277 lines
36 KiB
XML
277 lines
36 KiB
XML
<document id="4BFDD2FAA22B6742F5E6E688BB2C1578" ID-DOI="10.15298/rusentj.31.3.11" ID-ISSN="0132-8069" ID-Zenodo-Dep="13177632" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722592676046" checkinUser="felipe" docAuthor="Grichanov, I. Ya." docDate="2022" docId="815087FC7B28FFABFCBFF8A0E76C9C7D" docLanguage="en" docName="RussEntomolJ.31.3.293-296.pdf" docOrigin="Russian Entomological Journal 31 (3)" docSource="http://dx.doi.org/10.15298/rusentj.31.3.11" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Griphophanes cameroonensis Grichanov 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="295" masterDocId="7D69FF847B28FFA9FFEDFFAAE3299901" masterDocTitle="Discovery of the genus Griphophanes Grootaert et Meuffels in Cameroon (Diptera: Dolichopodidae)" masterLastPageNumber="296" masterPageNumber="293" pageNumber="293" updateTime="1722631785395" updateUser="ExternalLinkService">
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<mods:titleInfo id="E65DD84C74856D2496D45773B2E807F1">
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<mods:title id="562C237F2B75FA0745A14AC398F65529">Discovery of the genus Griphophanes Grootaert et Meuffels in Cameroon (Diptera: Dolichopodidae)</mods:title>
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<mods:namePart id="0F283B14204DA31E3C17158DD606D127">Grichanov, I. Ya.</mods:namePart>
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<mods:typeOfResource id="0684E4F8995F5FF548853B4906C52E45">text</mods:typeOfResource>
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<mods:title id="772B4838C6023B89B7DB7859E3D95E68">Russian Entomological Journal</mods:title>
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<mods:part id="4096B219991CE55213B25133858C2EE5">
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<mods:date id="935AA0D69C15D07F77941FE1BF9EBF2E">2022</mods:date>
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<mods:number id="2A7534FF12BB811FDE5ACBBF47D626AF">2022-09-30</mods:number>
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<mods:number id="23CF7B0E31FC58D85AFB2484D06AFF09">31</mods:number>
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<mods:start id="DBA10E64517C97C0ADDAE3729E7EFE7A">293</mods:start>
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<mods:classification id="CE34E4FCD7AB3513798499A4F5E70C37">journal article</mods:classification>
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<mods:identifier id="A0C561BB3E752D03266AAB07DD43F27D" type="DOI">10.15298/rusentj.31.3.11</mods:identifier>
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<mods:identifier id="047F1085B5C64828178A5CBE30C08D27" type="ISSN">0132-8069</mods:identifier>
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<treatment id="815087FC7B28FFABFCBFF8A0E76C9C7D" LSID="urn:lsid:plazi:treatment:815087FC7B28FFABFCBFF8A0E76C9C7D" httpUri="http://treatment.plazi.org/id/815087FC7B28FFABFCBFF8A0E76C9C7D" lastPageId="2" lastPageNumber="295" pageId="0" pageNumber="293">
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<subSubSection id="41E365617B28FFA9FCBFF8A0E6799E23" box="[850,1360,1802,1826]" pageId="0" pageNumber="293" type="nomenclature">
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<paragraph id="094636EA7B28FFA9FCBFF8A0E6799E23" blockId="0.[812,1399,1802,1960]" box="[850,1360,1802,1826]" pageId="0" pageNumber="293">
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<heading id="520E81867B28FFA9FCBFF8A0E6799E23" box="[850,1360,1802,1826]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="293" reason="8">
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<taxonomicName id="CEF94D697B28FFA9FCBFF8A0E6279E23" ID-CoL="9J774" authority="Grichanov" authorityName="Grichanov" authorityYear="2022" box="[850,1294,1802,1826]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="293" phylum="Arthropoda" rank="species" species="cameroonensis" status="sp. nov.">
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<emphasis id="3B8DEAF87B28FFA9FCBFF8A0E7BA9E23" box="[850,1171,1802,1826]" italics="true" pageId="0" pageNumber="293">Griphophanes cameroonensis</emphasis>
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Grichanov
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</taxonomicName>
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||
,
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<emphasis id="3B8DEAF87B28FFA9FAF6F8A0E6799E23" bold="true" box="[1307,1360,1802,1826]" pageId="0" pageNumber="293">
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<taxonomicNameLabel id="20BE57837B28FFA9FAF6F8A0E6799E23" box="[1307,1360,1802,1826]" pageId="0" pageNumber="293" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="41E365617B28FFA9FBF2F882E7AA9E41" box="[1055,1155,1832,1856]" pageId="0" pageNumber="293" type="description">
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<paragraph id="094636EA7B28FFA9FBF2F882E7AA9E41" blockId="0.[812,1399,1802,1960]" box="[1055,1155,1832,1856]" pageId="0" pageNumber="293">
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<figureCitation id="91C22A6F7B28FFA9FBF2F882E7569E41" box="[1055,1151,1832,1856]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="0" pageNumber="293">Figs 1–9</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="41E365617B28FFABFCBFF8E7E021987F" lastPageId="2" lastPageNumber="295" pageId="0" pageNumber="293" type="materials_examined">
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<paragraph id="094636EA7B28FFA9FCBFF8E7E65F9EA9" blockId="0.[812,1399,1802,1960]" pageId="0" pageNumber="293">
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MATERIAL.
