treatments-xml/data/81/7C/87/817C8781FFFCFFB9A2E2FBBBFD1AFD43.xml
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<document id="8207971232DF193ED644944B9AA97FA0" ID-CLB-Dataset="7506" ID-DOI="10.11646/zootaxa.4950.1.1" ID-GBIF-Dataset="30557468-7467-4c99-b6fc-806f4be784e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4643245" ID-ZooBank="B9578A01-9B27-40B9-BEF9-C6DEB714C652" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617001503997" checkinUser="plazi" docAuthor="Ramalho, Laís V., Moraes, Fernando C., Salgado, Leonardo T., Bastos, Alex C. &amp; Moura, Rodrigo L." docDate="2021" docId="817C8781FFFCFFB9A2E2FBBBFD1AFD43" docLanguage="en" docName="zootaxa.4950.1.1.pdf" docOrigin="Zootaxa 4950 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stephanollona domuspusilla Ramalho &amp; Moraes &amp; Salgado &amp; Bastos &amp; Moura 2021, n. sp." docType="treatment" docUuid="F5C3D8C0-EDD8-475F-B8D7-4F6E2EC037F1" docUuidSource="ZooBank" docVersion="7" lastPageNumber="26" masterDocId="7D45FFF9FFEAFFA0A275FF88FFFFFFEA" masterDocTitle="Bryozoa from the reefs off the Amazon River mouth: checklist, thirteen new species, and notes on their ecology and distribution" masterLastPageNumber="45" masterPageNumber="1" pageNumber="23" updateTime="1698935303943" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0CE662E5719547699F1B7E9F681F9388">Bryozoa from the reefs off the Amazon River mouth: checklist, thirteen new species, and notes on their ecology and distribution</mods:title>
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<mods:namePart id="7B4D5F919B9FDA1D5F757A618CFE3001">Ramalho, Laís V.</mods:namePart>
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<mods:affiliation id="D19D256C5BDBFE0C0EE85B8174BDCAE0">Diretoria de Pesquisa, Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rio de Janeiro-RJ, Brazil. &amp; laisvr 10 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1459 - 3954</mods:affiliation>
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<mods:namePart id="BF1C1AD9AC91EE184B36578688CF4B9C">Moraes, Fernando C.</mods:namePart>
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<mods:affiliation id="5908C5447465B3A6D68D3124E60E53CD">Diretoria de Pesquisa, Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rio de Janeiro-RJ, Brazil. &amp; fmoraes @ mn. ufrj. br; https: // orcid. org / 0000 - 0001 - 8251 - 6868</mods:affiliation>
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<mods:namePart id="96552D1739BC874CADDC1262AC539534">Salgado, Leonardo T.</mods:namePart>
<mods:nameIdentifier id="9F559B4B1D2B87D5F613804FF678B3F6" type="ORCID">0000-0003-3215-9633</mods:nameIdentifier>
<mods:affiliation id="BBBAE69E226008C9A92CA20A8668F7EB">Diretoria de Pesquisa, Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rio de Janeiro-RJ, Brazil. &amp; lsalgado @ jbrj. gov. br; https: // orcid. org / 0000 - 0003 - 3215 - 9633</mods:affiliation>
<mods:nameIdentifier id="084F0364127CE0B7F492C03FE2C54183" type="email">lsalgado@jbrj.gov.br</mods:nameIdentifier>
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<mods:namePart id="F314B8C32FF8955009513779AB4236D2">Bastos, Alex C.</mods:namePart>
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<mods:affiliation id="BD9AE8A212E1CB0668BC7390A0518B9E">Departamento de Oceanografia, Universidade Federal do Espírito Santo, Vitória-ES, Brazil. alexcardosobastos @ gmail. com; https: // orcid. org / 0000 - 0002 - 1272 - 1134</mods:affiliation>
<mods:nameIdentifier id="08B0AF29F8A7E1664B2056613B316994" type="email">alexcardosobastos@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A6274EE7CFE622F838818FB6CBBCE865">Moura, Rodrigo L.</mods:namePart>
<mods:nameIdentifier id="568419F1D871C38D0BBFFA248F08231F" type="ORCID">0000-0002-5597-6196</mods:nameIdentifier>
<mods:affiliation id="8131F913F0CB26A22E60640B05800380">Instituto de Biologia and SAGE / COPPE, Universidade Federal do Rio de Janeiro, Rio de Janeiro-RJ, Brazil. moura. uesc @ gmail. com; https: // orcid. org / 0000 - 0002 - 5597 - 6196</mods:affiliation>
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<mods:title id="A2D75F65D0F18450EE0FCB70DF95ECD4">Zootaxa</mods:title>
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<mods:date id="99445CB44A1779EE22F849ADCE4763BF">2021</mods:date>
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<treatment id="817C8781FFFCFFB9A2E2FBBBFD1AFD43" ID-DOI="http://doi.org/10.5281/zenodo.4643291" ID-GBIF-Taxon="180875116" ID-Zenodo-Dep="4643291" LSID="urn:lsid:zoobank.org:act:F5C3D8C0-EDD8-475F-B8D7-4F6E2EC037F1" httpUri="http://treatment.plazi.org/id/817C8781FFFCFFB9A2E2FBBBFD1AFD43" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
