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<mods:title id="EB7C68303B7FB8EB53389F074C276203">Crocodylomorph and dinosaur tracks from the lowermost Jurassic of Le Veillon (western France): ichnotaxonomic revision of the type material (Lapparent collection)</mods:title>
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<emphasis id="3BEEEACEDE1E7C1CFC9597CBDE43F966" bold="true" box="[859,1078,1687,1713]" italics="true" pageId="12" pageNumber="353">Batrachopus gilberti</emphasis>
<bibRefCitation id="6D0B4B2DDE1E7C1CFB8697CADFF9F978" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[1096,1420,1685,1712]" pageId="12" pageNumber="353" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967</bibRefCitation>
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(
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)
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<emphasis id="3BEEEACEDE1E7C1CFCE397B3D99BF8D0" box="[813,1006,1775,1799]" italics="true" pageId="12" pageNumber="353">Batrachopus gilberti</emphasis>
<bibRefCitation id="6D0B4B2DDE1E7C1CFC3897B3DF06F8D0" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[1014,1395,1775,1799]" pageId="12" pageNumber="353" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967: 11-13</bibRefCitation>
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, pl. I, fig. 4.
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. — Subjective junior synonym of
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<emphasis id="3BEEEACEDE1E7C1CFB7F9665DF14F886" box="[1201,1377,1849,1873]" italics="true" pageId="12" pageNumber="353">Batrachopus deweyi</emphasis>
(Hitchcock, 1843)
<bibRefCitation id="6D0B4B2DDE1E7C1CFC699608DE25F8BB" author="HITCHCOCK E." box="[935,1104,1876,1900]" pageId="12" pageNumber="353" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock, 1845</bibRefCitation>
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.
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EXAMINED MATERIAL. —
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Other specimens:
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,
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,
</materialsCitation>
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<specimenCode id="593C9EA7DE1E7C1CFCE5968FD9B4F83C" box="[811,961,2003,2027]" pageId="12" pageNumber="353">ULB-04C14_F</specimenCode>
,
</materialsCitation>
<materialsCitation id="B9F23C81DE1E7C1CFC05968FDE1BF83C" ID-GBIF-Occurrence="4873857313" box="[971,1134,2003,2027]" collectionCode="ULB" pageId="12" pageNumber="353" specimenCode="ULB-04C14_G" specimenCount="1">
<specimenCode id="593C9EA7DE1E7C1CFC05968FDE1CF83C" box="[971,1129,2003,2027]" pageId="12" pageNumber="353">ULB-04C14_G</specimenCode>
,
</materialsCitation>
<materialsCitation id="B9F23C81DE1E7C1CFBBA968FDF6CF83C" ID-GBIF-Occurrence="4873857318" box="[1140,1305,2003,2027]" collectionCode="ULB" pageId="12" pageNumber="353" specimenCode="ULB-04C14_H" specimenCount="1">
<specimenCode id="593C9EA7DE1E7C1CFBBA968FDF60F83C" box="[1140,1301,2003,2027]" pageId="12" pageNumber="353">ULB-04C14_H</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="5DE56654DE1F7C1DFF4A9504D928FB40" ID-DOI="http://doi.org/10.5281/zenodo.11501867" ID-Zenodo-Dep="11501867" httpUri="https://zenodo.org/record/11501867/files/figure.png" pageId="13" pageNumber="354" startId="13.[132,143,1112,1129]" targetBox="[166,1418,215,1069]" targetPageId="13" targetType="figure">
<paragraph id="092536DCDE1F7C1DFF4A9504D928FB40" blockId="13.[132,1455,1112,1175]" pageId="13" pageNumber="354">
<smallCapsWord id="0FC3A000DE1F7C1DFF4A9504DAE8FBBE" baselines="1125,1126" box="[132,157,1112,1129]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="13" pageNumber="354">FIG</smallCapsWord>
. 6. — “
<taxonomicName id="CE9A4D5FDE1F7C1DFF139504D809FBBE" authority="&quot; Lapparent &amp; Montenat, 1967" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[221,636,1112,1129]" genus="Anatopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="3BEEEACEDE1F7C1DFF139504DB08FBBE" box="[221,381,1112,1129]" italics="true" pageId="13" pageNumber="354">Anatopus palmatus</emphasis>
<bibRefCitation id="6D0B4B2DDE1F7C1DFE459504D809FBBE" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[395,636,1112,1129]" pageId="13" pageNumber="354" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967</bibRefCitation>
</taxonomicName>
, that is here invalidated:
<emphasis id="3BEEEACEDE1F7C1DFC859504D918FBBE" bold="true" box="[843,877,1112,1129]" pageId="13" pageNumber="354">A -C</emphasis>
, ULB-04C15_B (holotype of “
<taxonomicName id="CE9A4D5FDE1F7C1DFBA89504DEBCFBBE" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[1126,1225,1112,1129]" genus="Anatopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="3BEEEACEDE1F7C1DFBA89504DEBCFBBE" box="[1126,1225,1112,1129]" italics="true" pageId="13" pageNumber="354">A. palmatus</emphasis>
</taxonomicName>
”), photograph (
<emphasis id="3BEEEACEDE1F7C1DFA829504DF2CFBBE" bold="true" box="[1356,1369,1112,1129]" pageId="13" pageNumber="354">A</emphasis>
), interpretative sketch of
<bibRefCitation id="6D0B4B2DDE1F7C1DFEC59533DB8EFB57" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[267,507,1135,1152]" pageId="13" pageNumber="354" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat (1967</bibRefCitation>
: fig. 16A and pl.XII.3) (
<emphasis id="3BEEEACEDE1F7C1DFD749533D8B2FB57" bold="true" box="[698,711,1135,1152]" pageId="13" pageNumber="354">B</emphasis>
), and our interpretation (
<emphasis id="3BEEEACEDE1F7C1DFC599533D9D0FB57" bold="true" box="[919,933,1135,1152]" pageId="13" pageNumber="354">C</emphasis>
);
<emphasis id="3BEEEACEDE1F7C1DFC7B9533D9A0FB57" bold="true" box="[949,981,1135,1152]" pageId="13" pageNumber="354">D -F</emphasis>
, ULB-04C15_C, photograph (
<emphasis id="3BEEEACEDE1F7C1DFB1F9533DEAAFB57" bold="true" box="[1233,1247,1135,1152]" pageId="13" pageNumber="354">D</emphasis>
), interpretative sketch of
<bibRefCitation id="6D0B4B2DDE1F7C1DFF4A95DADB06FB40" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[132,371,1158,1175]" pageId="13" pageNumber="354" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat (1967</bibRefCitation>
: fig. 16B1) (
<emphasis id="3BEEEACEDE1F7C1DFE1895DADB97FB40" bold="true" box="[470,482,1158,1175]" pageId="13" pageNumber="354">E</emphasis>
), and our interpretation (
<emphasis id="3BEEEACEDE1F7C1DFD7E95DAD8CEFB40" bold="true" box="[688,699,1158,1175]" pageId="13" pageNumber="354">F</emphasis>
). Scale bars: 5 cm.