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<typeStatus id="D64288487B28FFA9FC26F8E7E7329E61" box="[971,1051,1869,1888]" pageId="0" pageNumber="293" type="holotype">Holotype</typeStatus>
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<specimenCount id="1FFFFD637B28FFA9FBF2F8E0E71A9E61" box="[1055,1075,1866,1888]" count="1" pageId="0" pageNumber="293" type="male">♂</specimenCount>
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,
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<materialsCitation id="B9913CB77B28FFA9FBD0F8E7E76F9EA9" collectingDate="2013-08-17" collectingDateMax="2013-08-19" collectingDateMin="2013-08-17" collectionCode="BMSA" collectorName="A. H. Kirk-Spriggs" country="Cameroon" elevation="1060" latitude="6.0837665" location="Mezam" longLatPrecision="1" longitude="10.124033" municipality="Bafut village" pageId="0" pageNumber="293" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="71EE767A7B28FFA9FBD0F8E7E7B09E61" box="[1085,1177,1869,1888]" name="Cameroon" pageId="0" pageNumber="293">Cameroon</collectingCountry>
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, Northwest Reg.,
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<location id="0C2660317B28FFA9FADDF8E7E65A9E61" LSID="urn:lsid:plazi:treatment:815087FC7B28FFABFCBFF8A0E76C9C7D:0C2660317B28FFA9FADDF8E7E65A9E61" box="[1328,1395,1869,1888]" country="Cameroon" latitude="6.0837665" longLatPrecision="1" longitude="10.124033" municipality="Bafut village" name="Mezam" pageId="0" pageNumber="293">Mezam</location>
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,
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<collectingMunicipality id="E922AC907B28FFA9FCC1F8CFE0899E79" box="[812,928,1893,1912]" pageId="0" pageNumber="293">Bafut village</collectingMunicipality>
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at
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<geoCoordinate id="6CCD502D7B28FFA9FC2CF8CFE7019E79" box="[961,1064,1893,1912]" degrees="6" direction="north" minutes="05.026" orientation="latitude" pageId="0" pageNumber="293" precision="1" value="6.0837665">6°05.026´N</geoCoordinate>
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,
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<geoCoordinate id="6CCD502D7B28FFA9FBD9F8CFE78D9E79" box="[1076,1188,1893,1912]" degrees="10" direction="east" minutes="07.442" orientation="longitude" pageId="0" pageNumber="293" precision="1" value="10.124033">10°07.442´E</geoCoordinate>
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,
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<quantity id="CE019B0F7B28FFA9FB5DF8CFE7DE9E79" box="[1200,1271,1893,1912]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="0" pageNumber="293" unit="m" value="1060.0">
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<elevation id="82D4D1D97B28FFA9FB5DF8CFE7DE9E79" box="[1200,1271,1893,1912]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="0" pageNumber="293" unit="m" value="1060.0">1060 m</elevation>
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</quantity>
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,
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<collectingMethod id="D0B84EFD7B28FFA9FAEFF8CFE65A9E79" box="[1282,1395,1893,1912]" pageId="0" pageNumber="293">Malaise trap</collectingMethod>
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, cultivated plot and degraded forest,
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<date id="7D47102A7B28FFA9FB98F8D7E6219E91" box="[1141,1288,1917,1936]" pageId="0" pageNumber="293" value="2013-08-17" valueMax="2013-08-19" valueMin="2013-08-17">
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<collectingDate id="6D03E9C27B28FFA9FB98F8D7E6219E91" box="[1141,1288,1917,1936]" pageId="0" pageNumber="293" value="2013-08-17" valueMax="2013-08-19" valueMin="2013-08-17">17–19.VIII.2013</collectingDate>
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</date>
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,
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<collectorName id="A40C533C7B28FFA9FAF8F8D7E0479EA9" pageId="0" pageNumber="293">A.H. Kirk-Spriggs</collectorName>
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leg. (
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<collectionCode id="6FE8AE2F7B28FFA9FC49F83FE0CF9EA9" box="[932,998,1941,1960]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="0" pageNumber="293">BMSA</collectionCode>
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).
