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<mods:title id="7862454FB6B0437D7FD1B28856187373">New Mesomantispinae (Insecta: Neuroptera: Mantispidae) from the Jurassic of Karatau, Kazakhstan</mods:title>
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<paragraph id="096D368DFFB4FF9AFF36CAE1DF66FF50" blockId="9.[151,362,151,176]" box="[151,362,151,176]" pageId="9" pageNumber="572">
<heading id="522581E1FFB4FF9AFF36CAE1DF66FF50" bold="true" box="[151,362,151,176]" fontSize="10" level="2" pageId="9" pageNumber="572" reason="2">
<taxonomicName id="CED24D0EFFB4FF9AFF36CAE1DF66FF50" authorityName="Makarkin" authorityYear="1996" box="[151,362,151,176]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">
<emphasis id="3BA6EA9FFFB4FF9AFF36CAE1DF66FF50" bold="true" box="[151,362,151,176]" pageId="9" pageNumber="572">Mesomantispinae</emphasis>
</taxonomicName>
</heading>
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<paragraph id="096D368DFFB4FF9AFF00CAA9DA0EFD80" blockId="9.[151,1436,223,608]" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF00CAA9DF6BFF18" authorityName="Makarkin" authorityYear="1996" box="[161,359,223,248]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
is composed of six genera and nine described species, and two indeterminate species one of
<taxonomicName id="CED24D0EFFB4FF9AFF36CB72DC51FEFC" ID-CoL="ZWC" authority="(Jepson et al., 2013)" authorityName="Jepson et al." authorityYear="2013" box="[151,605,260,284]" class="Insecta" family="Mantispidae" genus="Archaeodrepanicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFF36CB72DF7CFEFC" box="[151,368,260,284]" italics="true" pageId="9" pageNumber="572">Archaeodrepanicus</emphasis>
(
<bibRefCitation id="6D434B7CFFB4FF9AFEDECB72DC5AFEFC" box="[383,598,260,284]" pageId="9" pageNumber="572" refId="ref5832" refString="Jepson, J. E., Heads, S. W., Makarkin, V. N. &amp; Ren, D. (2013) New fossil Mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of North-Eastern China. Palaeontology, 56, 603 - 613. https: // doi. org / 10.1111 / pala. 12005" type="journal article">
Jepson
<emphasis id="3BA6EA9FFFB4FF9AFE74CB73DC19FEFC" box="[469,533,260,284]" italics="true" pageId="9" pageNumber="572">et al.,</emphasis>
2013
</bibRefCitation>
)
</taxonomicName>
and one
<taxonomicName id="CED24D0EFFB4FF9AFD64CB72DD87FEFD" authorityName="Makarkin" authorityYear="1996" box="[709,907,260,285]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
gen et sp. indet. (herein), the species are listed in
<tableCitation id="44500336FFB4FF9AFF15CB51DF04FEA0" box="[180,264,295,320]" captionStart="TABLE 1" captionStartId="9.[151,239,656,679]" captionText="TABLE 1. Fossil record of known Mesomantispinae, with locality, and age information" pageId="9" pageNumber="572">Table 1</tableCitation>
, other undescribed Mesozoic Chinese
<taxonomicName id="CED24D0EFFB4FF9AFD1FCB51DD8FFEA0" authorityName="Makarkin" authorityYear="1996" box="[702,899,295,320]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
fossils have also been discovered (
<bibRefCitation id="6D434B7CFFB4FF9AFAB0CB51DF0EFE84" pageId="9" pageNumber="572" refId="ref6332" refString="Makarkin, V. N., Yang, Q., Peng, Y. Y. &amp; Ren, D. (2012) A comparative overview of the neuropteran assemblage of the Lower Cretaceous Yixian Formation (China), with description of a new genus of Psychopsidae (Insecta: Neuroptera). Cretaceous Research, 35, 57 - 68. https: // doi. org / 10.1016 / j. cretres. 2011.11.013" type="journal article">
Makarkin
<emphasis id="3BA6EA9FFFB4FF9AFA29CB5FDEB8FE84" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
, 2012
</bibRefCitation>
). The subfamily represents one of the earliest records of the family
<taxonomicName id="CED24D0EFFB4FF9AFBBACB3ADAA5FE85" ID-CoL="624L3" authorityName="Leach" authorityYear="1815" box="[1051,1193,332,357]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="family">Mantispidae</taxonomicName>
, first recorded in the MiddleUpper Jurassic of Daohugou,
<collectingCountry id="71C5761DFFB4FF9AFDF6CB19DC92FE68" box="[599,670,367,392]" name="China" pageId="9" pageNumber="572">China</collectingCountry>
. The only older known mantispid is from the Lower Jurassic of
<collectingCountry id="71C5761DFFB4FF9AFF36CBE2DF0CFE4C" box="[151,256,404,428]" name="Germany" pageId="9" pageNumber="572">Germany</collectingCountry>
,
<taxonomicName id="CED24D0EFFB4FF9AFEACCBE2DC13FE4C" ID-CoL="726Y6" authorityName="Ansorge &amp; Schlüter" authorityYear="1990" box="[269,543,404,428]" class="Insecta" family="Mantispidae" genus="Liassochrysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="stigmatica">
<emphasis id="3BA6EA9FFFB4FF9AFEACCBE2DC13FE4C" box="[269,543,404,428]" italics="true" pageId="9" pageNumber="572">Liassochrysa stigmatica</emphasis>
</taxonomicName>
. To date mesomantispines have only been found in Asia (
<collectingCountry id="71C5761DFFB4FF9AFB1FCBE2DB09FE4D" box="[1214,1285,404,429]" name="China" pageId="9" pageNumber="572">China</collectingCountry>
,
