityName="Schreber" baseAuthorityYear="1774" box="[924,1321,660,686]" class="Mammalia" family="Nycteridae" genus="Nycteris" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="322" phylum="Chordata" rank="species" species="hispida">
<emphasis id="39A1EADFFF9EC070FC00FD6BFB95FD6A" box="[924,1116,660,686]" italics="true" pageId="10" pageNumber="322">Nycteris hispida</emphasis>
(Schreber, 1774)
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<subSubSection id="43CF6546FF9EC070FC6DFD4AFB1CFD0B" box="[1009,1237,693,719]" pageId="10" pageNumber="322" type="vernacular_names">
<paragraph id="0B6A36CDFF9EC070FC6DFD4AFB1CFD0B" blockId="10.[816,1351,660,746]" box="[1009,1237,693,719]" pageId="10" pageNumber="322">Hairy slit-faced bat</paragraph>
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<treatmentCitation id="8A7410DCFF9EC070FCACFD2AFB23FD2E" box="[816,1258,725,746]" pageId="10" pageNumber="322">
<taxonomicName id="CCD54D4EFF9EC070FCACFD2AFB23FD2E" authority="Schreber, 1774: 169" authorityName="Schreber" authorityPageNumber="169" authorityYear="1774" box="[816,1258,725,746]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="322" phylum="Chordata" rank="species" species="hispidus">
<emphasis id="39A1EADFFF9EC070FCACFD2AFC27FD2E" box="[816,1006,725,746]" italics="true" pageId="10" pageNumber="322">Vespertilio hispidus</emphasis>
<bibRefCitation id="6F444B3CFF9EC070FC68FD2AFB23FD2E" box="[1012,1258,725,746]" pageId="10" pageNumber="322">Schreber, 1774: 169, 188</bibRefCitation>
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;
<collectingCountry id="73C2765DFF9EC070FB68FD2AFA8AFD2E" box="[1268,1347,725,746]" name="Senegal" pageId="10" pageNumber="322">Senegal</collectingCountry>
.
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</paragraph>
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<subSubSection id="43CF6546FF9EC070FCACFCEAFA9DFC3E" pageId="10" pageNumber="322" type="materials_examined">
<paragraph id="0B6A36CDFF9EC070FCACFCEAFC19FCEB" blockId="10.[816,1429,789,1018]" box="[816,976,789,815]" pageId="10" pageNumber="322">
<emphasis id="39A1EADFFF9EC070FCACFCEAFC19FCEB" box="[816,976,789,815]" italics="true" pageId="10" pageNumber="322">New material</emphasis>
</paragraph>
<paragraph id="0B6A36CDFF9EC070FCCAFCC8FA9DFC3E" blockId="10.[816,1429,789,1018]" pageId="10" pageNumber="322">
<materialsCitation id="BBBD3C90FF9EC070FCCAFCC8FA8CFCB7" ID-GBIF-Occurrence="2813185310" collectingDate="4 August, 2012" collectionCode="HZM" collectorName="Previous &amp; Appendix I" county="Lekoumou" latitude="-2.7590833" location="Waterfall forest" longLatPrecision="1" longitude="13.6057" municipality="Waterfall forest" pageId="10" pageNumber="322" specimenCode="HZM.52.40188" specimenCount="1" specimenCount-male="1">
<specimenCode id="5B739EB6FF9EC070FCCAFCC8FBD9FC95" box="[854,1040,823,849]" pageId="10" pageNumber="322">HZM.52.40188</specimenCode>
,
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,
<collectingDate id="6F2FE9E5FF9EC070FBCCFCC8FADFFC95" box="[1104,1302,823,849]" pageId="10" pageNumber="322">
<date id="7F6B100DFF9EC070FBCCFCC8FB0DFC95" box="[1104,1220,823,849]" pageId="10" pageNumber="322">4 August</date>
, 2012
</collectingDate>
,
<location id="0E0A6016FF9EC070FAB4FCC8FCBFFCB7" LSID="urn:lsid:plazi:treatment:837C87DBFF9EC071FC00FD6BFD83FDE1:0E0A6016FF9EC070FAB4FCC8FCBFFCB7" county="Lekoumou" latitude="-2.7590833" longLatPrecision="1" longitude="13.6057" municipality="Waterfall forest" name="Waterfall forest" pageId="10" pageNumber="322">Waterfall forest</location>
,
<collectingCounty id="E20B4E41FF9EC070FC1FFCA6FBCCFCB7" box="[899,1029,857,883]" pageId="10" pageNumber="322">Lekoumou</collectingCounty>
,
<geoCoordinate id="6EE1500AFF9EC070FB8EFCA6FB53FCB7" box="[1042,1178,857,883]" degrees="2" direction="south" minutes="45.545" orientation="latitude" pageId="10" pageNumber="322" precision="1" value="-2.7590833">2°45.545S</geoCoordinate>
,
<geoCoordinate id="6EE1500AFF9EC070FB34FCA6FA88FCB7" box="[1192,1345,857,883]" degrees="13" direction="east" minutes="36.342" orientation="longitude" pageId="10" pageNumber="322" precision="1" value="13.6057">13°36.342E</geoCoordinate>
.
