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<document id="33D4540E40688A9011966E588E158471" ID-DOI="10.5281/zenodo.293907" ID-GBIF-Dataset="e8c1af78-5bb0-4705-9f21-7c15bd42137a" ID-ISSN="1175-5326" ID-Zenodo-Dep="293907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460215710041" checkinUser="plazi" docAuthor="Bosselaers, Jan, Dierick, Manuel, Cnudde, Veerle, Masschaele, Bert, Hoorebeke, Luc Van &amp; Jacobs, Patric" docDate="2010" docId="832C87E64C7FFF867EBFE985978EE80A" docLanguage="en" docName="zt02427p035.pdf" docOrigin="Zootaxa 2427" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Donuea collustrata Bosselaers &amp; Dierick, sp. nov." docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="7F15FF9E4C7DFF817E28ED77920EEC4D" masterDocTitle="High-resolution X-ray computed tomography of an extant new Donuea (Araneae: Liocranidae) species in Madagascan copal" masterLastPageNumber="35" masterPageNumber="25" pageNumber="27" updateTime="1698237010531" updateUser="plazi">
<mods:mods id="12ED511F77FE7FAE4A174E774E060E78" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="98B65FE343F300F853A1312D1891EFD8">High-resolution X-ray computed tomography of an extant new Donuea (Araneae: Liocranidae) species in Madagascan copal</mods:title>
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<mods:namePart id="0F9A41E4E72124668968E3D3C71E82FA">Bosselaers, Jan</mods:namePart>
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<mods:namePart id="2E8DCE2246522DBA9B930758DCB5A75C">Dierick, Manuel</mods:namePart>
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<mods:namePart id="906BC7A387172F500DCFFE6FA1F4461D">Cnudde, Veerle</mods:namePart>
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<mods:namePart id="3CEB41F7A2E003D877F31FB4D686FF51">Masschaele, Bert</mods:namePart>
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<mods:namePart id="F53F9AC2D6245F38C23E7661A8EC26FE">Hoorebeke, Luc Van</mods:namePart>
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<mods:namePart id="D5AAA457EC8DF86F068BC3FE5B4B6B22">Jacobs, Patric</mods:namePart>
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<mods:date id="C612CDFCF8B22F7FA4F951D17F680662">2010</mods:date>
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<treatment id="832C87E64C7FFF867EBFE985978EE80A" ID-DOI="http://doi.org/10.5281/zenodo.6201660" ID-GBIF-Taxon="159608007" ID-Zenodo-Dep="6201660" LSID="urn:lsid:plazi:treatment:832C87E64C7FFF867EBFE985978EE80A" httpUri="http://treatment.plazi.org/id/832C87E64C7FFF867EBFE985978EE80A" lastPageId="7" lastPageNumber="32" pageId="2" pageNumber="27">
<subSubSection id="439F657B4C7FFF837EBFE9859035E97F" pageId="2" pageNumber="27" type="nomenclature">
<paragraph id="0B3A36F04C7FFF837EBFE98590FEE941" blockId="2.[151,752,1266,1330]" box="[151,752,1266,1292]" pageId="2" pageNumber="27">
<heading id="5072819C4C7FFF837EBFE98590FEE941" bold="true" box="[151,752,1266,1292]" fontSize="11" level="1" pageId="2" pageNumber="27" reason="1">
<emphasis id="39F1EAE24C7FFF837EBFE98590FEE941" bold="true" box="[151,752,1266,1292]" pageId="2" pageNumber="27">
<taxonomicName id="CC854D734C7FFF837EBFE9859082E941" authority="Bosselaers &amp; Dierick" authorityName="Bosselaers &amp; Dierick" box="[151,652,1266,1292]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="collustrata" status="sp. nov.">
<emphasis id="39F1EAE24C7FFF837EBFE9859370E941" bold="true" box="[151,382,1266,1292]" italics="true" pageId="2" pageNumber="27">Donuea collustrata</emphasis>
Bosselaers &amp; Dierick
</taxonomicName>
<taxonomicNameLabel id="22C257994C7FFF837CBBE98590FEE941" box="[659,752,1266,1292]" pageId="2" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B3A36F04C7FFF837EBFE86F9035E97F" blockId="2.[151,752,1266,1330]" box="[151,571,1304,1330]" pageId="2" pageNumber="27">
<figureCitation id="93BE2A754C7FFF837EBFE86F9329E97F" box="[151,295,1304,1330]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="2" pageNumber="27">Figs. 1 AE</figureCitation>
; 2 AF; 3 AM; 4 AI.
</paragraph>
</subSubSection>
<subSubSection id="439F657B4C7FFF837EBFE8159744E9EF" pageId="2" pageNumber="27" type="materials_examined">
<paragraph id="0B3A36F04C7FFF837EBFE8159744E9EF" blockId="2.[151,1437,1378,2002]" pageId="2" pageNumber="27">
<emphasis id="39F1EAE24C7FFF837EBFE8159344E931" bold="true" box="[151,330,1378,1404]" pageId="2" pageNumber="27">
<typeStatus id="D43E88524C7FFF837EBFE81592DCE931" box="[151,210,1378,1404]" pageId="2" pageNumber="27">Type</typeStatus>
material.