</paragraph>
</caption>
<subSubSection id="41806557DE1F7C1DFF4A9588D970F83C" pageId="13" pageNumber="354" type="description">
<paragraph id="092536DCDE1F7C1DFF4A9588DB60FB3B" blockId="13.[130,777,1236,2028]" box="[132,277,1236,1262]" pageId="13" pageNumber="354">
<smallCapsWord id="0FC3A000DE1F7C1DFF4A9588DB60FB3B" baselines="1255,1255" box="[132,277,1236,1262]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="13" pageNumber="354">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="092536DCDE1F7C1DFF4A95AFD970F83C" blockId="13.[130,777,1236,2028]" pageId="13" pageNumber="354">
The material includes a single trackway (ULB-04C10_E;
<figureCitation id="91A12A59DE1F7C1DFF48944FDAB1FAF9" box="[134,196,1299,1326]" captionStart="FIG" captionStartId="14.[132,143,1940,1957]" captionTargetBox="[152,1425,215,1869]" captionTargetId="figure-18@14.[152,1418,209,1891]" captionTargetPageId="14" captionText="FIG. 7. — Batrachopus deweyi (Hitchcock,1843) Hitchcock, 1845: A-C, trackway ULB-04C10_E, photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C); D-F, pes/manus set 04C10_A_1 (holotype of “B. gilberti” that is here invalidated), photograph (D), DEM and false-colour depth map (E) and interpretative sketch (F); G-I, pes/manus set 04C14_C, photograph (G), DEM and false-colour depth map (H) and interpretative sketch (I); J-L, pes/manus set 04C14_C, photograph (J), DEM and false-colour depth map (K) and interpretative sketch (L). Scale bars represent: A-C, 10 cm; D-L, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11501871" httpUri="https://zenodo.org/record/11501871/files/figure.png" pageId="13" pageNumber="354">Fig. 7</figureCitation>
A-C) and several isolated pes/manus sets (
<figureCitation id="91A12A59DE1F7C1DFD5C944FD8A4FAF9" box="[658,721,1299,1326]" captionStart="FIG" captionStartId="14.[132,143,1940,1957]" captionTargetBox="[152,1425,215,1869]" captionTargetId="figure-18@14.[152,1418,209,1891]" captionTargetPageId="14" captionText="FIG. 7. — Batrachopus deweyi (Hitchcock,1843) Hitchcock, 1845: A-C, trackway ULB-04C10_E, photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C); D-F, pes/manus set 04C10_A_1 (holotype of “B. gilberti” that is here invalidated), photograph (D), DEM and false-colour depth map (E) and interpretative sketch (F); G-I, pes/manus set 04C14_C, photograph (G), DEM and false-colour depth map (H) and interpretative sketch (I); J-L, pes/manus set 04C14_C, photograph (J), DEM and false-colour depth map (K) and interpretative sketch (L). Scale bars represent: A-C, 10 cm; D-L, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11501871" httpUri="https://zenodo.org/record/11501871/files/figure.png" pageId="13" pageNumber="354">Fig. 7</figureCitation>
D-L;
<tableCitation id="44180367DE1F7C1DFF4A946FDABBFA9A" box="[132,206,1331,1357]" captionStart="TABLE" captionStartId="6.[134,143,220,237]" captionTargetPageId="6" captionText="TABLE 1. — Values measured on Hettangian crocodylomorph tracks from Le Veillon. Abbreviations: LM, LP, length of manus and pes tracks, respectively; WM, WP, width of manus and pes tracks, respectively; I-V, divarication angle between digit I and digit V; I-IV, divarication angle between digit I and digit IV (all in cm except I-IV and I-V in degrees)." httpUri="http://table.plazi.org/id/5DE56654DE147C16FF489180DBCEFECD" pageId="13" pageNumber="354" tableUuid="5DE56654DE147C16FF489180DBCEFECD">Table 1</tableCitation>
).The trackway is narrow,
<quantity id="CE629B39DE1F7C1DFE1D946FD866FA99" box="[467,531,1331,1358]" metricMagnitude="-1" metricUnit="m" metricValue="7.6" pageId="13" pageNumber="354" unit="cm" value="76.0">76 cm</quantity>
long and
<quantity id="CE629B39DE1F7C1DFDBA946FD8D2FA99" box="[628,679,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="354" unit="cm" value="8.0">8 cm</quantity>
wide and is composed of five consecutive pes/manus sets. Pes and manus track strides are 33.0-(35.5)-38.0 cm long. Pes track and manus track pace is 16.0-(17.7)-
<quantity id="CE629B39DE1F7C1DFE4494CFDB97FA79" box="[394,482,1427,1454]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" pageId="13" pageNumber="354" unit="cm" value="19.5">19.5 cm</quantity>
long. Pes and manus tracks show a small positive rotation angle relative to the trackway axis (15-(20)-25°). Pes and manus tracks show a pronounced heteropody. Pes imprints are digitigrade and functionally tetradactyl, longer than wide to as wide as long (2.7-(3.2)-
<quantity id="CE629B39DE1F7C1DFD73974FD972F9FA" box="[701,775,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="354" unit="cm" value="3.9">3.9 cm</quantity>
long and 2.1-(3.0)-
<quantity id="CE629B39DE1F7C1DFE9E976FDBE9F99A" box="[336,412,1587,1613]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="354" unit="cm" value="3.8">3.8 cm</quantity>
wide;
<tableCitation id="44180367DE1F7C1DFE28976ED841F99A" box="[486,564,1586,1613]" captionStart="TABLE" captionStartId="6.[134,143,220,237]" captionTargetPageId="6" captionText="TABLE 1. — Values measured on Hettangian crocodylomorph tracks from Le Veillon. Abbreviations: LM, LP, length of manus and pes tracks, respectively; WM, WP, width of manus and pes tracks, respectively; I-V, divarication angle between digit I and digit V; I-IV, divarication angle between digit I and digit IV (all in cm except I-IV and I-V in degrees)." httpUri="http://table.plazi.org/id/5DE56654DE147C16FF489180DBCEFECD" pageId="13" pageNumber="354" tableUuid="5DE56654DE147C16FF489180DBCEFECD">Table 1</tableCitation>
). The
<taxonomicName id="CE9A4D5FDE1F7C1DFDB3976ED8A3F99A" authority="PL" authorityName="PL" box="[637,726,1586,1613]" family="Chirotheriidae" genus="Dahutherium" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="ratio">ratio PL</taxonomicName>
