248 lines
30 KiB
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248 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e85571" ID-Pensoft-Pub="1864-8312-81-409" ID-Pensoft-UUID="1F7030AA7B0B54EAB1846C6B1339DB56" ID-ZooBank="764845DB2D5D4E7E9E12DCA816F5BCD8" ModsDocID="1864-8312-81-409" checkinTime="1682026883038" checkinUser="pensoft" docAuthor="Garcia, Andres F. & Villarreal, Osvaldo" docDate="2023" docId="7F1C8F4013E4560287118EDEBC27B8CF" docLanguage="en" docName="ArthropodSystPhyl 81: 409-437" docOrigin="Arthropod Systematics & amp; Phylogeny 81" docPubDate="2023-04-20" docSource="http://dx.doi.org/10.3897/asp.81.e85571" docTitle="Paravima totoro García & Villarreal 2023, sp. nov." docType="treatment" docUuid="06A646EB-4735-4078-B3A5-3A36A3ED964C" docUuidSource="ZooBank" docVersion="1" id="1F7030AA7B0B54EAB1846C6B1339DB56" lastPageNumber="409" masterDocId="1F7030AA7B0B54EAB1846C6B1339DB56" masterDocTitle="Looks are deceiving: a cladistic analysis, three new species, and a new diagnosis of Paravima Caporiacco, 1951 (Opiliones: Agoristenidae)" masterLastPageNumber="437" masterPageNumber="409" pageNumber="409" updateTime="1682026883038" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Looks are deceiving: a cladistic analysis, three new species, and a new diagnosis of Paravima Caporiacco, 1951 (Opiliones: Agoristenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Garcia, Andres F.</mods:namePart>
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<mods:affiliation>Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, Sao Cristovao, 20.940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">agarciarinc@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Villarreal, Osvaldo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5355-3723</mods:nameIdentifier>
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<mods:affiliation>Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, Sao Cristovao, 20.940 - 040, Rio de Janeiro, RJ, Brazil & Museo del Instituto de Zoologia Agricola, Facultad de Agronomia, Universidad Central de Venezuela, Maracay, Aragua, Venezuela</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>81</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>409</mods:start>
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<mods:end>437</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e85571</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e85571</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-81-409</mods:identifier>
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<mods:identifier type="ZooBank">764845DB2D5D4E7E9E12DCA816F5BCD8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1F7030AA7B0B54EAB1846C6B1339DB56</mods:identifier>
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<subSection pageId="0" pageNumber="409" type="3. results">
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<treatment LSID="urn:lsid:zoobank.org:act:06A646EB-4735-4078-B3A5-3A36A3ED964C" httpUri="http://treatment.plazi.org/id/7F1C8F4013E4560287118EDEBC27B8CF" lastPageNumber="409" pageId="0" pageNumber="409">
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<subSubSection pageId="0" pageNumber="409" type="nomenclature">
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<paragraph pageId="0" pageNumber="409">
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<taxonomicName LSID="https://zoobank.org/06A646EB-4735-4078-B3A5-3A36A3ED964C" authority="García & Villarreal, 2023" authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima totoro" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="totoro" status="sp. nov.">Paravima totoro</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="409">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="description">
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<paragraph pageId="0" pageNumber="409">
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">Figs 14</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">, 15</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Map showing the geographic distribution of Paravima in Northern South America. The background colored shapes refer to the biogeographic provinces of Morrone et al. (2022)." figureDoi="10.3897/asp.81.e85571.figure18" httpUri="https://binary.pensoft.net/fig/840450" pageId="0" pageNumber="409">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="diagnosis">
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<paragraph pageId="0" pageNumber="409">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="409">
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Mesotergal area III with paired dome-shaped tubercles (the remaining
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<taxonomicName authorityName="Caporiacco" authorityYear="1951" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="409">Paravima</emphasis>
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</taxonomicName>
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varies between acute spines, conical/mammilliform tubercles, or lack of tubercles). DS variegated, with mesotergal areas II-III darker in the center and gradually fading towards the laterals (in
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<taxonomicName lsidName="P. magistri" pageId="0" pageNumber="409" rank="species" species="magistri">
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<emphasis italics="true" pageId="0" pageNumber="409">P. magistri</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="409">sp. nov.</emphasis>
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variegated in the anterior region of DS; in the other