</emphasis>
<typeStatus id="D43E88524C7FFF837F7CE81593CFE931" box="[340,449,1378,1404]" pageId="2" pageNumber="27" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="739276604C7FFF837C38E8159091E931" box="[528,671,1378,1404]" name="Madagascar" pageId="2" pageNumber="27">Madagascar</collectingCountry>
, Foulpointe, forest on clay soil, beating vegetation,
<date id="7F3B10304C7FFF837AD6E815977DE931" box="[1278,1395,1378,1404]" pageId="2" pageNumber="27" value="1995-07">July 1995</date>
, A. Pauly leg. (
<collectionCode id="6D94AE354C7FFF837F33E8FF937DE9EF" box="[283,371,1416,1442]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="27">MRAC</collectionCode>
206172).
<typeStatus id="D43E88524C7FFF837FC3E8FF905CE9EF" box="[491,594,1416,1442]" pageId="2" pageNumber="27" type="paratype">Paratype</typeStatus>
male, in copal from the Sambava area, NE
<collectingCountry id="739276604C7FFF837A79E8FF96EEE9EF" box="[1105,1248,1416,1442]" name="Madagascar" pageId="2" pageNumber="27">Madagascar</collectingCountry>
,
<collectionCode id="6D94AE354C7FFF837AC4E8FF9749E9EF" box="[1260,1351,1416,1442]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="27">MRAC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="439F657B4C7FFF837EEEE8DA91B4EA7A" pageId="2" pageNumber="27" type="diagnosis">
<paragraph id="0B3A36F04C7FFF837EEEE8DA91B4EA7A" blockId="2.[151,1437,1378,2002]" pageId="2" pageNumber="27">
<emphasis id="39F1EAE24C7FFF837EEEE8DA9348E98A" bold="true" box="[198,326,1453,1479]" pageId="2" pageNumber="27">Diagnosis.</emphasis>
The species can be distinguished from the
<typeStatus id="D43E88524C7FFF837D62E8DA9173E98A" box="[842,893,1453,1479]" pageId="2" pageNumber="27">type</typeStatus>
species
<taxonomicName id="CC854D734C7FFF837DCAE8DA9743E98A" authority="Simon, 1903" authorityName="Simon" authorityYear="1903" box="[994,1357,1453,1479]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="39F1EAE24C7FFF837DCAE8DA96A5E98B" box="[994,1195,1453,1478]" italics="true" pageId="2" pageNumber="27">Donuea decorsei</emphasis>
<bibRefCitation id="6F144B014C7FFF837A9CE8DA9743E98A" author="Simon" box="[1204,1357,1453,1479]" pageId="2" pageNumber="34" refString="Simon, E. (1903) Descriptions d'arachnides nouveaux de Madagascar, faisant partie des collections du Museum. Bulletin du Museum d'histoire naturelle de Paris 9, 133 - 140." type="journal article" year="1903">Simon, 1903</bibRefCitation>
</taxonomicName>
, by its bifid RTA as opposed to a simple one (
<bibRefCitation id="6F144B014C7FFF837CAFE8A59169E9A1" author="Simon" box="[647,871,1490,1516]" pageId="2" pageNumber="34" refString="Simon, E. (1903) Descriptions d'arachnides nouveaux de Madagascar, faisant partie des collections du Museum. Bulletin du Museum d'histoire naturelle de Paris 9, 133 - 140." type="journal article" year="1903">Simon 1903: 138</bibRefCitation>
), its large, bifurcated MA, a simple, pointed conductor situated ventrally instead of dorsally of MA and its shorter embolus which does not describe a circular loop before passing behind the tegulum (
<figureCitation id="93BE2A754C7FFF837CFEEB6A9129EA7A" box="[726,807,1565,1591]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="2" pageNumber="27">Figs. 1</figureCitation>
F, 4 C, F, I).