<collectingCountry id="71C5761DFFB4FF9AFAB3CBE2DB94FE4D" box="[1298,1432,404,429]" name="Kazakhstan" pageId="9" pageNumber="572">Kazakhstan</collectingCountry>
, and Siberia). Another mantispid is also known in the
<collectingCountry id="71C5761DFFB4FF9AFD4ACBC1DD62FE30" box="[747,878,439,464]" name="Kazakhstan" pageId="9" pageNumber="572">Kazakhstan</collectingCountry>
fauna,
<taxonomicName id="CED24D0EFFB4FF9AFC61CBCEDB41FE30" ID-CoL="4MSD6" authority="Panfilov, 1980" authorityName="Panfilov" authorityYear="1980" box="[960,1357,439,464]" class="Insecta" family="Mantispidae" genus="Promantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="3BA6EA9FFFB4FF9AFC61CBCEDAACFE30" box="[960,1184,440,464]" italics="true" pageId="9" pageNumber="572">Promantispa similis</emphasis>
<bibRefCitation id="6D434B7CFFB4FF9AFB06CBC1DB41FE30" box="[1191,1357,439,464]" pageId="9" pageNumber="572" refId="ref6480" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. &amp; Pritykina, L. N. (Eds.), Mesozoic Fossil Insects. Nauka Dumka, Kiev, pp. 82 - 111. [in Russian]" type="book chapter">Panfilov, 1980</bibRefCitation>
</taxonomicName>
, which is thought to be sister group to
<emphasis id="3BA6EA9FFFB4FF9AFDA0CBAADCAAFE13" box="[513,678,476,500]" italics="true" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFDA0CBAADC99FE14" ID-CoL="6393J" authorityName="Ansorge &amp; Schlüter" authorityYear="1990" box="[513,661,476,500]" class="Insecta" family="Mantispidae" genus="Liassochrysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">Liassochrysa</taxonomicName>
(
</emphasis>
<bibRefCitation id="6D434B7CFFB4FF9AFD06CBAADDFBFE14" box="[679,1015,476,501]" pageId="9" pageNumber="572" refId="ref6555" refString="Wedmann, S. &amp; Makarkin, V. N. (2007) A new genus of Mantispidae (Insecta: Neuroptera) from the Eocene of Germany, with a review of the fossil record and palaeobiogeography of the family. Zoological Journal of the Linnean Society, 149, 701 - 716. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00273. x" type="journal article">Wedmann &amp; Makarkin, 2007</bibRefCitation>
;
<bibRefCitation id="6D434B7CFFB4FF9AFBA4CBAADAB7FE15" box="[1029,1211,476,501]" pageId="9" pageNumber="572" refId="ref6214" refString="Liu, X., Winterton, S. L., Wu, C., Piper, R. &amp; Ohl, M. (2015) A new genus of mantidflies discovered in the Oriental region, with a higher-level phylogeny of Mantispidae (Neuroptera) using DNA sequences and morphology. Systematic Entomology, 40, 183 - 206. http: // dx. doi. org / 10.1111 / syen. 12096" type="journal article">
Liu
<emphasis id="3BA6EA9FFFB4FF9AFB94CBABDA7CFE14" box="[1077,1136,476,500]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
, 2015
</bibRefCitation>
), both being placed within the extant subfamily
<taxonomicName id="CED24D0EFFB4FF9AFE70CB89DC6AFDF8" authorityName="Enderlein" authorityYear="1910" box="[465,614,511,536]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Drepanicinae">Drepanicinae</taxonomicName>
(
<bibRefCitation id="6D434B7CFFB4FF9AFDD4CB89DD2BFDF8" box="[629,807,511,536]" pageId="9" pageNumber="572" refId="ref6214" refString="Liu, X., Winterton, S. L., Wu, C., Piper, R. &amp; Ohl, M. (2015) A new genus of mantidflies discovered in the Oriental region, with a higher-level phylogeny of Mantispidae (Neuroptera) using DNA sequences and morphology. Systematic Entomology, 40, 183 - 206. http: // dx. doi. org / 10.1111 / syen. 12096" type="journal article">
Liu
<emphasis id="3BA6EA9FFFB4FF9AFD05C877DCD0FDF8" box="[676,732,512,536]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
, 2015
</bibRefCitation>
). The first record of mantispids in Europe and then their absence until the Eocene, may represent a lack of fossil localities, or specimens not yet discovered, or only sparse distribution or diversification compared to the diverse fauna observed in Asia.
</paragraph>
<caption id="5DAD6605FFB4FF9AFF36C8E6DA32FD47" box="[151,1086,656,679]" pageId="9" pageNumber="572">
<paragraph id="096D368DFFB4FF9AFF36C8E6DA32FD47" blockId="9.[151,1086,656,679]" box="[151,1086,656,679]" pageId="9" pageNumber="572">
<emphasis id="3BA6EA9FFFB4FF9AFF36C8E6DF06FD47" bold="true" box="[151,266,656,679]" pageId="9" pageNumber="572">TABLE 1.</emphasis>
Fossil record of known
<taxonomicName id="CED24D0EFFB4FF9AFDACC8E6DCC0FD47" authorityName="Makarkin" authorityYear="1996" box="[525,716,656,679]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
, with locality, and age information
</paragraph>
</caption>
<paragraph id="096D368DFFB4FF9AFF3EC8B5DAF9FB07" pageId="9" pageNumber="572">
<table id="7BD2C42DFFB4006CFF3EC8B4DB6AFAE6" box="[159,1382,706,1286]" gridcols="4" gridrows="13" pageId="9" pageNumber="572">
<tr id="B7E234CFFFB4006CFF3EC8B4DB6AFD38" box="[159,1382,706,728]" gridrow="0" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC8B4DC35FD38" box="[159,569,706,728]" gridcol="0" gridrow="0" pageId="9" pageNumber="572">Taxa</th>
<th id="F4335DB3FFB4006CFDF2C8B4DD43FD38" box="[595,847,706,728]" gridcol="1" gridrow="0" pageId="9" pageNumber="572">Deposit</th>