</materialsCitation>
<collectorName id="A620531BFF9EC070FAD1FCA6FC90FC51" pageId="10" pageNumber="322">Previous</collectorName>
records included in
<figureCitation id="93EE2A48FF9EC070FBD1FC84FB1CFC51" box="[1101,1237,891,917]" captionStart="APPENDIX" captionStartId="21.[99,120,486,512]" captionTargetPageId="21" captionText="APPENDIX I A list of the forty-three bat species currently recorded from Congo, including citations in the literature and locality data. CS (Conservation status NT: Near threatened; DD: Data deficient; LC: Least concern) is based on IUCN Red List of Threatened Species retrieved on 8 May, 2013. Two species listed here have not been assessed (na) by IUCN for their conservation status but rather have been included within other taxa: Triaenops afer in Triaenops persicus and Chaerephon leucogaster in Tadarida pumila. Superscripts associated with localities in column 4 refer to the superscripts for the literature cited in column 3" figureDoi="http://doi.org/10.5281/zenodo.3943589" httpUri="https://zenodo.org/record/3943589/files/figure.png" pageId="10" pageNumber="322">
<collectorName id="A620531BFF9EC070FBD1FC84FB1CFC51" box="[1101,1237,891,917]" pageId="10" pageNumber="322">Appendix I</collectorName>
</figureCitation>
; the known distribution corresponds to localities 15, 24, 26, 29, and
<quantity id="CC2D9B28FF9EC070FCACFC41FCA7FC1C" box="[816,878,958,984]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="10" pageNumber="322" unit="in" value="31.0">31 in</quantity>
<figureCitation id="93EE2A48FF9EC070FCE4FC41FC09FC1C" box="[888,960,958,984]" captionStart="FIG" captionStartId="3.[159,172,1686,1707]" captionTargetBox="[166,1418,221,1664]" captionTargetId="figure@3.[135,1537,218,1670]" captionTargetPageId="3" captionText="FIG. 1. Localities from which bats have been collected in Congo based on the literature and current study (see Appendix I). Size of red dot is proportional in area to the number of species collected from that locality. Habitat types are based on Bergmans (1979). Localities (listed in Appendix III): (1) Pointe-Noire; (2) Hinda; (3) Loandjili; (4) Tchissanga; (5) Koubotchi; (6) Ménengué; (7) NGongo; (8) Dinguembo; (9) Béna; (10) Goumina; (11) Sounda, Gorge of; (12) Makaba; (13) Bouloungi; (14) Dimonika; (15) Mayombe; (16) Bamba, col du Mont; (17ac) Loudima-Kimongo (includes Cave of Doumboula, Cave of Loudima, and Cave of Viaduc); (18a,b) Kila NTari and Mouyondzi; (19ah) Bangou Massif (includes Bouaboumpono, Cave of Kimanika, Cave of Matouridi, Meya / Cave of Meya-Nzouari, Cave of Meya-Nwadi, Mountembessa, Cave of Mpasa, and Cave of Mpoka); (20) Voka; (21) Kinkala; (22) Ile MBamou; (23) La Région du Pool (Stanley Pool); (24) Brazzaville; (25) Grand Bois; (26) Sibiti; (27) Komono; (28) Divenié; (29) Lekoumou; (30) Lesio-Louna Reserve; (31) Ngabé; (32) Plateau Konkouya; (33) Lékana; (34) Djambala; (35) Ewo; (36) Konda; (37) Ubangi (Oubangi); (38) Mouth of Ubangi River; (39) Kellé; (40) Odzala/Odzala National Park; (41) Kango; (42) Impfondo; (43) Epena. Scale as marked. C.A.R.= Central African Republic; D.R. Congo = Democratic Republic of Congo" figureDoi="http://doi.org/10.5281/zenodo.3943565" httpUri="https://zenodo.org/record/3943565/files/figure.png" pageId="10" pageNumber="322">Fig. 1</figureCitation>