</paragraph>
</subSubSection>
<subSubSection id="439F657B4C7FFF867EEEEB3593AAEE71" lastPageId="7" lastPageNumber="32" pageId="2" pageNumber="27" type="description">
<paragraph id="0B3A36F04C7FFF867EEEEB359089ED12" blockId="2.[151,1437,1378,2002]" lastBlockId="7.[151,1437,152,1095]" lastPageId="7" lastPageNumber="32" pageId="2" pageNumber="27">
<emphasis id="39F1EAE24C7FFF837EEEEB359350EA11" bold="true" box="[198,350,1602,1628]" pageId="2" pageNumber="27">Description.</emphasis>
Body length without chelicerae (
<typeStatus id="D43E88524C7FFF837CD0EB359168EA11" box="[760,870,1602,1628]" pageId="2" pageNumber="27" type="holotype">holotype</typeStatus>
): 4.34, with chelicerae: 5.76. Carapace length: 1.81; w: 1.55, oval, yellow brown, darker in eye region, and with two brown lateral bands (
<figureCitation id="93BE2A754C7FFF837AF6EB1F9780EACF" box="[1246,1422,1640,1666]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="2" pageNumber="27">Fig. 1 AB, D</figureCitation>
). Fovea short and deep, pale brown, length 0.08, anterior end 1.14 from front. Eight eyes in two rows, ringed with black, AER w: 0.82, recurved from above, slightly procurved from front, PER w: 0.89, procurved from above, strongly procurved from front (
<figureCitation id="93BE2A754C7FFF837C4FEBAF90F1EABF" box="[615,767,1752,1778]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="2" pageNumber="27">Figs. 1 A, D</figureCitation>
,
<figureCitation id="93BE2A754C7FFF837D25EBAF914DEABF" box="[781,835,1752,1778]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="2" pageNumber="27">3 M</figureCitation>
). MOQ depth: 0.32, anterior w: 0.37, posterior w: 0.42. All eyes subequal, ALE oval, others circular. All four median eyes with dark retina restricted to median 2/3. Anterior median eyes slightly closer to each other than to ALE. Eyes of PER equidistant, separated by about twice their diameter. ALE and PLE touching, on common tubercle. Clypeus vertical, 1/3 of diameter of AME. Chilum orange-brown, sclerotised, split (
<figureCitation id="93BE2A754C7FFF837CF6EA1A9140EBCA" box="[734,846,1901,1927]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="2" pageNumber="27">Fig. 3 M</figureCitation>
). Chelicerae large, yellow brown, elongated and pointing forward, parallel to the longitudinal axis of the body (
<figureCitation id="93BE2A754C7FFF837DBBEAE59623EBE1" box="[915,1069,1938,1964]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="2" pageNumber="27">Figs. 1 AD</figureCitation>
,
<figureCitation id="93BE2A754C7FFF837A12EAE59689EBE1" box="[1082,1159,1938,1964]" captionStart="FIGURE 2 A H" captionStartId="4.[151,259,1488,1512]" captionTargetBox="[151,1436,194,1466]" captionTargetId="figure@4.[151,1436,194,1466]" captionTargetPageId="4" captionText="FIGURE 2 A H. Donuea collustrata sp. n. A, copal piece holding specimen, after trimming; B, copal preserved specimen, do; C, copal specimen, ve; D, frontal view of copal specimen, showing opaque layer of air bubbles; E, leg spination scheme of holotype, legend of unfolded article below; F, copal specimen, reconstruction of habitus, do; G, right male palp of copal specimen, rl; H, left male palp of copal specimen, pl. Scale bars: B, C: 1.0; F: 1.5; H: 0.25." httpUri="https://zenodo.org/record/293909/files/figure.png" pageId="2" pageNumber="27">2 C, F</figureCitation>
,
<figureCitation id="93BE2A754C7FFF837ABCEAE596CBEBE1" box="[1172,1221,1938,1964]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="2" pageNumber="27">3 A</figureCitation>
). One large kneeshaped seta and a row of long, curved setae on ACR, cheliceral fangs brown. Six teeth on ACR, largest one closest to fang base, followed by an isolated very small tooth and two pairs of small teeth separated by a small distance. Seven teeth on PCR, a pair of medium sized teeth close to fang base, followed at some distance by five small teeth in a row. Sternum length: 1.05; w: 0.95, smooth, yellow, elongated and shield-shaped, with pale broad and blunt ICS between legs III and II-III, and thin, brown and pointed PCT facing all coxae (
<figureCitation id="93BE2A754C79FF857B44EBAC92E5EB5A" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="4" pageNumber="29">Fig. 1 C, E</figureCitation>
). Pleural bars (
<bibRefCitation id="6F144B014C79FF857F82EB8A90F3EB5A" author="Bosselaers" box="[426,765,1789,1815]" pageId="4" pageNumber="33" refString="Bosselaers, J. &amp; Jocque, R. (2002) Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae. Zoologica Scripta, 31, 241 - 270." type="journal article" year="2002">Bosselaers &amp; Jocqué 2002</bibRefCitation>
: 247, fig. 1P;
