273 lines
33 KiB
XML
273 lines
33 KiB
XML
<document id="8B23F51DAB143DAE9CB954081E8B356E" ID-DOI="10.11646/zootaxa.3986.4.1" ID-GBIF-Dataset="00f5fe41-ca13-41a4-baaa-02c4dd6aebdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="234891" ID-ZooBank="5D6CD3DC-46B9-47F0-B518-AE29234F96F4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461264179591" checkinUser="plazi" docAuthor="Petrolli, Marina G. & Benine, Ricardo C." docDate="2015" docId="7F3387F9AF79FFC6FF1CE792507BB88E" docLanguage="en" docName="zt03986p420.pdf" docOrigin="Zootaxa 3986 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moenkhausia alesis Petrolli & Benine, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="408" masterDocId="830AFF81AF7FFFC1FF8BE5455406BD4E" masterDocTitle="Description of three new species of Moenkhausia (Teleostei, Characiformes, Characidae) with the definition of the Moenkhausia jamesi species complex" masterLastPageNumber="420" masterPageNumber="401" pageNumber="407" updateTime="1698610128910" updateUser="plazi">
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<mods:titleInfo id="A94B778EC6B243A6D5BA784C19B717A0">
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<mods:title id="ED974970F8A72BED64C0517DDBB5B490">Description of three new species of Moenkhausia (Teleostei, Characiformes, Characidae) with the definition of the Moenkhausia jamesi species complex</mods:title>
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<mods:namePart id="DF06C2E2356F68B6A159849CE2BB7C0A">Petrolli, Marina G.</mods:namePart>
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<mods:roleTerm id="0C0D1A5B0DC47F05EE1D72271E209307">Author</mods:roleTerm>
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<mods:namePart id="3981F0DFA8C95AFBEC5B23F5308F7C2E">Benine, Ricardo C.</mods:namePart>
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<mods:typeOfResource id="2E996E60B5638D589B12BADF37F8FBF2">text</mods:typeOfResource>
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<mods:title id="707567B9BB0C3B9172992FC431D081E3">Zootaxa</mods:title>
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<mods:part id="6B90422E2FC3FA64629BA000D44F1559">
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<mods:date id="AE90FAFCEDEC519343DA124329AFAFB2">2015</mods:date>
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<mods:number id="91207833A7055F18B8B36A7BB87B92A9">3986</mods:number>
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<mods:detail id="293B9C0DEA61948547EA15E786CE18BE" type="issue">
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<mods:classification id="79A49C4DA6B7D6E74F94EE321DF979B0">journal article</mods:classification>
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<mods:identifier id="A1D048F4CE9408F53EF7A22C8AB43F5E" type="DOI">10.11646/zootaxa.3986.4.1</mods:identifier>
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<mods:identifier id="4EA081462EE31E266A7F13B3FC52216C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="7F3387F9AF79FFC6FF1CE792507BB88E" ID-DOI="http://doi.org/10.5281/zenodo.5620460" ID-GBIF-Taxon="119637459" ID-Zenodo-Dep="5620460" LSID="urn:lsid:plazi:treatment:7F3387F9AF79FFC6FF1CE792507BB88E" httpUri="http://treatment.plazi.org/id/7F3387F9AF79FFC6FF1CE792507BB88E" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
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<subSubSection id="BF806564AF79FFC7FF1CE7925586BE5D" pageId="6" pageNumber="407" type="nomenclature">
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<paragraph id="F72536EFAF79FFC7FF1CE7925623BFBC" blockId="6.[151,549,727,787]" box="[151,549,727,754]" pageId="6" pageNumber="407">
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<heading id="AC6D8183AF79FFC7FF1CE7925623BFBC" bold="true" box="[151,549,727,754]" fontSize="11" level="1" pageId="6" pageNumber="407" reason="1">
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<emphasis id="C5EEEAFDAF79FFC7FF1CE7925623BFBC" bold="true" box="[151,549,727,754]" pageId="6" pageNumber="407">
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<taxonomicName id="309A4D6CAF79FFC7FF1CE7925580BFBF" box="[151,390,727,753]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="alesis" status="sp. nov.">
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<emphasis id="C5EEEAFDAF79FFC7FF1CE7925580BFBF" bold="true" box="[151,390,727,753]" italics="true" pageId="6" pageNumber="407">Moenkhausia alesis</emphasis>