. This is one of the most common nycterids in Africa (
<bibRefCitation id="6F444B3CFF9EC070FC6EFC1FFA81FC3E" author="Happold and Happold" box="[1010,1352,992,1018]" firstAuthor="Happold" pageId="10" pageNumber="322" refId="ref13898" refString="HAPPOLD, M., and D. C. D. HAPPOLD (eds.). 2013. Mammals of Africa Volume IV: Hedgehogs, shrews, and bats. Bloomsbury Publishing, London, 800 pp." type="book" year="2013">Happold and Happold, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="43CF6546FF9EC071FCACFBDBFDC8FEB9" lastPageId="11" lastPageNumber="323" pageId="10" pageNumber="322" type="description">
<paragraph id="0B6A36CDFF9EC070FCACFBDBFC72FBFA" blockId="10.[816,1429,1060,1458]" box="[816,955,1060,1086]" pageId="10" pageNumber="322">
<emphasis id="39A1EADFFF9EC070FCACFBDBFC72FBFA" box="[816,955,1060,1086]" italics="true" pageId="10" pageNumber="322">Description</emphasis>
</paragraph>
<paragraph id="0B6A36CDFF9EC071FCCAFBB9FDC8FEB9" blockId="10.[816,1429,1060,1458]" lastBlockId="11.[159,771,220,381]" lastPageId="11" lastPageNumber="323" pageId="10" pageNumber="322">
A very small species of slit-faced bat with a forearm length of
<quantity id="CC2D9B28FF9EC070FC4AFB98FB8BFB45" box="[982,1090,1127,1153]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="10" pageNumber="322" unit="mm" value="35.8">35.8 mm</quantity>
. The face is characterised by a noseleaf which comprises two flaps of skin, which give the impression of a slit running down the length of the muzzle. For
<taxonomicName id="CCD54D4EFF9EC070FB89FB32FBBEFB23" box="[1045,1143,1229,1255]" class="Mammalia" family="Nycteridae" genus="Nycteris" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="322" phylum="Chordata" rank="genus">
<emphasis id="39A1EADFFF9EC070FB89FB32FBBEFB23" box="[1045,1143,1229,1255]" italics="true" pageId="10" pageNumber="322">Nycteris</emphasis>
</taxonomicName>
, the ears of this species are relatively short and the tragus has a smooth, curved, outer margin (
<figureCitation id="93EE2A48FF9EC070FBDEFAEFFB6DFAEE" box="[1090,1188,1296,1322]" captionStart="FIG" captionStartId="10.[159,172,1966,1987]" captionTargetBox="[364,1229,1530,1948]" captionTargetId="figure@10.[322,1290,1525,1958]" captionTargetPageId="10" captionText="FIG. 7. Nycteris hispida, HZM.52.40188. (A): Anterior, left mandible; (B): left, upper incisors and canine; (C): right tragus; (D): baculum (dorsal view [left] and lateral view [right]). Scales for A, B, and D = 1 mm; scale for C = 2 mm" figureDoi="http://doi.org/10.5281/zenodo.3943577" httpUri="https://zenodo.org/record/3943577/files/figure.png" pageId="10" pageNumber="322">Fig. 7C</figureCitation>
). The skull is short (damaged in this specimen) and the upper toothrow (CM
<superScript id="FCA09B85FF9EC070FCEAFAAFFC49FAA6" attach="left" box="[886,896,1360,1378]" fontSize="8" pageId="10" pageNumber="322">3</superScript>
) is
<quantity id="CC2D9B28FF9EC070FC28FAABFBE8FAAA" box="[948,1057,1364,1390]" metricMagnitude="-3" metricUnit="m" metricValue="5.58" pageId="10" pageNumber="322" unit="mm" value="5.58">5.58 mm</quantity>
in length. Both the upper and lower incisors are trifid (