<bibRefCitation id="6F144B014C79FF857D98EB8A9642EB5A" author="Simon" box="[944,1100,1789,1815]" pageId="4" pageNumber="34" refString="Simon, E. (1892) Histoire Naturelle des Araignees. Tome 1. Roret, Paris. 1084 pp." type="book" year="1892">Simon 1892</bibRefCitation>
: 11, fig. 29) yellow-brown continuous strips, weakly protruding between coxae. Labium brown, subtrapezoidal, as long as it is wide at its base, with white, thickened anterior rim. Endites subrectangular, with apical hair tuft and serrula, no oblique depression (
<figureCitation id="93BE2A754C79FF857F01EA1293BDEB32" box="[297,435,1893,1919]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="4" pageNumber="29">Fig. 1 C, E</figureCitation>
). Abdomen cream, unicolorous ventrally and with two lateral, purple brown bands dorsally (
<figureCitation id="93BE2A754C79FF857F23EAFF93CFEBEF" box="[267,449,1928,1954]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="4" pageNumber="29">Fig. 1 AB, D</figureCitation>
). ALS contiguous, conical, with short, blunt apical segment. PMS very thin and slender, PLS subcylindrical, mottled with grey, with pale, rounded apical segment (
<figureCitation id="93BE2A754C79FF857A51EADC971EEB88" box="[1145,1296,1963,1989]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="4" pageNumber="29">Figs. 1 BD</figureCitation>
,
<figureCitation id="93BE2A754C79FF857B34EADC9742EB88" box="[1308,1356,1963,1989]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="4" pageNumber="29">3 A</figureCitation>
). Legs yellow, fe, ti and mt ringed with grey basally and/or terminally (
<figureCitation id="93BE2A754C79FF857DABEABA9618EBAA" box="[899,1046,1997,2023]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="4" pageNumber="29">Figs. 1 AC</figureCitation>
,
<figureCitation id="93BE2A754C79FF857A09EABA9681EBAA" box="[1057,1167,1997,2023]" captionStart="FIGURE 2 A H" captionStartId="4.[151,259,1488,1512]" captionTargetBox="[151,1436,194,1466]" captionTargetId="figure@4.[151,1436,194,1466]" captionTargetPageId="4" captionText="FIGURE 2 A H. Donuea collustrata sp. n. A, copal piece holding specimen, after trimming; B, copal preserved specimen, do; C, copal specimen, ve; D, frontal view of copal specimen, showing opaque layer of air bubbles; E, leg spination scheme of holotype, legend of unfolded article below; F, copal specimen, reconstruction of habitus, do; G, right male palp of copal specimen, rl; H, left male palp of copal specimen, pl. Scale bars: B, C: 1.0; F: 1.5; H: 0.25." httpUri="https://zenodo.org/record/293909/files/figure.png" pageId="4" pageNumber="29">2 BC, F</figureCitation>
). No trochanter notch, no retrocoxal hymen (
<bibRefCitation id="6F144B014C7AFF867FB2EDEF9022ECFF" author="Raven" box="[410,556,152,178]" pageId="7" pageNumber="34" refString="Raven, R. (1998) Revision of the Australian genera of the Miturgidae with a preview of their relationships. In: XIVth International Congress of Arachnology, Abstracts, p. 31." type="book chapter" year="1998">Raven 1998</bibRefCitation>
;
<bibRefCitation id="6F144B014C7AFF867C11EDEF917AECFF" author="Bosselaers" box="[569,884,152,178]" pageId="7" pageNumber="33" refString="Bosselaers, J. &amp; Jocque, R. (2002) Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae. Zoologica Scripta, 31, 241 - 270." type="journal article" year="2002">Bosselaers &amp; Jocqué 2002</bibRefCitation>
), patellar indentation long and narrow, ¾ of pa length. Metatarsi III and IV with sparse ve terminal preening brush. Tarsi with two claws and dense claw tufts. Leg formula 1423. Leg spination (
<figureCitation id="93BE2A754C7AFF867C02EDAA9083ECBA" box="[554,653,221,247]" captionStart="FIGURE 2 A H" captionStartId="4.[151,259,1488,1512]" captionTargetBox="[151,1436,194,1466]" captionTargetId="figure@4.[151,1436,194,1466]" captionTargetPageId="4" captionText="FIGURE 2 A H. Donuea collustrata sp. n. A, copal piece holding specimen, after trimming; B, copal preserved specimen, do; C, copal specimen, ve; D, frontal view of copal specimen, showing opaque layer of air bubbles; E, leg spination scheme of holotype, legend of unfolded article below; F, copal specimen, reconstruction of habitus, do; G, right male palp of copal specimen, rl; H, left male palp of copal specimen, pl. Scale bars: B, C: 1.0; F: 1.5; H: 0.25." httpUri="https://zenodo.org/record/293909/files/figure.png" pageId="7" pageNumber="32">Fig. 2 E</figureCitation>
): fe: palp do 0-1-1 rlv 1-1-1-1-1-0; I pl 0-1-1-1 do 0-1-1-1; II pl 0- 0-0-1 do 0-1-1-1; III do 0-1-1-2; IV do 1-1-1-1-2-2; pa: palp pl 1; ti: palp pl 2-0-0; I pl 0-1-1 ve 2-2-2-2-2; II ve 2-2-2-2; III rl 0-1-0 plv 1-1-0; IV rl 0-1-1-0 plv 1-1-0 rlv 0-0-1; mt: I pl 1-0-0 ve 2-2-2; II ve 2-2-0; III pl 0- 1-0 rl 1-0-0 ve 2-0-0; IV rl 0-0-1 ve 2-0-2.