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<typeStatus id="D64288487B28FFA9FC1BF83FE76F9EA9" box="[1014,1094,1941,1960]" pageId="0" pageNumber="293" type="holotype">Holotype</typeStatus>
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</materialsCitation>
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in good condition.
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||
<typeStatus id="D64288487B28FFA9FB14F83FE6619EA9" box="[1273,1352,1941,1960]" pageId="0" pageNumber="293" type="paratype">Paratype</typeStatus>
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:
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||
<specimenCount id="1FFFFD637B28FFA9FABFF83FE6589EA9" box="[1362,1393,1938,1960]" count="1" pageId="0" pageNumber="293" type="male">1♂</specimenCount>
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,
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||
</paragraph>
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||
<paragraph id="094636EA7B29FFABFF50FF58E1789867" blockId="1.[189,880,240,266]" lastBlockId="2.[189,776,315,2063]" lastPageId="2" lastPageNumber="295" pageId="1" pageNumber="294">
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||
<materialsCitation id="B9913CB77B29FFABFF50FF58E1649867" collectingDate="2013-08-17" collectingDateMax="2013-08-19" collectingDateMin="2013-08-17" collectionCode="BMSA" collectorName="I. Ya. Grichanov" country="Cameroon" elevation="1060" lastPageId="2" lastPageNumber="295" latitude="6.0837665" location="Mezam" longLatPrecision="1" longitude="10.124033" municipality="Bafut village" pageId="1" pageNumber="294" specimenCount="1">
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294
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<collectorName id="A40C533C7B29FFA8FD2EFF5AE0599809" box="[707,880,240,264]" pageId="1" pageNumber="294">I.Ya. Grichanov</collectorName>
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same data as holotype; terminalia dissected and stored in glycerin in microvial pinned with the specimen (
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<collectionCode id="6FE8AE2F7B2AFFABFDEBFEF9E1619867" box="[518,584,339,358]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="295">BMSA</collectionCode>
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||
)
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</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="094636EA7B29FFA8FCCEFE00E01B9E8C" pageId="1" pageNumber="294">4 5 6 1 2 3 7 8 9</paragraph>
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<caption id="5D8666627B29FFA8FF0FF81BE65F9111" pageId="1" pageNumber="294" startId="1.[226,263,1969,1988]" targetType="figure">
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<paragraph id="094636EA7B29FFA8FF0FF81BE6769EDD" blockId="1.[189,1398,1969,2065]" pageId="1" pageNumber="294">
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Figs 1–9.
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<taxonomicName id="CEF94D697B29FFA8FED7F81BE1159EC5" authorityName="Grichanov" authorityYear="2022" box="[314,572,1969,1988]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="cameroonensis" status="sp. nov.">
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<emphasis id="3B8DEAF87B29FFA8FED7F81BE1159EC5" box="[314,572,1969,1988]" italics="true" pageId="1" pageNumber="294">Griphophanes cameroonensis</emphasis>
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</taxonomicName>
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,
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<emphasis id="3B8DEAF87B29FFA8FDA5F81BE15A9EC5" bold="true" box="[584,627,1969,1988]" pageId="1" pageNumber="294">
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||
<taxonomicNameLabel id="20BE57837B29FFA8FDA5F81BE15A9EC5" box="[584,627,1969,1988]" pageId="1" pageNumber="294" rank="species">sp.n.</taxonomicNameLabel>
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||
</emphasis>
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||
, male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view).
|
||
</paragraph>
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<paragraph id="094636EA7B29FFA8FF0FF84CE65F9111" blockId="1.[189,1398,1969,2065]" pageId="1" pageNumber="294">
|
||
Рис. 1–9.