/PW varies from 0.8 to 1.4. On pes tracks, impressions of digits are clearly marked, straight to curved, and short to quite elongated. Their apices are rounded and can sometimes bear tiny marks of claws. On pes tracks, impression of digit III is always the longest, and impression of digit I the smallest (
<figureCitation id="91A12A59DE1F7C1DFDF8978ED804F93B" box="[566,625,1746,1772]" captionStart="FIG" captionStartId="14.[132,143,1940,1957]" captionTargetBox="[152,1425,215,1869]" captionTargetId="figure-18@14.[152,1418,209,1891]" captionTargetPageId="14" captionText="FIG. 7. — Batrachopus deweyi (Hitchcock,1843) Hitchcock, 1845: A-C, trackway ULB-04C10_E, photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C); D-F, pes/manus set 04C10_A_1 (holotype of “B. gilberti” that is here invalidated), photograph (D), DEM and false-colour depth map (E) and interpretative sketch (F); G-I, pes/manus set 04C14_C, photograph (G), DEM and false-colour depth map (H) and interpretative sketch (I); J-L, pes/manus set 04C14_C, photograph (J), DEM and false-colour depth map (K) and interpretative sketch (L). Scale bars represent: A-C, 10 cm; D-L, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11501871" httpUri="https://zenodo.org/record/11501871/files/figure.png" pageId="13" pageNumber="354">Fig. 7</figureCitation>
D-L). Track of digit II is often the second longest. The divarication angle of digits I-IV strongly varies (36-(64)-82°).On pes tracks,the traces of digits II and III are the deepest. Pes tracks can show a short and large plantar-like impression.Manus tracks are pentadactyl and smaller than pes tracks, wider than long to as long as wide, 1.1-(1.8)-
<quantity id="CE629B39DE1F7C1DFF2B96CDDB45F87B" box="[229,304,1937,1964]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="354" unit="cm" value="2.4">2.4 cm</quantity>
long and 1.6-(1.8)-
<quantity id="CE629B39DE1F7C1DFE3896CDD83EF87B" box="[502,587,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="354" unit="mm" value="2.2">2.2 mm</quantity>
wide (ML/MW = 0.7-(0.)-1.9). On manus tracks, digit imprints are very short and rounded. When preserved, digit V is oriented posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="41806557DE1F7C1FFCE39588D8F8FA79" lastPageId="15" lastPageNumber="356" pageId="13" pageNumber="354" type="discussion">
<paragraph id="092536DCDE1F7C1DFCE39588D9E4FB3B" blockId="13.[811,1457,1236,2028]" box="[813,913,1236,1262]" pageId="13" pageNumber="354">
<smallCapsWord id="0FC3A000DE1F7C1DFCE39588D9E4FB3B" baselines="1255,1255" box="[813,913,1236,1262]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="13" pageNumber="354">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="092536DCDE1F7C1DFCE395AFDE78F97A" blockId="13.[811,1457,1236,2028]" pageId="13" pageNumber="354">
These tracks were first ascribed to
<taxonomicName id="CE9A4D5FDE1F7C1DFB4295AFDF56FADA" box="[1164,1315,1267,1293]" class="Mammalia" family="Dugongidae" genus="Cheirotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sirenia" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFB4295AFDF56FADA" box="[1164,1315,1267,1293]" italics="true" pageId="13" pageNumber="354">Cheirotherium</emphasis>
</taxonomicName>
(synonym of
<taxonomicName id="CE9A4D5FDE1F7C1DFCE5944FDE30FAF9" authority="Kaup, 1835" authorityName="Kaup" authorityYear="1835" box="[811,1093,1299,1326]" class="Mammalia" family="Chirotheriidae" genus="Chirotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sirenia" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFCE5944FD9CCFAFA" box="[811,953,1299,1325]" italics="true" pageId="13" pageNumber="354">Chirotherium</emphasis>
Kaup, 1835
</taxonomicName>
) by
<bibRefCitation id="6D0B4B2DDE1F7C1DFBB69448DF07FAF9" author="BESSONNAT G. &amp; LAPPARENT A. F. DE &amp; MONTENAT C. &amp; TERS M." box="[1144,1394,1299,1326]" pageId="13" pageNumber="354" pagination="5324 - 5326" refId="ref16499" refString="BESSONNAT G., LAPPARENT A. F. DE, MONTENAT C. &amp; TERS M. 1965. - Decouverte de nombreuses empreintes de pas de reptiles dans le Lias inferieur de la cote de Vendee. Comptes rendus hebdomadaires des seances de l'Academie des sciences 260: 5324 - 5326. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 4020 b / f 699. item" type="journal article" year="1965">
Bessonnat
<emphasis id="3BEEEACEDE1F7C1DFB229448DF54FAFA" box="[1260,1313,1299,1325]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
(1965)
</bibRefCitation>
. This Triassic ichnogenus differs from the tracks from Le Veillon in showing larger tracks, pentadactyl pes tracks with a trace of digit V much more proximal than the group I-IV. Based on the following morphological characters, tracks from Le Veillon can be confidently ascribed to
<taxonomicName id="CE9A4D5FDE1F7C1DFB7894EFD916FA39" authority="Hitchcock, 1845" authorityName="Hitchcock" authorityYear="1845" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="354" phylum="Arthropoda" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFB7894EFDF47FA1A" box="[1206,1330,1459,1485]" italics="true" pageId="13" pageNumber="354">Batrachopus</emphasis>
<bibRefCitation id="6D0B4B2DDE1F7C1DFAF794EFD916FA39" author="HITCHCOCK E." pageId="13" pageNumber="354" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock, 1845</bibRefCitation>
</taxonomicName>
(see the emended diagnosis in
<bibRefCitation id="6D0B4B2DDE1F7C1DFB7D948FDFD5FA3A" author="OLSEN P. E. &amp; PADIAN K." box="[1203,1440,1491,1517]" pageId="13" pageNumber="354" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian 1986</bibRefCitation>
): small quadrupedal trackway; pes tracks are digitigrade and functionally tetradactyl; digit III of the pes is the longest and digit I the shortest; pentadactyl and digitigrade manus, usually rotated so that digit II points forward, digit IV points laterally, and digit V points posteriorly; and manus much smaller than the pes.