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="DEDD5786AF79FFC7FE1EE79D5623BFBC" box="[405,549,728,754]" pageId="6" pageNumber="407" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="F72536EFAF79FFC7FF1CE7BF5586BE5D" blockId="6.[151,549,727,787]" box="[151,384,762,787]" pageId="6" pageNumber="407">
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(
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<figureCitation id="6FA12A6AAF79FFC7FF14E7BF54EBBE5D" box="[159,237,762,787]" captionStart="FIGURE 5" captionStartId="6.[151,250,1534,1556]" captionTargetBox="[226,1360,948,1513]" captionTargetId="figure@6.[226,1360,948,1513]" captionTargetPageId="6" captionText="FIGURE 5. Moenkhausia alesis, MZUSP 117139, holotype, 52.6 mm SL." httpUri="https://zenodo.org/record/234896/files/figure.png" pageId="6" pageNumber="407">Figs. 5</figureCitation>
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,
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<figureCitation id="6FA12A6AAF79FFC7FF71E7BF550FBE5D" box="[250,265,762,787]" captionStart="FIGURE 9" captionStartId="15.[151,250,1554,1576]" captionTargetBox="[174,1419,325,1476]" captionTargetId="figure@15.[151,1436,265,1533]" captionTargetPageId="15" captionText="FIGURE 9. Dentary of Moenkhausia jamesi species complex. (a) M. jamesi, MZUSP 49507, 54.0 mm SL; (b) M. justae, INPA 21489, 47.0 mm SL; (c) M. ischyognatha, LBP 15948, 48.2 mm SL; (d) M. alesis, INPA 48444, 50.4 mm SL; (e) M. sthenosthoma, LBP 19956, 46.0 mm SL; and (f) M. sthenosthoma, LBP 19958, 63.4 mm SL. Scale bars = 1 mm." httpUri="https://zenodo.org/record/234900/files/figure.png" pageId="6" pageNumber="407">9</figureCitation>
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d,
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<tableCitation id="BA180354AF79FFC7FEAEE7BF5573BE5D" box="[293,373,762,787]" captionStart="TABLE 1" captionStartId="4.[119,204,573,595]" captionTargetBox="[119,1855,670,1647]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data for Moenkhausia ischyognatha sp. n., Moenkhausia alesis sp. n., and Moenkhausia sthenosthoma sp. n. N includes holotype. Range includes holotype and paratypes." httpUri="http://table.plazi.org/id/A3E56667AF7BFFC5FFFCE7785561BF20" pageId="6" pageNumber="407" tableUuid="A3E56667AF7BFFC5FFFCE7785561BF20">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="BF806564AF79FFC7FF1CE60750E5BA40" pageId="6" pageNumber="407" type="materials_examined">
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<paragraph id="F72536EFAF79FFC7FF1CE6075168BE31" blockId="6.[151,1436,834,895]" pageId="6" pageNumber="407">
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<typeStatus id="2821884DAF79FFC7FF1CE6075502BE15" box="[151,260,834,859]" pageId="6" pageNumber="407" type="holotype">
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<emphasis id="C5EEEAFDAF79FFC7FF1CE6075502BE15" bold="true" box="[151,260,834,859]" pageId="6" pageNumber="407">Holotype</emphasis>
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</typeStatus>
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.
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<collectionCode id="918BAE2AAF79FFC7FE99E6065569BE15" box="[274,367,835,859]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="407">MZUSP</collectionCode>
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117139,
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<quantity id="30629B0AAF79FFC7FE5DE607563CBE14" box="[470,570,834,859]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" pageId="6" pageNumber="407" unit="mm" value="52.6">52.6 mm</quantity>
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SL, Tocantins, Conceição de Tocantins; rio Palmas at TO 0 50, near to Chuva de Manga locality,
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<geoCoordinate id="92AE5028AF79FFC7FEE6E62355E0BE31" box="[365,486,870,895]" direction="south" orientation="latitude" pageId="6" pageNumber="407" precision="15" value="-12.418612">12°25’7”S</geoCoordinate>
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,
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<geoCoordinate id="92AE5028AF79FFC7FE79E6235686BE31" box="[498,640,870,895]" direction="west" orientation="longitude" pageId="6" pageNumber="407" precision="15" value="-47.19778">47°11’52”W</geoCoordinate>
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, J.L. Birindelli; F.C. Dagosta; M.V. Loeb; C. Santos; 0
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<date id="8324102FAF79FFC7FB63E622516CBE31" box="[1256,1386,871,895]" pageId="6" pageNumber="407" value="2012-12-01">1 Dec 2012</date>
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.