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||
<taxonomicName id="CEF94D697B29FFA8FED2F84FE16B9EF9" authorityName="Grichanov" authorityYear="2022" box="[319,578,2021,2040]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="cameroonensis" status="sp. nov.">
|
||
<emphasis id="3B8DEAF87B29FFA8FED2F84FE16B9EF9" box="[319,578,2021,2040]" italics="true" pageId="1" pageNumber="294">Griphophanes cameroonensis</emphasis>
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||
</taxonomicName>
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||
,
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||
<emphasis id="3B8DEAF87B29FFA8FDBDF84FE1529EF9" bold="true" box="[592,635,2021,2040]" pageId="1" pageNumber="294">
|
||
<taxonomicNameLabel id="20BE57837B29FFA8FDBDF84FE1529EF9" box="[592,635,2021,2040]" pageId="1" pageNumber="294" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу).
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||
</paragraph>
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||
</caption>
|
||
<paragraph id="094636EA7B2AFFABFF0FFEC1E021987F" blockId="2.[189,776,315,2063]" box="[226,776,361,383]" pageId="2" pageNumber="295">
|
||
<materialsCitation id="B9913CB77B2AFFABFF0FFEC1E02D987F" box="[226,772,361,383]" collectingDate="2013-08-17" collectingDateMax="2013-08-19" collectingDateMin="2013-08-17" collectionCode="BMSA" collectorName="A. H. Kirk-Spriggs" country="Cameroon" elevation="1060" latitude="6.0837665" location="Mezam" longLatPrecision="1" longitude="10.124033" municipality="Bafut village" pageId="2" pageNumber="295" specimenCount="1" specimenCount-female="1">
|
||
<typeStatus id="D64288487B2AFFABFF0FFEC1E2FE987F" box="[226,471,363,382]" isOtherTypeStatus="true" pageId="2" pageNumber="295">ADDITIONAL MATERIAL</typeStatus>
|
||
.
|
||
<specimenCount id="1FFFFD637B2AFFABFE0DFEC1E2DE987E" box="[480,503,361,383]" count="1" pageId="2" pageNumber="295" type="female">1♀</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="6FE8AE2F7B2AFFABFD50FEC1E1D7987F" box="[701,766,363,382]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="295">BMSA</collectionCode>
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||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41E365617B2AFFABFF0FFE29E7679D07" pageId="2" pageNumber="295" type="description">
|
||
<paragraph id="094636EA7B2AFFABFF0FFE29E24F9A5E" blockId="2.[189,776,315,2063]" pageId="2" pageNumber="295">
|
||
DESCRIPTION.
|
||
<emphasis id="3B8DEAF87B2AFFABFE66FE29E2969899" bold="true" box="[395,447,387,408]" pageId="2" pageNumber="295">Male</emphasis>
|
||
(
|
||
<figureCitation id="91C22A6F7B2AFFABFE21FE29E12D9899" box="[460,516,387,409]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Fig. 1</figureCitation>
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||
). Head (
|
||
<figureCitation id="91C22A6F7B2AFFABFDB7FE29E1BC9899" box="[602,661,387,409]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Fig. 2</figureCitation>
|
||
). Frons and face with black ground colour; frons brownish pollinose; face whitish pollinose; one vertical seta at upper corner of frons; one hairlike postvertical seta positioned behind vertical seta; ocellar setae long; two short fine postocellars; postocular setae all dark, slightly increasing in length downward; eyes with short ommatrichia; face under antennae 1.5 times as wide as height of postpedicel, slightly narrowing towards clypeus; antenna black, slightly longer than head height; scape small, vase-like; pedicel larger, globular, with ring of short setulae and 1–2 elongate apicodorsal setae; postpedicel triangular, as long as high at base, densely haired; stylus middorsal, nearly 4× as long as main segments of antenna combined, pubescent, with short thick segment 1 and filiform segment 2; ratio of lengths of scape to pedicel to postpedicel to stylomere 1 to stylomere 2, 4/4/8/4/54; palpus and proboscis small, blackish-brown, palpus oval, with black seta and dark pubescence.
