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<document id="76222C006E0BEAE19122BDD876976186" ID-DOI="10.26107/RBZ-2020-0054" ID-ISSN="2345-7600" ID-Zenodo-Dep="5344486" ID-ZooBank="7B13EA09-398D-4AB6-8358-308694D8CCCD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630198330262" checkinUser="diego" docAuthor="Páll-Gergely, Barna, Hunyadi, András &amp; Hausdorf, Bernhard" docDate="2020" docId="8069CD03FFD4FF9EFC27B4AE5D906236" docLanguage="en" docName="RBZ-2020-0054.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Landouria intha Páll-Gergely &amp; Hunyadi &amp; Hausdorf 2020, new species" docType="treatment" docVersion="3" lastPageNumber="383" masterDocId="7C50B57BFFD5FF9AFFE1B07358536A6E" masterDocTitle="A new species of Landouria Godwin-Austen, 1918 (Gastropoda: Stylommatophora: Camaenidae) from Myanmar" masterLastPageNumber="386" masterPageNumber="379" pageNumber="380" updateTime="1699183233406" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6598A50154359E19CD2DE0BB4268C2BA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BA5EFEBEC89DE734F932729376C540DE">
<mods:title id="9936DCDF63F483AB305F8473CDAB2D7D">A new species of Landouria Godwin-Austen, 1918 (Gastropoda: Stylommatophora: Camaenidae) from Myanmar</mods:title>
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<mods:name id="6FD3978824C154149A8C874F4CC07482" type="personal">
<mods:role id="C0B8120026C4A7DF5BC876C0AF9E5C59">
<mods:roleTerm id="5C7EA70979D48A2656646314CA13BB52">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0A51E7B371CF239747767EF383AF709">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="5A20B6B94E972F0C50B4D34368E752FB" type="personal">
<mods:role id="F71A310C35B302A477AA28AE7203D597">
<mods:roleTerm id="68B0FC04F7818925C3D15D9DA9D1D58D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2910B5525F1352A87F522B6B32E131B">Hunyadi, András</mods:namePart>
</mods:name>
<mods:name id="ABCCA8EBD2E3E17654DE4F6B86551B92" type="personal">
<mods:role id="A1378F1A07183644284C8CB602B79767">
<mods:roleTerm id="DF37E828244D66ABBCA12C6AC1DB0D80">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="55D863933F1398DC81DAB2853674988C">Hausdorf, Bernhard</mods:namePart>
</mods:name>
<mods:typeOfResource id="2F736DDCC41A224981A3F6829E10059D">text</mods:typeOfResource>
<mods:relatedItem id="BF5FBBB0AC0C1EE211AFEA95EF10D550" type="host">
<mods:titleInfo id="A366C5B7183BD0FBC0A6073BB7C8A33E">
<mods:title id="AFEC866A71A7A5945B4011ACD705897C">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="841CC7AD0B30AF07407173F7114A306A">
<mods:date id="50779219D6F7E6627CBAA5E4C9835D14">2020</mods:date>
<mods:detail id="6F97DC2CF63689E74DF135EF8F11FAE5" type="pubDate">
<mods:number id="EE106BCACA267D7AC22ED9E4C64ECF73">2020-05-14</mods:number>
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<mods:detail id="03B4F8D2B3A9A8031DA68EF96C0A419E" type="volume">
<mods:number id="CDAD039BDED4E6F4948E396ED790DF4B">68</mods:number>
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<mods:start id="D38E5F18DC2463A36179B81E4E9CF086">379</mods:start>
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<mods:classification id="0B37E2A32F81F68E58E6437FFCAA310C">journal article</mods:classification>
<mods:identifier id="17734B878EBE7E2F2088255F28FB2ED8" type="DOI">10.26107/RBZ-2020-0054</mods:identifier>
<mods:identifier id="5CF7FAD8D919EC091D9831564C83FD73" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DBBF8731E6C1F6909EFDDEA8397572DE" type="Zenodo-Dep">5344486</mods:identifier>
<mods:identifier id="FE0C2EFBA7059B74C56F233962220688" type="ZooBank">7B13EA09-398D-4AB6-8358-308694D8CCCD</mods:identifier>
</mods:mods>
<treatment id="8069CD03FFD4FF9EFC27B4AE5D906236" LSID="urn:lsid:plazi:treatment:8069CD03FFD4FF9EFC27B4AE5D906236" httpUri="http://treatment.plazi.org/id/8069CD03FFD4FF9EFC27B4AE5D906236" lastPageId="4" lastPageNumber="383" pageId="1" pageNumber="380">
<subSubSection id="40DA2F9EFFD4FF9BFC27B4AE5D416E9B" box="[966,1298,1245,1269]" pageId="1" pageNumber="380" type="nomenclature">
<paragraph id="087F7C15FFD4FF9BFC27B4AE5D416E9B" blockId="1.[966,1298,1245,1301]" box="[966,1298,1245,1269]" pageId="1" pageNumber="380">
<heading id="5337CB79FFD4FF9BFC27B4AE5D416E9B" bold="true" box="[966,1298,1245,1269]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="380" reason="2">
<emphasis id="3AB4A007FFD4FF9BFC27B4AE5D416E9B" bold="true" box="[966,1298,1245,1269]" pageId="1" pageNumber="380">
<taxonomicName id="CFC00796FFD4FF9BFC27B4AE5C2D6E9B" ID-CoL="7RTHJ" authority="Páll-Gergely &amp; Hunyadi &amp; Hausdorf, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Hausdorf" authorityYear="2020" box="[966,1150,1245,1269]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="380" phylum="Mollusca" rank="species" species="intha" status="sp. nov.">
<emphasis id="3AB4A007FFD4FF9BFC27B4AE5C2D6E9B" bold="true" box="[966,1150,1245,1269]" italics="true" pageId="1" pageNumber="380">Landouria intha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="21871D7CFFD4FF9BFB6CB4AE5D416E9B" box="[1165,1298,1245,1269]" pageId="1" pageNumber="380" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="40DA2F9EFFD4FF9BFBD1B48E5CFB6F7B" box="[1072,1192,1277,1301]" pageId="1" pageNumber="380" type="description">
<paragraph id="087F7C15FFD4FF9BFBD1B48E5CFB6F7B" blockId="1.[966,1298,1245,1301]" box="[1072,1192,1277,1301]" pageId="1" pageNumber="380">