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</paragraph>
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<caption id="A3E56667AF79FFC7FF1CE0BB578CBB5A" box="[151,906,1534,1556]" httpUri="https://zenodo.org/record/234896/files/figure.png" pageId="6" pageNumber="407" targetBox="[226,1360,948,1513]" targetPageId="6">
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<paragraph id="F72536EFAF79FFC7FF1CE0BB578CBB5A" blockId="6.[151,906,1534,1556]" box="[151,906,1534,1556]" pageId="6" pageNumber="407">
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<emphasis id="C5EEEAFDAF79FFC7FF1CE0BB5514BB5A" bold="true" box="[151,274,1534,1556]" pageId="6" pageNumber="407">FIGURE 5.</emphasis>
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<taxonomicName id="309A4D6CAF79FFC7FE92E0BA55E4BB5A" box="[281,482,1535,1556]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="alesis">
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<emphasis id="C5EEEAFDAF79FFC7FE92E0BA55E4BB5A" box="[281,482,1535,1556]" italics="true" pageId="6" pageNumber="407">Moenkhausia alesis</emphasis>
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</taxonomicName>
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, MZUSP 117139, holotype, 52.6 mm SL.
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</paragraph>
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</caption>
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<paragraph id="F72536EFAF79FFC7FF4CE30450E5BA40" blockId="6.[151,1437,1601,2022]" pageId="6" pageNumber="407">
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<typeStatus id="2821884DAF79FFC7FF4CE3045539BB14" box="[199,319,1601,1626]" pageId="6" pageNumber="407" type="paratype">
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<emphasis id="C5EEEAFDAF79FFC7FF4CE3045539BB14" bold="true" box="[199,319,1601,1626]" pageId="6" pageNumber="407">Paratypes</emphasis>
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</typeStatus>
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.
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<collectionCode id="918BAE2AAF79FFC7FEC6E30755ACBB14" box="[333,426,1602,1626]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="407">MZUSP</collectionCode>
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113845, 2,
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<quantity id="30629B0AAF79FFC7FDB9E30456D0BB17" box="[562,726,1601,1626]" metricMagnitude="-2" metricUnit="m" metricValue="5.035" metricValueMax="5.56" metricValueMin="4.51" pageId="6" pageNumber="407" unit="mm" value="50.35" valueMax="55.6" valueMin="45.1">45.1–55.6 mm</quantity>
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||
SL, same data as
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<typeStatus id="2821884DAF79FFC7FC2EE304500DBB14" box="[933,1035,1601,1626]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
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.
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<collectionCode id="918BAE2AAF79FFC7FB9DE3075053BB14" box="[1046,1109,1602,1626]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="6" pageNumber="407">INPA</collectionCode>
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48444, 6 (1 c&s),
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<quantity id="30629B0AAF79FFC7FAA7E30454C5BB33" metricMagnitude="-2" metricUnit="m" metricValue="4.99" metricValueMax="5.51" metricValueMin="4.47" pageId="6" pageNumber="407" unit="mm" value="49.9" valueMax="55.1" valueMin="44.7">44.7–55.1 mm</quantity>
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SL, Caseara, Parque Estadual do Cantão, E.G. Ferreira, J. A. Zuanon & G. M. Santos,
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<date id="8324102FAF79FFC7FB32E3235134BB30" box="[1209,1330,1638,1662]" pageId="6" pageNumber="407" value="2000-05">May 2000</date>
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.