<th id="F4335DB3FFB4006CFCC9C8B4DA2DFD38" box="[872,1057,706,728]" gridcol="2" gridrow="0" pageId="9" pageNumber="572">Age</th>
<th id="F4335DB3FFB4006CFB96C8B4DB6AFD38" box="[1079,1382,706,728]" gridcol="3" gridrow="0" pageId="9" pageNumber="572">Reference</th>
</tr>
<tr id="B7E234CFFFB4006CFF3EC89ADB6AFCE2" box="[159,1382,748,770]" gridrow="1" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC89ADC35FCE2" box="[159,569,748,770]" gridcol="0" gridrow="1" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3EC89ADFA1FCE2" ID-CoL="5VPSG" box="[159,429,748,770]" class="Insecta" family="Mantispidae" genus="Archaeodrepanicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="nuddsi">
<emphasis id="3BA6EA9FFFB4FF9AFF3EC89ADFA1FCE2" box="[159,429,748,770]" italics="true" pageId="9" pageNumber="572">Archaeodrepanicus nuddsi</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2C89ADD43FCE2" box="[595,847,748,770]" gridcol="1" gridrow="1" pageId="9" pageNumber="572">Yixian Formation, China</td>
<td id="F4335DB3FFB4006CFCC9C89ADA2DFCE2" box="[872,1057,748,770]" gridcol="2" gridrow="1" pageId="9" pageNumber="572">Lower Cretaceous</td>
<td id="F4335DB3FFB4006CFB96C89ADB6AFCE2" box="[1079,1382,748,770]" gridcol="3" gridrow="1" pageId="9" pageNumber="572">
Jepson
<emphasis id="3BA6EA9FFFB4FF9AFB23C89BDAB0FCE2" box="[1154,1212,748,770]" italics="true" pageId="9" pageNumber="572">et al.,</emphasis>
2013
</td>
</tr>
<tr id="B7E234CFFFB4006CFF3EC963DB6AFCCB" box="[159,1382,789,811]" gridrow="2" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC963DC35FCCB" box="[159,569,789,811]" gridcol="0" gridrow="2" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3EC960DEF1FCCB" ID-CoL="G7PT" box="[159,253,790,811]" class="Insecta" family="Mantispidae" genus="Archaeodrepanicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="acutus">
<emphasis id="3BA6EA9FFFB4FF9AFF3EC960DEF1FCCB" box="[159,253,790,811]" italics="true" pageId="9" pageNumber="572">A. acutus</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2C963DD43FCCB" box="[595,847,789,811]" gridcol="1" gridrow="2" pageId="9" pageNumber="572">Yixian Formation, China</td>
<td id="F4335DB3FFB4006CFCC9C963DA2DFCCB" box="[872,1057,789,811]" gridcol="2" gridrow="2" pageId="9" pageNumber="572">Lower Cretaceous</td>
<td id="F4335DB3FFB4006CFB96C963DB6AFCCB" box="[1079,1382,789,811]" gridcol="3" gridrow="2" pageId="9" pageNumber="572">
Jepson
<emphasis id="3BA6EA9FFFB4FF9AFB23C960DAB0FCCB" box="[1154,1212,789,811]" italics="true" pageId="9" pageNumber="572">et al.,</emphasis>
2013
</td>
</tr>
<tr id="B7E234CFFFB4006CFF3EC948DB6AFCB5" box="[159,1382,830,853]" gridrow="3" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC948DC35FCB5" box="[159,569,830,853]" gridcol="0" gridrow="3" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3EC948DF68FCB4" ID-CoL="ZWC" box="[159,356,830,852]" class="Insecta" family="Mantispidae" genus="Archaeodrepanicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFF3EC948DF68FCB4" box="[159,356,830,852]" italics="true" pageId="9" pageNumber="572">Archaeodrepanicus</emphasis>
</taxonomicName>
sp.
</th>
<td id="F4335DB3FFB4006CFDF2C948DD43FCB5" box="[595,847,830,853]" gridcol="1" gridrow="3" pageId="9" pageNumber="572">Yixian Formation, China</td>
<td id="F4335DB3FFB4006CFCC9C948DA2DFCB5" box="[872,1057,830,853]" gridcol="2" gridrow="3" pageId="9" pageNumber="572">Lower Cretaceous</td>
<td id="F4335DB3FFB4006CFB96C948DB6AFCB5" box="[1079,1382,830,853]" gridcol="3" gridrow="3" pageId="9" pageNumber="572">
Jepson
<emphasis id="3BA6EA9FFFB4FF9AFB23C936DAB0FCB4" box="[1154,1212,830,853]" italics="true" pageId="9" pageNumber="572">et al.,</emphasis>
2013
</td>
</tr>
<tr id="B7E234CFFFB4006CFF3EC91EDB6AFC9E" box="[159,1382,872,894]" gridrow="4" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC91EDC35FC9E" box="[159,569,872,894]" gridcol="0" gridrow="4" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3EC91FDFE5FC9E" ID-CoL="6YR9F" box="[159,489,872,894]" class="Insecta" family="Mantispidae" genus="Sinomesomantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="microdentata">
<emphasis id="3BA6EA9FFFB4FF9AFF3EC91FDFE5FC9E" box="[159,489,872,894]" italics="true" pageId="9" pageNumber="572">Sinomesomantispa microdentata</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2C91EDD43FC9E" box="[595,847,872,894]" gridcol="1" gridrow="4" pageId="9" pageNumber="572">Yixian Formation, China</td>
<td id="F4335DB3FFB4006CFCC9C91EDA2DFC9E" box="[872,1057,872,894]" gridcol="2" gridrow="4" pageId="9" pageNumber="572">Lower Cretaceous</td>
<td id="F4335DB3FFB4006CFB96C91EDB6AFC9E" box="[1079,1382,872,894]" gridcol="3" gridrow="4" pageId="9" pageNumber="572">
Jepson
<emphasis id="3BA6EA9FFFB4FF9AFB23C91FDAB0FC9E" box="[1154,1212,872,894]" italics="true" pageId="9" pageNumber="572">et al.,</emphasis>
2013
</td>
</tr>
<tr id="B7E234CFFFB4006CFF3EC9E7DB6AFC47" box="[159,1382,913,935]" gridrow="5" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC9E7DC35FC47" box="[159,569,913,935]" gridcol="0" gridrow="5" pageId="9" pageNumber="572">