(
<figureCitation id="90FB6090FFD4FF9BFBD9B48E5CF36F7B" box="[1080,1184,1277,1301]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[113,148,2051,2073]" captionStartId-1="4.[136,171,1148,1170]" captionStartId-2="5.[113,148,1991,2013]" captionTargetBox-0="[113,1451,230,2035]" captionTargetBox-1="[133,1476,192,1129]" captionTargetBox-2="[113,1451,262,1976]" captionTargetId-0="figure-10@3.[113,1451,230,2035]" captionTargetId-1="figure-7@4.[133,1484,184,1129]" captionTargetId-2="figure-10@5.[113,1451,262,1976]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 2. Holotype of Landouria intha, new species (HNHM 104442). Scale bar for A = 10 mm, B–E = 1 mm." captionText-1="Fig. 3. Reproductive anatomy of the holotype of Landouria intha, new species (HNHM 104442). Abbreviations: a, atrium; ag, albumen gland; b, bursa of the bursa copulatrix; bb, base of the peduncle of bursa copulatrix; e, epiphallus; f, flagellum; p, penis; ps, penial sheath (indicated with grey shading); rm, retractor muscle; so, spermoviduct; v, vagina; vd, vas deferens. Asterisks indicate originally conjoined ends. Scale bar = 5 mm." captionText-2="Fig. 4. Inner structure of genital organs of Landouria intha, new species (holotype, HNHM 104442). A, penis; B, penial papilla; C, penial papilla opened; D, vagina and base of bursa copulatrix; E, flagellum." figureDoi-0="http://doi.org/10.5281/zenodo.5344494" figureDoi-1="http://doi.org/10.5281/zenodo.5344496" figureDoi-2="http://doi.org/10.5281/zenodo.5344498" httpUri-0="https://zenodo.org/record/5344494/files/figure.png" httpUri-1="https://zenodo.org/record/5344496/files/figure.png" httpUri-2="https://zenodo.org/record/5344498/files/figure.png" pageId="1" pageNumber="380">Figs. 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="40DA2F9EFFD4FF9BFCCCB54E5D486C3B" pageId="1" pageNumber="380" type="materials_examined">
<paragraph id="087F7C15FFD4FF9BFCCCB54E5D486C3B" blockId="1.[811,1452,1341,1621]" pageId="1" pageNumber="380">
<emphasis id="3AB4A007FFD4FF9BFCCCB54E5B806F3B" bold="true" box="[813,979,1341,1365]" pageId="1" pageNumber="380">Type material.</emphasis>
<materialsCitation id="B8A87648FFD4FF9BFC39B54E5CB46F9B" collectingDate="2018-10-04" collectionCode="HNHM" collectorName="A. Hunyadi &amp; J. U. Otani" country="Myanmar" county="Pinlaung" elevation="700" latitude="19.97547" location="Nam Pam" longLatPrecision="1" longitude="96.67076" municipality="Laneli Bridge" pageId="1" pageNumber="380" specimenCode="HNHM 104442" specimenCount="1" stateProvince="Shan" typeStatus="holotype">
<typeStatus id="D77BC2B7FFD4FF9BFC39B54E5C686F3B" box="[984,1083,1341,1365]" pageId="1" pageNumber="380" type="holotype">Holotype</typeStatus>
(D:
<quantity id="CF38D1F0FFD4FF9BFB88B54E5C9A6F3B" box="[1129,1225,1341,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="1" pageNumber="380" unit="mm" value="19.4">19.4 mm</quantity>
, H:
<quantity id="CF38D1F0FFD4FF9BFB12B54E5D166F3B" box="[1267,1349,1341,1365]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="380" unit="mm" value="6.8">6.8 mm</quantity>
) (
<specimenCode id="5866D46EFFD4FF9BFAB9B54E5B2C6F1B" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="380">HNHM 104442</specimenCode>
),
<collectingCountry id="70D73C85FFD4FF9BFC71B52D5BA96F18" box="[912,1018,1374,1398]" name="Myanmar" pageId="1" pageNumber="380">Myanmar</collectingCountry>
,
<collectingRegion id="CA04B2F7FFD4FF9BFBE2B52E5C256F1B" box="[1027,1142,1373,1397]" country="Myanmar" name="Shan" pageId="1" pageNumber="380">Shan State</collectingRegion>
,
<collectingCounty id="E11E0499FFD4FF9BFB9EB52E5C8D6F1B" box="[1151,1246,1373,1397]" pageId="1" pageNumber="380">Pinlaung</collectingCounty>
centre, SSW ca.
<quantity id="CF38D1F0FFD4FF9BFA71B52E5B1E6FFB" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="380" unit="km" value="29.0">29 km</quantity>
–
<collectingMunicipality id="E81BE66FFFD4FF9BFC8BB50E5C526FFB" box="[874,1025,1405,1429]" pageId="1" pageNumber="380">Laneli Bridge</collectingMunicipality>
,
<location id="0D1F2ACEFFD4FF9BFBEDB50D5C296FFB" LSID="urn:lsid:plazi:treatment:8069CD03FFD4FF9EFC27B4AE5D906236:0D1F2ACEFFD4FF9BFBEDB50D5C296FFB" box="[1036,1146,1406,1429]" country="Myanmar" county="Pinlaung" latitude="19.97547" longLatPrecision="1" longitude="96.67076" municipality="Laneli Bridge" name="Nam Pam" pageId="1" pageNumber="380" stateProvince="Shan">Nam Pam</location>
, right side of rd., near “War Lee Kwey Cave (Resurgence)”,
<elevation id="83ED9B26FFD4FF9BFB70B5EE5D406FDB" box="[1169,1299,1437,1461]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="380" unit="m" value="700.0">
<quantity id="CF38D1F0FFD4FF9BFB70B5EE5C8B6FDB" box="[1169,1240,1437,1461]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="380" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="6DF41AD2FFD4FF9BFAC2B5EE5DF46FDB" box="[1315,1447,1437,1461]" degrees="19.97547" direction="north" orientation="latitude" pageId="1" pageNumber="380" precision="1" value="19.97547">19.97547°N</geoCoordinate>
,
<geoCoordinate id="6DF41AD2FFD4FF9BFCCCB5CE5BF86FBB" box="[813,939,1469,1493]" degrees="96.67076" direction="east" orientation="longitude" pageId="1" pageNumber="380" precision="1" value="96.67076">96.67076°E</geoCoordinate>
(locality codes: 2018/31 and 20181004C), coll.