|
||
</paragraph>
|
||
<paragraph id="094636EA7B2AFFABFF0FFCC9E02F9AFF" blockId="2.[189,776,315,2063]" pageId="2" pageNumber="295">
|
||
Thorax. Bluish-black, grey pollinose, with dark bristles; mesonotum flattened posteriorly; 6 (2+4) dorsocentrals, with 6
|
||
<superScript id="FE8C9BA27B2AFFABFF24FC32E3FD9AA5" attach="left" box="[201,212,920,932]" fontSize="5" pageId="2" pageNumber="295">th</superScript>
|
||
seta shifted towards lateral margin and 1
|
||
<superScript id="FE8C9BA27B2AFFABFDBAFC32E1499AA5" attach="left" box="[599,608,920,932]" fontSize="5" pageId="2" pageNumber="295">st</superScript>
|
||
seta rather small; short acrostichals in two regular rows; scutellum with one pair of strong setae and one pair of lateral setae; upper propleuron without setae; proepisternum with 1 strong seta.
|
||
</paragraph>
|
||
<paragraph id="094636EA7B2AFFABFF0FFBAEE02E9F7F" blockId="2.[189,776,315,2063]" pageId="2" pageNumber="295">
|
||
Legs. Mostly black; knees, tibiae and basitarsi blackish brown; all tarsi simple, with 5
|
||
<superScript id="FE8C9BA27B2AFFABFE15FBB7E12A9D28" attach="left" box="[504,515,1053,1065]" fontSize="5" pageId="2" pageNumber="295">th</superScript>
|
||
segment flattened dorsoventrally, with well developed pulvilli and claws; fore and mid coxae with dark anterior and apical setae; hind coxa with 1 dark bristle at middle; fore leg (
|
||
<figureCitation id="91C22A6F7B2AFFABFE29FBC4E2D69D82" box="[452,511,1134,1156]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Fig. 4</figureCitation>
|
||
) without strong setae; fore femur ventrally, fore tibia dorsally and ventrally, fore tarsus dorsally and laterally with rows of elongate setulae, nearly as long as diameter of corresponding podomeres; length ratio of fore tibia to tarsus (segments from first to fifth), 66/32/16/11/ 8/12. Mid femur simple, bearing 1 strong but short anterior subapical bristle and 2 rows of ventral hairs, half as long as height of femur; mid tibia simple, with 1 anterodorsal and 1 posterodorsal setae at basal third, 1 posterodorsal seta at distal third and 5 apical setae; no ventral setae; mid tarsus simple; segments 1–4 each with short apical spinules; segments 4–5 with elongate setulae; length ratio of mid tibia to tarsus (segments from first to fifth), 87/40/22/13/8/8; hind femur with 1 anterior subapical bristle, with elongate dorsal setae on basal half; hind tibia (
|
||
<figureCitation id="91C22A6F7B2AFFABFE8FFA49E2B69CF8" box="[354,415,1507,1529]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Fig. 5</figureCitation>
|
||
) simple, with 1 strong anterodorsal bristle at basal third, dorsal row of 6–8 short setae and with strong and long, curved subapical dorsal bristle, 3/4 as long as hind basitarsus; hind basitarsus with 1 short apicoventral seta, with very small basal tooth posteriorly; length ratio of hind tibia to tarsus (segments from first to fifth), 91/27/28/17/10/11.