</paragraph>
<caption id="5FFA66784C7EFF827EBFEACE96A0EBBD" httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="3" pageNumber="28" targetBox="[151,1436,197,1951]" targetPageId="3">
<paragraph id="0B3A36F04C7EFF827EBFEACE96A0EBBD" blockId="3.[151,1436,1977,2032]" pageId="3" pageNumber="28">
<emphasis id="39F1EAE24C7EFF827EBFEACE935FEB9C" bold="true" box="[151,337,1977,2001]" pageId="3" pageNumber="28">FIGURE 1 AE.</emphasis>
<taxonomicName id="CC854D734C7EFF827F70EACE9029EB9D" box="[344,551,1977,2000]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="collustrata" status="sp. nov.">
<emphasis id="39F1EAE24C7EFF827F70EACE9029EB9D" box="[344,551,1977,2000]" italics="true" pageId="3" pageNumber="28">Donuea collustrata</emphasis>
</taxonomicName>
<emphasis id="39F1EAE24C7EFF827C05EACE9066EB9C" bold="true" box="[557,616,1977,2001]" pageId="3" pageNumber="28">
<taxonomicNameLabel id="22C257994C7EFF827C05EACE9066EB9C" box="[557,616,1977,2001]" pageId="3" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="39F1EAE24C7EFF827CC8EACE90FCEB9C" bold="true" box="[736,754,1977,2001]" pageId="3" pageNumber="28">F.</emphasis>
<taxonomicName id="CC854D734C7EFF827CD3EACE91BEEB9D" box="[763,944,1977,2000]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="39F1EAE24C7EFF827CD3EACE91BEEB9D" box="[763,944,1977,2000]" italics="true" pageId="3" pageNumber="28">Donuea decorsei</emphasis>
</taxonomicName>
, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5.
</paragraph>
</caption>
<caption id="5FFA66784C79FF857EBFE8A79725EA0A" httpUri="https://zenodo.org/record/293909/files/figure.png" pageId="4" pageNumber="29" targetBox="[151,1436,194,1466]" targetPageId="4">
<paragraph id="0B3A36F04C79FF857EBFE8A79725EA0A" blockId="4.[151,1436,1488,1607]" pageId="4" pageNumber="29">
<emphasis id="39F1EAE24C79FF857EBFE8A79350E9A5" bold="true" box="[151,350,1488,1512]" pageId="4" pageNumber="29">FIGURE 2 AH.</emphasis>
<taxonomicName id="CC854D734C79FF857F4FE8A7904CE9AA" box="[359,578,1488,1511]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="collustrata" status="sp. nov.">
<emphasis id="39F1EAE24C79FF857F4FE8A7904CE9AA" box="[359,578,1488,1511]" italics="true" pageId="4" pageNumber="29">Donuea collustrata</emphasis>
</taxonomicName>
<emphasis id="39F1EAE24C79FF857C61E8A79084E9A5" bold="true" box="[585,650,1488,1512]" pageId="4" pageNumber="29">
<taxonomicNameLabel id="22C257994C79FF857C61E8A79084E9A5" box="[585,650,1488,1512]" pageId="4" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A, copal piece holding specimen, after trimming; B, copal preserved specimen, do; C, copal specimen, ve; D, frontal view of copal specimen, showing opaque layer of air bubbles; E, leg spination scheme of holotype, legend of unfolded article below; F, copal specimen, reconstruction of habitus, do; G, right male palp of copal specimen, rl; H, left male palp of copal specimen, pl. Scale bars: B, C: 1.0; F: 1.5; H: 0.25.
</paragraph>
</caption>
<caption id="5FFA66784C78FF847EBFEBAE90E7EBDD" httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="5" pageNumber="30" targetBox="[151,1436,194,1731]" targetPageId="5">
<paragraph id="0B3A36F04C78FF847EBFEBAE90E7EBDD" blockId="5.[151,1436,1753,1936]" pageId="5" pageNumber="30">
<emphasis id="39F1EAE24C78FF847EBFEBAE9354EABC" bold="true" box="[151,346,1753,1777]" pageId="5" pageNumber="30">FIGURE 3 AM.</emphasis>
<taxonomicName id="CC854D734C78FF847F48EBAE9021EABD" box="[352,559,1753,1776]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="collustrata" status="sp. nov.">
<emphasis id="39F1EAE24C78FF847F48EBAE9021EABD" box="[352,559,1753,1776]" italics="true" pageId="5" pageNumber="30">Donuea collustrata</emphasis>
</taxonomicName>
<emphasis id="39F1EAE24C78FF847C1DEBAE907FEABC" bold="true" box="[565,625,1753,1777]" pageId="5" pageNumber="30">
<taxonomicNameLabel id="22C257994C78FF847C1DEBAE907FEABC" box="[565,625,1753,1777]" pageId="5" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened.