</paragraph>
<paragraph id="092536DCDE1F7C1FFC8D97EED863FEA6" blockId="13.[811,1457,1236,2028]" lastBlockId="15.[131,778,215,1454]" lastPageId="15" lastPageNumber="356" pageId="13" pageNumber="354">
The gross morphology of
<taxonomicName id="CE9A4D5FDE1F7C1DFBA097EEDE84F91B" authorityName="Hitchcock" authorityYear="1845" box="[1134,1265,1714,1740]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="354" phylum="Arthropoda" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFBA097EEDE84F91B" box="[1134,1265,1714,1740]" italics="true" pageId="13" pageNumber="354">Batrachopus</emphasis>
</taxonomicName>
is close to those of
<taxonomicName id="CE9A4D5FDE1F7C1DFC83978EDE1EF93A" authority="Hitchcock, 1858" authorityName="Hitchcock" authorityYear="1858" box="[845,1131,1746,1773]" class="Reptilia" family="Batrachopodidae" genus="Antipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFC83978ED9D5F93B" box="[845,928,1746,1772]" italics="true" pageId="13" pageNumber="354">Antipus</emphasis>
<bibRefCitation id="6D0B4B2DDE1F7C1DFC64978EDE1EF93A" author="HITCHCOCK E." box="[938,1131,1746,1773]" pageId="13" pageNumber="354" refId="ref17969" refString="HITCHCOCK E. 1858. - Ichnology of New England: a Report on the Sandstone of the Connecticut Valley Especially its Fossil Footmarks, Made to the Government of the Commonwealth of Massachusetts. William White, Boston, 220 p." type="book" year="1858">Hitchcock, 1858</bibRefCitation>
</taxonomicName>
and
<emphasis id="3BEEEACEDE1F7C1DFB64978EDF24F93B" box="[1194,1361,1746,1772]" italics="true" pageId="13" pageNumber="354">Crocodylopodus</emphasis>
Fuentes Vidarte &amp; Meijide Calvo, 1999.
<bibRefCitation id="6D0B4B2DDE1F7C1DFB6897AEDFC4F8DB" author="OLSEN P. E. &amp; PADIAN K." box="[1190,1457,1778,1804]" pageId="13" pageNumber="354" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986)</bibRefCitation>
considered
<taxonomicName id="CE9A4D5FDE1F7C1DFC67964ED98FF8FB" authorityName="Hitchcock" authorityYear="1858" box="[937,1018,1810,1836]" class="Reptilia" family="Batrachopodidae" genus="Antipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFC67964ED98FF8FB" box="[937,1018,1810,1836]" italics="true" pageId="13" pageNumber="354">Antipus</emphasis>
</taxonomicName>
as a subjective synonym of
<taxonomicName id="CE9A4D5FDE1F7C1DFAE7964EDFDDF8FB" authorityName="Hitchcock" authorityYear="1845" box="[1321,1448,1810,1836]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="354" phylum="Arthropoda" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFAE7964EDFDDF8FB" box="[1321,1448,1810,1836]" italics="true" pageId="13" pageNumber="354">Batrachopus</emphasis>
</taxonomicName>
. However,
<bibRefCitation id="6D0B4B2DDE1F7C1DFC6E966DDE21F89C" author="COOMBS W. P. &amp; JR" box="[928,1108,1841,1867]" pageId="13" pageNumber="354" pagination="327 - 331" refId="ref16734" refString="COOMBS W. P. JR 1996. - Redescription of the ichnospecies Antipus flexiloquus Hitchcock, from the Early Jurassic of the Connecticut Valley. Journal of Paleontology 70: 327 - 331. https: // doi. org / 10.1017 / s 0022336000023416" type="journal article" year="1996">Coombs (1996)</bibRefCitation>
and
<bibRefCitation id="6D0B4B2DDE1F7C1DFB5D966DDFC4F89B" author="LOCKLEY M. G. &amp; MEYER C." box="[1171,1457,1841,1868]" pageId="13" pageNumber="354" pagination="167 - 178" refId="ref18264" refString="LOCKLEY M. G. &amp; MEYER C. 2004. - Crocodylomorph trackways from the Jurassic to Early Cretaceous of North America and Europe: implications for ichnotaxonomy. Ichnos 11: 167 - 178. https: // doi. org / 10.1080 / 10420940490428832" type="journal article" year="2004">Lockley &amp; Meyer (2004)</bibRefCitation>
rejected this proposition. According to
<bibRefCitation id="6D0B4B2DDE1F7C1DFB25960DD90DF85C" author="LOCKLEY M. G. &amp; MEYER C." pageId="13" pageNumber="354" pagination="167 - 178" refId="ref18264" refString="LOCKLEY M. G. &amp; MEYER C. 2004. - Crocodylomorph trackways from the Jurassic to Early Cretaceous of North America and Europe: implications for ichnotaxonomy. Ichnos 11: 167 - 178. https: // doi. org / 10.1080 / 10420940490428832" type="journal article" year="2004">Lockley &amp; Meyer (2004)</bibRefCitation>
,
<taxonomicName id="CE9A4D5FDE1F7C1DFC48962ED9A2F85B" authorityName="Hitchcock" authorityYear="1858" box="[902,983,1906,1932]" class="Reptilia" family="Batrachopodidae" genus="Antipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFC48962ED9A2F85B" box="[902,983,1906,1932]" italics="true" pageId="13" pageNumber="354">Antipus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="CE9A4D5FDE1F7C1DFBAB962DDE91F85C" authorityName="Hitchcock" authorityYear="1845" box="[1125,1252,1905,1931]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="354" phylum="Arthropoda" rank="genus">
<emphasis id="3BEEEACEDE1F7C1DFBAB962DDE91F85C" box="[1125,1252,1905,1931]" italics="true" pageId="13" pageNumber="354">Batrachopus</emphasis>
</taxonomicName>
in showing: much longer impressions of digits on the manus tracks; slender, tapering, quite curvated digit tracks with higher divarication angles and narrow claw impressions. The tracks from Le Veillon clearly differ from
<taxonomicName id="CE9A4D5FDE1D7C1FFE179184D85EFF25" authorityName="Hitchcock" authorityYear="1858" box="[473,555,216,242]" class="Reptilia" family="Batrachopodidae" genus="Antipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="15" pageNumber="356" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE1D7C1FFE179184D85EFF25" box="[473,555,216,242]" italics="true" pageId="15" pageNumber="356">Antipus</emphasis>
</taxonomicName>
in showing shorter blunt-toed digits.