<figureCitation id="93EE2A48FF9EC070FBCAFA89FAC8FA54" box="[1110,1281,1398,1424]" captionStart="FIG" captionStartId="10.[159,172,1966,1987]" captionTargetBox="[364,1229,1530,1948]" captionTargetId="figure@10.[322,1290,1525,1958]" captionTargetPageId="10" captionText="FIG. 7. Nycteris hispida, HZM.52.40188. (A): Anterior, left mandible; (B): left, upper incisors and canine; (C): right tragus; (D): baculum (dorsal view [left] and lateral view [right]). Scales for A, B, and D = 1 mm; scale for C = 2 mm" figureDoi="http://doi.org/10.5281/zenodo.3943577" httpUri="https://zenodo.org/record/3943577/files/figure.png" pageId="10" pageNumber="322">Fig. 7A and B</figureCitation>
). The posterior lower premolar is small (
<figureCitation id="93EE2A48FF9EC070FB14FA67FB2EFA76" box="[1160,1255,1432,1458]" captionStart="FIG" captionStartId="10.[159,172,1966,1987]" captionTargetBox="[364,1229,1530,1948]" captionTargetId="figure@10.[322,1290,1525,1958]" captionTargetPageId="10" captionText="FIG. 7. Nycteris hispida, HZM.52.40188. (A): Anterior, left mandible; (B): left, upper incisors and canine; (C): right tragus; (D): baculum (dorsal view [left] and lateral view [right]). Scales for A, B, and D = 1 mm; scale for C = 2 mm" figureDoi="http://doi.org/10.5281/zenodo.3943577" httpUri="https://zenodo.org/record/3943577/files/figure.png" pageId="10" pageNumber="322">Fig. 7A</figureCitation>
). The baculum is 3.0 mm in length and characteristically hooked at the tip, with a slender, curved shaft and a slightly bilobate base (
<figureCitation id="93EE2A48FF9FC071FED3FEE0FE79FEFD" box="[335,432,287,313]" captionStart="FIG" captionStartId="10.[159,172,1966,1987]" captionTargetBox="[364,1229,1530,1948]" captionTargetId="figure@10.[322,1290,1525,1958]" captionTargetPageId="10" captionText="FIG. 7. Nycteris hispida, HZM.52.40188. (A): Anterior, left mandible; (B): left, upper incisors and canine; (C): right tragus; (D): baculum (dorsal view [left] and lateral view [right]). Scales for A, B, and D = 1 mm; scale for C = 2 mm" figureDoi="http://doi.org/10.5281/zenodo.3943577" httpUri="https://zenodo.org/record/3943577/files/figure.png" pageId="11" pageNumber="323">Fig. 7D</figureCitation>
); for comparison with other African
<taxonomicName id="CCD54D4EFF9FC071FE9DFEBEFEAAFE9F" box="[257,355,321,347]" class="Mammalia" family="Nycteridae" genus="Nycteris" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="323" phylum="Chordata" rank="genus">
<emphasis id="39A1EADFFF9FC071FE9DFEBEFEAAFE9F" box="[257,355,321,347]" italics="true" pageId="11" pageNumber="323">Nycteris</emphasis>
</taxonomicName>
taxa see
<bibRefCitation id="6F444B3CFF9FC071FE48FEBEFD06FE9F" author="Thomas" box="[468,719,321,347]" etAl="et al." firstAuthor="Thomas" pageId="11" pageNumber="323" pagination="17 - 31" refId="ref14653" refString="THOMAS, N. M., D. L. HARRISON, and P. J. J. BATES. 1994. A study of the baculum in the genus Nycteris (Mammalia, Chiroptera, Nycteridae) with consideration of its taxonomic importance. Bonner zoologische Beitrage, 45: 17 - 31." type="journal article" year="1994">
Thomas
<emphasis id="39A1EADFFF9FC071FDA0FEBEFDB1FE9F" box="[572,632,321,347]" italics="true" pageId="11" pageNumber="323">et al.</emphasis>