</paragraph>
</caption>
<caption id="5FFA66784C7BFF877EBFEAEC93CBEBBF" httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="6" pageNumber="31" targetBox="[318,1274,178,1926]" targetPageId="6">
<paragraph id="0B3A36F04C7BFF877EBFEAEC93CBEBBF" blockId="6.[151,1436,1947,2034]" pageId="6" pageNumber="31">
<emphasis id="39F1EAE24C7BFF877EBFEAEC9344EBFE" bold="true" box="[151,330,1947,1971]" pageId="6" pageNumber="31">FIGURE 4 AI.</emphasis>
<taxonomicName id="CC854D734C7BFF877F79EAEC9011EBFF" box="[337,543,1947,1970]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="collustrata" status="sp. nov.">
<emphasis id="39F1EAE24C7BFF877F79EAEC9011EBFF" box="[337,543,1947,1970]" italics="true" pageId="6" pageNumber="31">Donuea collustrata</emphasis>
</taxonomicName>
<emphasis id="39F1EAE24C7BFF877C0DEAEC906FEBFE" bold="true" box="[549,609,1947,1971]" pageId="6" pageNumber="31">
<taxonomicNameLabel id="22C257994C7BFF877C0DEAEC906FEBFE" box="[549,609,1947,1971]" pageId="6" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male palp. AC, rl; DF, ve; GI, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25.
</paragraph>
</caption>
<paragraph id="0B3A36F04C7AFF867EECEC1F96F1EE5A" blockId="7.[151,1437,152,1095]" pageId="7" pageNumber="32">
<collectingCountry id="739276604C7AFF867EECEC1F930EEDCF" box="[196,256,360,386]" name="Malta" pageId="7" pageNumber="32">Male</collectingCountry>
palp with a simple, blunt PTA (
<figureCitation id="93BE2A754C7AFF867C52EC1F9120EDCF" box="[634,814,360,386]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="7" pageNumber="32">Figs. 3 D, E, L</figureCitation>
,
<figureCitation id="93BE2A754C7AFF867D12EC1F9176EDCF" box="[826,888,360,386]" captionStart="FIGURE 4 A I" captionStartId="6.[151,255,1947,1971]" captionTargetBox="[318,1274,178,1926]" captionTargetId="figure@6.[305,1282,168,1938]" captionTargetPageId="6" captionText="FIGURE 4 A I. Donuea collustrata sp. n., male palp. A C, rl; D F, ve; G I, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25." httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="7" pageNumber="32">4 F, I</figureCitation>
) and a bifid RTA consisting of a long, pointed do part and a cup-shaped ve part (
<figureCitation id="93BE2A754C7AFF867C0AECFA90EAEDEA" box="[546,740,397,423]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="7" pageNumber="32">
Figs.
<date id="7F3B10304C7AFF867C4CECFA908BEDEA" box="[612,645,397,423]" pageId="7" pageNumber="32">3 I</date>
, J, K, L
</figureCitation>
,
<figureCitation id="93BE2A754C7AFF867CD8ECFA914FEDEA" box="[752,833,397,423]" captionStart="FIGURE 4 A I" captionStartId="6.[151,255,1947,1971]" captionTargetBox="[318,1274,178,1926]" captionTargetId="figure@6.[305,1282,168,1938]" captionTargetPageId="6" captionText="FIGURE 4 A I. Donuea collustrata sp. n., male palp. A C, rl; D F, ve; G I, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25." httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="7" pageNumber="32">4 AC</figureCitation>
). Palpal tarsus elongated and piriform, with a large, bifid, flag-shaped MA (
<figureCitation id="93BE2A754C7AFF867F93ECC59040ED81" box="[443,590,434,460]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="7" pageNumber="32">Figs. 3 CF</figureCitation>
,
<figureCitation id="93BE2A754C7AFF867C73ECC590A8ED81" box="[603,678,434,460]" captionStart="FIGURE 4 A I" captionStartId="6.[151,255,1947,1971]" captionTargetBox="[318,1274,178,1926]" captionTargetId="figure@6.[305,1282,168,1938]" captionTargetPageId="6" captionText="FIGURE 4 A I. Donuea collustrata sp. n., male palp. A C, rl; D F, ve; G I, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25." httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="7" pageNumber="32">4 DI</figureCitation>
), a simple, blunt-tipped conductor situated ventrally of the pl branch of the MA (
<figureCitation id="93BE2A754C7AFF867F52ECAF9029EDBF" box="[378,551,472,498]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="7" pageNumber="32">Figs. 3 CF, L</figureCitation>
,
<figureCitation id="93BE2A754C7AFF867C1BECAF9077EDBF" box="[563,633,472,498]" captionStart="FIGURE 4 A I" captionStartId="6.[151,255,1947,1971]" captionTargetBox="[318,1274,178,1926]" captionTargetId="figure@6.[305,1282,168,1938]" captionTargetPageId="6" captionText="FIGURE 4 A I. Donuea collustrata sp. n., male palp. A C, rl; D F, ve; G I, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25." httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="7" pageNumber="32">4 DI</figureCitation>