<subSubSection id="41CF651CFFFCFFB6A2E2FBBBFCC6FBA4" box="[151,825,1075,1102]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="096A3697FFFCFFB6A2E2FBBBFCC6FBA4" blockId="22.[151,825,1075,1139]" box="[151,825,1075,1102]" pageId="22" pageNumber="23">
<heading id="522281FBFFFCFFB6A2E2FBBBFCC6FBA4" bold="true" box="[151,825,1075,1102]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="3BA1EA85FFFCFFB6A2E2FBBBFCC6FBA4" bold="true" box="[151,825,1075,1102]" pageId="22" pageNumber="23">
<taxonomicName id="CED54D14FFFCFFB6A2E2FBBBFD11FBA4" authority="Ramalho &amp; Moraes" authorityName="Ramalho &amp; Moraes &amp; Salgado &amp; Bastos &amp; Moura" authorityYear="2021" box="[151,750,1075,1102]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="23" phylum="Bryozoa" rank="species" species="domuspusilla" status="sp. nov.">
<emphasis id="3BA1EA85FFFCFFB6A2E2FBBBFE0FFBA4" bold="true" box="[151,496,1075,1102]" italics="true" pageId="22" pageNumber="23">Stephanollona domuspusilla</emphasis>
Ramalho &amp; Moraes
</taxonomicName>
<taxonomicNameLabel id="209257FEFFFCFFB6A083FBBCFCC6FBA4" box="[758,825,1076,1102]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="41CF651CFFFCFFB6A2E2FBD0FE70FB99" box="[151,399,1112,1139]" pageId="22" pageNumber="23" type="description">
<paragraph id="096A3697FFFCFFB6A2E2FBD0FE70FB99" blockId="22.[151,825,1075,1139]" box="[151,399,1112,1139]" pageId="22" pageNumber="23">
(
<figureCitation id="91EE2A12FFFCFFB6A2EAFBD0FED6FB99" box="[159,297,1112,1139]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Figs 11D, E</figureCitation>
;
<figureCitation id="91EE2A12FFFCFFB6A340FBD0FE79FB99" box="[309,390,1112,1139]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">12A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="41CF651CFFFCFFB6A2E2FB28FAA7FB35" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="096A3697FFFCFFB6A2E2FB28FAA7FB35" blockId="22.[151,1437,1184,2038]" pageId="22" pageNumber="23">
<emphasis id="3BA1EA85FFFCFFB6A2E2FB28FE7FFB50" bold="true" box="[151,384,1184,1210]" pageId="22" pageNumber="23">Material examined.</emphasis>
<materialsCitation id="B9BD3CCAFFFCFFB6A3F2FB28FAA7FB35" ID-GBIF-Occurrence="3068349310" collectingDate="2014-09-26" collectorName="Fernando Moraes &amp; Rodrigo Moura" country="Brazil" elevation="90" latitude="3.590445" location="Cruzeiro do Sul" longLatPrecision="1" longitude="-49.126713" municipality="Sta" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
<typeStatus id="D66E8835FFFCFFB6A3F2FB28FE0BFB50" box="[391,500,1184,1210]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
: MNRJBRY-1543:
<collectingCountry id="71C27607FFFCFFB6A0BBFB28FCEAFB50" box="[718,789,1184,1210]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
,
<collectingRegion id="CB11F875FFFCFFB6A16BFB28FC56FB51" box="[798,937,1184,1211]" country="Brazil" name="Amapa" pageId="22" pageNumber="23">Amapá state</collectingRegion>
(
<collectingMunicipality id="E90EACEDFFFCFFB6A1CCFB28FC1FFB50" box="[953,992,1184,1210]" pageId="22" pageNumber="23">Sta</collectingMunicipality>
#3,
<geoCoordinate id="6CE15050FFFCFFB6A672FB28FB59FB51" box="[1031,1190,1184,1211]" degrees="03" direction="north" minutes="35.4267" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="3.590445">03°35.4267N</geoCoordinate>
<geoCoordinate id="6CE15050FFFCFFB6A6C3FB28FA91FB51" box="[1206,1390,1184,1211]" degrees="049" direction="west" minutes="07.6028" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-49.126713">049°07.6028W</geoCoordinate>
),
<quantity id="CE2D9B72FFFCFFB6A7F5FB28FF50FB35" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="23" unit="m" value="90.0">
<elevation id="82F8D1A4FFFCFFB6A7F5FB28FF50FB35" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="23" unit="m" value="90.0">90 m</elevation>
</quantity>
, on sponge,
<date id="7D6B1057FFFCFFB6A34EFB4CFDE7FB34" box="[315,536,1220,1246]" pageId="22" pageNumber="23" value="2014-09-26">
<collectingDate id="6D2FE9BFFFFCFFB6A34EFB4CFDE7FB34" box="[315,536,1220,1246]" pageId="22" pageNumber="23" value="2014-09-26">26 September 2014</collectingDate>
</date>
, collected by
<collectorName id="A4205341FFFCFFB6A0C7FB4CFC88FB35" box="[690,887,1220,1247]" pageId="22" pageNumber="23">Fernando Moraes</collectorName>
&amp;
<collectorName id="A4205341FFFCFFB6A1EEFB4CFBB7FB35" box="[923,1096,1220,1247]" pageId="22" pageNumber="23">Rodrigo Moura</collectorName>
(NHo
<location id="0C0A604CFFFCFFB6A6E3FB4CFAB4FB35" LSID="urn:lsid:plazi:treatment:817C8781FFFCFFB9A2E2FBBBFD1AFD43:0C0A604CFFFCFFB6A6E3FB4CFAB4FB35" box="[1174,1355,1220,1247]" country="Brazil" latitude="3.590445" longLatPrecision="1" longitude="-49.126713" municipality="Sta" name="Cruzeiro do Sul" pageId="22" pageNumber="23" stateProvince="Amapa">Cruzeiro do Sul</location>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="41CF651CFFFCFFB6A2B2FB60FDAAFACC" pageId="22" pageNumber="23" type="etymology">