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<taxonomicName authorityName="Caporiacco" authorityYear="1951" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="409">Paravima</emphasis>
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</taxonomicName>
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species uniformly variegated).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="description">
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<paragraph pageId="0" pageNumber="409">Description.</paragraph>
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<paragraph pageId="0" pageNumber="409">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="409">Holotype</emphasis>
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(MNRJ 9255*). Measurements: CL: 1.1, AL: 1.5, CW: 2.0, AW: 2.3, BaCh: 0.3, IOD: 0.9; Leg I (Tr: 0.3, Fe: 3.2, Pa: 0.3, Ti: 2.0, Mt: 4.5, Ta: 1.6); Leg II (Tr: 0.3, Fe: 8.2, Pa: 0.7, Ti: 6.0, Mt: 10.1, Ta: 3.4); Leg III (Tr: 0.5, Fe: 6.7, Pa: 1.0, Ti: 3.1, Mt: 7.0, Ta: 1.7); Leg IV (Tr: 0.6, Fe: 10.8, Pa: 1.1, Ti: 5.0, Mt: 10.0, Ta: 2.7). -
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<emphasis italics="true" pageId="0" pageNumber="409">Dorsum</emphasis>
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: Anterior and lateral margins of DS smooth. Ocularium smooth (Figs
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14C, E</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15A</figureCitation>
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). Mesotergum delimited, divided into four areas. Area I divided into two halves, each one with one tubercle; area II-IV undivided; II invading I, and with a pair of small tubercles close to the medial axis of the body (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15A</figureCitation>
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); III with a pair of high paramedian domed tubercles (Figs
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14C, E</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15B</figureCitation>
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); IV with four small tubercles. Posterior border of scutum straight, with a row of small tubercles. Free tergites and anal operculum with some tubercles (Figs
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14D</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15A</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Venter</emphasis>
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: Coxae I-IV with some granules (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14D</figureCitation>
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). Coxa I with one medial tubercle on the anterior margin, a group of five tubercles on the proximal region, three large tubercles reaching the anterodistal margin, and two tubercles close to the posterodistal region (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15C</figureCitation>
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); coxa II longer than coxa I; coxa III longer than I and II; coxa IV backward projected (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14D</figureCitation>
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). Sternites and anal operculum with a few small tubercles. Stigmatic area smooth. Stigmata large, oval and transverse (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14D</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Chelicerae</emphasis>
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: Segment I rectangular, with well-marked bulla (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15A, B</figureCitation>
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), one small ectal subdistal tubercle, and two tubercles on the proximal border (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14B</figureCitation>
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). Chelicera swollen (Figs
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14C, E</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15B</figureCitation>
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). Hand with one tubercle at the mesal subapical region, a group of nine transversal tubercles going from ectal to mesal region, and a group of setiferous tubercles close to the base of the fingers (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15D</figureCitation>
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). Fixed finger with some teeth on the inner surface. Movable finger with one trapezoid, sub-basal tooth and with the inner surface dentate (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Phylogenetic relationship of Leiosteninae based on cladistic analyses using morphological data (in the left tree the terminals of Globibuninae and Agoristeninae are not shown; Globibuninae (Rivetinus) and Agoristeninae branches are condensed; the right tree is a continuation of the left tree). Sensitivity plots (' Navajo rugs') indicate the recovery of the nodes in the analyses under implied weights with K values of 1, 2, 4 and 8. Clear circles represent homoplastic synapomorphies and filled circles represent non-homoplastic synapomorphies." figureDoi="10.3897/asp.81.e85571.figure16" httpUri="https://binary.pensoft.net/fig/840448" pageId="0" pageNumber="409">16D</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Pedipalps</emphasis>