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<collectionCode id="918BAE2AAF79FFC7FAB4E323519ABB30" box="[1343,1436,1638,1662]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="407">MZUSP</collectionCode>
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52354, 10 of 20,
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<quantity id="30629B0AAF79FFC7FEFBE3CC561FBBEF" box="[368,537,1673,1698]" metricMagnitude="-2" metricUnit="m" metricValue="5.58" metricValueMax="5.77" metricValueMin="5.39" pageId="6" pageNumber="407" unit="mm" value="55.8" valueMax="57.7" valueMin="53.9">53.9–57.7 mm</quantity>
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SL, Goiás; Rio Araguaia, at beach between Bandeirantes and Luis Alves;
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<geoCoordinate id="92AE5028AF79FFC7FF1CE3EB551BBB88" box="[151,285,1710,1734]" direction="south" orientation="latitude" pageId="6" pageNumber="407" precision="15" value="-13.666667">13°40’00”S</geoCoordinate>
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,
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<geoCoordinate id="92AE5028AF79FFC7FEA7E3E855BDBB88" box="[300,443,1709,1734]" direction="west" orientation="longitude" pageId="6" pageNumber="407" precision="15" value="-50.8">50°48’00”W</geoCoordinate>
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||
, Coleção Rio Araguaia, Jul/
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<date id="8324102FAF79FFC7FD74E3E85772BB88" box="[767,884,1709,1734]" pageId="6" pageNumber="407" value="1997-08">Aug 1997</date>
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||
. LBP 18905, 1,
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||
<quantity id="30629B0AAF79FFC7FBB9E3E8509EBB8B" box="[1074,1176,1709,1734]" metricMagnitude="-2" metricUnit="m" metricValue="5.84" pageId="6" pageNumber="407" unit="mm" value="58.4">58.4 mm</quantity>
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||
SL, Goiás, Aragarças, beach of rio Araguaia;
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||
<geoCoordinate id="92AE5028AF79FFC7FE2BE3945643BBA4" box="[416,581,1745,1770]" direction="south" orientation="latitude" pageId="6" pageNumber="407" precision="1" value="-15.891691">15°53'30.09"S</geoCoordinate>
|
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,
|
||
<geoCoordinate id="92AE5028AF79FFC7FDD9E394570FBBA7" box="[594,777,1745,1770]" direction="west" orientation="longitude" pageId="6" pageNumber="407" precision="1" value="-52.249916">52°14'59.70" W</geoCoordinate>
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||
, P.C. Vênere, no date.
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||
<collectionCode id="918BAE2AAF79FFC7FB9BE397506BBBA4" box="[1040,1133,1746,1770]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="407">MZUSP</collectionCode>
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||
52347, 12, 52.0–
|
||
<quantity id="30629B0AAF79FFC7FABCE394519BBBA7" box="[1335,1437,1745,1770]" metricMagnitude="-2" metricUnit="m" metricValue="5.74" pageId="6" pageNumber="407" unit="mm" value="57.4">57.4 mm</quantity>
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||
SL, Mato Grosso; Rio Araguaia, Fio Velasco,
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||
<geoCoordinate id="92AE5028AF79FFC7FD10E3B05724BA40" box="[667,802,1781,1806]" direction="south" orientation="latitude" pageId="6" pageNumber="407" precision="15" value="-12.882778">12°52’58”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="92AE5028AF79FFC7FCA5E3B057BBBA40" box="[814,957,1781,1806]" direction="west" orientation="longitude" pageId="6" pageNumber="407" precision="15" value="-50.501945">50°30’07”W</geoCoordinate>
|
||
, R. S.A. Matias,
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||
<date id="8324102FAF79FFC7FBF7E3B050D9BA40" box="[1148,1247,1781,1806]" pageId="6" pageNumber="407" value="1997-07">Jul 1997</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BF806564AF79FFC6FF4CE25C5164BDB6" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="diagnosis">
|
||
<paragraph id="F72536EFAF79FFC6FF4CE25C5164BDB6" blockId="6.[151,1437,1601,2022]" lastBlockId="7.[151,1437,151,1472]" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
|
||
<emphasis id="C5EEEAFDAF79FFC7FF4CE25C553CBA7C" bold="true" box="[199,314,1817,1842]" pageId="6" pageNumber="407">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="309A4D6CAF79FFC7FEC3E25C5623BA7C" box="[328,549,1817,1842]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="alesis">
|
||
<emphasis id="C5EEEAFDAF79FFC7FEC3E25C5623BA7C" box="[328,549,1817,1842]" italics="true" pageId="6" pageNumber="407">Moenkhausia alesis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="309A4D6CAF79FFC7FC95E25F578ABA7F" box="[798,908,1818,1841]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="jamesi">
|
||
<emphasis id="C5EEEAFDAF79FFC7FC95E25F578ABA7F" box="[798,908,1818,1841]" italics="true" pageId="6" pageNumber="407">M. jamesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="309A4D6CAF79FFC7FC11E25F5052BA7C" box="[922,1108,1817,1842]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ischyognatha">
|
||
<emphasis id="C5EEEAFDAF79FFC7FC11E25F5052BA7C" box="[922,1108,1817,1842]" italics="true" pageId="6" pageNumber="407">M. ischyognatha</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="309A4D6CAF79FFC7FB1AE25F5156BA7C" box="[1169,1360,1817,1842]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="sthenosthoma">
|
||
<emphasis id="C5EEEAFDAF79FFC7FB1AE25F5156BA7C" box="[1169,1360,1817,1842]" italics="true" pageId="6" pageNumber="407">M. sthenosthoma</emphasis>
|
||
</taxonomicName>
|
||
by the number of scale rows between lateral line and dorsal-fin origin (eight to nine
|
||
<emphasis id="C5EEEAFDAF79FFC7FB9CE27B5028BA1B" box="[1047,1070,1854,1877]" italics="true" pageId="6" pageNumber="407">vs</emphasis>
|
||
seven in the last three). Differs from
|
||
<taxonomicName id="309A4D6CAF79FFC7FF5EE2275596BA34" box="[213,400,1889,1914]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ischyognatha">
|
||
<emphasis id="C5EEEAFDAF79FFC7FF5EE22754EABA37" box="[213,236,1890,1913]" italics="true" pageId="6" pageNumber="407">M</emphasis>
|
||
.