<paragraph id="096A3697FFFCFFB6A2B2FB60FDAAFACC" blockId="22.[151,1437,1184,2038]" pageId="22" pageNumber="23">
<emphasis id="3BA1EA85FFFCFFB6A2B2FB60FEB1FAE8" bold="true" box="[199,334,1256,1282]" pageId="22" pageNumber="23">Etymology.</emphasis>
From the Latin noun
<emphasis id="3BA1EA85FFFCFFB6A031FB60FD72FAE8" box="[580,653,1256,1282]" italics="true" pageId="22" pageNumber="23">domus</emphasis>
(meaning home) plus the adjective
<emphasis id="3BA1EA85FFFCFFB6A655FB60FB93FAE8" box="[1056,1132,1256,1282]" italics="true" pageId="22" pageNumber="23">pusilla</emphasis>
, -
<emphasis id="3BA1EA85FFFCFFB6A6F1FB61FB62FAE8" box="[1156,1181,1257,1282]" italics="true" pageId="22" pageNumber="23">us</emphasis>
, -
<emphasis id="3BA1EA85FFFCFFB6A6C0FB61FB28FAE8" box="[1205,1239,1257,1282]" italics="true" pageId="22" pageNumber="23">um</emphasis>
(meaning small), referring to the small size of the zooids.
</paragraph>
</subSubSection>
<subSubSection id="41CF651CFFFCFFB6A2B2FAB8FAB4FA5C" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="096A3697FFFCFFB6A2B2FAB8FAB4FA5C" blockId="22.[151,1437,1184,2038]" pageId="22" pageNumber="23">
<emphasis id="3BA1EA85FFFCFFB6A2B2FAB8FEBDFAA0" bold="true" box="[199,322,1328,1354]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Autozooids with tubercular, imperforate frontal shield; 57 oral spines, the most proximal pair lanceolate; orifice with distal border denticulate and U-shaped sinus constricted by small lateral teeth; condyles projecting beyond the sinus border. Two kinds of adventitious avicularia: small, single and rounded suboral avicularium with serrated rostrum, and another large, elongate avicularium, triangular with rounded tip, placed laterally.
</paragraph>
</subSubSection>
<subSubSection id="41CF651CFFFCFFB6A2B2FA48FE63F81C" pageId="22" pageNumber="23" type="description">
<paragraph id="096A3697FFFCFFB6A2B2FA48FE63F81C" blockId="22.[151,1437,1184,2038]" pageId="22" pageNumber="23">
<emphasis id="3BA1EA85FFFCFFB6A2B2FA48FEA6FA30" bold="true" box="[199,345,1472,1498]" pageId="22" pageNumber="23">Description.</emphasis>
Colony encrusting, multiserial, uni- to multilaminar (
<figureCitation id="91EE2A12FFFCFFB6A1B9FA48FBC8FA30" box="[972,1079,1472,1498]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Fig. 11D</figureCitation>
). Autozooids small, oval to irregularly polygonal, longer than wide [L 317369441 (SD 35, N 11); W 242302
<quantity id="CE2D9B72FFFCFFB6A639FA6CFB5BFA15" box="[1100,1188,1508,1535]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" pageId="22" pageNumber="23" unit="mm" value="360.0">360 µm</quantity>
(SD 42, N 11)]; frontal shield tubercular, imperforate except for some (57) scattered medium to large sized [1424
<quantity id="CE2D9B72FFFCFFB6A69DF980FACDF9C9" box="[1256,1330,1544,1571]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="22" pageNumber="23" unit="mm" value="39.0">39 µm</quantity>
diameter (SD 6, N 27)], rounded, marginal areolar pores (
<figureCitation id="91EE2A12FFFCFFB6A0B1F9A4FCAEF9AD" box="[708,849,1580,1607]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Figs 11D, E</figureCitation>
,
<figureCitation id="91EE2A12FFFCFFB6A12AF9A4FC4BF9AD" box="[863,948,1580,1607]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">12A, B</figureCitation>
). Orifice cleithridiate, longer than wide [L 116133147 (SD 10, N 11); W 101111
<quantity id="CE2D9B72FFFCFFB6A00DF9D8FD31F981" box="[632,718,1616,1643]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="22" pageNumber="23" unit="mm" value="118.0">118 µm</quantity>
(SD 7, N 11)], with 57 (usually six) hollow spines [1326
<quantity id="CE2D9B72FFFCFFB6A7F4F9D8FF42F965" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="22" pageNumber="23" unit="mm" value="35.0">35 µm</quantity>
diameter (SD 6, N 17)], four visible in ovicellate zooids; the most proximal pair of spines lanceolate, the others spear-shaped with irregular calcification (
<figureCitation id="91EE2A12FFFCFFB6A0E1F910FD03F958" box="[660,764,1688,1714]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Figs 11E</figureCitation>
,
<figureCitation id="91EE2A12FFFCFFB6A17DF910FCA3F959" box="[776,860,1688,1715]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">12A, B</figureCitation>
); distal border denticulate with 1418 small beads; U-shaped sinus constricted by small lateral teeth; two long downward facing condyles projecting beyond the sinus border (
<figureCitation id="91EE2A12FFFCFFB6A29AF968FEA8F910" box="[239,343,1760,1786]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Figs 11E</figureCitation>
,
<figureCitation id="91EE2A12FFFCFFB6A316F968FE49F911" box="[355,438,1760,1787]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">12A, B</figureCitation>
); one zooid with a broader and shallower sinus observed (