|
||
</paragraph>
|
||
<paragraph id="094636EA7B2AFFABFF0FF929E02C9E55" blockId="2.[189,776,315,2063]" pageId="2" pageNumber="295">
|
||
Wing (
|
||
<figureCitation id="91C22A6F7B2AFFABFEC4F929E24E9F98" box="[297,359,1667,1689]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Fig. 6</figureCitation>
|
||
). Hyaline, with greyish tinge, veins dark brown; costa simple, with simple costal setulae; ratio of part of costa between R
|
||
<subScript id="957D34AF7B2AFFABFE98F962E2A29FD5" attach="left" box="[373,395,1736,1748]" fontSize="5" pageId="2" pageNumber="295">2+3</subScript>
|
||
and R
|
||
<subScript id="957D34AF7B2AFFABFE27F962E2C99FD5" attach="left" box="[458,480,1736,1748]" fontSize="5" pageId="2" pageNumber="295">4+5</subScript>
|
||
to that between R
|
||
<subScript id="957D34AF7B2AFFABFD7DF962E18F9FD5" attach="left" box="[656,678,1736,1748]" fontSize="5" pageId="2" pageNumber="295">4+5</subScript>
|
||
and M
|
||
<subScript id="957D34AF7B2AFFABFD06F962E0299FD5" attach="right" box="[747,768,1736,1748]" fontSize="5" pageId="2" pageNumber="295">1+2</subScript>
|
||
, 5/4; R
|
||
<subScript id="957D34AF7B2AFFABFF14F949E2279FEE" attach="left" box="[249,270,1763,1775]" fontSize="5" pageId="2" pageNumber="295">2+3</subScript>
|
||
and R
|
||
<subScript id="957D34AF7B2AFFABFEA3F948E24A9FEF" attach="left" box="[334,355,1762,1774]" fontSize="5" pageId="2" pageNumber="295">4+5</subScript>
|
||
slightly diverging towards wing apex; R
|
||
<subScript id="957D34AF7B2AFFABFD1CF948E02F9FEF" attach="left" box="[753,774,1762,1774]" fontSize="5" pageId="2" pageNumber="295">4+5</subScript>
|
||
and M
|
||
<subScript id="957D34AF7B2AFFABFF13F957E23D9E08" attach="left" box="[254,276,1789,1801]" fontSize="5" pageId="2" pageNumber="295">1+2</subScript>
|
||
slightly convex anteriorly, parallel in distal half; ratio of
|
||
<emphasis id="3B8DEAF87B2AFFABFEE7F8A3E26C9E1F" box="[266,325,1801,1822]" italics="true" pageId="2" pageNumber="295">dm–m</emphasis>
|
||
to apical part of M
|
||
<subScript id="957D34AF7B2AFFABFDEFF8B2E1209E25" attach="right" box="[514,521,1816,1828]" fontSize="5" pageId="2" pageNumber="295">4</subScript>
|
||
, 21/35; anal vein distinct, not reaching wing margin; anal lobe well developed; lower calypter small, brown, with black setae; halter knob yellow.
|
||
</paragraph>
|
||
<paragraph id="094636EA7B2AFFABFF0FF8F3E65E9A16" blockId="2.[189,776,315,2063]" lastBlockId="2.[812,1399,315,1404]" pageId="2" pageNumber="295">
|
||
Abdomen. Black, cylindrical, with black hairs and marginal setae; tergum 6 setose; sterna 5 and 6 weakly sclerotized. Segment 7 weakly setose, as long as tergum 6, with tergum and sternum distinct; segment 8 black, large, setose, with ventral inverted “Y” shaped sclerotized corner; hypopygium (
|
||
<figureCitation id="91C22A6F7B2AFFABFF16F874E2799EF2" box="[251,336,2014,2036]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Figs 8–9</figureCitation>
|
||
) black, with brown cercus; epandrium ovoid, asymmetrical, longer than high, convex ventrally in middle, somewhat widening distally; foramen lateral, positioned in basal half of epandrium; hypandrium fused with epandrium at base with strongly sclerotized ring, long, cylindric, with big and broad hook distally; phallus thin and long, concealed within hypandrium; epandrial lobes asymmetrical; left lobe strongly projected distad, flat, leaflike, bearing 2 strong apicoventral setae of unequal length; right epandrial lobe divided into two strong unequal hooks directed inside, with 1 thick and long and 1 short setae at apex of distal hook; additional thin epandrial process with 1 apical seta adjacent both left and right lobes; surstyli symmetrical, adjacent to cerci, with dorsal arm twice shorter than ventral one; dorsal arm finger-like, with 1 dorsal seta and few minute hairs at apex; ventral arm of surstylus broad to apex, with 1 thick and long inner bristle at middle of ventral margin, 1 middorsal seta, few minute hairs at apex and short but thick apical seta; cercus small, elongate-triangular, much shorter than surstylus, regularly covered with long setae and microscopic hairs.