|
||
<emphasis id="C5EEEAFDAF79FFC7FF70E2245596BA34" box="[251,400,1889,1914]" italics="true" pageId="6" pageNumber="407">ischyognatha</emphasis>
|
||
</taxonomicName>
|
||
by its greater body depth (43.5–54.6% in SL
|
||
<emphasis id="C5EEEAFDAF79FFC7FC11E22757B7BA37" box="[922,945,1890,1913]" italics="true" pageId="6" pageNumber="407">vs</emphasis>
|
||
37.0–43.0% in SL in
|
||
<taxonomicName id="309A4D6CAF79FFC7FB27E2275161BA34" box="[1196,1383,1889,1914]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ischyognatha">
|
||
<emphasis id="C5EEEAFDAF79FFC7FB27E2275161BA34" box="[1196,1383,1889,1914]" italics="true" pageId="6" pageNumber="407">M. ischyognatha</emphasis>
|
||
</taxonomicName>
|
||
), by its greater head depth (35.9–43.2% in SL vs 29.3–32.6% in SL in
|
||
<taxonomicName id="309A4D6CAF79FFC7FC27E2C3506DBAD0" box="[940,1131,1925,1950]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ischyognatha">
|
||
<emphasis id="C5EEEAFDAF79FFC7FC27E2C3506DBAD0" box="[940,1131,1925,1950]" italics="true" pageId="6" pageNumber="407">M. ischyognatha</emphasis>
|
||
</taxonomicName>
|
||
), and by its lower caudal peduncle length (3.8–7.5% in SL
|
||
<emphasis id="C5EEEAFDAF79FFC7FDAEE2EF563ABA8F" box="[549,572,1962,1985]" italics="true" pageId="6" pageNumber="407">vs</emphasis>
|
||
8.3–11.2% in SL in
|
||
<emphasis id="C5EEEAFDAF79FFC7FCB2E2EF5006BA8C" box="[825,1024,1961,1986]" italics="true" pageId="6" pageNumber="407">
|
||
<taxonomicName id="309A4D6CAF79FFC7FCB2E2EF57FEBA8C" box="[825,1016,1961,1986]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ischyognatha">M. ischyognatha</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="309A4D6CAF79FFC7FB98E2EC50F3BA8C" box="[1043,1269,1961,1986]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="alesis">
|
||
<emphasis id="C5EEEAFDAF79FFC7FB98E2EC50F3BA8C" box="[1043,1269,1961,1986]" italics="true" pageId="6" pageNumber="407">Moenkhausia alesis</emphasis>
|
||
</taxonomicName>
|
||
is additionaly distinguished from
|
||
<taxonomicName id="309A4D6CAF79FFC7FEE4E28A55DBBAAB" box="[367,477,1998,2022]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="jamesi">
|
||
<emphasis id="C5EEEAFDAF79FFC7FEE4E28A55DBBAAB" box="[367,477,1998,2022]" italics="true" pageId="6" pageNumber="407">M. jamesi</emphasis>
|
||
</taxonomicName>
|
||
by the number of scale rows between lateral line and midventral scale series (seven to eight
|
||
<emphasis id="C5EEEAFDAF78FFC6FF5DE5DC54EBBDFE" box="[214,237,153,176]" italics="true" pageId="7" pageNumber="408">vs</emphasis>
|
||
six in
|
||
<taxonomicName id="309A4D6CAF78FFC6FEB1E5DC55AFBDFE" box="[314,425,153,176]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="jamesi">
|
||
<emphasis id="C5EEEAFDAF78FFC6FEB1E5DC55AFBDFE" box="[314,425,153,176]" italics="true" pageId="7" pageNumber="408">M. jamesi</emphasis>
|
||
</taxonomicName>
|
||
) and by the number of principal dentary teeth (four
|
||
<emphasis id="C5EEEAFDAF78FFC6FC7DE5DC500BBDFE" box="[1014,1037,153,176]" italics="true" pageId="7" pageNumber="408">vs</emphasis>
|
||
five in
|
||
<taxonomicName id="309A4D6CAF78FFC6FBEEE5DC50D2BDFE" box="[1125,1236,153,176]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="jamesi">
|
||