<figureCitation id="91EE2A12FFFCFFB6A630F968FB56F910" box="[1093,1193,1760,1786]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Fig. 11E</figureCitation>
). Two
<typeStatus id="D66E8835FFFCFFB6A680F969FACFF911" box="[1269,1328,1761,1787]" pageId="22" pageNumber="23">types</typeStatus>
of avicularia: (1) single, small [L 4257
<quantity id="CE2D9B72FFFCFFB6A07EF88CFDABF8F5" box="[523,596,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="22" pageNumber="23" unit="mm" value="73.0">73 µm</quantity>
(SD 10, N 7)], rounded avicularium placed suborally (rarely laterally) and oriented laterally, usually inclined in respect to the frontal wall; rostrum rounded, serrated, and crossbar complete without columella (
<figureCitation id="91EE2A12FFFCFFB6A303F8C4FE1FF88C" box="[374,480,1868,1894]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Figs 11E</figureCitation>
,
<figureCitation id="91EE2A12FFFCFFB6A39BF8C4FDBBF88D" box="[494,580,1868,1895]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">12A, B</figureCitation>
); (2) single large, elongate [L 187203
<quantity id="CE2D9B72FFFCFFB6A663F8C4FB90F88D" box="[1046,1135,1868,1895]" metricMagnitude="-1" metricUnit="m" metricValue="2.19" pageId="22" pageNumber="23" unit="mm" value="219.0">219 µm</quantity>
(SD 16, N 3)], somewhat triangular avicularium, placed laterally to orifice; rostrum triangular with rounded and slightly hooked tip, slightly curved to one side; crossbar complete without columella (
<figureCitation id="91EE2A12FFFCFFB6A159F81CFC6AF844" box="[812,917,1940,1966]" captionStart="FIGURE 11" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[235,1353,182,1810]" captionTargetId="figure-18@23.[234,1353,181,1810]" captionTargetPageId="23" captionText="FIGURE 11. AC. Stephanollona aff. contracta (Waters, 1899) (MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="22" pageNumber="23">Figs 11E</figureCitation>
,
<figureCitation id="91EE2A12FFFCFFB6A1D6F81CFC28F844" box="[931,983,1940,1966]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">12A</figureCitation>
). Ovicell hyperstomial, wider than long [L 146149151 (SD 4, N 2); W 181186
<quantity id="CE2D9B72FFFCFFB6A00FF830FD30F839" box="[634,719,1976,2003]" metricMagnitude="-1" metricUnit="m" metricValue="1.91" pageId="22" pageNumber="23" unit="mm" value="191.0">191 µm</quantity>
(SD 7, N 2)], with smooth ooecial surface similar to the zooidal frontal wall (
<figureCitation id="91EE2A12FFFCFFB6A35DF854FE70F81C" box="[296,399,2012,2038]" captionStart="FIGURE 12" captionStartId="24.[151,250,1620,1644]" captionTargetBox="[151,1435,181,1596]" captionTargetId="figure-32@24.[151,1436,181,1596]" captionTargetPageId="24" captionText="FIGURE 12. A, B. Stephanollona domuspusilla Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1543).A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF. Pourtalesella duoavicularia Ramalho &amp; Moraes n. sp. (Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="22" pageNumber="23">Fig. 12B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="5DAA661FFFFDFFB7A2E2F8A2FE40F81D" ID-DOI="http://doi.org/10.5281/zenodo.4643277" ID-Zenodo-Dep="4643277" httpUri="https://zenodo.org/record/4643277/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1834,1858]" targetBox="[235,1353,182,1810]" targetPageId="23">
<paragraph id="096A3697FFFDFFB7A2E2F8A2FE40F81D" blockId="23.[151,1437,1834,2039]" pageId="23" pageNumber="24">
<emphasis id="3BA1EA85FFFDFFB7A2E2F8A2FEE6F8A8" bold="true" box="[151,281,1834,1858]" pageId="23" pageNumber="24">FIGURE 11</emphasis>
. AC.
<taxonomicName id="CED54D14FFFDFFB7A311F8A2FCD8F8A8" authority="(Waters, 1899)" baseAuthorityName="Waters" baseAuthorityYear="1899" box="[356,807,1834,1858]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="24" phylum="Bryozoa" rank="species" species="contracta">
<emphasis id="3BA1EA85FFFDFFB7A311F8A2FE09F8A8" box="[356,502,1834,1858]" italics="true" pageId="23" pageNumber="24">Stephanollona</emphasis>
aff.
<emphasis id="3BA1EA85FFFDFFB7A052F8A3FD77F8A8" box="[551,648,1835,1858]" italics="true" pageId="23" pageNumber="24">contracta</emphasis>
(
<bibRefCitation id="6D444B66FFFDFFB7A0E2F8A2FCDFF8A8" author="Waters, A. W." box="[663,800,1834,1858]" pageId="23" pageNumber="24" pagination="6 - 16" refId="ref31089" refString="Waters, A. W. (1899) Bryozoa from Madeira. Journal of the Royal Microscopical Society, 1899, 6 - 16. https: // doi. org / 10.1111 / j. 1365 - 2818.1899. tb 00139. x" type="journal article" year="1899">Waters, 1899</bibRefCitation>
)
</taxonomicName>
(MNRJBRY-1515). A. General view of the colony. B. Distal part of a zooid with elongate, latero-oral avicularium and orifice with six spines and a shallow, broad, almost rectangular sinus. C. Zooids bearing rounded avicularia with serrated rostrum and ovicell with a frontal, uncalcified area. D, E.