<emphasis id="3BA6EA9FFFB4FF9AFF3EC9E7DFD5FC47" box="[159,473,913,935]" italics="true" pageId="9" pageNumber="572">
Undescribed
<taxonomicName id="CED24D0EFFB4FF9AFE87C9E4DFD5FC47" authorityName="Makarkin" authorityYear="1996" box="[294,473,914,935]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
</emphasis>
</th>
<td id="F4335DB3FFB4006CFDF2C9E7DD43FC47" box="[595,847,913,935]" gridcol="1" gridrow="5" pageId="9" pageNumber="572">Yixian Formation, China</td>
<td id="F4335DB3FFB4006CFCC9C9E7DA2DFC47" box="[872,1057,913,935]" gridcol="2" gridrow="5" pageId="9" pageNumber="572">Lower Cretaceous</td>
<td id="F4335DB3FFB4006CFB96C9E7DB6AFC47" box="[1079,1382,913,935]" gridcol="3" gridrow="5" pageId="9" pageNumber="572">
Makarkin
<emphasis id="3BA6EA9FFFB4FF9AFB01C9E4DADFFC47" box="[1184,1235,913,935]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
, 2012
</td>
</tr>
<tr id="B7E234CFFFB4006CFF3EC9CCDB6AFC0F" box="[159,1382,954,1007]" gridrow="6" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3EC9CCDC35FC0F" box="[159,569,954,1007]" gridcol="0" gridrow="6" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3EC9CDDF8BFC30" ID-CoL="424HG" box="[159,391,954,976]" class="Insecta" family="Mantispidae" genus="Mesomantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="sibirica">
<emphasis id="3BA6EA9FFFB4FF9AFF3EC9CDDF8BFC30" box="[159,391,954,976]" italics="true" pageId="9" pageNumber="572">Mesomantispa sibirica</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2C9CCDD43FC0F" box="[595,847,954,1007]" gridcol="1" gridrow="6" pageId="9" pageNumber="572">Baissa, Sibiria</td>
<td id="F4335DB3FFB4006CFCC9C9CCDA2DFC0F" box="[872,1057,954,1007]" gridcol="2" gridrow="6" pageId="9" pageNumber="572">Lower Cretaceous</td>
<td id="F4335DB3FFB4006CFB96C9CCDB6AFC0F" box="[1079,1382,954,1007]" gridcol="3" gridrow="6" pageId="9" pageNumber="572">Makarkin, 1997; Wedmann &amp; Makarkin 2007</td>
</tr>
<tr id="B7E234CFFFB4006CFF3ECE74DB6AFBF8" box="[159,1382,1026,1048]" gridrow="7" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3ECE74DC35FBF8" box="[159,569,1026,1048]" gridcol="0" gridrow="7" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3ECE75DFA3FBF8" ID-CoL="3R2Q5" baseAuthorityName="Khramov" baseAuthorityYear="2013" box="[159,431,1027,1048]" class="Insecta" family="Mantispidae" genus="Karataumantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="carnaria">
<emphasis id="3BA6EA9FFFB4FF9AFF3ECE75DFA3FBF8" box="[159,431,1027,1048]" italics="true" pageId="9" pageNumber="572">Karataumantispa carnaria</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2CE74DD43FBF8" box="[595,847,1026,1048]" gridcol="1" gridrow="7" pageId="9" pageNumber="572">Karatau, Kazakhstan</td>
<td id="F4335DB3FFB4006CFCC9CE74DA2DFBF8" box="[872,1057,1026,1048]" gridcol="2" gridrow="7" pageId="9" pageNumber="572">Upper Jurassic</td>
<td id="F4335DB3FFB4006CFB96CE74DB6AFBF8" box="[1079,1382,1026,1048]" gridcol="3" gridrow="7" pageId="9" pageNumber="572">Khramov 2013; Jepson, 2015</td>
</tr>
<tr id="B7E234CFFFB4006CFF3ECE5ADB6AFBA2" box="[159,1382,1068,1090]" gridrow="8" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3ECE5ADC35FBA2" box="[159,569,1068,1090]" gridcol="0" gridrow="8" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3ECE5ADFADFBA2" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[159,417,1068,1090]" class="Palaeacanthocephala" family="Pomphorhynchidae" genus="Longicollum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echinorhynchida" pageId="9" pageNumber="572" phylum="Acanthocephala" rank="species" species="benmaddoxi">
<emphasis id="3BA6EA9FFFB4FF9AFF3ECE5ADFADFBA2" box="[159,417,1068,1090]" italics="true" pageId="9" pageNumber="572">Longicollum benmaddoxi</emphasis>
</taxonomicName>
<emphasis id="3BA6EA9FFFB4FF9AFE06CE5ADC35FBA2" bold="true" box="[423,569,1068,1090]" pageId="9" pageNumber="572">gen et sp. nov</emphasis>
</th>
<td id="F4335DB3FFB4006CFDF2CE5ADD43FBA2" box="[595,847,1068,1090]" gridcol="1" gridrow="8" pageId="9" pageNumber="572">Karatau, Kazakhstan</td>
<td id="F4335DB3FFB4006CFCC9CE5ADA2DFBA2" box="[872,1057,1068,1090]" gridcol="2" gridrow="8" pageId="9" pageNumber="572">Upper Jurassic</td>
<td id="F4335DB3FFB4006CFB96CE5ADB6AFBA2" box="[1079,1382,1068,1090]" gridcol="3" gridrow="8" pageId="9" pageNumber="572">Herein</td>
</tr>
<tr id="B7E234CFFFB4006CFF3ECE23DB6AFB8B" box="[159,1382,1109,1131]" gridrow="9" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3ECE23DC35FB8B" box="[159,569,1109,1131]" gridcol="0" gridrow="9" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3ECE20DF5EFB8B" authorityName="Makarkin" authorityYear="1996" box="[159,338,1110,1131]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">
<emphasis id="3BA6EA9FFFB4FF9AFF3ECE20DF5EFB8B" box="[159,338,1110,1131]" italics="true" pageId="9" pageNumber="572">Mesomantispinae</emphasis>
</taxonomicName>
gen. et sp. indet.
</th>
<td id="F4335DB3FFB4006CFDF2CE23DD43FB8B" box="[595,847,1109,1131]" gridcol="1" gridrow="9" pageId="9" pageNumber="572">Karatau, Kazakhstan</td>
<td id="F4335DB3FFB4006CFCC9CE23DA2DFB8B" box="[872,1057,1109,1131]" gridcol="2" gridrow="9" pageId="9" pageNumber="572">Upper Jurassic</td>
<td id="F4335DB3FFB4006CFB96CE23DB6AFB8B" box="[1079,1382,1109,1131]" gridcol="3" gridrow="9" pageId="9" pageNumber="572">Herein</td>
</tr>
<tr id="B7E234CFFFB4006CFF3ECE08DB6AFB75" box="[159,1382,1150,1173]" gridrow="10" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3ECE08DC35FB75" box="[159,569,1150,1173]" gridcol="0" gridrow="10" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3ECE08DFADFB75" ID-CoL="4B9M7" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" baseAuthorityName="Khramov" baseAuthorityYear="2013" box="[159,417,1150,1173]" class="Insecta" family="Mantispidae" genus="Ovalofemora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="3BA6EA9FFFB4FF9AFF3ECE08DFADFB75" box="[159,417,1150,1173]" italics="true" pageId="9" pageNumber="572">Ovalofemora monstruosa</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2CE08DD43FB75" box="[595,847,1150,1173]" gridcol="1" gridrow="10" pageId="9" pageNumber="572">Karatau, Kazakhstan</td>
<td id="F4335DB3FFB4006CFCC9CE08DA2DFB75" box="[872,1057,1150,1173]" gridcol="2" gridrow="10" pageId="9" pageNumber="572">Upper Jurassic</td>
<td id="F4335DB3FFB4006CFB96CE08DB6AFB75" box="[1079,1382,1150,1173]" gridcol="3" gridrow="10" pageId="9" pageNumber="572">Khramov 2013; Jepson 2015</td>
</tr>
<tr id="B7E234CFFFB4006CFF3ECEDEDB6AFB5E" box="[159,1382,1192,1214]" gridrow="11" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3ECEDEDC35FB5E" box="[159,569,1192,1214]" gridcol="0" gridrow="11" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3ECEDFDF1BFB5E" ID-CoL="4B9M6" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[159,279,1192,1214]" class="Insecta" family="Mantispidae" genus="Ovalofemora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="abbottae">
<emphasis id="3BA6EA9FFFB4FF9AFF3ECEDFDF1BFB5E" box="[159,279,1192,1214]" italics="true" pageId="9" pageNumber="572">O. abbottae</emphasis>
</taxonomicName>
<emphasis id="3BA6EA9FFFB4FF9AFEBDCEDEDFA2FB5E" bold="true" box="[284,430,1192,1214]" pageId="9" pageNumber="572">gen et sp. nov</emphasis>
</th>
<td id="F4335DB3FFB4006CFDF2CEDEDD43FB5E" box="[595,847,1192,1214]" gridcol="1" gridrow="11" pageId="9" pageNumber="572">Karatau, Kazakhstan</td>
<td id="F4335DB3FFB4006CFCC9CEDEDA2DFB5E" box="[872,1057,1192,1214]" gridcol="2" gridrow="11" pageId="9" pageNumber="572">Upper Jurassic</td>
<td id="F4335DB3FFB4006CFB96CEDEDB6AFB5E" box="[1079,1382,1192,1214]" gridcol="3" gridrow="11" pageId="9" pageNumber="572">Herein</td>
</tr>
<tr id="B7E234CFFFB4006CFF3ECEA7DB6AFAE6" box="[159,1382,1233,1286]" gridrow="12" pageId="9" pageNumber="572">
<th id="F4335DB3FFB4006CFF3ECEA7DC35FAE6" box="[159,569,1233,1286]" gridcol="0" gridrow="12" pageId="9" pageNumber="572">
<taxonomicName id="CED24D0EFFB4FF9AFF3ECEA7DF8AFB07" ID-CoL="VVK8" box="[159,390,1233,1255]" class="Insecta" family="Mantispidae" genus="Clavifemora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="rotundata">
<emphasis id="3BA6EA9FFFB4FF9AFF3ECEA7DF8AFB07" box="[159,390,1233,1255]" italics="true" pageId="9" pageNumber="572">Clavifemora rotundata</emphasis>
</taxonomicName>
</th>
<td id="F4335DB3FFB4006CFDF2CEA7DD43FAE6" box="[595,847,1233,1286]" gridcol="1" gridrow="12" pageId="9" pageNumber="572">Jiulongshan Formation, Daohugou, China</td>
<td id="F4335DB3FFB4006CFCC9CEA7DA2DFAE6" box="[872,1057,1233,1286]" gridcol="2" gridrow="12" pageId="9" pageNumber="572">MidUpper Jurassic</td>
<td id="F4335DB3FFB4006CFB96CEA7DB6AFAE6" box="[1079,1382,1233,1286]" gridcol="3" gridrow="12" pageId="9" pageNumber="572">
Jepson
<emphasis id="3BA6EA9FFFB4FF9AFB23CEA4DAB0FB07" box="[1154,1212,1233,1255]" italics="true" pageId="9" pageNumber="572">et al.,</emphasis>
2013
</td>
</tr>
</table>
</paragraph>
<paragraph id="096D368DFFB4FF9AFF66CF3CDA54FA13" blockId="9.[151,1437,1354,1883]" pageId="9" pageNumber="572">
The presence of raptorial forelegs on the mesomantispines suggest a predatory lifestyle, as observed in the extant mantispids. A major difference however is the absence of a large sub-basal spine on the fore femur, which is considered to be a plesiomorphic condition of
<taxonomicName id="CED24D0EFFB4FF9AFD04CFE4DD3FFA4B" ID-CoL="624L3" authorityName="Leach" authorityYear="1815" box="[677,819,1426,1451]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="family">Mantispidae</taxonomicName>
. The adaptive significance of the large sub-basal spine is unclear, but it cannot be excluded that its presence facilitates capturing of prey, so in this respect members of the extant subfamilies of
<taxonomicName id="CED24D0EFFB4FF9AFE28CFACDC1FFA13" ID-CoL="624L3" authorityName="Leach" authorityYear="1815" box="[393,531,1498,1523]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="family">Mantispidae</taxonomicName>
may have had an advantage over mesomantispines.