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: Longer than DS length, smooth. Tr ventrally with one subapical setiferous tubercle. Fe with a ventromesal row of five setiferous tubercles (the basalmost larger than the distalmost), and one large ventroectal setiferous tubercle in the distal portion (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14C, D</figureCitation>
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). Pa with one large mesal setiferous tubercle. Ti ectal iII, mesal IIi. Ta ectal IIi, mesal IIi (Figs
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14B, D</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Legs</emphasis>
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: Legs I-IV smooth. Leg I filiform, the rest, getting steadily thicker from leg II to IV (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. Habitus: A Panoramic view; B dorsal view; C lateral view; D ventral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409">14A</figureCitation>
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). Fe IV four times DS length. Tarsal counts: 6(3)-6(3)/?-14(3)/7-7/7-7. -
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<emphasis italics="true" pageId="0" pageNumber="409">Penis</emphasis>
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: LP short and depressed, half-moon shaped, with anterolateral corners dorsally pointed (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15E-G</figureCitation>
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). Malleus carrying three pairs of branched MS-A and one pair of branched MS-B (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15G</figureCitation>
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). MS-D1-2 long, located in a vertical line on a keel between the LP and the base of the stylus (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15E, G</figureCitation>
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). MS-E2 large and branched (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15F, G</figureCitation>
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). Stylus sinuous, surpassing the lamina parva, narrower at distal region, tip irregular (Figs
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Paravima totoro sp. nov. (MNRJ 9255 *) male holotype. A Habitus, dorsal view; B habitus, lateral view; C left coxa I, ventral view; D left chelicera, frontal view. Apical part of the penis in E dorsal view, F ventral view, and G lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409">15E-G</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Color</emphasis>
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(in ethanol): Carapace reticulated Dark Brown (59) on Deep Orange Yellow (51). Mesotergum, posterior border and free tergites Dark Brown (59). Pedipalps, chelicerae and legs Vivid Orange Yellow (66). -
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<emphasis italics="true" pageId="0" pageNumber="409">Female</emphasis>
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: Unknown.
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</paragraph>
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|
<caption doi="10.3897/asp.81.e85571.figure14" httpUri="https://binary.pensoft.net/fig/840446" pageId="0" pageNumber="409" start="Figure 14" startId="F14">
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<paragraph pageId="0" pageNumber="409">
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<emphasis bold="true" pageId="0" pageNumber="409">Figure 14.</emphasis>
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<taxonomicName authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima totoro" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="totoro">
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<emphasis italics="true" pageId="0" pageNumber="409">Paravima totoro</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="409">sp. nov.</emphasis>
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(MNRJ 9255*) male holotype. Habitus:
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<emphasis bold="true" pageId="0" pageNumber="409">A</emphasis>
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Panoramic view;
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<emphasis bold="true" pageId="0" pageNumber="409">B</emphasis>
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dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="409">C</emphasis>
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lateral view;
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<emphasis bold="true" pageId="0" pageNumber="409">D</emphasis>
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ventral view;
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<emphasis bold="true" pageId="0" pageNumber="409">E</emphasis>
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anterior view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.81.e85571.figure15" httpUri="https://binary.pensoft.net/fig/840447" pageId="0" pageNumber="409" start="Figure 15" startId="F15">
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|
<paragraph pageId="0" pageNumber="409">
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<emphasis bold="true" pageId="0" pageNumber="409">Figure 15.</emphasis>
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<taxonomicName authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima totoro" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="totoro">
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<emphasis italics="true" pageId="0" pageNumber="409">Paravima totoro</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="409">sp. nov.</emphasis>
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(MNRJ 9255*) male holotype.