<taxonomicName id="CED54D14FFFDFFB7A697F8FAFE2DF844" authority="Ramalho &amp; Moraes" authorityName="Ramalho &amp; Moraes &amp; Salgado &amp; Bastos &amp; Moura" authorityYear="2021" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="24" phylum="Bryozoa" rank="species" species="domuspusilla" status="sp. nov.">
<emphasis id="3BA1EA85FFFDFFB7A697F8FAFEFAF844" italics="true" pageId="23" pageNumber="24">Stephanollona domuspusilla</emphasis>
Ramalho &amp; Moraes
</taxonomicName>
<emphasis id="3BA1EA85FFFDFFB7A3A2F81EFDEFF844" bold="true" box="[471,528,1942,1966]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="209257FEFFFDFFB7A3A2F81EFDEFF844" box="[471,528,1942,1966]" pageId="23" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Holotype MNRJBRY-1543). D. Colony showing irregular growth pattern. E. Autozooids showing distal lanceolate and spear-shaped oral spines, and one zooid with a broader and shallow sinus. Scale bars: A, 500 µm; B, C, 100 µm; D, E, 200 µm.
</paragraph>
</caption>
<caption id="5DAA661FFFF2FFB8A2E2F9DCFCD8F8AE" ID-DOI="http://doi.org/10.5281/zenodo.4643281" ID-Zenodo-Dep="4643281" httpUri="https://zenodo.org/record/4643281/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1620,1644]" targetBox="[151,1435,181,1596]" targetPageId="24">
<paragraph id="096A3697FFF2FFB8A2E2F9DCFCD8F8AE" blockId="24.[151,1437,1620,1860]" pageId="24" pageNumber="25">
<emphasis id="3BA1EA85FFF2FFB8A2E2F9DCFEE0F986" bold="true" box="[151,287,1620,1644]" pageId="24" pageNumber="25">FIGURE 12.</emphasis>
A, B.
<taxonomicName id="CED54D14FFF2FFB8A315F9DCFCB3F981" authority="Ramalho &amp; Moraes" authorityName="Ramalho &amp; Moraes &amp; Salgado &amp; Bastos &amp; Moura" authorityYear="2021" box="[352,844,1620,1644]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="25" phylum="Bryozoa" rank="species" species="domuspusilla" status="sp. nov.">
<emphasis id="3BA1EA85FFF2FFB8A315F9DCFD80F986" box="[352,639,1620,1644]" italics="true" pageId="24" pageNumber="25">Stephanollona domuspusilla</emphasis>
Ramalho &amp; Moraes
</taxonomicName>
<emphasis id="3BA1EA85FFF2FFB8A127F9DCFC74F986" bold="true" box="[850,907,1620,1644]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="209257FEFFF2FFB8A127F9DCFC74F986" box="[850,907,1620,1644]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Holotype MNRJBRY-1543). A. Autozooids having orifice with U-shaped sinus, six oral spines, a suboral avicularium with rounded and serrated rostrum, and a different type of avicularium (lower right) that is elongate, more or less triangular. B. Ovicellate zooid with suboral avicularium. CF.
<taxonomicName id="CED54D14FFF2FFB8A738F915FDBCF93D" authority="Ramalho &amp; Moraes" authorityName="Ramalho &amp; Moraes &amp; Salgado &amp; Bastos &amp; Moura" authorityYear="2021" class="Gymnolaemata" family="Celleporidae" genus="Pourtalesella" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="25" phylum="Bryozoa" rank="species" species="duoavicularia" status="sp. nov.">
<emphasis id="3BA1EA85FFF2FFB8A738F915FE90F932" italics="true" pageId="24" pageNumber="25">Pourtalesella duoavicularia</emphasis>
Ramalho &amp; Moraes
</taxonomicName>
<emphasis id="3BA1EA85FFF2FFB8A03EF948FD79F932" bold="true" box="[587,646,1728,1752]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="209257FEFFF2FFB8A03EF948FD79F932" box="[587,646,1728,1752]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Holotype MNRJBRY-1443). C. General view of the colony. D. Autozooids showing two types of avicularia (see arrows), and orifice surrounded by a raised peristome. E. Ovicell with uncalcified, frontal area and two different kinds of avicularia (arrow). F. Part of a colony showing irregular growth form, two kinds of avicularia (arrow) and ovicells. Scale bars: A, B, DF, 100 µm; C, 500 µm.