(1994)
</bibRefCitation>
and
<bibRefCitation id="6F444B3CFF9FC071FF03FE9CFE34FEB9" author="Happold and Happold" box="[159,509,355,381]" firstAuthor="Happold" pageId="11" pageNumber="323" refId="ref13898" refString="HAPPOLD, M., and D. C. D. HAPPOLD (eds.). 2013. Mammals of Africa Volume IV: Hedgehogs, shrews, and bats. Bloomsbury Publishing, London, 800 pp." type="book" year="2013">Happold and Happold (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="5FAA6645FF9EC070FF03F851FAE5F824" ID-DOI="http://doi.org/10.5281/zenodo.3943577" ID-Zenodo-Dep="3943577" httpUri="https://zenodo.org/record/3943577/files/figure.png" pageId="10" pageNumber="322" targetBox="[364,1229,1530,1948]" targetPageId="10">
<paragraph id="0B6A36CDFF9EC070FF03F851FAE5F824" blockId="10.[159,1429,1966,2016]" pageId="10" pageNumber="322">
FIG. 7.
<emphasis id="39A1EADFFF9EC070FF71F851FE5FF807" box="[237,406,1966,1987]" italics="true" pageId="10" pageNumber="322">
<taxonomicName id="CCD54D4EFF9EC070FF71F851FE5BF807" baseAuthorityName="Schreber" baseAuthorityYear="1774" box="[237,402,1966,1987]" class="Mammalia" family="Nycteridae" genus="Nycteris" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="322" phylum="Chordata" rank="species" species="hispida">Nycteris hispida</taxonomicName>
,
</emphasis>
HZM.52.40188. (A): Anterior, left mandible; (B): left, upper incisors and canine; (C): right tragus; (D): baculum (dorsal view [left] and lateral view [right]). Scales for A, B, and D = 1 mm; scale for C = 2 mm
</paragraph>
</caption>
<subSubSection id="43CF6546FF9FC071FF03FE59FD83FDE1" pageId="11" pageNumber="323" type="discussion">
<paragraph id="0B6A36CDFF9FC071FF03FE59FEADFE04" blockId="11.[159,771,422,549]" box="[159,356,422,448]" pageId="11" pageNumber="323">
<emphasis id="39A1EADFFF9FC071FF03FE59FEADFE04" box="[159,356,422,448]" italics="true" pageId="11" pageNumber="323">Taxonomic notes</emphasis>
</paragraph>
<paragraph id="0B6A36CDFF9FC071FF59FE37FD83FDE1" blockId="11.[159,771,422,549]" pageId="11" pageNumber="323">
Specimens from
<collectingCountry id="73C2765DFF9FC071FE10FE37FE13FE26" box="[396,474,456,482]" name="Democratic Republic of the Congo" pageId="11" pageNumber="323">Congo</collectingCountry>
are referred to the nominate subspecies (
<bibRefCitation id="6F444B3CFF9FC071FEEFFE16FF14FDE1" author="Van Cakenberghe and De Vree" firstAuthor="Van Cakenberghe" pageId="11" pageNumber="323" pagination="299 - 332" refId="ref14805" refString="VAN CAKENBERGHE, V., and F. DE VREE. 1993. Systematics of African Nycteris (Mammalia: Chiroptera) Part II. The Nycteris hispida group. Bonner zoologische Beitrage, 44: 299 - 332." type="journal article" year="1993">Van Cakenberghe and De Vree, 1993</bibRefCitation>
;
<bibRefCitation id="6F444B3CFF9FC071FF76FDF4FDF7FDE1" author="Happold and Happold" box="[234,574,523,549]" firstAuthor="Happold" pageId="11" pageNumber="323" refId="ref13898" refString="HAPPOLD, M., and D. C. D. HAPPOLD (eds.). 2013. Mammals of Africa Volume IV: Hedgehogs, shrews, and bats. Bloomsbury Publishing, London, 800 pp." type="book" year="2013">Happold and Happold, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>