</paragraph>
</subSubSection>
<subSubSection id="41C86506FFB4FF99FF66CF88DB75FF50" lastPageId="10" lastPageNumber="573" pageId="9" pageNumber="572" type="description">
<paragraph id="096D368DFFB4FF9AFF66CF88DFA6F9BF" blockId="9.[151,1437,1354,1883]" pageId="9" pageNumber="572">
Nothing is known about the larvae of
<taxonomicName id="CED24D0EFFB4FF9AFDD6CF88DD31F9F7" authorityName="Makarkin" authorityYear="1996" box="[631,829,1534,1559]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">mesomantispinae</taxonomicName>
, therefore no interpretations about larval lifestyle can be made, such as whether they had a relationship with spiders, as observed in modern
<taxonomicName id="CED24D0EFFB4FF9AFB39CC54DB2EF9DB" authorityName="Leach" authorityYear="1815" box="[1176,1314,1570,1595]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mantispinae">Mantispinae</taxonomicName>
and some extinct taxa (
<bibRefCitation id="6D434B7CFFB4FF9AFE89CC30DF92F9BF" box="[296,414,1606,1631]" pageId="9" pageNumber="572" refId="ref6449" refString="Ohl, M. (2004) Annotated catalog of the Mantispidae of the world (Neuroptera). Contribution on Entomology, International, 5, 133 - 242." type="journal article">Ohl, 2004</bibRefCitation>
).
</paragraph>
<paragraph id="096D368DFFB4FF9AFF66CC1CDD9EF8BB" blockId="9.[151,1437,1354,1883]" pageId="9" pageNumber="572">
The genera of
<taxonomicName id="CED24D0EFFB4FF9AFECBCC1CDC3CF963" authorityName="Makarkin" authorityYear="1996" box="[362,560,1642,1667]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
are mostly differentiated on their body morphology, for example the structure of their raptorial forelegs, which is quite variable, ranging from clubbed, oval, to slender. Variation is also observed with the length of the pronotum, which ranges from relatively short to elongate. The wing venation on the whole is similar between the genera, however, some venation characters can be used to group genera together. Examples of this are the crossveins within the CuP area, separating
<taxonomicName id="CED24D0EFFB4FF9AFD5ECC8DDDAFF8F2" authorityName="Makarkin" authorityYear="1997" box="[767,931,1787,1810]" class="Insecta" family="Mantispidae" genus="Mesomantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFD5ECC8DDDAFF8F2" box="[767,931,1787,1810]" italics="true" pageId="9" pageNumber="572">Mesomantispa</emphasis>
</taxonomicName>
from the other genera, and also the structure of CuP, whether it is pectinate (
<taxonomicName id="CED24D0EFFB4FF9AFDEBCD68DCDAF8D6" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[586,726,1822,1846]" class="Insecta" family="Mantispidae" genus="Clavifemora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFDEBCD68DCDAF8D6" box="[586,726,1822,1846]" italics="true" pageId="9" pageNumber="572">Clavifemora</emphasis>
</taxonomicName>
,
<taxonomicName id="CED24D0EFFB4FF9AFD53CD69DDC8F8D6" box="[754,964,1823,1846]" class="Insecta" family="Mantispidae" genus="Sinomesomantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFD53CD69DDC8F8D6" box="[754,964,1823,1846]" italics="true" pageId="9" pageNumber="572">Sinomesomantispa</emphasis>
</taxonomicName>
,
<taxonomicName id="CED24D0EFFB4FF9AFC41CD68DAB5F8D6" box="[992,1209,1822,1846]" class="Insecta" family="Mantispidae" genus="Archaeodrepanicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFC41CD68DAB5F8D6" box="[992,1209,1822,1846]" italics="true" pageId="9" pageNumber="572">Archaeodrepanicus</emphasis>
</taxonomicName>
) or non-pectinate (
<taxonomicName id="CED24D0EFFB4FF9AFF01CD35DC0AF8BB" authority="Ovalofemora" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[160,518,1859,1883]" class="Insecta" family="Mantispidae" genus="Karataumantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3BA6EA9FFFB4FF9AFF01CD35DF68F8BA" box="[160,356,1859,1882]" italics="true" pageId="9" pageNumber="572">Karataumantispa</emphasis>
,
<emphasis id="3BA6EA9FFFB4FF9AFED3CD35DC0AF8BB" box="[370,518,1859,1883]" italics="true" pageId="9" pageNumber="572">Ovalofemora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209557E4FFB4FF9AFDACCD34DC7AF8BB" box="[525,630,1858,1883]" pageId="9" pageNumber="572" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFDACCD34DC7AF8BB" bold="true" box="[525,630,1858,1883]" pageId="9" pageNumber="572">gen. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="CED24D0EFFB4FF9AFD25CD35DD1DF8BB" authority="Jepson &amp; Khramov &amp; Ohl, 2018" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[644,785,1859,1883]" class="Palaeacanthocephala" family="Pomphorhynchidae" genus="Longicollum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echinorhynchida" pageId="9" pageNumber="572" phylum="Acanthocephala" rank="genus" status="gen. nov.">
<emphasis id="3BA6EA9FFFB4FF9AFD25CD35DD1DF8BB" box="[644,785,1859,1883]" italics="true" pageId="9" pageNumber="572">Longicollum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209557E4FFB4FF9AFCB9CD34DD8DF8BB" box="[792,897,1858,1883]" pageId="9" pageNumber="572" rank="genus">
<emphasis id="3BA6EA9FFFB4FF9AFCB9CD34DD8DF8BB" bold="true" box="[792,897,1858,1883]" pageId="9" pageNumber="572">gen. nov.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
<paragraph id="096D368DFFB7FF99FF36CAE1DB75FF50" blockId="10.[151,1401,151,176]" box="[151,1401,151,176]" pageId="10" pageNumber="573">
Below is a key of
<taxonomicName id="CED24D0EFFB7FF99FEC3CAE1DC24FF50" authorityName="Makarkin" authorityYear="1996" box="[354,552,151,176]" class="Insecta" family="Mantispidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="subFamily" subFamily="Mesomantispinae">Mesomantispinae</taxonomicName>
genera, updated from
<bibRefCitation id="6D434B7CFFB7FF99FC85CAEEDDCEFF50" box="[804,962,151,176]" pageId="10" pageNumber="573" refId="ref5897" refString="Jepson, J. E. (2015) A review of the current state of knowledge of fossil Mantispidae (Insecta: Neuroptera). Zootaxa, 3964, 419 - 432. https: // doi. org / 10.11646 / zootaxa. 3964.4.2" type="journal article">Jepson (2015)</bibRefCitation>
to include the newly described genera:
</paragraph>
</subSubSection>
<subSubSection id="41C86506FFB7FF99FF36CAA9DB97FCBD" pageId="10" pageNumber="573" type="key">
<key id="7D4361BCFFB7FF99FF36CAA9DB97FCBD" pageId="10" pageNumber="573">
<keyStep id="B2262E28FFB7FF99FF36CAA9DB90FECF" pageId="10" pageNumber="573">
<paragraph id="096D368DFFB7FF99FF36CAA9DB99FEF2" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CAA9DB99FEF2" pageId="10" pageNumber="573">
1. Crossveins between branches of CuP present; area posterior to CuA approximately half the width of the wing......................................................................................................
<taxonomicName id="CED24D0EFFB7FF99FAA5CA8BDB99FEF2" ID-CoL="5PTV" authorityName="Makarkin" authorityYear="1997" box="[1284,1429,253,274]" class="Insecta" family="Mantispidae" genus="Mesomantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FAA5CA8BDB99FEF2" box="[1284,1429,253,274]" italics="true" pageId="10" pageNumber="573">Mesomantispa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="096D368DFFB7FF99FF36CB6CDB90FECF" blockId="10.[151,1436,223,861]" box="[151,1436,281,303]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CB6CDB90FECF" box="[151,1436,281,303]" pageId="10" pageNumber="573">- Crossveins between branches of CuP absent; area posterior to CuA less than half the width of the wing................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="B2262E28FFB7FF99FF36CB41DB99FE67" pageId="10" pageNumber="573">
<paragraph id="096D368DFFB7FF99FF36CB41DB90FEAD" blockId="10.[151,1436,223,861]" box="[151,1436,311,333]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CB41DB90FEAD" box="[151,1436,311,333]" pageId="10" pageNumber="573">2. Forewing CuP pectinate, prostrate setae on foretibia not hooked, foretibia and foretarsus combined longer than forefemur... 3</keyLead>
</paragraph>
<paragraph id="096D368DFFB7FF99FF36CB23DB99FE67" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CB23DB99FE67" pageId="10" pageNumber="573">- Forewing CuP non-pectinate, prostrate setae on foretibia hooked, foretibia and foretarsus combined longer than forefemur...................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="B2262E28FFB7FF99FF36CBF9DB97FE3F" pageId="10" pageNumber="573">
<paragraph id="096D368DFFB7FF99FF36CBF9DB97FE22" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CBF9DB97FE22" pageId="10" pageNumber="573">
3. Forelegs greatly enlarged to give a club-like appearance (
<bibRefCitation id="6D434B7CFFB7FF99FCA4CBF9DDB3FE45" box="[773,959,399,421]" pageId="10" pageNumber="573" refId="ref5832" refString="Jepson, J. E., Heads, S. W., Makarkin, V. N. &amp; Ren, D. (2013) New fossil Mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of North-Eastern China. Palaeontology, 56, 603 - 613. https: // doi. org / 10.1111 / pala. 12005" type="journal article">
Jepson
<emphasis id="3BA6EA9FFFB7FF99FCEECBE6DD75FE45" box="[847,889,399,421]" italics="true" pageId="10" pageNumber="573">et al</emphasis>
., 2013
</bibRefCitation>
: fig. 7); forewing colour pattern consists of spots...........................................................................................
<taxonomicName id="CED24D0EFFB7FF99FA82CBDADB97FE22" ID-CoL="3QP8" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[1315,1435,428,450]" class="Insecta" family="Mantispidae" genus="Clavifemora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FA82CBDADB97FE22" box="[1315,1435,428,450]" italics="true" pageId="10" pageNumber="573">Clavifemora</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="096D368DFFB7FF99FF36CBBCDB97FE3F" blockId="10.[151,1436,223,861]" box="[151,1435,457,479]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CBBCDB97FE3F" box="[151,1435,457,479]" pageId="10" pageNumber="573">- Forelegs not club-like in appearance; no spots on forewings.................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="B2262E28FFB7FF99FF36CB91DB99FD4D" pageId="10" pageNumber="573">
<paragraph id="096D368DFFB7FF99FF36CB91DB9AFDB5" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36CB91DB9AFDB5" pageId="10" pageNumber="573">
4. Wide fore femur (
<quantity id="CE2A9B68FFB7FF99FE2CCB91DFD5FE1C" box="[397,473,487,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="573" unit="mm" value="1.7">1.7 mm</quantity>
) and very small fore femoral spines (&lt;
<quantity id="CE2A9B68FFB7FF99FCEDCB91DDC1FE1C" box="[844,973,487,509]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="10" pageNumber="573" unit="mm" value="0.13" valueMax="0.16" valueMin="0.1">0.10.16 mm</quantity>
long) (
<bibRefCitation id="6D434B7CFFB7FF99FBB3CB91DAC7FE1D" box="[1042,1227,487,509]" pageId="10" pageNumber="573" refId="ref5832" refString="Jepson, J. E., Heads, S. W., Makarkin, V. N. &amp; Ren, D. (2013) New fossil Mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of North-Eastern China. Palaeontology, 56, 603 - 613. https: // doi. org / 10.1111 / pala. 12005" type="journal article">
Jepson
<emphasis id="3BA6EA9FFFB7FF99FBFACB9EDA80FE1D" box="[1115,1164,487,509]" italics="true" pageId="10" pageNumber="573">et al.</emphasis>
, 2013
</bibRefCitation>
: Fig. 6B); costal margin concave before mid-point of wing; costal space of forewing sharply narrowed towards apex; RP with 9 branches, simple for most of their length; forewing colour pattern consists of a distal distinct thick dark band (
<bibRefCitation id="6D434B7CFFB7FF99FBF9C857DB1CFDD7" box="[1112,1296,545,567]" pageId="10" pageNumber="573" refId="ref5832" refString="Jepson, J. E., Heads, S. W., Makarkin, V. N. &amp; Ren, D. (2013) New fossil Mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of North-Eastern China. Palaeontology, 56, 603 - 613. https: // doi. org / 10.1111 / pala. 12005" type="journal article">
Jepson
<emphasis id="3BA6EA9FFFB7FF99FB00C854DADEFDD7" box="[1185,1234,545,567]" italics="true" pageId="10" pageNumber="573">et al.</emphasis>
, 2013
</bibRefCitation>
: fig. 6A).........................................................................................