<emphasis id="C5EEEAFDAF78FFC6FBEEE5DC50D2BDFE" box="[1125,1236,153,176]" italics="true" pageId="7" pageNumber="408">M. jamesi</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="6FA12A6AAF78FFC6FB66E5D25137BDFE" box="[1261,1329,151,176]" captionStart="FIGURE 9" captionStartId="15.[151,250,1554,1576]" captionTargetBox="[174,1419,325,1476]" captionTargetId="figure@15.[151,1436,265,1533]" captionTargetPageId="15" captionText="FIGURE 9. Dentary of Moenkhausia jamesi species complex. (a) M. jamesi, MZUSP 49507, 54.0 mm SL; (b) M. justae, INPA 21489, 47.0 mm SL; (c) M. ischyognatha, LBP 15948, 48.2 mm SL; (d) M. alesis, INPA 48444, 50.4 mm SL; (e) M. sthenosthoma, LBP 19956, 46.0 mm SL; and (f) M. sthenosthoma, LBP 19958, 63.4 mm SL. Scale bars = 1 mm." httpUri="https://zenodo.org/record/234900/files/figure.png" pageId="7" pageNumber="408">Fig. 9</figureCitation>
|
||
a, d). It is distinguished from
|
||
<taxonomicName id="309A4D6CAF78FFC6FEF8E5F855DABD9A" box="[371,476,189,212]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="justae">
|
||
<emphasis id="C5EEEAFDAF78FFC6FEF8E5F855DABD9A" box="[371,476,189,212]" italics="true" pageId="7" pageNumber="408">M. justae</emphasis>
|
||
</taxonomicName>
|
||
by the number of teeth in the maxilla (edentulous
|
||
<emphasis id="C5EEEAFDAF78FFC6FB90E5F85034BD9A" box="[1051,1074,189,212]" italics="true" pageId="7" pageNumber="408">vs</emphasis>
|
||
one tooth with five cusps in
|
||
<emphasis id="C5EEEAFDAF78FFC6FAF6E5F854ECBDB9" italics="true" pageId="7" pageNumber="408">
|
||
<taxonomicName id="309A4D6CAF78FFC6FAF6E5F854D8BDB9" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="justae">M. justae</taxonomicName>
|
||
),
|
||
</emphasis>
|
||
and by the number of cusps on the fourth dentary tooth (three to five
|
||
<emphasis id="C5EEEAFDAF78FFC6FC7CE5A45008BDB6" box="[1015,1038,225,248]" italics="true" pageId="7" pageNumber="408">vs</emphasis>
|
||
six in
|
||
<taxonomicName id="309A4D6CAF78FFC6FBD1E5A450C4BDB6" box="[1114,1218,225,248]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="justae">
|
||
<emphasis id="C5EEEAFDAF78FFC6FBD1E5A450C4BDB6" box="[1114,1218,225,248]" italics="true" pageId="7" pageNumber="408">M. justae</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="6FA12A6AAF78FFC6FB50E59A512EBDB6" box="[1243,1320,223,248]" captionStart="FIGURE 9" captionStartId="15.[151,250,1554,1576]" captionTargetBox="[174,1419,325,1476]" captionTargetId="figure@15.[151,1436,265,1533]" captionTargetPageId="15" captionText="FIGURE 9. Dentary of Moenkhausia jamesi species complex. (a) M. jamesi, MZUSP 49507, 54.0 mm SL; (b) M. justae, INPA 21489, 47.0 mm SL; (c) M. ischyognatha, LBP 15948, 48.2 mm SL; (d) M. alesis, INPA 48444, 50.4 mm SL; (e) M. sthenosthoma, LBP 19956, 46.0 mm SL; and (f) M. sthenosthoma, LBP 19958, 63.4 mm SL. Scale bars = 1 mm." httpUri="https://zenodo.org/record/234900/files/figure.png" pageId="7" pageNumber="408">Figs. 9</figureCitation>
|
||
b, d).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BF806564AF78FFC6FF4CE44154F4B87E" pageId="7" pageNumber="408" type="description">
|
||
<paragraph id="F72536EFAF78FFC6FF4CE4415190BCE3" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">
|
||
<emphasis id="C5EEEAFDAF78FFC6FF4CE4415557BC53" bold="true" box="[199,337,260,285]" pageId="7" pageNumber="408">Description</emphasis>
|
||
. Morphometric data summarized in
|
||
<tableCitation id="BA180354AF78FFC6FC8AE441575CBC52" box="[769,858,260,285]" captionStart="TABLE 1" captionStartId="4.[119,204,573,595]" captionTargetBox="[119,1855,670,1647]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data for Moenkhausia ischyognatha sp. n., Moenkhausia alesis sp. n., and Moenkhausia sthenosthoma sp. n. N includes holotype. Range includes holotype and paratypes." httpUri="http://table.plazi.org/id/A3E56667AF7BFFC5FFFCE7785561BF20" pageId="7" pageNumber="408" tableUuid="A3E56667AF7BFFC5FFFCE7785561BF20">Table 1</tableCitation>