|
||
</paragraph>
|
||
<paragraph id="094636EA7B2AFFABFCBFFCB6E6759A4D" blockId="2.[812,1399,315,1404]" pageId="2" pageNumber="295">Measurements (mm). Body length 2, antenna length 0.7, wing length 2.3, wing width 0.8, hypopygium length 0.5.</paragraph>
|
||
<paragraph id="094636EA7B2AFFABFCBFFCFBE7679D07" blockId="2.[812,1399,315,1404]" pageId="2" pageNumber="295">
|
||
<emphasis id="3B8DEAF87B2AFFABFCBFFCFBE0B49A67" bold="true" box="[850,925,849,870]" pageId="2" pageNumber="295">Female</emphasis>
|
||
(
|
||
<figureCitation id="91C22A6F7B2AFFABFC46FCFBE72A9A67" box="[939,1027,849,871]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Figs 3, 7</figureCitation>
|
||
). Similar to male except lacking male secondary sexual characters. Body length
|
||
<quantity id="CE019B0F7B2AFFABFB3DFCC1E6369A80" box="[1232,1311,875,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="295" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, antenna length
|
||
<quantity id="CE019B0F7B2AFFABFC9CFC2CE0969A9A" box="[881,959,902,924]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="295" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, wing length
|
||
<quantity id="CE019B0F7B2AFFABFBAAFC2CE7BE9A9A" box="[1095,1175,902,923]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="295" unit="mm" value="1.9">1.9 mm</quantity>
|
||
. Face wider than male face, brownish pollinose; postpedicel higher than long (11/ 7), tibiae and basitarsi yellow; ratio of part of costa between R
|
||
<subScript id="957D34AF7B2AFFABFCD1FC4FE07B9AF0" attach="left" box="[828,850,997,1009]" fontSize="5" pageId="2" pageNumber="295">2+3</subScript>
|
||
and R
|
||
<subScript id="957D34AF7B2AFFABFC7CFC4FE08E9AF0" attach="left" box="[913,935,997,1009]" fontSize="5" pageId="2" pageNumber="295">4+5</subScript>
|
||
to that between R
|
||
<subScript id="957D34AF7B2AFFABFBB5FC4FE7449AF0" attach="left" box="[1112,1133,997,1009]" fontSize="5" pageId="2" pageNumber="295">4+5</subScript>
|
||
and M
|
||
<subScript id="957D34AF7B2AFFABFB5FFC4FE7E19AF0" attach="right" box="[1202,1224,997,1009]" fontSize="5" pageId="2" pageNumber="295">1+2</subScript>
|
||
, 2/1; ratio of
|
||
<emphasis id="3B8DEAF87B2AFFABFAA0FC7CE0149D07" italics="true" pageId="2" pageNumber="295">dm– m</emphasis>
|
||
to apical part of M
|
||
<subScript id="957D34AF7B2AFFABFC11FBAAE72A9D0D" attach="right" box="[1020,1027,1024,1036]" fontSize="5" pageId="2" pageNumber="295">4</subScript>
|
||
, 19/42.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41E365617B2AFFABFCBFFBA1E0479D3D" pageId="2" pageNumber="295" type="etymology">
|
||
<paragraph id="094636EA7B2AFFABFCBFFBA1E0479D3D" blockId="2.[812,1399,315,1404]" pageId="2" pageNumber="295">ETYMOLOGY: The species is named for the country of origin.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41E365617B2AFFABFCBFFBEBE75C9D57" box="[850,1141,1089,1110]" pageId="2" pageNumber="295" type="distribution">
|
||
<paragraph id="094636EA7B2AFFABFCBFFBEBE75C9D57" blockId="2.[812,1399,315,1404]" box="[850,1141,1089,1110]" pageId="2" pageNumber="295">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="71EE767A7B2AFFABFBE7FBEBE75B9D57" box="[1034,1138,1089,1110]" name="Cameroon" pageId="2" pageNumber="295">Cameroon</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41E365617B2AFFABFCBFFBF1E76C9C7D" pageId="2" pageNumber="295" type="diagnosis">
|
||
<paragraph id="094636EA7B2AFFABFCBFFBF1E76C9C7D" blockId="2.[812,1399,315,1404]" pageId="2" pageNumber="295">
|
||
DIAGNOSIS.