<collectorName id="A53519C3FFD4FF9BFCCCB5AD5BF66F9B" box="[813,933,1501,1526]" pageId="1" pageNumber="380">A. Hunyadi</collectorName>
&amp;
<collectorName id="A53519C3FFD4FF9BFC22B5AD5C7C6F9B" box="[963,1071,1501,1525]" pageId="1" pageNumber="380">J.U. Otani</collectorName>
,
<date id="7C7E5AD5FFD4FF9BFBD6B5AE5CB46F9B" box="[1079,1255,1501,1525]" pageId="1" pageNumber="380" value="2018-10-04">
<collectingDate id="6C3AA33DFFD4FF9BFBD6B5AE5CB46F9B" box="[1079,1255,1501,1525]" pageId="1" pageNumber="380" value="2018-10-04">04 October 2018</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="B8A87648FFD4FF9BFB0EB5AD5BDC6C5B" collectingDate="2018-10-04" collectionCode="HNHM" collectorName="A. Hunyadi &amp; J. U. Otani" country="Myanmar" county="Pinlaung" elevation="700" latitude="19.97547" location="Nam Pam" longLatPrecision="1" longitude="96.67076" municipality="Laneli Bridge" pageId="1" pageNumber="380" specimenCode="HNHM 104443" specimenCount="1" stateProvince="Shan" typeStatus="paratype">
<typeStatus id="D77BC2B7FFD4FF9BFB0EB5AD5D196F98" box="[1263,1354,1502,1526]" pageId="1" pageNumber="380" type="paratype">Paratype</typeStatus>
(juvenile shell and body in ethanol) (
<specimenCode id="5866D46EFFD4FF9BFBBBB58E5D596C7B" box="[1114,1290,1533,1557]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="380">HNHM 104443</specimenCode>
), same data as holotype
</materialsCitation>
;
<typeStatus id="D77BC2B7FFD4FF9BFC7CB66D5C556C58" box="[925,1030,1566,1590]" pageId="1" pageNumber="380" type="paratype">Paratypes</typeStatus>
(9 empty shells) (HA) (D:
<quantity id="CF38D1F0FFD4FF9BFAA1B66E5B0A6C3B" metricMagnitude="-2" metricUnit="m" metricValue="1.775" metricValueMax="1.88" metricValueMin="1.67" pageId="1" pageNumber="380" unit="mm" value="17.75" valueMax="18.8" valueMin="16.7">16.7–18.8 mm</quantity>
, H:
<quantity id="CF38D1F0FFD4FF9BFC68B64E5C5D6C3B" box="[905,1038,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.5" metricValueMin="5.1" pageId="1" pageNumber="380" unit="mm" value="5.8" valueMax="6.5" valueMin="5.1">5.1–6.5 mm</quantity>
), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="40DA2F9EFFD4FF98FCCCB60E59236DA1" lastPageId="2" lastPageNumber="381" pageId="1" pageNumber="380" type="description">
<paragraph id="087F7C15FFD4FF98FCCCB60E5B556F01" blockId="1.[812,1452,1661,2101]" lastBlockId="2.[135,775,1335,1391]" lastPageId="2" lastPageNumber="381" pageId="1" pageNumber="380">
<emphasis id="3AB4A007FFD4FF9BFCCCB60E5C6F6CFB" bold="true" box="[813,1084,1661,1685]" pageId="1" pageNumber="380">Description of the shell</emphasis>
(
<figureCitation id="90FB6090FFD4FF9BFBAFB60E5CF46CFB" box="[1102,1191,1661,1685]" captionStart="Fig" captionStartId="3.[113,148,2051,2073]" captionTargetBox="[113,1451,230,2035]" captionTargetId="figure-10@3.[113,1451,230,2035]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Landouria intha, new species (HNHM 104442). Scale bar for A = 10 mm, B–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5344494" httpUri="https://zenodo.org/record/5344494/files/figure.png" pageId="1" pageNumber="380">Fig. 2A</figureCitation>
). D:
<quantity id="CF38D1F0FFD4FF9BFB03B60E5DD06CFB" box="[1250,1411,1661,1685]" metricMagnitude="-2" metricUnit="m" metricValue="1.8049999999999997" metricValueMax="1.94" metricValueMin="1.67" pageId="1" pageNumber="380" unit="mm" value="18.049999999999997" valueMax="19.4" valueMin="16.7">16.7–19.4 mm</quantity>
, H:
<quantity id="CF38D1F0FFD4FF9BFCCCB6EE5BE06CDB" box="[813,947,1693,1717]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.8" metricValueMin="5.1" pageId="1" pageNumber="380" unit="mm" value="5.949999999999999" valueMax="6.8" valueMin="5.1">5.1–6.8 mm</quantity>
. Colour light reddish brown. Shell depressed, dorsal side flat, consisting of 5.25–5.5 slightly convex whorls. Body whorl with strong, sharp keel, with slight furrow below and above keel. Protoconch (
<figureCitation id="90FB6090FFD4FF9BFB58B68E5D5D6D7B" box="[1209,1294,1789,1813]" captionStart="Fig" captionStartId="3.[113,148,2051,2073]" captionTargetBox="[113,1451,230,2035]" captionTargetId="figure-10@3.[113,1451,230,2035]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Landouria intha, new species (HNHM 104442). Scale bar for A = 10 mm, B–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5344494" httpUri="https://zenodo.org/record/5344494/files/figure.png" pageId="1" pageNumber="380">Fig. 2B</figureCitation>
) consisting of 1.75 whorls, almost smooth, with very fine radial ribbing and slight indication of spiral striation along the suture. Teleoconch roughly, irregularly wrinkled without any traces of spiral striation. Radial wrinkles with additional, scale-like, triangular periostracal folds (scales) (
<figureCitation id="90FB6090FFD4FF9BFB22B7EE5D6A6DDB" box="[1219,1337,1949,1973]" captionStart="Fig" captionStartId="3.[113,148,2051,2073]" captionTargetBox="[113,1451,230,2035]" captionTargetId="figure-10@3.[113,1451,230,2035]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Landouria intha, new species (HNHM 104442). Scale bar for A = 10 mm, B–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5344494" httpUri="https://zenodo.org/record/5344494/files/figure.png" pageId="1" pageNumber="380">Fig. 2C–E</figureCitation>
). Aperture quadrangular due to the prominent keel, upper insertion of peristome slightly descending. Peristome lighter in colour than the rest of the shell, strongly expanded, slightly reflected, discontinuous; parietal callus only indicated by slight transparent calcareous layer. Umbilicus open, relatively wide, funnel-shaped, occupying ca. one third of the shell’s width.