<emphasis id="3BEEEACEDE1D7C1FFE9E91ABDB87FEC6" box="[336,498,247,273]" italics="true" pageId="15" pageNumber="356">Crocodylopodus</emphasis>
differs from tracks of Le Veillon in showing more slender and divergent digits, less outward rotation of the pes and more outward rotation of the manus (
<bibRefCitation id="6D0B4B2DDE1D7C1FFEC8900BD872FEA6" author="LOCKLEY M. G. &amp; MEYER C." box="[262,519,343,369]" pageId="15" pageNumber="356" pagination="167 - 178" refId="ref18264" refString="LOCKLEY M. G. &amp; MEYER C. 2004. - Crocodylomorph trackways from the Jurassic to Early Cretaceous of North America and Europe: implications for ichnotaxonomy. Ichnos 11: 167 - 178. https: // doi. org / 10.1080 / 10420940490428832" type="journal article" year="2004">Lockley &amp; Meyer 2004</bibRefCitation>
).
</paragraph>
<caption id="5DE56654DE1C7C1EFF4A96C8DF71F83E" ID-DOI="http://doi.org/10.5281/zenodo.11501871" ID-Zenodo-Dep="11501871" httpUri="https://zenodo.org/record/11501871/files/figure.png" pageId="14" pageNumber="355" startId="14.[132,143,1940,1957]" targetBox="[152,1425,215,1869]" targetPageId="14" targetType="figure">
<paragraph id="092536DCDE1C7C1EFF4A96C8DF71F83E" blockId="14.[132,1455,1940,2025]" pageId="14" pageNumber="355">
<smallCapsWord id="0FC3A000DE1C7C1EFF4A96C8DAE8F873" baselines="1953,1953" box="[132,157,1940,1957]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="14" pageNumber="355">FIG</smallCapsWord>
. 7. —
<taxonomicName id="CE9A4D5FDE1C7C1EFF1D96C8D8D7F872" authority="(Hitchcock, 1843) Hitchcock, 1845" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" box="[211,674,1940,1957]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="deweyi">
<emphasis id="3BEEEACEDE1C7C1EFF1D96C8DB09F872" box="[211,380,1940,1957]" italics="true" pageId="14" pageNumber="355">Batrachopus deweyi</emphasis>
(Hitchcock,1843)
<bibRefCitation id="6D0B4B2DDE1C7C1EFDD896C8D8D7F872" author="HITCHCOCK E." box="[534,674,1940,1957]" pageId="14" pageNumber="355" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock, 1845</bibRefCitation>
</taxonomicName>
:
<emphasis id="3BEEEACEDE1C7C1EFD6796C8D8BEF872" bold="true" box="[681,715,1940,1957]" pageId="14" pageNumber="355">A -C</emphasis>
, trackway ULB-04C10_E, photograph (
<emphasis id="3BEEEACEDE1C7C1EFBDE96C8DE68F872" bold="true" box="[1040,1053,1940,1957]" pageId="14" pageNumber="355">A</emphasis>
), DEM and false-colour depth map (
<emphasis id="3BEEEACEDE1C7C1EFA8696C8DF20F872" bold="true" box="[1352,1365,1940,1957]" pageId="14" pageNumber="355">B</emphasis>
) and interpretative sketch (
<emphasis id="3BEEEACEDE1C7C1EFEDD96F7DB54F86B" bold="true" box="[275,289,1963,1980]" pageId="14" pageNumber="355">C</emphasis>
);
<emphasis id="3BEEEACEDE1C7C1EFEFE96F7DB25F86B" bold="true" box="[304,336,1963,1980]" pageId="14" pageNumber="355">D -F</emphasis>
, pes/manus set 04C10_A_1 (holotype of “
<taxonomicName id="CE9A4D5FDE1C7C1EFD6096F7D888F86B" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[686,765,1963,1980]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis id="3BEEEACEDE1C7C1EFD6096F7D888F86B" box="[686,765,1963,1980]" italics="true" pageId="14" pageNumber="355">B. gilberti</emphasis>
</taxonomicName>
” that is here invalidated), photograph (
<emphasis id="3BEEEACEDE1C7C1EFBF196F7DE38F86B" bold="true" box="[1087,1101,1963,1980]" pageId="14" pageNumber="355">D</emphasis>
), DEM and false-colour depth map (
<emphasis id="3BEEEACEDE1C7C1EFAB496F7DFF3F86B" bold="true" box="[1402,1414,1963,1980]" pageId="14" pageNumber="355">E</emphasis>
) and interpretative sketch (
<emphasis id="3BEEEACEDE1C7C1EFEF4969EDB30F804" bold="true" box="[314,325,1986,2003]" pageId="14" pageNumber="355">F</emphasis>
);
<emphasis id="3BEEEACEDE1C7C1EFE9B969EDB05F804" bold="true" box="[341,368,1986,2003]" pageId="14" pageNumber="355">G -I</emphasis>
, pes/manus set 04C14_C, photograph (
<emphasis id="3BEEEACEDE1C7C1EFD0F969ED8BAF804" bold="true" box="[705,719,1986,2003]" pageId="14" pageNumber="355">G</emphasis>
), DEM and false-colour depth map (
<emphasis id="3BEEEACEDE1C7C1EFBCD969EDE64F804" bold="true" box="[1027,1041,1986,2003]" pageId="14" pageNumber="355">H</emphasis>
) and interpretative sketch (
<emphasis id="3BEEEACEDE1C7C1EFB38969EDE89F804" bold="true" box="[1270,1276,1986,2003]" pageId="14" pageNumber="355">I</emphasis>
);
<emphasis id="3BEEEACEDE1C7C1EFAC5969EDF5DF804" bold="true" box="[1291,1320,1986,2003]" pageId="14" pageNumber="355">J -L</emphasis>
, pes/manus set 04C14_C, photograph (
<emphasis id="3BEEEACEDE1C7C1EFE869684DB27F83E" bold="true" box="[328,338,2008,2025]" pageId="14" pageNumber="355">J</emphasis>
), DEM and false-colour depth map (
<emphasis id="3BEEEACEDE1C7C1EFD4E9684D8F8F83E" bold="true" box="[640,653,2008,2025]" pageId="14" pageNumber="355">K</emphasis>
) and interpretative sketch (
<emphasis id="3BEEEACEDE1C7C1EFCBE9684D90EF83E" bold="true" box="[880,891,2008,2025]" pageId="14" pageNumber="355">L</emphasis>
). Scale bars represent: A-C, 10 cm; D-L, 1 cm.