<taxonomicName id="CED24D0EFFB7FF99FB43C849DB9AFDB5" ID-CoL="7HF3" box="[1250,1430,575,597]" class="Insecta" family="Mantispidae" genus="Sinomesomantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FB43C849DB9AFDB5" box="[1250,1430,575,597]" italics="true" pageId="10" pageNumber="573">Sinomesomantispa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="096D368DFFB7FF99FF36C82BDB99FD4D" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36C82BDB99FD4D" pageId="10" pageNumber="573">
- Fore femur more slender (
<quantity id="CE2A9B68FFB7FF99FE46C82ADC41FD92" box="[487,589,604,626]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="10" pageNumber="573" unit="mm" value="0.95" valueMax="1.0" valueMin="0.9">0.91 mm</quantity>
) and relatively larger femoral spines (
<quantity id="CE2A9B68FFB7FF99FC6BC82ADA41FD92" box="[970,1101,604,626]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="10" pageNumber="573" unit="mm" value="0.175" valueMax="0.25" valueMin="0.1">0.10.25 mm</quantity>
long); costal margin not concave before mid-point of wing; RP with 6 branches, some branches forked before mid-point of wing; forewing colour pattern consists of 3 stripes (
<bibRefCitation id="6D434B7CFFB7FF99FE26C8E1DC33FD4D" box="[391,575,663,685]" pageId="10" pageNumber="573" refId="ref5832" refString="Jepson, J. E., Heads, S. W., Makarkin, V. N. &amp; Ren, D. (2013) New fossil Mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of North-Eastern China. Palaeontology, 56, 603 - 613. https: // doi. org / 10.1111 / pala. 12005" type="journal article">
Jepson
<emphasis id="3BA6EA9FFFB7FF99FE71C8EEDC0DFD4D" box="[464,513,663,685]" italics="true" pageId="10" pageNumber="573">et al.</emphasis>
, 2013
</bibRefCitation>
: fig 1A).................................................
<taxonomicName id="CED24D0EFFB7FF99FB7AC8E1DB99FD4D" ID-CoL="ZWC" box="[1243,1429,663,685]" class="Insecta" family="Mantispidae" genus="Archaeodrepanicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FB7AC8E1DB99FD4D" box="[1243,1429,663,685]" italics="true" pageId="10" pageNumber="573">Archaeodrepanicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B2262E28FFB7FF99FF36C8C2DB97FCE5" pageId="10" pageNumber="573">
<paragraph id="096D368DFFB7FF99FF36C8C2DB9AFD08" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36C8C2DB9AFD08" pageId="10" pageNumber="573">
5. Length to width ratio of forefemur (at widest part) is less than 3, forefemur oval shaped, forecoxa, foretrochanter stout, all covered in setae........................................................................
<taxonomicName id="CED24D0EFFB7FF99FB16C8A7DB3AFD07" ID-CoL="6B3K" authority="Jepson &amp; Khramov &amp; Ohl, 2018" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[1207,1334,721,743]" class="Insecta" family="Mantispidae" genus="Ovalofemora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3BA6EA9FFFB7FF99FB16C8A7DB3AFD07" box="[1207,1334,721,743]" italics="true" pageId="10" pageNumber="573">Ovalofemora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209557E4FFB7FF99FA9DC8A4DB9AFD08" box="[1340,1430,722,744]" pageId="10" pageNumber="573" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FA9DC8A4DB9AFD08" bold="true" box="[1340,1430,722,744]" pageId="10" pageNumber="573">gen. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="096D368DFFB7FF99FF36C899DB97FCE5" blockId="10.[151,1436,223,861]" box="[151,1435,751,773]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36C899DB97FCE5" box="[151,1435,751,773]" pageId="10" pageNumber="573">- Length to width ratio of forefemur (at widest part) is 3 or more, some setae........................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="B2262E28FFB7FF99FF36C97ADB97FCBD" pageId="10" pageNumber="573">
<paragraph id="096D368DFFB7FF99FF36C97ADB90FCC2" blockId="10.[151,1436,223,861]" box="[151,1436,780,802]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36C97ADB90FCC2" box="[151,1436,780,802]" pageId="10" pageNumber="573">
6. Forefemur and forecoxa slender, pronotum slightly elongate 1.1x longer than wide, MP deeply forked.....
<taxonomicName id="CED24D0EFFB7FF99FB55C97BDB90FCC2" ID-CoL="57HF" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[1268,1436,781,802]" class="Insecta" family="Mantispidae" genus="Karataumantispa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FB55C97BDB90FCC2" box="[1268,1436,781,802]" italics="true" pageId="10" pageNumber="573">Karataumantispa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="096D368DFFB7FF99FF36C95CDB97FCBD" blockId="10.[151,1436,223,861]" pageId="10" pageNumber="573">
<keyLead id="B22395B8FFB7FF99FF36C95CDB97FCBD" pageId="10" pageNumber="573">
- Forefemur widened at proximal end, with spines at distal apex, pronotum very elongate 2.8x longer than wide, MP simple most of length forking distally..........................................................
<taxonomicName id="CED24D0EFFB7FF99FB60C931DB36FCBD" ID-CoL="5GD3" authority="Jepson &amp; Khramov &amp; Ohl, 2018" authorityName="Jepson &amp; Khramov &amp; Ohl" authorityYear="2018" box="[1217,1338,839,861]" class="Palaeacanthocephala" family="Pomphorhynchidae" genus="Longicollum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echinorhynchida" pageId="10" pageNumber="573" phylum="Acanthocephala" rank="genus" status="gen. nov.">
<emphasis id="3BA6EA9FFFB7FF99FB60C931DB36FCBD" box="[1217,1338,839,861]" italics="true" pageId="10" pageNumber="573">Longicollum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209557E4FFB7FF99FAE3C931DB97FCBD" box="[1346,1435,839,861]" pageId="10" pageNumber="573" rank="genus">
<emphasis id="3BA6EA9FFFB7FF99FAE3C931DB97FCBD" bold="true" box="[1346,1435,839,861]" pageId="10" pageNumber="573">gen. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>