</paragraph>
<caption id="5CBF2C9DFFD7FF98FF66B4C35D906F6E" ID-DOI="http://doi.org/10.5281/zenodo.5344492" ID-Zenodo-Dep="5344492" httpUri="https://zenodo.org/record/5344492/files/figure.png" pageId="2" pageNumber="381" startId="2.[135,170,1200,1222]" targetBox="[136,1473,192,1182]" targetPageId="2">
<paragraph id="087F7C15FFD7FF98FF66B4C35D906F6E" blockId="2.[135,1475,1200,1280]" pageId="2" pageNumber="381">
Fig. 1. Bayesian 50% majority rule consensus tree based on mitochondrial 16S rDNA sequences of
<taxonomicName id="CFC00796FFD7FF98FB88B4C35D4C6EA8" box="[1129,1311,1200,1222]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="3AB4A007FFD7FF98FB88B4C35C9D6EAB" box="[1129,1230,1200,1221]" italics="true" pageId="2" pageNumber="381">Landouria</emphasis>
species
</taxonomicName>
, and
<taxonomicName id="CFC00796FFD7FF98FAB7B4C358846E8D" authorityName="Minato" authorityYear="1978" class="Gastropoda" family="Camaenidae" genus="Mandarina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="species" species="suenoae">
<emphasis id="3AB4A007FFD7FF98FAB7B4C358846E8D" italics="true" pageId="2" pageNumber="381">Mandarina suenoae</emphasis>
</taxonomicName>
and
<taxonomicName id="CFC00796FFD7FF98FEECB4BE59B66E8C" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1850" box="[269,485,1229,1250]" class="Gastropoda" family="Camaenidae" genus="Euhadra" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="species" species="peliomphala">
<emphasis id="3AB4A007FFD7FF98FEECB4BE59B66E8C" box="[269,485,1229,1250]" italics="true" pageId="2" pageNumber="381">Euhadra peliomphala</emphasis>
</taxonomicName>
as outgroups. Values at the nodes represent Bayesian posterior probabilities (left) and maximum likelihood bootstrap values (right). Numbers after taxon names correspond to museum catalogue numbers or GenBank accession numbers.
</paragraph>
</caption>
<paragraph id="087F7C15FFD7FF98FF69B5E459236DA1" blockId="2.[135,776,1431,2000]" pageId="2" pageNumber="381">
<emphasis id="3AB4A007FFD7FF98FF69B5E459E46FC1" bold="true" box="[136,439,1431,1455]" pageId="2" pageNumber="381">Description of the genitalia</emphasis>
(
<figureCitation id="90FB6090FFD7FF98FE25B5E45A5D6FC1" box="[452,526,1431,1455]" captionStart="Fig" captionStartId="4.[136,171,1148,1170]" captionTargetBox="[133,1476,192,1129]" captionTargetId="figure-7@4.[133,1484,184,1129]" captionTargetPageId="4" captionText="Fig. 3. Reproductive anatomy of the holotype of Landouria intha, new species (HNHM 104442). Abbreviations: a, atrium; ag, albumen gland; b, bursa of the bursa copulatrix; bb, base of the peduncle of bursa copulatrix; e, epiphallus; f, flagellum; p, penis; ps, penial sheath (indicated with grey shading); rm, retractor muscle; so, spermoviduct; v, vagina; vd, vas deferens. Asterisks indicate originally conjoined ends. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5344496" httpUri="https://zenodo.org/record/5344496/files/figure.png" pageId="2" pageNumber="381">Figs. 3</figureCitation>
,
<figureCitation id="90FB6090FFD7FF98FDF9B5E45A7B6FC1" box="[536,552,1431,1455]" captionStart="Fig" captionStartId="5.[113,148,1991,2013]" captionTargetBox="[113,1451,262,1976]" captionTargetId="figure-10@5.[113,1451,262,1976]" captionTargetPageId="5" captionText="Fig. 4. Inner structure of genital organs of Landouria intha, new species (holotype, HNHM 104442). A, penis; B, penial papilla; C, penial papilla opened; D, vagina and base of bursa copulatrix; E, flagellum." figureDoi="http://doi.org/10.5281/zenodo.5344498" httpUri="https://zenodo.org/record/5344498/files/figure.png" pageId="2" pageNumber="381">4</figureCitation>
). Atrium short. Penis very long, without penial caecum, with membranaceous sheath around very distal part of penis. Penis internally (
<figureCitation id="90FB6090FFD7FF98FD3DB5A458FA6C61" captionStart="Fig" captionStartId="5.[113,148,1991,2013]" captionTargetBox="[113,1451,262,1976]" captionTargetId="figure-10@5.[113,1451,262,1976]" captionTargetPageId="5" captionText="Fig. 4. Inner structure of genital organs of Landouria intha, new species (holotype, HNHM 104442). A, penis; B, penial papilla; C, penial papilla opened; D, vagina and base of bursa copulatrix; E, flagellum." figureDoi="http://doi.org/10.5281/zenodo.5344498" httpUri="https://zenodo.org/record/5344498/files/figure.png" pageId="2" pageNumber="381">Fig. 4A</figureCitation>
) with a few fine longitudinal folds that become slightly stronger towards penial verge. Penial verge (
<figureCitation id="90FB6090FFD7FF98FD64B6645AAC6C41" box="[645,767,1559,1583]" captionStart="Fig" captionStartId="5.[113,148,1991,2013]" captionTargetBox="[113,1451,262,1976]" captionTargetId="figure-10@5.[113,1451,262,1976]" captionTargetPageId="5" captionText="Fig. 4. Inner structure of genital organs of Landouria intha, new species (holotype, HNHM 104442). A, penis; B, penial papilla; C, penial papilla opened; D, vagina and base of bursa copulatrix; E, flagellum." figureDoi="http://doi.org/10.5281/zenodo.5344498" httpUri="https://zenodo.org/record/5344498/files/figure.png" pageId="2" pageNumber="381">Fig. 4B, C</figureCitation>
) small, rather conical, slightly irregular, opens terminally. Epiphallus approximately as long as penis, internally also with weak longitudinal folds. Retractor muscle slender, inserts at the middle of epiphallus. Flagellum (
<figureCitation id="90FB6090FFD7FF98FDD7B6E45ADB6CC1" box="[566,648,1687,1711]" captionStart="Fig" captionStartId="5.[113,148,1991,2013]" captionTargetBox="[113,1451,262,1976]" captionTargetId="figure-10@5.[113,1451,262,1976]" captionTargetPageId="5" captionText="Fig. 4. Inner structure of genital organs of Landouria intha, new species (holotype, HNHM 104442). A, penis; B, penial papilla; C, penial papilla opened; D, vagina and base of bursa copulatrix; E, flagellum." figureDoi="http://doi.org/10.5281/zenodo.5344498" httpUri="https://zenodo.org/record/5344498/files/figure.png" pageId="2" pageNumber="381">Fig. 4E</figureCitation>
) very long, broadest at its base, with a few nodes at the basal part, and multiple horizontally arranged nodes at the terminal part.