</paragraph>
</caption>
<paragraph id="092536DCDE1D7C1FFF55902BD8F8FA79" blockId="15.[131,778,215,1454]" pageId="15" pageNumber="356">
Based on ULB-04C10_
<bibRefCitation id="6D0B4B2DDE1D7C1FFE55902BD970FE46" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[411,773,375,401]" pageId="15" pageNumber="356" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">A, Lapparent &amp; Montenat (1967)</bibRefCitation>
erected
<taxonomicName id="CE9A4D5FDE1D7C1FFF1490CBDB33FE67" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[218,326,406,432]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis id="3BEEEACEDE1D7C1FFF1490CBDB33FE67" box="[218,326,406,432]" italics="true" pageId="15" pageNumber="356">B. gilberti</emphasis>
</taxonomicName>
. The validity of the diverse ichnospecies of
<taxonomicName id="CE9A4D5FDE1D7C1FFF6E90EADB6AFE07" authorityName="Hitchcock" authorityYear="1845" box="[160,287,438,464]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="3BEEEACEDE1D7C1FFF6E90EADB6AFE07" box="[160,287,438,464]" italics="true" pageId="15" pageNumber="356">Batrachopus</emphasis>
</taxonomicName>
was strongly debated (
<bibRefCitation id="6D0B4B2DDE1D7C1FFDDA90EAD88AFE07" author="OLSEN P. E. &amp; PADIAN K." box="[532,767,438,465]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian 1986</bibRefCitation>
;
<bibRefCitation id="6D0B4B2DDE1D7C1FFF4A908ADB43FE26" author="RAINFORTH E. C." box="[132,310,470,497]" pageId="15" pageNumber="356" refId="ref19604" refString="RAINFORTH E. C. 2005. - Ichnotaxonomy of the Fossil Footprints of the Connecticut Valley (Early Jurassic, Newark Supergroup, Connecticut and Massachusetts). Columbia University, New York, 1301 p." type="book" year="2005">Rainforth 2005</bibRefCitation>
,
<bibRefCitation id="6D0B4B2DDE1D7C1FFE89908ADBF7FE27" author="RAINFORTH E. C." box="[327,386,470,496]" pageId="15" pageNumber="356" pagination="49 - 59" refId="ref19642" refString="RAINFORTH E. C. 2007. - Ichnotaxonomic updates from the Newark Supergroup, in Contributions to the Paleontology of New Jersey (II): Field Guide and Proceedings. Geological Association of New Jersey, Trenton: 49 - 59." type="book chapter" year="2007">2007</bibRefCitation>
). In their revision of
<taxonomicName id="CE9A4D5FDE1D7C1FFDB1908ADBE6FDC7" authority=", Olsen &amp; Padian (1986)" authorityName=", Olsen &amp; Padian" authorityYear="1986" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="3BEEEACEDE1D7C1FFDB1908AD975FE27" box="[639,768,470,496]" italics="true" pageId="15" pageNumber="356">Batrachopus</emphasis>
,
<bibRefCitation id="6D0B4B2DDE1D7C1FFF4A90AADBE6FDC7" author="OLSEN P. E. &amp; PADIAN K." box="[132,403,502,528]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986)</bibRefCitation>
</taxonomicName>
considered only three valid ichnospecies for this ichnogenus and emended the diagnosis of each of them:
<taxonomicName id="CE9A4D5FDE1D7C1FFE86936BDAB4FDA6" authority="(Hitchcock, 1843) Hitchcock, 1845" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">
<emphasis id="3BEEEACEDE1D7C1FFE86936BDBDBFD87" box="[328,430,566,592]" italics="true" pageId="15" pageNumber="356">B. deweyi</emphasis>
(Hitchcock, 1843)
<bibRefCitation id="6D0B4B2DDE1D7C1FFD41936ADAB4FDA6" author="HITCHCOCK E." pageId="15" pageNumber="356" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock, 1845</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="CE9A4D5FDE1D7C1FFF02930BD8A2FDA6" authority="(Hitchcock, 1841) Hitchcock 1845" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1841" box="[204,727,598,625]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="3BEEEACEDE1D7C1FFF02930BDB33FDA7" box="[204,326,598,624]" italics="true" pageId="15" pageNumber="356">B. parvulus</emphasis>
(
<bibRefCitation id="6D0B4B2DDE1D7C1FFE98930AD867FDA7" author="HITCHCOCK E." box="[342,530,598,624]" pageId="15" pageNumber="356" refId="ref17893" refString="HITCHCOCK E. 1841. - Final Report on the Geology of Massachusetts. J. H. Butler, Northampton, 831 p." type="book" year="1841">Hitchcock, 1841</bibRefCitation>
)
<bibRefCitation id="6D0B4B2DDE1D7C1FFDEA930AD8A2FDA6" author="HITCHCOCK E." box="[548,727,598,625]" pageId="15" pageNumber="356" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock 1845</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="CE9A4D5FDE1D7C1FFF48932BDB2AFD47" authority="Lull, 1904" authorityName="Lull" authorityYear="1904" box="[134,351,630,656]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="3BEEEACEDE1D7C1FFF48932BDA91FD47" box="[134,228,630,656]" italics="true" pageId="15" pageNumber="356">B. dispar</emphasis>
<bibRefCitation id="6D0B4B2DDE1D7C1FFF27932ADB2AFD47" author="LULL R. S." box="[233,351,630,656]" pageId="15" pageNumber="356" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull, 1904</bibRefCitation>
</taxonomicName>
. Based on the following characters, the tracks from Le Veillon can be confidently ascribed to
<taxonomicName id="CE9A4D5FDE1D7C1FFD7893CADAD5FD18" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">
<emphasis id="3BEEEACEDE1D7C1FFD7893CADAD5FD18" italics="true" pageId="15" pageNumber="356">B. deweyi</emphasis>
</taxonomicName>
(see emended diagnosis in
<bibRefCitation id="6D0B4B2DDE1D7C1FFE0993E9D8C3FD18" author="OLSEN P. E. &amp; PADIAN K." box="[455,694,693,720]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian 1986</bibRefCitation>
): range size of pes length (between 2 and
<quantity id="CE629B39DE1D7C1FFE279389D857FD27" box="[489,546,725,752]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="356" unit="cm" value="6.0">6 cm</quantity>
); manus tracks about 75% of the length of the pes; on pes tracks, imprints of digit II and IV are sub-equal in length. We consider “
<taxonomicName id="CE9A4D5FDE1D7C1FFD5C924AD889FCF8" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[658,764,789,815]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis id="3BEEEACEDE1D7C1FFD5C924AD889FCF8" box="[658,764,789,815]" italics="true" pageId="15" pageNumber="356">B. gilberti</emphasis>
</taxonomicName>
” as a junior synonym of
<taxonomicName id="CE9A4D5FDE1D7C1FFE4D926ADB93FC98" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" box="[387,486,821,847]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">
<emphasis id="3BEEEACEDE1D7C1FFE4D926ADB93FC98" box="[387,486,821,847]" italics="true" pageId="15" pageNumber="356">B. deweyi</emphasis>
</taxonomicName>
.