</paragraph>
</caption>
<subSubSection id="41CF651CFFF2FFB9A2B2F8FAFD1AFD43" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="096A3697FFF2FFB9A2B2F8FAFD9DFE38" blockId="24.[151,1436,1906,2041]" lastBlockId="25.[151,1437,151,681]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="3BA1EA85FFF2FFB8A2B2F8FAFEC4F866" bold="true" box="[199,315,1906,1932]" pageId="24" pageNumber="25">Remarks.</emphasis>
The two most similar species recorded in
<collectingCountry id="71C27607FFF2FFB8A162F8FAFCA4F866" box="[791,859,1906,1932]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
are
<taxonomicName id="CED54D14FFF2FFB8A1F9F8FAFA63F866" authority="Vieira et al., 2010 b" authorityName="Vieira" authorityYear="2010" box="[908,1436,1906,1933]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="25" phylum="Bryozoa" rank="species" species="arborescens">
<emphasis id="3BA1EA85FFF2FFB8A1F9F8FAFB42F866" box="[908,1213,1906,1932]" italics="true" pageId="24" pageNumber="25">Stephanollona arborescens</emphasis>
<bibRefCitation id="6D444B66FFF2FFB8A6B1F8FAFA63F866" author="Vieira, L. M. &amp; Gordon, D. P. &amp; Souza, F. B. C. &amp; Haddad, M. A." box="[1220,1436,1906,1933]" pageId="24" pageNumber="25" pagination="1 - 53" refId="ref30797" refString="Vieira, L. M., Gordon, D. P., Souza, F. B. C. &amp; Haddad, M. A. (2010 b) New and little-known cheilostomatous Bryozoa from the south and southeastern Brazilian continental shelf and slope. Zootaxa, 2722 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 2722.1.1" type="journal article" year="2010">
Vieira
<emphasis id="3BA1EA85FFF2FFB8A765F8FBFABEF866" box="[1296,1345,1906,1932]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
., 2010b
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="CED54D14FFF2FFB8A2B3F81EFDE5F85A" authority="Vieira et al., 2010 b" authorityName="Vieira" authorityYear="2010" box="[198,538,1942,1969]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="25" phylum="Bryozoa" rank="species" species="angusta">
<emphasis id="3BA1EA85FFF2FFB8A2B3F81EFEC4F85A" box="[198,315,1942,1968]" italics="true" pageId="24" pageNumber="25">S. angusta</emphasis>
<bibRefCitation id="6D444B66FFF2FFB8A334F81EFDE5F85A" author="Vieira, L. M. &amp; Gordon, D. P. &amp; Souza, F. B. C. &amp; Haddad, M. A." box="[321,538,1942,1969]" pageId="24" pageNumber="25" pagination="1 - 53" refId="ref30797" refString="Vieira, L. M., Gordon, D. P., Souza, F. B. C. &amp; Haddad, M. A. (2010 b) New and little-known cheilostomatous Bryozoa from the south and southeastern Brazilian continental shelf and slope. Zootaxa, 2722 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 2722.1.1" type="journal article" year="2010">
Vieira
<emphasis id="3BA1EA85FFF2FFB8A3F9F81FFE43F85A" box="[396,444,1942,1968]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
., 2010b
</bibRefCitation>
</taxonomicName>
. The former species has the same small, rounded avicularium and six spines, but differs from
<taxonomicName id="CED54D14FFF2FFB8A353F832FD41F83F" authority="Ramalho &amp; Moraes" authorityName="Ramalho &amp; Moraes &amp; Salgado &amp; Bastos &amp; Moura" authorityYear="2021" box="[294,702,1978,2005]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="25" phylum="Bryozoa" rank="species" species="domuspusilla" status="sp. nov.">
<emphasis id="3BA1EA85FFF2FFB8A353F832FE28F83E" box="[294,471,1978,2004]" italics="true" pageId="24" pageNumber="25">S. domuspusilla</emphasis>
Ramalho &amp; Moraes
</taxonomicName>
<emphasis id="3BA1EA85FFF2FFB8A0B0F832FCFAF83E" bold="true" box="[709,773,1978,2004]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="209257FEFFF2FFB8A0B0F832FCFAF83E" box="[709,773,1978,2004]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in having larger zooids (L 346468667, W 259346
<quantity id="CE2D9B72FFF2FFB8A706F832FF3FF813" metricMagnitude="-1" metricUnit="m" metricValue="4.45" pageId="24" pageNumber="25" unit="mm" value="445.0">445 µm</quantity>
), smaller orifice wider than long (L 8094111, W 93103
<quantity id="CE2D9B72FFF2FFB8A114F856FC46F813" box="[865,953,2014,2041]" metricMagnitude="-1" metricUnit="m" metricValue="1.17" pageId="24" pageNumber="25" unit="mm" value="117.0">117 µm</quantity>
), condyles encroaching on the sinus (in the new species the condyles slightly surpass the opening), small avicularium usually proximo-laterally placed and proximo-laterally directed (in the new species the avicularium is suboral and laterally directed), and occasionally elongate avicularia rarely associated with the orifice (while in the new species it is lateral to orifice), and the rostrum is not hooked and the lateral margins are almost straight (see fig. 87, 89 in
<bibRefCitation id="6D444B66FFF3FFB9A191FE8BFB45FEF4" author="Vieira, L. M. &amp; Gordon, D. P. &amp; Souza, F. B. C. &amp; Haddad, M. A." box="[996,1210,259,286]" pageId="25" pageNumber="26" pagination="1 - 53" refId="ref30797" refString="Vieira, L. M., Gordon, D. P., Souza, F. B. C. &amp; Haddad, M. A. (2010 b) New and little-known cheilostomatous Bryozoa from the south and southeastern Brazilian continental shelf and slope. Zootaxa, 2722 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 2722.1.1" type="journal article" year="2010">
Vieira
<emphasis id="3BA1EA85FFF3FFB9A645FE8BFB9EFEF7" box="[1072,1121,259,285]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2010b
</bibRefCitation>
).
<taxonomicName id="CED54D14FFF3FFB9A6BBFE8BFF2BFEAA" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="species" species="angusta">
<emphasis id="3BA1EA85FFF3FFB9A6BBFE8BFF2BFEAA" italics="true" pageId="25" pageNumber="26">Stephanollona angusta</emphasis>
</taxonomicName>
has two or more avicularia per autozooid, small, rounded avicularium with smooth rostrum placed proximolaterally to the orifice or proximally to the large avicularia.
<taxonomicName id="CED54D14FFF3FFB9A135FEC3FAA1FE8F" authority="Yang et al., 2018 b" authorityName="Yang" authorityYear="2018" box="[832,1374,331,357]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="species" species="boreopacifica">
<emphasis id="3BA1EA85FFF3FFB9A135FEC3FB7AFE8F" box="[832,1157,331,357]" italics="true" pageId="25" pageNumber="26">Stephanollona boreopacifica</emphasis>
<bibRefCitation id="6D444B66FFF3FFB9A6F9FEC3FAA1FE8F" author="Yang, H. J. &amp; Seo, J. E. &amp; Gordon, D. P." box="[1164,1374,331,357]" pageId="25" pageNumber="26" pagination="493 - 518" refId="ref31532" refString="Yang, H. J., Seo, J. E. &amp; Gordon, D. P. (2018 b) Sixteen new generic records of Korean Bryozoa from southern coastal waters and Jeju Island, East China Sea: evidence of tropical affinities. Zootaxa, 4422 (4), 493 - 518. https: // doi. org / 10.11646 / zootaxa. 4422.4.3" type="journal article" year="2018">
Yang
<emphasis id="3BA1EA85FFF3FFB9A6BBFEC3FAFEFE8F" box="[1230,1281,331,357]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
., 2018b
</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="71C27607FFF3FFB9A2E2FEE7FF21FE63" box="[151,222,367,393]" name="South Korea" pageId="25" pageNumber="26">Korea</collectingCountry>
, resembles the new species in having U-shaped sinus and rounded small avicularia with serrated borders, but can be distinguished by the narrower orifice (L 103138, W
<quantity id="CE2D9B72FFF3FFB9A146FE1AFC57FE47" box="[819,936,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="8.5" metricValueMin="7.0" pageId="25" pageNumber="26" unit="mm" value="77.5" valueMax="85.0" valueMin="70.0">7085 µm</quantity>
), and wider, more elongate avicularia associated with the ovicell (
<bibRefCitation id="6D444B66FFF3FFB9A3FEFE3FFDA9FE38" author="Yang, H. J. &amp; Seo, J. E. &amp; Gordon, D. P." box="[395,598,439,466]" pageId="25" pageNumber="26" pagination="493 - 518" refId="ref31532" refString="Yang, H. J., Seo, J. E. &amp; Gordon, D. P. (2018 b) Sixteen new generic records of Korean Bryozoa from southern coastal waters and Jeju Island, East China Sea: evidence of tropical affinities. Zootaxa, 4422 (4), 493 - 518. https: // doi. org / 10.11646 / zootaxa. 4422.4.3" type="journal article" year="2018">
Yang
<emphasis id="3BA1EA85FFF3FFB9A3B9FE3FFE02FE3B" box="[460,509,439,465]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2018b
</bibRefCitation>
).
</paragraph>
<paragraph id="096A3697FFF3FFB9A2B2FE53FD1AFD43" blockId="25.[151,1437,151,681]" pageId="25" pageNumber="26">
Several species included in
<taxonomicName id="CED54D14FFF3FFB9A38CFE53FD64FE1F" authorityName="Duvergier" authorityYear="1921" box="[505,667,475,501]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="genus">
<emphasis id="3BA1EA85FFF3FFB9A38CFE53FD64FE1F" box="[505,667,475,501]" italics="true" pageId="25" pageNumber="26">Stephanollona</emphasis>
</taxonomicName>
have broad, shallow orificial sinus, a feature characteristic of
<taxonomicName id="CED54D14FFF3FFB9A732FE53FE5AFDF3" authority="Vigneaux, 1949" authorityName="Vigneaux" authorityYear="1949" class="Gymnolaemata" family="Sfeniellidae" genus="Stephanollina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="genus">
<emphasis id="3BA1EA85FFF3FFB9A732FE53FF19FDF3" italics="true" pageId="25" pageNumber="26">Stephanollina</emphasis>
<bibRefCitation id="6D444B66FFF3FFB9A29AFE77FE5AFDF3" author="Vigneaux, M." box="[239,421,511,537]" pageId="25" pageNumber="26" pagination="1 - 155" refId="ref31056" refString="Vigneaux, M. (1949) Revision des bryozoaires neogenes du Bassin d'Aquitaine et essai de classification. Memoires de la Societe Geologique de France, 60, 1 - 155." type="journal article" year="1949">Vigneaux, 1949</bibRefCitation>
</taxonomicName>
, previously a subgenus of
<taxonomicName id="CED54D14FFF3FFB9A0A2FE77FC86FDF3" authorityName="Duvergier" authorityYear="1921" box="[727,889,511,537]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="genus">
<emphasis id="3BA1EA85FFF3FFB9A0A2FE77FC86FDF3" box="[727,889,511,537]" italics="true" pageId="25" pageNumber="26">Stephanollona</emphasis>
</taxonomicName>
, that
<bibRefCitation id="6D444B66FFF3FFB9A1C9FE77FB9AFDF0" author="Gordon, D. P." box="[956,1125,511,538]" pageId="25" pageNumber="26" pagination="65 - 72" refId="ref25487" refString="Gordon, D. P. (2006) Sfeniella, nom. nov. pro Sphenella Duvergier 1924, and Sfeniellidae, a new family of umbonuloid-shielded cheilostomes. Courier Forschungsinstitut Senckenberg, 2006, 65 - 72." type="journal article" year="2006">Gordon (2006)</bibRefCitation>
suggested to elevate to the rank of genus in the new family
<taxonomicName id="CED54D14FFF3FFB9A07EFDABFCC6FDD7" authority="Gordon, 2006" authorityName="Gordon" authorityYear="2006" box="[523,825,547,573]" class="Gymnolaemata" family="Sfeniellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="family">
Sfeniellidae
<bibRefCitation id="6D444B66FFF3FFB9A0ECFDABFCC6FDD7" author="Gordon, D. P." box="[665,825,547,573]" pageId="25" pageNumber="26" pagination="65 - 72" refId="ref25487" refString="Gordon, D. P. (2006) Sfeniella, nom. nov. pro Sphenella Duvergier 1924, and Sfeniellidae, a new family of umbonuloid-shielded cheilostomes. Courier Forschungsinstitut Senckenberg, 2006, 65 - 72." type="journal article" year="2006">Gordon, 2006</bibRefCitation>
</taxonomicName>
. More recently,
<bibRefCitation id="6D444B66FFF3FFB9A184FDABFACEFDD7" author="Di Martino, E. &amp; Taylor, P. D." box="[1009,1329,547,573]" pageId="25" pageNumber="26" pagination="783 - 796" refId="ref24863" refString="Di Martino, E. &amp; Taylor, P. D. (2017) Some Miocene cheilostome bryozoan genera of Michel Vigneaux - systematic revision and scanning electron microscopic study. Geodiversitas, 39, 783 - 796. https: // doi. org / 10.5252 / g 2017 n 4 a 7" type="journal article" year="2017">Di Martino &amp; Taylor (2017)</bibRefCitation>
followed
<bibRefCitation id="6D444B66FFF3FFB9A2E2FDCFFEADFD88" author="Gordon, D. P." box="[151,338,583,610]" pageId="25" pageNumber="26" pagination="65 - 72" refId="ref25487" refString="Gordon, D. P. (2006) Sfeniella, nom. nov. pro Sphenella Duvergier 1924, and Sfeniellidae, a new family of umbonuloid-shielded cheilostomes. Courier Forschungsinstitut Senckenberg, 2006, 65 - 72." type="journal article" year="2006">Gordons (2006)</bibRefCitation>
suggestions and included
<taxonomicName id="CED54D14FFF3FFB9A00EFDCFFCE8FD8B" authorityName="Vigneaux" authorityYear="1949" box="[635,791,583,609]" class="Gymnolaemata" family="Sfeniellidae" genus="Stephanollina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="genus">
<emphasis id="3BA1EA85FFF3FFB9A00EFDCFFCE8FD8B" box="[635,791,583,609]" italics="true" pageId="25" pageNumber="26">Stephanollina</emphasis>
</taxonomicName>
in
<taxonomicName id="CED54D14FFF3FFB9A149FDCFFC3DFD8B" authorityName="Gordon" authorityYear="2006" box="[828,962,583,609]" class="Gymnolaemata" family="Sfeniellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="family">Sfeniellidae</taxonomicName>
modifying both the diagnoses of the family and of the genus
<taxonomicName id="CED54D14FFF3FFB9A3F1FDE3FDDFFD6F" authorityName="Vigneaux" authorityYear="1949" box="[388,544,619,645]" class="Gymnolaemata" family="Sfeniellidae" genus="Stephanollina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="genus">
<emphasis id="3BA1EA85FFF3FFB9A3F1FDE3FDDFFD6F" box="[388,544,619,645]" italics="true" pageId="25" pageNumber="26">Stephanollina</emphasis>
</taxonomicName>
, selecting
<taxonomicName id="CED54D14FFF3FFB9A0EEFDE3FC6BFD6F" baseAuthorityName="Lamouroux" baseAuthorityYear="1816" box="[667,916,619,645]" class="Gymnolaemata" family="Sfeniellidae" genus="Stephanollina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="species" species="dentata">
<emphasis id="3BA1EA85FFF3FFB9A0EEFDE3FC6BFD6F" box="[667,916,619,645]" italics="true" pageId="25" pageNumber="26">Stephanollina dentata</emphasis>
</taxonomicName>
as the
<typeStatus id="D66E8835FFF3FFB9A19DFDE3FBE7FD6F" box="[1000,1048,619,645]" pageId="25" pageNumber="26">type</typeStatus>
species. Based on this, species of
<taxonomicName id="CED54D14FFF3FFB9A2E2FD07FEC6FD43" authorityName="Duvergier" authorityYear="1921" box="[151,313,655,681]" class="Gymnolaemata" family="Phidoloporidae" genus="Stephanollona" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="genus">
<emphasis id="3BA1EA85FFF3FFB9A2E2FD07FEC6FD43" box="[151,313,655,681]" italics="true" pageId="25" pageNumber="26">Stephanollona</emphasis>
</taxonomicName>
with shallow sinus must be reviewed.
</paragraph>
</subSubSection>
</treatment>
</document>