<collectingRegion id="CA04B2F7FFD7FF98FF69B68B58E26D61" box="[136,177,1784,1807]" country="Hungary" name="Vas" pageId="2" pageNumber="381">Vas</collectingRegion>
deferens narrow and long. Vagina slightly longer than penis, internally with longitudinal, occasionally serrate folds (
<figureCitation id="90FB6090FFD7FF98FF6EB74458BA6D21" box="[143,233,1847,1871]" captionStart="Fig" captionStartId="5.[113,148,1991,2013]" captionTargetBox="[113,1451,262,1976]" captionTargetId="figure-10@5.[113,1451,262,1976]" captionTargetPageId="5" captionText="Fig. 4. Inner structure of genital organs of Landouria intha, new species (holotype, HNHM 104442). A, penis; B, penial papilla; C, penial papilla opened; D, vagina and base of bursa copulatrix; E, flagellum." figureDoi="http://doi.org/10.5281/zenodo.5344498" httpUri="https://zenodo.org/record/5344498/files/figure.png" pageId="2" pageNumber="381">Fig. 4D</figureCitation>
). Peduncle of bursa copulatrix subdivided into a strongly thickened base, internally with elevated, irregular, longitudinal folds, and a slender, much longer proximal part; spermoviduct longer then vagina, albumen gland also very long, elongated.
</paragraph>
</subSubSection>
<subSubSection id="40DA2F9EFFD7FF9EFF69B7845C3D6F96" lastPageId="4" lastPageNumber="383" pageId="2" pageNumber="381" type="diagnosis">
<paragraph id="087F7C15FFD7FF98FF69B7845CF96FE1" blockId="2.[136,775,2039,2095]" lastBlockId="2.[835,1474,1335,1423]" pageId="2" pageNumber="381">
<emphasis id="3AB4A007FFD7FF98FF69B78459DA6261" bold="true" box="[136,393,2039,2063]" pageId="2" pageNumber="381">Differential diagnosis.</emphasis>
This new species differs from all
<taxonomicName id="CFC00796FFD7FF98FF69B864588F6240" authorityName="Albers" authorityYear="1850" box="[136,220,2071,2094]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="genus">
<emphasis id="3AB4A007FFD7FF98FF69B864588F6240" box="[136,220,2071,2094]" italics="true" pageId="2" pageNumber="381">Aegista</emphasis>
</taxonomicName>
,
<taxonomicName id="CFC00796FFD7FF98FF0EB86459366241" authorityName="Godwin-Austen" authorityYear="1918" box="[239,357,2071,2095]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="genus">
<emphasis id="3AB4A007FFD7FF98FF0EB86459366241" box="[239,357,2071,2095]" italics="true" pageId="2" pageNumber="381">Landouria</emphasis>
</taxonomicName>
, and
<taxonomicName id="CFC00796FFD7FF98FE4BB8645AC36241" box="[426,656,2071,2095]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="3AB4A007FFD7FF98FE4BB8645A676241" box="[426,564,2071,2095]" italics="true" pageId="2" pageNumber="381">Plectotropis</emphasis>
species
</taxonomicName>
described from
<collectingCountry id="70D73C85FFD7FF98FC60B54B5BBA6F3E" box="[897,1001,1336,1360]" name="Myanmar" pageId="2" pageNumber="381">Myanmar</collectingCountry>
and
<collectingCountry id="70D73C85FFD7FF98FBC2B5445C096F21" box="[1059,1114,1335,1359]" name="India" pageId="2" pageNumber="381">India</collectingCountry>
by the flat apical side, the wide umbilicus, and the relatively large, permanent periostracal folds covering the entire surface.
</paragraph>
<paragraph id="087F7C15FFD7FF98FCA2B5C45D1D6CE1" blockId="2.[835,1475,1463,1679]" pageId="2" pageNumber="381">
<materialsCitation id="B8A87648FFD7FF98FCA2B5C45D196CE1" collectionCode="NHMUK" collectorName="Godwin-Austen" location="Assam" pageId="2" pageNumber="381" specimenCount="3" stateProvince="Assam" typeStatus="syntype">
The most similar species is
<taxonomicName id="CFC00796FFD7FF98FB8FB5C45D916FA1" authority="Gude, 1914" authorityName="Gude" authorityYear="1914" box="[1134,1474,1463,1487]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="species" species="nutans">
<emphasis id="3AB4A007FFD7FF98FB8FB5C45D6D6FA1" box="[1134,1342,1463,1487]" italics="true" pageId="2" pageNumber="381">Plectotropis nutans</emphasis>
Gude, 1914
</taxonomicName>
(described in
<bibRefCitation id="6C5101E4FFD7FF98FC02B5A45C2B6F81" author="Gude GK" box="[995,1144,1495,1519]" pageId="2" pageNumber="381" pagination="52 - 57" refId="ref3472" refString="Gude GK (1914 a) Descriptions of new species of helicoids from the Indian Region. Proceedings of the Malacological Society of London, 11 (1): 52 - 57." type="journal article" year="1914">Gude, 1914a</bibRefCitation>
;
<figureCitation id="90FB6090FFD7FF98FB66B5A45C9F6F81" box="[1159,1228,1495,1519]" captionStart="Fig" captionStartId="6.[135,170,1380,1402]" captionTargetBox="[138,1472,189,1365]" captionTargetId="figure-22@6.[138,1472,189,1365]" captionTargetPageId="6" captionText="Fig. 5. Syntype of Plectotropis nutans Gude, 1914, NHMUK 1906.3.3.68. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5344500" httpUri="https://zenodo.org/record/5344500/files/figure.png" pageId="2" pageNumber="381">Fig. 5</figureCitation>
) (material examined:
<specimenCount id="1EC6B79CFFD7FF98FCA2B5845BE96C7E" box="[835,954,1527,1552]" pageId="2" pageNumber="381" type="generic" typeStatus="syntypes">3 syntypes</specimenCount>
[
<collectionCode id="6ED1E4D0FFD7FF98FC2BB58B5C606C61" box="[970,1075,1528,1551]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="381" type="Museum">NHMUK</collectionCode>
1906.3.3.68], Habiang, Garo Hills,
<collectingRegion id="CA04B2F7FFD7FF98FCA2B6645BDD6C41" box="[835,910,1559,1583]" country="India" name="Assam" pageId="2" pageNumber="381">Assam</collectingRegion>
, coll.