<taxonomicName id="CE9A4D5FDE1D7C1FFE3B926AD818FC98" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1841" box="[501,621,821,847]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="3BEEEACEDE1D7C1FFE3B926AD818FC98" box="[501,621,821,847]" italics="true" pageId="15" pageNumber="356">B. parvulus</emphasis>
</taxonomicName>
(see emended diagnosis in
<bibRefCitation id="6D0B4B2DDE1D7C1FFEC09209D86FFCB8" author="OLSEN P. E. &amp; PADIAN K." box="[270,538,853,879]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986))</bibRefCitation>
differs from tracks of Le Veillon in showing a very short impression of digit I on pes tracks. According to
<bibRefCitation id="6D0B4B2DDE1D7C1FFE4A92C9D849FC67" author="RAINFORTH E. C." box="[388,572,917,944]" pageId="15" pageNumber="356" refId="ref19604" refString="RAINFORTH E. C. 2005. - Ichnotaxonomy of the Fossil Footprints of the Connecticut Valley (Early Jurassic, Newark Supergroup, Connecticut and Massachusetts). Columbia University, New York, 1301 p." type="book" year="2005">Rainforth (2005)</bibRefCitation>
,
<taxonomicName id="CE9A4D5FDE1D7C1FFD8792CAD8CAFC78" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1841" box="[585,703,917,943]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="3BEEEACEDE1D7C1FFD8792CAD8CAFC78" box="[585,703,917,943]" italics="true" pageId="15" pageNumber="356">B. parvulus</emphasis>
</taxonomicName>
can be synonymised with the
<typeStatus id="D621887EDE1D7C1FFEB892E9DBD0FC18" box="[374,421,949,975]" pageId="15" pageNumber="356">type</typeStatus>
ichnospecies
<emphasis id="3BEEEACEDE1D7C1FFDF492E9D973FC18" box="[570,774,949,975]" italics="true" pageId="15" pageNumber="356">
<taxonomicName id="CE9A4D5FDE1D7C1FFDF492E9D8D4FC18" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" box="[570,673,949,975]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">B. deweyi</taxonomicName>
.
<taxonomicName id="CE9A4D5FDE1D7C1FFD6492E9D973FC18" authorityName="Lull" authorityYear="1904" box="[682,774,949,975]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="dispar">B. dispar</taxonomicName>
</emphasis>
(see emended diagnosis in
<bibRefCitation id="6D0B4B2DDE1D7C1FFE609288D8CEFC39" author="OLSEN P. E. &amp; PADIAN K." box="[430,699,980,1007]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986))</bibRefCitation>
differs from tracks of Le Veillon in showing a stronger heteropody.
<bibRefCitation id="6D0B4B2DDE1D7C1FFF4A9548DB36FBF8" author="RAINFORTH E. C." box="[132,323,1044,1071]" pageId="15" pageNumber="356" refId="ref19604" refString="RAINFORTH E. C. 2005. - Ichnotaxonomy of the Fossil Footprints of the Connecticut Valley (Early Jurassic, Newark Supergroup, Connecticut and Massachusetts). Columbia University, New York, 1301 p." type="book" year="2005">Rainforth (2005)</bibRefCitation>
considered three additional valid ichnospecies:
<taxonomicName id="CE9A4D5FDE1D7C1FFF139569D8F6FB99" authority="(Hitchcock, 1858) Lull, 1904" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1858" box="[221,643,1076,1103]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="3BEEEACEDE1D7C1FFF139569DB43FB99" box="[221,310,1076,1102]" italics="true" pageId="15" pageNumber="356">B. bellus</emphasis>
(
<bibRefCitation id="6D0B4B2DDE1D7C1FFE8B9568D874FB98" author="HITCHCOCK E." box="[325,513,1076,1103]" pageId="15" pageNumber="356" refId="ref17969" refString="HITCHCOCK E. 1858. - Ichnology of New England: a Report on the Sandstone of the Connecticut Valley Especially its Fossil Footmarks, Made to the Government of the Commonwealth of Massachusetts. William White, Boston, 220 p." type="book" year="1858">Hitchcock, 1858</bibRefCitation>
)
<bibRefCitation id="6D0B4B2DDE1D7C1FFDDE9568D8F6FB99" author="LULL R. S." box="[528,643,1076,1102]" pageId="15" pageNumber="356" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull, 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="CE9A4D5FDE1D7C1FFD5C9569DBB0FBB9" authority="(Hitchcock, 1865) Lull, 1904" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1865" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="3BEEEACEDE1D7C1FFD5C9569D972FB99" box="[658,775,1076,1102]" italics="true" pageId="15" pageNumber="356">B. gracilior</emphasis>
(Hitchcock, 1865)
<bibRefCitation id="6D0B4B2DDE1D7C1FFE9A9508DBB0FBB9" author="LULL R. S." box="[340,453,1108,1134]" pageId="15" pageNumber="356" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull, 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="CE9A4D5FDE1D7C1FFE1C9509DA8EFB59" authority="(Hitchcock, 1848) Lull, 1904" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1848" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="15" pageNumber="356" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="3BEEEACEDE1D7C1FFE1C9509D842FBB9" box="[466,567,1108,1134]" italics="true" pageId="15" pageNumber="356">B. gracilis</emphasis>
(Hitchcock, 1848)
<bibRefCitation id="6D0B4B2DDE1D7C1FFF4D9528DA8EFB59" author="LULL R. S." box="[131,251,1140,1166]" pageId="15" pageNumber="356" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull, 1904</bibRefCitation>
</taxonomicName>
. It contrasts with
<bibRefCitation id="6D0B4B2DDE1D7C1FFE069528D8BAFB59" author="OLSEN P. E. &amp; PADIAN K." box="[456,719,1140,1166]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986)</bibRefCitation>
who subjectively synonymised these ichnospecies in
<emphasis id="3BEEEACEDE1D7C1FFD5495C9D97FFB79" box="[666,778,1172,1198]" italics="true" pageId="15" pageNumber="356">
<taxonomicName id="CE9A4D5FDE1D7C1FFD5495C9D970FB79" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" box="[666,773,1172,1198]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">B. deweyi</taxonomicName>
.
</emphasis>
According to
<bibRefCitation id="6D0B4B2DDE1D7C1FFEEA95E8DB93FB18" author="RAINFORTH E. C." box="[292,486,1204,1231]" pageId="15" pageNumber="356" refId="ref19604" refString="RAINFORTH E. C. 2005. - Ichnotaxonomy of the Fossil Footprints of the Connecticut Valley (Early Jurassic, Newark Supergroup, Connecticut and Massachusetts). Columbia University, New York, 1301 p." type="book" year="2005">Rainforth (2005)</bibRefCitation>
,
<taxonomicName id="CE9A4D5FDE1D7C1FFE3995E9D821FB19" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1858" box="[503,596,1204,1230]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="3BEEEACEDE1D7C1FFE3995E9D821FB19" box="[503,596,1204,1230]" italics="true" pageId="15" pageNumber="356">B. bellus</emphasis>
</taxonomicName>
cannot be synonymised with
<taxonomicName id="CE9A4D5FDE1D7C1FFEE39588DBE4FB39" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" box="[301,401,1236,1262]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">
<emphasis id="3BEEEACEDE1D7C1FFEE39588DBE4FB39" box="[301,401,1236,1262]" italics="true" pageId="15" pageNumber="356">B. deweyi</emphasis>
</taxonomicName>
because it differs from this ichnospecies in showing tridactyl pes (not concordant with the sketch of
<taxonomicName id="CE9A4D5FDE1D7C1FFF229448DB31FAFA" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1858" box="[236,324,1299,1325]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="3BEEEACEDE1D7C1FFF229448DB31FAFA" box="[236,324,1299,1325]" italics="true" pageId="15" pageNumber="356">B. bellus</emphasis>
</taxonomicName>
in fig. 20.2B of
<bibRefCitation id="6D0B4B2DDE1D7C1FFE39944FD976FAFA" author="OLSEN P. E. &amp; PADIAN K." box="[503,771,1299,1326]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986))</bibRefCitation>
.
<bibRefCitation id="6D0B4B2DDE1D7C1FFF4A946FDB36FA9A" author="RAINFORTH E. C." box="[132,323,1331,1357]" pageId="15" pageNumber="356" pagination="49 - 59" refId="ref19642" refString="RAINFORTH E. C. 2007. - Ichnotaxonomic updates from the Newark Supergroup, in Contributions to the Paleontology of New Jersey (II): Field Guide and Proceedings. Geological Association of New Jersey, Trenton: 49 - 59." type="book chapter" year="2007">Rainforth (2007)</bibRefCitation>
tentatively synonymised
<taxonomicName id="CE9A4D5FDE1D7C1FFDAE9468D8A2FA9A" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1865" box="[608,727,1331,1357]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="3BEEEACEDE1D7C1FFDAE9468D8A2FA9A" box="[608,727,1331,1357]" italics="true" pageId="15" pageNumber="356">B. gracilior</emphasis>
</taxonomicName>
and
<taxonomicName id="CE9A4D5FDE1D7C1FFF4A9408DA9DFABA" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1848" box="[132,232,1363,1389]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="15" pageNumber="356" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="3BEEEACEDE1D7C1FFF4A9408DA9DFABA" box="[132,232,1363,1389]" italics="true" pageId="15" pageNumber="356">B. gracilis</emphasis>
</taxonomicName>
.
<bibRefCitation id="6D0B4B2DDE1D7C1FFF3A940FDB99FABA" author="OLSEN P. E. &amp; PADIAN K." box="[244,492,1363,1389]" pageId="15" pageNumber="356" pagination="259 - 273" refId="ref19377" refString="OLSEN P. E. &amp; PADIAN K. 1986. - Earliest records of Batrachopus from the southwestern United States, and a revision of some Early Mesozoic crocodylomorph ichnogenera in PADIAN K. (ed), The Beginning of the Age of Dinosaurs. Cambridge University Press: 259 - 273." type="book chapter" year="1986">Olsen &amp; Padian (1986)</bibRefCitation>
explained that “apart from its smaller, longer pace and the lack of impressions of digit V in the pes”
<taxonomicName id="CE9A4D5FDE1D7C1FFED494C8DBF4FA7A" authorityName="Lull" authorityYear="1904" baseAuthorityName="Hitchcock" baseAuthorityYear="1848" box="[282,385,1427,1453]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="15" pageNumber="356" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="3BEEEACEDE1D7C1FFED494C8DBF4FA7A" box="[282,385,1427,1453]" italics="true" pageId="15" pageNumber="356">B. gracilis</emphasis>
</taxonomicName>
is identical to
<taxonomicName id="CE9A4D5FDE1D7C1FFDEC94C8D8F3FA7A" authorityName="Hitchcock" authorityYear="1845" baseAuthorityName="Hitchcock" baseAuthorityYear="1843" box="[546,646,1427,1453]" class="Insecta" family="Romaleidae" genus="Batrachopus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="deweyi">
<emphasis id="3BEEEACEDE1D7C1FFDEC94C8D8F3FA7A" box="[546,646,1427,1453]" italics="true" pageId="15" pageNumber="356">B. deweyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>