|
||
. Largest specimen examined
|
||
<quantity id="30629B0AAF78FFC6FB33E4415127BC52" box="[1208,1313,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="5.77" pageId="7" pageNumber="408" unit="mm" value="57.7">57.7 mm</quantity>
|
||
SL. Body compressed and deep. Greatest body depth slightly anterior to dorsal-fin origin. Dorsal profile of head slightly convex; straight to slightly concave along the occipital spine; slightly convex to convex from tip of supraoccipital spine to end of dorsal-fin base; straight to slightly convex from end of dorsal fin up to end of adipose fin; caudal peduncle concave in dorsal and ventral margins; ventral profile slightly convex from tip of snout to end of anal fin.
|
||
</paragraph>
|
||
<paragraph id="F72536EFAF78FFC6FF4CE4F256EFBFE6" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">Mouth terminal. Maxilla only reaching the vertical through anterior margin, and not trespassing anterior third of second infraorbital. Premaxillary teeth in two rows. Inner row with five tetracuspidate (symphyseal) or pentacuspidate teeth with median cusp pronounced, the first two teeth from the symphysis with cusps arranged in a pronounced arch when examined from a ventral view; outer row with four to five pentacuspid teeth; edentulous maxilla. Dentary bearing four pentacuspidate teeth with central cusp longest followed by three to four distinctly small conical or tricuspidate teeth. First two or three dentary teeth from the symphysis with cusps arranged in a pronounced arch when examined from a dorsal view.</paragraph>
|
||
<paragraph id="F72536EFAF78FFC6FF4CE7F157B9BE5B" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">Dorsal-fin rays ii,9. Pectoral-fin rays i,11(2), 12*(23), 13(7). Tip of pectoral fin extends slightly beyond anterior insertion of pelvic fin. Pelvic-fin rays i,7. Tip of adpressed pelvic fin not reaching anal fin. Anal-fin rays iv,28(5), 29*(11), 30(8), 31(6), 32(2) rays; caudal fin forked with i,9,8,i.</paragraph>
|
||
<paragraph id="F72536EFAF78FFC6FF4CE65A5163BEEB" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">Scales cycloid. Lateral line with 36(11), 37*(13), 38(6), 39(1) perforated scales; Scale rows between lateral line and dorsal-fin origin 8*(30), 9(2). Scale rows between lateral line and midventral scale series 7*(23) to 8*(9). Circumpeduncular scale rows 13(1), 14*(22), 15(2), 16(2), 17(1), 18(4). Scale sheath along anal-fin base 7–15*, in 1–3* series, covering base of anteriormost rays. Small scales covering proximal two-third of caudal-fin lobes.</paragraph>
|
||
<paragraph id="F72536EFAF78FFC6FF4CE6EA55D8BEA2" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">First gill arch with 13(9), 14(9)*,15(1) gill rakers on lower limb and 9(5), 10(11)*, 11(3) on upper limb. Total vertebrae 32. Supraneurals 4.</paragraph>
|
||
<paragraph id="F72536EFAF78FFC6FF4CE6B254F4B87E" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">
|
||
<emphasis id="C5EEEAFDAF78FFC6FF4CE6B25593B95E" bold="true" box="[199,405,1015,1040]" pageId="7" pageNumber="408">Color in alcohol.</emphasis>
|
||