<collectorName id="A53519C3FFD7FF98FC2FB6645C2E6C41" box="[974,1149,1559,1583]" pageId="2" pageNumber="381">Godwin-Austen</collectorName>
), which has a slightly elevated spire resulting in a low conical shape, a much narrower umbilicus, and no deciduous periostracal folds, but is finely spirally striated and irregularly, finely wrinkled
</materialsCitation>
.
</paragraph>
<paragraph id="087F7C15FFD7FF9EFCA2B6C45AF16F19" blockId="2.[834,1476,1719,2095]" lastBlockId="4.[136,775,1312,1400]" lastPageId="4" lastPageNumber="383" pageId="2" pageNumber="381">
<taxonomicName id="CFC00796FFD7FF98FCA2B6C45D026CA1" authority="(Theobald, 1859)" baseAuthorityName="Theobald" baseAuthorityYear="1859" box="[835,1361,1719,1743]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="381" phylum="Mollusca" rank="species" species="akowtongensis">
<emphasis id="3AB4A007FFD7FF98FCA2B6C45C2D6CA1" box="[835,1150,1719,1743]" italics="true" pageId="2" pageNumber="381">Plectotropis akowtongensis</emphasis>
(
<bibRefCitation id="6C5101E4FFD7FF98FB70B6C45D1A6CA1" author="Theobald W" box="[1169,1353,1719,1743]" pageId="2" pageNumber="381" pagination="305 - 309" refId="ref4580" refString="Theobald W (1859) Descriptions of some new Burmese and Indian Helicidae with remarks on some previously described species. Journal of the Asiatic Society of Bengal, 28 (4): 305 - 309." type="journal article" year="1859">Theobald, 1859</bibRefCitation>
)
</taxonomicName>
(material examined: 6 shells [NHMUK 1906.2.2.130],
<collectingRegion id="CA04B2F7FFD7FF98FADCB6AB5D266C9E" box="[1341,1397,1752,1776]" country="Myanmar" name="Bago" pageId="2" pageNumber="381">Pegu</collectingRegion>
, Thyet Myo, coll. W.T. Blanford; 2 shells [NHMUK],
<collectingRegion id="CA04B2F7FFD7FF98FAD0B68B5D3A6D7E" box="[1329,1385,1784,1808]" country="Myanmar" name="Bago" pageId="2" pageNumber="381">Pegu</collectingRegion>
; 5 shells [NHMUK 1909.3.15.24], Akoutong,
<collectingRegion id="CA04B2F7FFD7FF98FB24B76B5CAE6D5E" box="[1221,1277,1816,1840]" country="Myanmar" name="Bago" pageId="2" pageNumber="381">Pegu</collectingRegion>
; 1 shell, [NHMUK 1910.6.33.1921],
<collectingRegion id="CA04B2F7FFD7FF98FBE8B74B5C106D3E" box="[1033,1091,1848,1872]" country="Myanmar" name="Bago" pageId="2" pageNumber="381">Pegu</collectingRegion>
; 1 shell [NHMUK 1903.7.1.394], Akoukthoung, on
<collectingRegion id="CA04B2F7FFD7FF98FBEAB7245C3D6D01" box="[1035,1134,1879,1903]" country="Myanmar" name="Ayeyawady" pageId="2" pageNumber="381">Irrawady</collectingRegion>
,
<collectingRegion id="CA04B2F7FFD7FF98FB9AB72B5CE06D1E" box="[1147,1203,1880,1904]" country="Myanmar" name="Bago" pageId="2" pageNumber="381">Pegu</collectingRegion>
, coll. Godwin-Austen; 2 shells [NHMUK], Thayet Myo, Burmah, coll. Salisbury) is much smaller, paler in colour, has a low dome-shaped dorsal side and a narrower umbilicus, but is somewhat similar in sculpture. Namely, the shell surface is very finely scaly. Fine periostracal folds, which are nevertheless much smaller than those of the new species, are also present near the suture and the keel in fresh shells. Even in live collected specimens the folds were largely lost, indicating that these are not permanent in
<taxonomicName id="CFC00796FFD1FF9EFE94B5135AC86F16" baseAuthorityName="Theobald" baseAuthorityYear="1859" box="[373,667,1376,1400]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="akowtongensis">
<emphasis id="3AB4A007FFD1FF9EFE94B5135AC86F16" box="[373,667,1376,1400]" italics="true" pageId="4" pageNumber="383">Plectotropis akowtongensis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="5CBF2C9DFFD6FF99FF90B8705CF66276" ID-DOI="http://doi.org/10.5281/zenodo.5344494" ID-Zenodo-Dep="5344494" box="[113,1189,2051,2073]" httpUri="https://zenodo.org/record/5344494/files/figure.png" pageId="3" pageNumber="382" startId="3.[113,148,2051,2073]" targetBox="[113,1451,230,2035]" targetPageId="3">
<paragraph id="087F7C15FFD6FF99FF90B8705CF66276" blockId="3.[113,1189,2051,2073]" box="[113,1189,2051,2073]" pageId="3" pageNumber="382">
Fig. 2. Holotype of
<taxonomicName id="CFC00796FFD6FF99FED6B87059876276" authority="Páll-Gergely &amp; Hunyadi &amp; Hausdorf, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Hausdorf" authorityYear="2020" box="[311,468,2051,2072]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="382" phylum="Mollusca" rank="species" species="intha" status="sp. nov.">
<emphasis id="3AB4A007FFD6FF99FED6B87059876276" box="[311,468,2051,2072]" italics="true" pageId="3" pageNumber="382">Landouria intha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="21871D7CFFD6FF99FE03B8705A046277" box="[482,599,2051,2073]" pageId="3" pageNumber="382" rank="species">new species</taxonomicNameLabel>
(HNHM 104442). Scale bar for A = 10 mm, B–E = 1 mm.
</paragraph>
</caption>
<caption id="5CBF2C9DFFD1FF9EFF69B40F59296E86" ID-DOI="http://doi.org/10.5281/zenodo.5344496" ID-Zenodo-Dep="5344496" httpUri="https://zenodo.org/record/5344496/files/figure.png" pageId="4" pageNumber="383" startId="4.[136,171,1148,1170]" targetBox="[133,1476,192,1129]" targetPageId="4">
<paragraph id="087F7C15FFD1FF9EFF69B40F59296E86" blockId="4.[136,1476,1148,1257]" pageId="4" pageNumber="383">
Fig. 3. Reproductive anatomy of the holotype of
<taxonomicName id="CFC00796FFD1FF9EFD8EB40F5B5F6EFF" authority="Páll-Gergely &amp; Hunyadi &amp; Hausdorf, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Hausdorf" authorityYear="2020" box="[623,780,1148,1169]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="intha" status="sp. nov.">
<emphasis id="3AB4A007FFD1FF9EFD8EB40F5B5F6EFF" box="[623,780,1148,1169]" italics="true" pageId="4" pageNumber="383">Landouria intha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="21871D7CFFD1FF9EFCFBB40E5BDC6EFC" box="[794,911,1148,1170]" pageId="4" pageNumber="383" rank="species">new species</taxonomicNameLabel>
(HNHM 104442). Abbreviations: a, atrium; ag, albumen gland; b, bursa of the bursa copulatrix; bb, base of the peduncle of bursa copulatrix; e, epiphallus; f, flagellum; p, penis; ps, penial sheath (indicated with grey shading); rm, retractor muscle; so, spermoviduct; v, vagina; vd, vas deferens. Asterisks indicate originally conjoined ends. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="087F7C15FFD1FF9EFF69B5D359AE6C19" blockId="4.[136,775,1440,1656]" pageId="4" pageNumber="383">
<taxonomicName id="CFC00796FFD1FF9EFF69B5D35A566FD6" authority="(Theobald, 1859)" baseAuthorityName="Theobald" baseAuthorityYear="1859" box="[136,517,1440,1464]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="phayrei">
<emphasis id="3AB4A007FFD1FF9EFF69B5D3596A6FD6" box="[136,313,1440,1464]" italics="true" pageId="4" pageNumber="383">Aegista phayrei</emphasis>
(
<bibRefCitation id="6C5101E4FFD1FF9EFEABB5D359AE6FD6" author="Theobald W" box="[330,509,1440,1464]" pageId="4" pageNumber="383" pagination="305 - 309" refId="ref4580" refString="Theobald W (1859) Descriptions of some new Burmese and Indian Helicidae with remarks on some previously described species. Journal of the Asiatic Society of Bengal, 28 (4): 305 - 309." type="journal article" year="1859">Theobald, 1859</bibRefCitation>
)
</taxonomicName>
(material examined: 4 shells [NHMUK 1906.2.2.274], Ava, Myaleit doung, coll. Godwin-Austen; 1 shell [NHMUK], Ava, Burmah, coll. Godwin-Austen; 6 shells [NHMUK], Ava, coll. H.F./W.F. Blanford) also has an elevated spire, much narrower umbilicus, and blunter keel, but its sculpture is roughly, irregularly covered by scale scars.
</paragraph>
<paragraph id="087F7C15FFD1FF9EFCA2B5535C3D6F96" blockId="4.[834,1474,1312,1528]" pageId="4" pageNumber="383">
The Chinese and Japanese
<taxonomicName id="CFC00796FFD1FF9EFB9AB5535D0E6F56" box="[1147,1373,1312,1336]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="3AB4A007FFD1FF9EFB9AB5535D516F56" box="[1147,1282,1312,1336]" italics="true" pageId="4" pageNumber="383">Plectotropis</emphasis>
species
</taxonomicName>
(e.g.,
<taxonomicName id="CFC00796FFD1FF9EFA4AB5525C096F36" authority="(Martens, 1881)" baseAuthorityName="Martens" baseAuthorityYear="1881" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="gerlachi">
<emphasis id="3AB4A007FFD1FF9EFA4AB5525BCE6F36" italics="true" pageId="4" pageNumber="383">P. gerlachi</emphasis>
(
<bibRefCitation id="6C5101E4FFD1FF9EFC4EB5335C016F36" author="Martens E von" box="[943,1106,1344,1368]" pageId="4" pageNumber="383" refId="ref3852" refString="Martens E von (1881) Conchologische Mittheilungen als Fortsetzung der Novitates Conchologicae. Volume 2. Theodor Fischer, Cassel, 101 pp., viii pls." type="book" year="1881">Martens, 1881</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="CFC00796FFD1FF9EFB88B5325BA76F16" authority="(Schmacker &amp; O. Boettger, 1890))" authorityName="" baseAuthorityName="Schmacker &amp; O. Boettger" baseAuthorityYear="1890" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="vulgivaga">
<emphasis id="3AB4A007FFD1FF9EFB88B5325CA76F36" box="[1129,1268,1344,1368]" italics="true" pageId="4" pageNumber="383">P. vulgivaga</emphasis>
(Schmacker &amp; O. Boettger, 1890))
</taxonomicName>
are strikingly similar to the new species in terms of shell shape and sculpture. However, the Japanese/ Taiwanese species usually have prominent, long periostracal folds on the keel, whereas the new species is uniformly scaled on its entire surface.
</paragraph>
</subSubSection>
<subSubSection id="40DA2F9EFFD1FF9EFCA2B6535D906236" pageId="4" pageNumber="383" type="etymology">
<paragraph id="087F7C15FFD1FF9EFCA2B6535CE56C16" blockId="4.[835,1475,1568,1656]" pageId="4" pageNumber="383">
<emphasis id="3AB4A007FFD1FF9EFCA2B6535B966C56" bold="true" box="[835,965,1568,1592]" pageId="4" pageNumber="383">Etymology.</emphasis>
The specific epithet refers to the Intha people inhabiting the environment of the closely situated Inle lake. It is used as a noun in apposition.
</paragraph>
<paragraph id="087F7C15FFD1FF9EFF69B6D35D916D56" blockId="4.[135,1476,1696,2136]" pageId="4" pageNumber="383">
<taxonomicName id="CFC00796FFD1FF9EFF69B6D359846CD6" authority="(Benson, 1859)" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[136,471,1696,1720]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="oldhami">
<emphasis id="3AB4A007FFD1FF9EFF69B6D359616CD6" box="[136,306,1696,1720]" italics="true" pageId="4" pageNumber="383">Aegista oldhami</emphasis>
(
<bibRefCitation id="6C5101E4FFD1FF9EFEDFB6D359836CD6" author="Benson WH" box="[318,464,1696,1720]" pageId="4" pageNumber="383" pagination="184 - 188" refId="ref3255" refString="Benson WH (1859) Descriptions of new species of Helix, Streptaxis, and Vitrina, collected by Mr. W. Theobald, jun. in Burmah, the Khasia Hills, and Hindustan. Annals and Magazine of Natural History, 3 (3): 184 - 188." type="journal article" year="1859">Benson, 1859</bibRefCitation>
)
</taxonomicName>
(photos examined:
<specimenCount id="1EC6B79CFFD1FF9EFD42B6D35B546CD6" box="[675,775,1696,1720]" pageId="4" pageNumber="383" type="generic" typeStatus="syntype">1 syntype</specimenCount>
[UMZC I_104645], Ava) is smaller and lighter in colour,
<emphasis id="3AB4A007FFD1FF9EFC07B6CC5D736CB9" bold="true" box="[998,1312,1727,1751]" pageId="4" pageNumber="383">ACKNOWLEDGEMENTS</emphasis>
has a slightly elevated spire, blunter keel, and its dorsal side is finely scaly, while the ventral side is practically smooth. We are grateful to Jonathan Ablett (NHM) for granting access to the collection of the NHM, to Harold Taylor
</paragraph>
<paragraph id="087F7C15FFD1FF9EFF69B7335D906236" blockId="4.[135,1476,1696,2136]" pageId="4" pageNumber="383">
<taxonomicName id="CFC00796FFD1FF9EFF69B73359846D36" authority="(Benson, 1836)" baseAuthorityName="Benson" baseAuthorityYear="1836" box="[136,471,1856,1880]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="tapeina">
<emphasis id="3AB4A007FFD1FF9EFF69B733597D6D39" box="[136,302,1856,1879]" italics="true" pageId="4" pageNumber="383">Aegista tapeina</emphasis>
(
<bibRefCitation id="6C5101E4FFD1FF9EFEDAB73359836D36" author="Benson WH" box="[315,464,1856,1880]" pageId="4" pageNumber="383" pagination="350 - 358" refId="ref3217" refString="Benson WH (1836) Descriptive catalogue of terrestrial and fluviatile Testacea, chiefly from the North-East frontier of Bengal. Journal of the Asiatic Society of Bengal, 5 (54): 350 - 358." type="journal article" year="1836">Benson, 1836</bibRefCitation>
)
</taxonomicName>
(material examined: 2 shells (NHM) for providing photographs of
<taxonomicName id="CFC00796FFD1FF9EFB06B7335DE86D36" authorityName="Gude" authorityYear="1914" box="[1255,1467,1856,1880]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="nutans">
<emphasis id="3AB4A007FFD1FF9EFB06B7335DE86D36" box="[1255,1467,1856,1880]" italics="true" pageId="4" pageNumber="383">Plectotropis nutans</emphasis>
</taxonomicName>
, [NHMUK], E. Frontier of Bengal;
<specimenCount id="1EC6B79CFFD1FF9EFE16B7135A6C6D16" box="[503,575,1888,1912]" pageId="4" pageNumber="383" type="adult">2 adult</specimenCount>
+
<specimenCount id="1EC6B79CFFD1FF9EFDBBB7135A906D16" box="[602,707,1888,1912]" pageId="4" pageNumber="383" type="juvenile">2 juvenile</specimenCount>
shells and to Tom White (NHM) for providing photos of
<taxonomicName id="CFC00796FFD1FF9EFA95B7135D976D19" authorityName="Albers" authorityYear="1850" box="[1396,1476,1888,1911]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="genus">
<emphasis id="3AB4A007FFD1FF9EFA95B7135D976D19" box="[1396,1476,1888,1911]" italics="true" pageId="4" pageNumber="383">Aegista</emphasis>
</taxonomicName>
[NHMUK], Theria Ghat, coll. Godwin-Austen) has a slightly
<taxonomicName id="CFC00796FFD1FF9EFCA2B7F35BC96DF6" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[835,922,1920,1944]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="383" phylum="Mollusca" rank="species" species="oldhami">
<emphasis id="3AB4A007FFD1FF9EFCA2B7F35BC96DF6" box="[835,922,1920,1944]" italics="true" pageId="4" pageNumber="383">oldhami</emphasis>
</taxonomicName>
. We thank Jennifer Lauschke (Zoological Museum, elevated spire, a narrower umbilicus and a blunter keel than
<collectingRegion id="CA04B2F7FFD1FF9EFCA2B7D35BF96DD6" box="[835,938,1952,1976]" country="Germany" name="Hamburg" pageId="4" pageNumber="383">Hamburg</collectingRegion>
) for performing DNA extraction and PCR. This the new species. Sculpture of the first sample is very finely study was supported by the MTA (Hungarian Academy of scaled, whereas that of the second sample is matte, finely Sciences) Premium Post Doctorate Research Program and the reticulated on the ventral side, and has some scales inside SYNTHESYS Project (GB-TAF-2523). We are also indebted the umbilicus and near the keel and suture. to The Biodiversity Heritage Library for the multitude of rare literature available to us (www.biodiversitylibrary.org).
</paragraph>
</subSubSection>
</treatment>
</document>
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