) and a long embolus emerging from below the rl branch of the MA, proceeding basally and subsequently passing behind the tegulum (
<figureCitation id="93BE2A754C7AFF867DB5EC8A9678EE5A" box="[925,1142,509,535]" captionStart="FIGURE 3 A M" captionStartId="5.[151,255,1753,1777]" captionTargetBox="[151,1436,194,1731]" captionTargetId="figure@5.[151,1436,194,1731]" captionTargetPageId="5" captionText="FIGURE 3 A M. Donuea collustrata sp. n., copal specimen, X-ray CT images. A, habitus, ve; B, transversal section of right male palp, showing conductor (green) and embolus (pink) originating from behind MA (blue); C, frontal view of right male palp, colour coding as in B; D, left male palp, pl; E, left male palp, pl-ve; F, left male palp, ve; G, left male palp, rl-ve; H, left male palp, rl; I, right male palp, rl-do view, showing embolus, PTA and RTA; J, right male palp, rl-ve, showing bifid RTA; K, right male palp, rl; L, caudal view of right male palp, showing RTA (left) and PTA (right); M, frontal view of eye region, chilum artificially darkened." httpUri="https://zenodo.org/record/293910/files/figure.png" pageId="7" pageNumber="32">Figs. 3 BC, GK</figureCitation>
,
<figureCitation id="93BE2A754C7AFF867AAAEC8A96FCEE5A" box="[1154,1266,509,535]" captionStart="FIGURE 4 A I" captionStartId="6.[151,255,1947,1971]" captionTargetBox="[318,1274,178,1926]" captionTargetId="figure@6.[305,1282,168,1938]" captionTargetPageId="6" captionText="FIGURE 4 A I. Donuea collustrata sp. n., male palp. A C, rl; D F, ve; G I, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25." httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="7" pageNumber="32">4 AC, F</figureCitation>
).
</paragraph>
<paragraph id="0B3A36F04C7AFF867EECEF5593AAEE71" blockId="7.[151,1437,152,1095]" box="[196,420,546,572]" pageId="7" pageNumber="32">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="439F657B4C7AFF867EEEEF3090B5EECA" pageId="7" pageNumber="32" type="etymology">
<paragraph id="0B3A36F04C7AFF867EEEEF3090B5EECA" blockId="7.[151,1437,152,1095]" pageId="7" pageNumber="32">
<emphasis id="39F1EAE24C7AFF867EEEEF30935DEE2C" bold="true" box="[198,339,583,609]" pageId="7" pageNumber="32">Etymology.</emphasis>
The species epithet
<taxonomicName id="CC854D734C7AFF867C6DEF3F90CAEE2C" box="[581,708,584,609]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="collustrata">
<emphasis id="39F1EAE24C7AFF867C6DEF3F90CAEE2C" box="[581,708,584,609]" italics="true" pageId="7" pageNumber="32">collustrata</emphasis>
</taxonomicName>
, irradiated, refers to the X-ray tomography technique used as an aid in the description of the present species.
</paragraph>
</subSubSection>
<subSubSection id="439F657B4C7AFF867EEEEFE5978EE80A" pageId="7" pageNumber="32" type="discussion">
<paragraph id="0B3A36F04C7AFF867EEEEFE59082EFB1" blockId="7.[151,1437,152,1095]" pageId="7" pageNumber="32">
<emphasis id="39F1EAE24C7AFF867EEEEFE5934EEEE1" bold="true" box="[198,320,658,684]" pageId="7" pageNumber="32">Remarks.</emphasis>
The specimen preserved in copal clearly is conspecific with the field-captured
<typeStatus id="D43E88524C7AFF867ACDEFE5975EEEE1" box="[1253,1360,658,684]" pageId="7" pageNumber="32" type="holotype">holotype</typeStatus>
. It has the same size (body length 4.21) and general morphology, and both specimens are identical with respect to a number of diagnostic details. Leg spination of the copal specimen is identical to the
<typeStatus id="D43E88524C7AFF867A55EFAA96EAEEBA" box="[1149,1252,733,759]" pageId="7" pageNumber="32" type="holotype">holotype</typeStatus>
although a few spines are missing, probably having been lost when the animal got caught in the sticky resin (
<figureCitation id="93BE2A754C7AFF867AD6EE75979EEF51" box="[1278,1424,770,796]" captionStart="FIGURE 2 A H" captionStartId="4.[151,259,1488,1512]" captionTargetBox="[151,1436,194,1466]" captionTargetId="figure@4.[151,1436,194,1466]" captionTargetPageId="4" captionText="FIGURE 2 A H. Donuea collustrata sp. n. A, copal piece holding specimen, after trimming; B, copal preserved specimen, do; C, copal specimen, ve; D, frontal view of copal specimen, showing opaque layer of air bubbles; E, leg spination scheme of holotype, legend of unfolded article below; F, copal specimen, reconstruction of habitus, do; G, right male palp of copal specimen, rl; H, left male palp of copal specimen, pl. Scale bars: B, C: 1.0; F: 1.5; H: 0.25." httpUri="https://zenodo.org/record/293909/files/figure.png" pageId="7" pageNumber="32">Figs. 2 EF</figureCitation>