|
||
<taxonomicName id="CEF94D697B2AFFABFC0EFBF1E65E9D70" authority="Grichanov" authorityName="Grichanov" authorityYear="2022" box="[995,1399,1115,1137]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="cameroonensis" status="sp. nov.">
|
||
<emphasis id="3B8DEAF87B2AFFABFC0EFBF1E62E9D70" box="[995,1287,1115,1137]" italics="true" pageId="2" pageNumber="295">Griphophanes cameroonensis</emphasis>
|
||
Grichanov
|
||
</taxonomicName>
|
||
<emphasis id="3B8DEAF87B2AFFABFCC1FBDCE0749D8A" bold="true" box="[812,861,1142,1163]" pageId="2" pageNumber="295">
|
||
<taxonomicNameLabel id="20BE57837B2AFFABFCC1FBDCE0749D8A" box="[812,861,1142,1163]" pageId="2" pageNumber="295" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is very close to
|
||
<taxonomicName id="CEF94D697B2AFFABFBFEFBDCE63D9D8D" authority="Grichanov" authorityName="Grichanov" authorityYear="2010" box="[1043,1300,1142,1164]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="congoensis">
|
||
<emphasis id="3B8DEAF87B2AFFABFBFEFBDCE7899D8D" box="[1043,1184,1142,1164]" italics="true" pageId="2" pageNumber="295">G. congoensis</emphasis>
|
||
Grichanov
|
||
</taxonomicName>
|
||
, differing from the latter in morphology of male genitalia. Ventral arm of male surstylus is angular at middle, pointed apically in
|
||
<taxonomicName id="CEF94D697B2AFFABFAB2FB06E0BE9DDD" authorityName="Grichanov" authorityYear="2010" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="congoensis">
|
||
<emphasis id="3B8DEAF87B2AFFABFAB2FB06E0BE9DDD" italics="true" pageId="2" pageNumber="295">G. congoensis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="6D684B1B7B2AFFABFC45FB6CE77E9DDA" author="Grichanov I. Ya." box="[936,1111,1222,1244]" pageId="2" pageNumber="295" pagination="1 - 20" refId="ref2458" refString="Grichanov I. Ya. 2010. Discovery of Griphophanes Grootaert & Meuffels and Nepalomyia Hollis in the Afrotropical Region with a key to Afrotropical genera of Peloropeodinae (Diptera: Dolichopodidae) // Zootaxa. Vol. 2668. P. 1 - 20." type="book chapter" year="2010">Grichanov, 2010</bibRefCitation>
|
||
:
|
||
<figureCitation id="91C22A6F7B2AFFABFB88FB6CE7EA9DDD" box="[1125,1219,1222,1244]" captionStart="Figs 1–9" captionStartId="1.[226,263,1969,1988]" captionText="Figs 1–9. Griphophanes cameroonensis, sp.n., male holotype (1–2, 4–9), female (3, 7): 1 — habitus; 2–3 — head; 4 — fore tibia and tarsus, dorsal view; 5— hind tibia and tarsus, lateral view; 6–7 —wing; 8–9 — hypopygium (8 — left lateral view; 9 — ventral view). Рис. 1–9. Griphophanes cameroonensis, sp.n., голотип, самец (1–2, 4–9), самка (3, 7): 1 — внеШний вид; 2–3 — голова; 4 — переднЯЯ голень и лапка, сверху; 5— ЗаднЯЯ голень и лапка, сбоку; 6–7 — крыло; 8–9 — гипопигий (8 — слева сбоку; 9 — сниЗу)." pageId="2" pageNumber="295">Figs 5–7</figureCitation>
|
||
], whereas ventral arm of surstylus is straight, broad to apex in the new species. Male cercus is rounded in
|
||
<taxonomicName id="CEF94D697B2AFFABFBC7FB56E79B9C10" authorityName="Grichanov" authorityYear="2010" box="[1066,1202,1276,1297]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="congoensis">
|
||
<emphasis id="3B8DEAF87B2AFFABFBC7FB56E79B9C10" box="[1066,1202,1276,1297]" italics="true" pageId="2" pageNumber="295">G. congoensis</emphasis>
|
||
</taxonomicName>
|
||
, but triangular in
|
||
<emphasis id="3B8DEAF87B2AFFABFAB2FB56E0ED9C2D" italics="true" pageId="2" pageNumber="295">
|
||
<taxonomicName id="CEF94D697B2AFFABFAB2FB56E0E99C2D" authorityName="Grichanov" authorityYear="2022" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="cameroonensis">G. cameroonensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
A female described has striking difference from males in wing vein length ratios and may belong to another species (females of the two other Afrotropical species are currently unknown).
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||
</paragraph>
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||
</subSubSection>
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||
</treatment>
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||
</document> |