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<emphasis bold="true" pageId="0" pageNumber="409">A</emphasis>
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Habitus, dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="409">B</emphasis>
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habitus, lateral view;
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<emphasis bold="true" pageId="0" pageNumber="409">C</emphasis>
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|
left coxa I, ventral view;
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<emphasis bold="true" pageId="0" pageNumber="409">D</emphasis>
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|
left chelicera, frontal view. Apical part of the penis in
|
|
<emphasis bold="true" pageId="0" pageNumber="409">E</emphasis>
|
|
dorsal view,
|
|
<emphasis bold="true" pageId="0" pageNumber="409">F</emphasis>
|
|
ventral view, and
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|
<emphasis bold="true" pageId="0" pageNumber="409">G</emphasis>
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|
lateral view. Scale bars: A, B = 1 mm; C, D = 0,5 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="etymology">
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<paragraph pageId="0" pageNumber="409">Derivatio nominis.</paragraph>
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|
<paragraph pageId="0" pageNumber="409">Totoro is a character in the Japanese animatedfantasy film My Neighbor Totoro (directed by Hayao Miyazaki and animated by Studio Ghibli), being a friendly wood spirit in post-war rural Japan. For us, the paramedian armature of the new species resembles the ears of the charismatic Totoro. We take advantage of exalting the excellent work of Studio Ghibli with this tribute. Noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="distribution">
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|
<paragraph pageId="0" pageNumber="409">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="409">
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|
Known just from the type locality, PN Henri Pittier, Aragua state, in the Venezuelan biogeographic province (
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|
<bibRefCitation DOI="https://doi.org/10.1590/0001-3765202220211167" author="Morrone, JJ" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="409" refId="B34" refString="Morrone, JJ, Escalante, T, Rodriguez-Tapia, G, Carmona, A, Arana, M, Mercado-Gomez, JD, 2022. Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias 94: e20211167. https://doi.org/10.1590/0001-3765202220211167" title="Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias 94: e 20211167." url="https://doi.org/10.1590/0001-3765202220211167" year="2022">Morrone et al. 2022</bibRefCitation>
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|
) (Fig.
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|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Map showing the geographic distribution of Paravima in Northern South America. The background colored shapes refer to the biogeographic provinces of Morrone et al. (2022)." figureDoi="10.3897/asp.81.e85571.figure18" httpUri="https://binary.pensoft.net/fig/840450" pageId="0" pageNumber="409">18</figureCitation>
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|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="409" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="409">Material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="409">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="409">Type material</emphasis>
|
|
:
|
|
<materialsCitation collectingDate="1983-03-31" collectorName="C. Bordon" country="VENEZUELA" elevation="1200" location="Rancho Grande" municipality="Parque Nacional Henri Pittier" specimenCount="♂" stateProvince="Aragua" typeStatus="Holotype">
|
|
<typeStatus>Holotype</typeStatus>
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|
<specimenCount>♂</specimenCount>
|
|
,
|
|
<collectingCountry name="Venezuela">VENEZUELA</collectingCountry>
|
|
,
|
|
<collectingRegion country="Venezuela" name="Aragua">Aragua</collectingRegion>
|
|
,
|
|
<collectingMunicipality>Parque Nacional Henri Pittier</collectingMunicipality>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:7F1C8F4013E4560287118EDEBC27B8CF:BDCE5B20C5A0DEBA8A8D9DED4982241C" country="VENEZUELA" municipality="Parque Nacional Henri Pittier" name="Rancho Grande" stateProvince="Aragua">Rancho Grande</location>
|
|
| [10.34947° -67.6843°] |
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
|
|
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
|
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</quantity>
|
|
|
|
|
<collectingDate value="1983-03-31">31.iii.1983</collectingDate>
|
|
,
|
|
<collectorName>
|
|
C.
|
|
<normalizedToken originalValue="Bordón">Bordon</normalizedToken>
|
|
</collectorName>
|
|
leg. (MNRJ 9255*).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</subSection>
|
|
</document> |