). The copal specimen has lost most of its original colour, the abdomen being partly transparent and showing two dark spots in the anterior half, presumably corresponding with the book lungs. However, the leg annulations have been mostly preserved and coincide with those of the field-captured animal (
<figureCitation id="93BE2A754C7AFF867AC5EE05978FEFC1" box="[1261,1409,882,908]" captionStart="FIGURE 1 A E" captionStartId="3.[151,255,1977,2001]" captionTargetBox="[151,1436,197,1951]" captionTargetId="figure@3.[151,1436,197,1953]" captionTargetPageId="3" captionText="FIGURE 1 A E. Donuea collustrata sp. n., holotype. F. Donuea decorsei, holotype, MNHN. A, habitus, do; B, habitus, lat.; C, habitus, ve; D, body, do; E, prosoma, ve; F, left male palp, ve. Scale bars: D, E: 1.0; F: 0.5." httpUri="https://zenodo.org/record/293908/files/figure.png" pageId="7" pageNumber="32">Figs. 1 AC</figureCitation>
,
<figureCitation id="93BE2A754C7AFF867BA5EE0592FEEFFF" captionStart="FIGURE 2 A H" captionStartId="4.[151,259,1488,1512]" captionTargetBox="[151,1436,194,1466]" captionTargetId="figure@4.[151,1436,194,1466]" captionTargetPageId="4" captionText="FIGURE 2 A H. Donuea collustrata sp. n. A, copal piece holding specimen, after trimming; B, copal preserved specimen, do; C, copal specimen, ve; D, frontal view of copal specimen, showing opaque layer of air bubbles; E, leg spination scheme of holotype, legend of unfolded article below; F, copal specimen, reconstruction of habitus, do; G, right male palp of copal specimen, rl; H, left male palp of copal specimen, pl. Scale bars: B, C: 1.0; F: 1.5; H: 0.25." httpUri="https://zenodo.org/record/293909/files/figure.png" pageId="7" pageNumber="32">2 B, C, F</figureCitation>
). Most significantly, palpal structure of both specimens is identical in minute detail, as can be proven by juxtaposing stereomicroscope observations of the field-captured
<typeStatus id="D43E88524C7AFF867DEBEECA9625EF9A" box="[963,1067,957,983]" pageId="7" pageNumber="32" type="holotype">holotype</typeStatus>
with X-ray micro-CT scans of the copal preserved specimen (
<figureCitation id="93BE2A754C7AFF867C29EE95908EEFB1" box="[513,640,994,1020]" captionStart="FIGURE 4 A I" captionStartId="6.[151,255,1947,1971]" captionTargetBox="[318,1274,178,1926]" captionTargetId="figure@6.[305,1282,168,1938]" captionTargetPageId="6" captionText="FIGURE 4 A I. Donuea collustrata sp. n., male palp. A C, rl; D F, ve; G I, pl; A, D, G, X-ray CT images of right palp of copal specimen, inverted; B, E, H, stereomicroscope photographs of left palp of holotype; C, F, I, drawings of left palp of holotype. Scale bar: 0.25." httpUri="https://zenodo.org/record/293911/files/figure.png" pageId="7" pageNumber="32">Fig. 4 AI</figureCitation>
).
</paragraph>
<paragraph id="0B3A36F04C7AFF867EECE97F978EE80A" blockId="7.[151,1437,152,1095]" pageId="7" pageNumber="32">
The assignment of
<taxonomicName id="CC854D734C7AFF867F8EE97F900DE86C" box="[422,515,1032,1057]" class="Arachnida" family="Liocranidae" genus="Donuea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="39F1EAE24C7AFF867F8EE97F900DE86C" box="[422,515,1032,1057]" italics="true" pageId="7" pageNumber="32">Donuea</emphasis>
</taxonomicName>
to
<taxonomicName id="CC854D734C7AFF867C02E97F90B8E86F" box="[554,694,1032,1058]" class="Arachnida" family="Liocranidae" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="32" phylum="Arthropoda" rank="family">Liocranidae</taxonomicName>
is debatable, and it can indeed be argued that the genus would be better placed in
<taxonomicName id="CC854D734C7AFF867F5EE95A9007E80A" box="[374,521,1069,1095]" class="Arachnida" family="Clubionidae" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="32" phylum="Arthropoda" rank="family">Clubionidae</taxonomicName>
. However, a formal transfer is outside the scope of the present contribution.
</paragraph>
</subSubSection>
</treatment>
</document>