Overall coloration slightly silvery or pale yellow. Field of few dark chromatophores on upper lip and maxilla. Infraorbital and opercular series silvery due to the presence of guanine pigmentation. Dark chromatophores more densely concentrated along entire dorsal midline. Sparsely spread dark chromatophores dorsal of horizontal skeletogenous septum. Dark line over horizontal skeletogenous septum. Conspicuous silver midlateral stripe extending from posterior margin of opercle to base of median caudal fin-rays. Irregularly shaped, humeral mark located over fourth to seventh lateral-line scales and extending vertically over four-five horizontal scale rows above and up to one horizontal scale row below lateral line. Paired fins and anal fin hyaline. Round dark spot at the base of the caudal-fin rays formed by few chromatophores. Adipose with very few dark chromatophores (
|
||
<figureCitation id="6FA12A6AAF78FFC6FF14E05254E2B87E" box="[159,228,1303,1328]" captionStart="FIGURE 5" captionStartId="6.[151,250,1534,1556]" captionTargetBox="[226,1360,948,1513]" captionTargetId="figure@6.[226,1360,948,1513]" captionTargetPageId="6" captionText="FIGURE 5. Moenkhausia alesis, MZUSP 117139, holotype, 52.6 mm SL." httpUri="https://zenodo.org/record/234896/files/figure.png" pageId="7" pageNumber="408">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BF806564AF78FFC6FF4CE07954F4B836" pageId="7" pageNumber="408" type="distribution">
|
||
<paragraph id="F72536EFAF78FFC6FF4CE07954F4B836" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">
|
||
<emphasis id="C5EEEAFDAF78FFC6FF4CE079555FB81B" bold="true" box="[199,345,1340,1365]" pageId="7" pageNumber="408">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="309A4D6CAF78FFC6FEECE0795640B81B" box="[359,582,1340,1365]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="alesis">
|
||
<emphasis id="C5EEEAFDAF78FFC6FEECE0795640B81B" box="[359,582,1340,1365]" italics="true" pageId="7" pageNumber="408">Moenkhausia alesis</emphasis>
|
||
</taxonomicName>
|
||
occurs in the upper and middle portions of Rio Araguaia and Rio Tocantins (
|
||
<figureCitation id="6FA12A6AAF78FFC6FF14E01A54E3B836" box="[159,229,1375,1400]" captionStart="FIGURE 4" captionStartId="5.[151,250,1096,1118]" captionTargetBox="[161,1428,203,1066]" captionTargetId="figure@5.[151,1436,195,1074]" captionTargetPageId="5" captionText="FIGURE 4. Partial map of South America indicating the distribuition of Moenkhausia jamesi species complex and M. justae. Red circle = M. jamesi; Blue circle = M. ischyognatha; Orange circle = M. alesis; Light blue circle = M. sthenosthoma; Yellow circle = M. justae." httpUri="https://zenodo.org/record/234895/files/figure.png" pageId="7" pageNumber="408">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BF806564AF78FFC6FF4CE0C1507BB88E" pageId="7" pageNumber="408" type="etymology">
|
||
<paragraph id="F72536EFAF78FFC6FF4CE0C1507BB88E" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="408">
|
||
<emphasis id="C5EEEAFDAF78FFC6FF4CE0C15548B8D5" bold="true" box="[199,334,1412,1437]" pageId="7" pageNumber="408">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="309A4D6CAF78FFC6FDAFE0C15662B8D3" box="[548,612,1412,1437]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="alesis">
|
||
<emphasis id="C5EEEAFDAF78FFC6FDAFE0C15662B8D3" box="[548,612,1412,1437]" italics="true" pageId="7" pageNumber="408">alesis</emphasis>
|
||
</taxonomicName>
|
||
is from the Greek meaning grinder, in reference to the robust teeth of the premaxilla and dentary, a diagnostic feature of the
|
||
<taxonomicName id="309A4D6CAF78FFC6FD44E0E257B0B88E" box="[719,950,1447,1472]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="jamesi">
|
||
<emphasis id="C5EEEAFDAF78FFC6FD44E0E257B0B88E" box="[719,950,1447,1472]" italics="true" pageId="7" pageNumber="408">Moenkhausia jamesi</emphasis>
|
||
</taxonomicName>
|
||
species complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |