1216 lines
131 KiB
XML
1216 lines
131 KiB
XML
<document id="6089BE91DE3F0708ED4C3FBB1743271E" ID-CLB-Dataset="29253" ID-DOI="10.11646/zootaxa.4496.1.31" ID-GBIF-Dataset="1ac9832a-bf80-404d-8103-e4379206ade4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446937" ID-ZooBank="9E12F8F2-4FAF-4DE6-98F1-F63D3FE2AA60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747957001" checkinUser="plazi" docAuthor="Fragoso, Carlos & Rojas, Patricia" docDate="2018" docId="7F0C87C1FFFE8E6EFF0FFDD412E3FD06" docLanguage="en" docName="zootaxa.4496.1.31.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lavellodrilus sheylae Fragoso & Rojas 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="428" masterDocId="8335FFB9FFF98E60FF98FFC91606FFAE" masterDocTitle="New Diplotrema and Lavellodrilus earthworm species from southern Mexico (Annelida, Crassiclitellata, Acanthodrilidae)" masterLastPageNumber="430" masterPageNumber="414" pageNumber="421" updateTime="1698566828741" updateUser="ExternalLinkService">
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<mods:titleInfo id="C44CADB5F96C57C6DD4D1F5DAE19817A">
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<mods:title id="330F6EEE7DABD1BBC1F09E3B03B2BCA8">New Diplotrema and Lavellodrilus earthworm species from southern Mexico (Annelida, Crassiclitellata, Acanthodrilidae)</mods:title>
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<mods:name id="E6C3D0FCF1683F2BEEFCC789891DF2CE" type="personal">
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<mods:namePart id="6852AF9F650C081EF183A7D81BBB200C">Fragoso, Carlos</mods:namePart>
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<mods:name id="7DF8C358557206FA68E94DA7100B1A83" type="personal">
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<mods:namePart id="B4E37903426DF1B0C0F944A708CF37CF">Rojas, Patricia</mods:namePart>
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<mods:typeOfResource id="AA8A45210AC354767399111AB56EFA86">text</mods:typeOfResource>
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<mods:title id="F2B691996F333AEF63F387429B761F59">Zootaxa</mods:title>
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<mods:part id="3BCAB978770FCF5E29F245D4D05C252F">
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<mods:date id="16BE3DCB83D298DC4F89ECFF30831CC2">2018</mods:date>
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<mods:number id="9B80304AE1753C74AB42CE8A8C087AD6">2018-10-05</mods:number>
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<mods:number id="A35CE52B3651BECE9BA3D0528390AAFC">4496</mods:number>
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<mods:detail id="FCE228647D8F2262B085DA875767B46F" type="issue">
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<mods:number id="F9E6A747FA32E5ABDEB3FCA8B2881B0C">1</mods:number>
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<mods:classification id="379D27EFF88CE88443DA570E337FF22D">journal article</mods:classification>
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<mods:identifier id="6BE9C83F56B19251D7760308A31BE97D" type="DOI">10.11646/zootaxa.4496.1.31</mods:identifier>
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<treatment id="7F0C87C1FFFE8E6EFF0FFDD412E3FD06" ID-DOI="http://doi.org/10.5281/zenodo.5950038" ID-GBIF-Taxon="148639960" ID-Zenodo-Dep="5950038" LSID="urn:lsid:plazi:treatment:7F0C87C1FFFE8E6EFF0FFDD412E3FD06" httpUri="http://treatment.plazi.org/id/7F0C87C1FFFE8E6EFF0FFDD412E3FD06" lastPageId="14" lastPageNumber="428" pageId="7" pageNumber="421">
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<subSubSection id="BFBF655CFFFE8E67FF0FFDD4174CFDF6" pageId="7" pageNumber="421" type="nomenclature">
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<paragraph id="F71A36D7FFFE8E67FF0FFDD417FAFD99" blockId="7.[151,508,541,600]" box="[151,508,541,567]" pageId="7" pageNumber="421">
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<heading id="AC5281BBFFFE8E67FF0FFDD417FAFD99" bold="true" box="[151,508,541,567]" fontSize="11" level="1" pageId="7" pageNumber="421" reason="1">
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<emphasis id="C5D1EAC5FFFE8E67FF0FFDD417FAFD99" bold="true" box="[151,508,541,567]" pageId="7" pageNumber="421">
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||
<taxonomicName id="30A54D54FFFE8E67FF0FFDD4179EFD99" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[151,408,541,567]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="421" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFFE8E67FF0FFDD4179EFD99" bold="true" box="[151,408,541,567]" italics="true" pageId="7" pageNumber="421">Lavellodrilus sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFFE8E67FE06FDD417FAFD99" box="[414,508,541,567]" pageId="7" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFFE8E67FF0FFDF6174CFDF6" blockId="7.[151,508,541,600]" box="[151,330,575,600]" pageId="7" pageNumber="421">
|
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(
|
||
<figureCitation id="6F9E2A52FFFE8E67FF06FDF6170CFDF6" box="[158,266,575,600]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">FigurES 3</figureCitation>
|
||
,
|
||
<figureCitation id="6F9E2A52FFFE8E67FE8FFD891721FDF6" box="[279,295,576,600]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="7" pageNumber="421">4</figureCitation>
|
||
,
|
||
<figureCitation id="6F9E2A52FFFE8E67FEABFDF61747FDF6" box="[307,321,575,600]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">5</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFBF655CFFFE8E67FF0FFD4E12F9FBDB" pageId="7" pageNumber="421" type="materials_examined">
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<paragraph id="F71A36D7FFFE8E67FF0FFD4E151EFC6E" blockId="7.[151,1437,647,2005]" pageId="7" pageNumber="421">
|
||
<emphasis id="C5D1EAC5FFFE8E67FF0FFD4E17B7FD0E" bold="true" box="[151,433,647,672]" pageId="7" pageNumber="421">Localities and material.</emphasis>
|
||
<collectingRegion id="3561F835FFFE8E67FE20FD4E1417FD0E" box="[440,529,647,672]" country="Mexico" name="Mexico" pageId="7" pageNumber="421">
|
||
<collectingCountry id="8FB27647FFFE8E67FE20FD4E1417FD0E" box="[440,529,647,672]" name="Mexico" pageId="7" pageNumber="421">MExico</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingRegion id="3561F835FFFE8E67FD84FD4E147AFD0E" box="[540,636,647,672]" country="Mexico" name="Tabasco" pageId="7" pageNumber="421">TabaSco</collectingRegion>
|
||
:
|
||
<emphasis id="C5D1EAC5FFFE8E67FD1EFD4E1492FD0E" bold="true" box="[646,660,647,672]" pageId="7" pageNumber="421">1</emphasis>
|
||
) RanchEria of San JoSE PuyacatEngo, municipality of TEapa, km 7 of road TEapa-VicEntE
|
||
<collectingRegion id="3561F835FFFE8E67FEE0FD6517E6FD6A" box="[376,480,684,708]" country="Mexico" name="Guerrero" pageId="7" pageNumber="421">GuErrEro</collectingRegion>
|
||
, foothillS of CErro Madrigal, 50 yr traditional Cocoa plantation without intEnSivE uSE of agrochEmicalS, cloSE to rivEr PuyacatEngo at
|
||
<quantity id="305D9B32FFFE8E67FD5FFD19152DFD49" box="[711,811,720,744]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="7" pageNumber="421" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
|
||
dEpth,
|
||
<geoCoordinate id="92915010FFFE8E67FC1DFD061228FD46" box="[901,1070,719,744]" direction="north" orientation="latitude" pageId="7" pageNumber="421" precision="1" value="17.526918">17°31'36.91''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="92915010FFFE8E67FBA4FD0612EEFD46" box="[1084,1256,719,744]" direction="west" orientation="longitude" pageId="7" pageNumber="421" precision="1" value="-92.92895">92°55'44.23''W</geoCoordinate>
|
||
,
|
||
<quantity id="305D9B32FFFE8E67FB6FFD191335FD49" box="[1271,1331,720,744]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="421" unit="m" value="80.0">80 m</quantity>
|
||
aSl, four clitEllatE adultS
|
||
<date id="831B1017FFFE8E67FED2FD3D17CBFCA3" box="[330,461,756,781]" pageId="7" pageNumber="421" value="2002-06-30">06/30/2002</date>
|
||
, ShEyla UribE;
|
||
<emphasis id="C5D1EAC5FFFE8E67FDE4FD3D1495FCA3" bold="true" box="[636,659,756,781]" pageId="7" pageNumber="421">2)</emphasis>
|
||
RanchEria of San JoSE PuyacatEngo, municipality of TEapa, km 7 of road TEapa-VicEntE
|
||
<collectingRegion id="3561F835FFFE8E67FE12FCD117F4FC9E" box="[394,498,792,816]" country="Mexico" name="Guerrero" pageId="7" pageNumber="421">GuErrEro</collectingRegion>
|
||
, high landS of CErro Madrigal, diSturbEd tropical forESt at
|
||
<quantity id="305D9B32FFFE8E67FB44FCD11340FC81" box="[1244,1350,792,816]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="7" pageNumber="421" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
|
||
dEpth,
|
||
<geoCoordinate id="92915010FFFE8E67FF0FFCF51739FCFA" box="[151,319,828,852]" direction="north" orientation="latitude" pageId="7" pageNumber="421" precision="1" value="17.525892">17°31'33.21''N</geoCoordinate>
|
||
, and
|
||
<geoCoordinate id="92915010FFFE8E67FE1CFCF51436FCFB" box="[388,560,828,853]" direction="west" orientation="longitude" pageId="7" pageNumber="421" precision="1" value="-92.925125">92°55'30.47''W</geoCoordinate>
|
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,
|
||
<quantity id="305D9B32FFFE8E67FDD8FCF5148AFCFA" box="[576,652,828,852]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="421" unit="m" value="240.0">240 m</quantity>
|
||
aSl, onE clitEllatE adult
|
||
<date id="831B1017FFFE8E67FC37FCF51236FCFB" box="[943,1072,828,853]" pageId="7" pageNumber="421" value="2002-06-11">11/06/2002</date>
|
||
, ShEyla UribE;
|
||
<emphasis id="C5D1EAC5FFFE8E67FB74FCF51302FCFB" bold="true" box="[1260,1284,828,853]" pageId="7" pageNumber="421">3)</emphasis>
|
||
Comalcalco, Municipality of Comalcalco,
|
||
<quantity id="305D9B32FFFE8E67FE7BFCA91434FCD6" box="[483,562,863,888]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="421" unit="km" value="1.2">1.2 km</quantity>
|
||
South of town, on thE road to JoSE Maria Pino SuarEz, Cocoa plantation with intEnSivE uSE of agrochEmicalS for morE than 10 yEarS at
|
||
<quantity id="305D9B32FFFE8E67FCBAFC4D1582FC32" box="[802,900,900,924]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="7" pageNumber="421" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
|
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dEpth,
|
||
<geoCoordinate id="92915010FFFE8E67FC42FC4D1272FC33" box="[986,1140,900,925]" direction="north" orientation="latitude" pageId="7" pageNumber="421" precision="1" value="18.226612">18°13'35.8''N</geoCoordinate>
|
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,
|
||
<geoCoordinate id="92915010FFFE8E67FB19FC4D1321FC32" box="[1153,1319,900,924]" direction="west" orientation="longitude" pageId="7" pageNumber="421" precision="1" value="-93.22586">93°13' 33.1''W</geoCoordinate>
|
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,
|
||
<quantity id="305D9B32FFFE8E67FAACFC4D1368FC32" box="[1332,1390,900,924]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="421" unit="m" value="13.0">13 m</quantity>
|
||
aSl, thrEE clitEllatE adultS,
|
||
<date id="831B1017FFFE8E67FE16FC611416FC6E" box="[398,528,936,960]" pageId="7" pageNumber="421" value="2004-01-30">01/30/2004</date>
|
||
, ShEyla UribE. (
|
||
<figureCitation id="6F9E2A52FFFE8E67FD5DFC6E150FFC6E" box="[709,777,935,960]" captionStart="FIGURE 6" captionStartId="14.[151,250,1463,1485]" captionTargetBox="[309,1278,738,1379]" captionTargetId="figure@14.[309,1278,735,1441]" captionTargetPageId="14" captionText="FIGURE 6. Geographic distribution of Diplotrema chajulensis sp. nov. and Lavellodrilus sheylae sp. nov." httpUri="https://zenodo.org/record/1446949/files/figure.png" pageId="7" pageNumber="421">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFFE8E67FF5FFC051762FBA6" blockId="7.[151,1437,647,2005]" pageId="7" pageNumber="421">
|
||
<emphasis id="C5D1EAC5FFFE8E67FF5FFC05173DFC4B" bold="true" box="[199,315,972,997]" pageId="7" pageNumber="421">Holotype.</emphasis>
|
||
EntirE clitEllatE adult with gEnital markS (GM) and pEnial SEtaE (PS) collEctEd in locality
|
||
<emphasis id="C5D1EAC5FFFE8E67FAAFFC051343FC4B" bold="true" box="[1335,1349,972,997]" pageId="7" pageNumber="421">1</emphasis>
|
||
,
|
||
<date id="831B1017FFFE8E67FACCFC0516D2FBA6" pageId="7" pageNumber="421" value="2002-06-30">06/30/ 2002</date>
|
||
: IEOL 4831.
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFFE8E67FF5FFBDD12F9FBDB" blockId="7.[151,1437,647,2005]" pageId="7" pageNumber="421">
|
||
<emphasis id="C5D1EAC5FFFE8E67FF5FFBDD1740FB83" bold="true" box="[199,326,1044,1069]" pageId="7" pageNumber="421">Paratypes.</emphasis>
|
||
ThrEE EntirE clitEllatE adultS from locality
|
||
<emphasis id="C5D1EAC5FFFE8E67FCBCFBDD1534FB83" bold="true" box="[804,818,1044,1069]" pageId="7" pageNumber="421">1</emphasis>
|
||
:
|
||
<date id="831B1017FFFE8E67FCD8FBDD15C2FB83" box="[832,964,1044,1069]" pageId="7" pageNumber="421" value="2002-06-30">06/30/2002</date>
|
||
: IEOL 4830, IEOL 4832, IEOL 4837; onE EntirE clitEllatE adult from locality
|
||
<emphasis id="C5D1EAC5FFFE8E67FDB0FBFE1430FBFE" bold="true" box="[552,566,1079,1104]" pageId="7" pageNumber="421">2</emphasis>
|
||
:
|
||
<date id="831B1017FFFE8E67FDDFFBFE14CCFBFE" box="[583,714,1079,1104]" pageId="7" pageNumber="421" value="2002-06-11">11/06/2002</date>
|
||
: IEOL 4832; thrEE clitEllatE adultS from locality
|
||
<emphasis id="C5D1EAC5FFFE8E67FB65FBFE130DFBFE" bold="true" box="[1277,1291,1079,1104]" pageId="7" pageNumber="421">3</emphasis>
|
||
<date id="831B1017FFFE8E67FA8CFBF1139EFBFE" box="[1300,1432,1080,1104]" pageId="7" pageNumber="421" value="2004-01-30">01/30/2004</date>
|
||
: onE cut in two piEcES, IEOL 4833, and two almoSt cut at clitEllum lEvEl, IEOL 4834 and IEOL 4835.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFBF655CFFFE8E6DFF5FFBB615DAFA6F" lastPageId="13" lastPageNumber="427" pageId="7" pageNumber="421" type="description">
|
||
<paragraph id="F71A36D7FFFE8E67FF5FFBB6179EF87A" blockId="7.[151,1437,647,2005]" pageId="7" pageNumber="421">
|
||
<emphasis id="C5D1EAC5FFFE8E67FF5FFBB61757FB36" bold="true" box="[199,337,1151,1176]" pageId="7" pageNumber="421">Description</emphasis>
|
||
.
|
||
<emphasis id="C5D1EAC5FFFE8E67FEC7FBB617C9FB36" bold="true" box="[351,463,1151,1176]" pageId="7" pageNumber="421">External.</emphasis>
|
||
LEngth
|
||
<quantity id="305D9B32FFFE8E67FDB3FBB614BFFB39" box="[555,697,1151,1176]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" metricValueMax="3.0" metricValueMin="1.76" pageId="7" pageNumber="421" unit="mm" value="23.8" valueMax="30.0" valueMin="17.6">17.6–30 mm</quantity>
|
||
(avEragE 23.5, n=8),
|
||
<typeStatus id="281E8875FFFE8E67FC31FBB6120DFB36" box="[937,1035,1151,1176]" pageId="7" pageNumber="421" type="holotype">holotypE</typeStatus>
|
||
<quantity id="305D9B32FFFE8E67FB8AFBB61262FB39" box="[1042,1124,1151,1176]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="421" unit="mm" value="25.0">25 mm</quantity>
|
||
. Width, middlE body:
|
||
<quantity id="305D9B32FFFE8E67FAC5FB4916F7FB12" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="11.0" metricValueMin="8.8" pageId="7" pageNumber="421" unit="mm" value="0.99" valueMax="1.1" valueMin="0.88">0.88– 1.1 mm</quantity>
|
||
(avEragE= 1, n= 8);
|
||
<typeStatus id="281E8875FFFE8E67FE74FB6D1448FB13" box="[492,590,1188,1213]" pageId="7" pageNumber="421" type="holotype">holotypE</typeStatus>
|
||
<quantity id="305D9B32FFFE8E67FDC1FB6D14B1FB12" box="[601,695,1188,1212]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="421" unit="mm" value="1.1">1.1 mm</quantity>
|
||
. NumbEr of SEgmEntS 79–87 (avEragE= 83, n= 8);
|
||
<typeStatus id="281E8875FFFE8E67FA8BFB6D1373FB13" box="[1299,1397,1188,1213]" pageId="7" pageNumber="421" type="holotype">holotypE</typeStatus>
|
||
87 SEgmEntS. ThE majority of SEgmEntS widEr than long (
|
||
<figureCitation id="6F9E2A52FFFE8E67FD62FB0E156FFB4E" box="[762,873,1223,1248]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig 3A,D</figureCitation>
|
||
); aftEr clitEllum SomE SEgmEntS almoSt aS long aS widE. PigmEnt abSEnt. ProStomium prolobic, invaginatEd in thE majority of individualS. SEcondary furrowS barEly viSiblE; if prESEnt onE poStSEtal aftEr clitEllum. SEtaE Eight pEr SEgmEnt, viSiblE from
|
||
<emphasis id="C5D1EAC5FFFE8E67FBCDFAC61265FA86" bold="true" box="[1109,1123,1295,1320]" pageId="7" pageNumber="421">2</emphasis>
|
||
(
|
||
<figureCitation id="6F9E2A52FFFE8E67FBECFAC612EFFA86" box="[1140,1257,1295,1320]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig. 3A,D</figureCitation>
|
||
); cloSEly pairEd throughout. SEtal formula (avEragES, n=3) (
|
||
<emphasis id="C5D1EAC5FFFE8E67FD3CFAFD1555FAE2" box="[676,851,1332,1356]" italics="true" pageId="7" pageNumber="421">aa:ab:bc:cd:dd</emphasis>
|
||
) at
|
||
<emphasis id="C5D1EAC5FFFE8E67FC16FAFD15ACFAE3" bold="true" box="[910,938,1332,1357]" pageId="7" pageNumber="421">10</emphasis>
|
||
: 5.2:1:6.2:0.8:15 and 1.2
|
||
<emphasis id="C5D1EAC5FFFE8E67FB44FAFD12FEFAE2" box="[1244,1272,1332,1356]" italics="true" pageId="7" pageNumber="421">dd</emphasis>
|
||
=1/2
|
||
<emphasis id="C5D1EAC5FFFE8E67FAB4FAFD1339FAE5" box="[1324,1343,1332,1355]" italics="true" pageId="7" pageNumber="421">C</emphasis>
|
||
; at
|
||
<emphasis id="C5D1EAC5FFFE8E67FAE0FAFD1392FAE3" bold="true" box="[1400,1428,1332,1357]" pageId="7" pageNumber="421">30</emphasis>
|
||
: 5.9:1:6:1.2:19.1 and 1.4
|
||
<emphasis id="C5D1EAC5FFFE8E67FE3BFA9117B9FADE" box="[419,447,1368,1392]" italics="true" pageId="7" pageNumber="421">dd</emphasis>
|
||
=1/2
|
||
<emphasis id="C5D1EAC5FFFE8E67FE6BFA911400FAC1" box="[499,518,1368,1391]" italics="true" pageId="7" pageNumber="421">C</emphasis>
|
||
; tEn SEgmEntS bEforE anuS: 6.4:1:6:1.2:17 and 1.3
|
||
<emphasis id="C5D1EAC5FFFE8E67FBAEFA911254FADE" box="[1078,1106,1368,1392]" italics="true" pageId="7" pageNumber="421">dd</emphasis>
|
||
=1/2
|
||
<emphasis id="C5D1EAC5FFFE8E67FB1DFA91129EFAC1" box="[1157,1176,1368,1391]" italics="true" pageId="7" pageNumber="421">C</emphasis>
|
||
. PEnial SEtaE in
|
||
<emphasis id="C5D1EAC5FFFE8E67FAC8FA9E136AFADE" bold="true" box="[1360,1388,1367,1392]" pageId="7" pageNumber="421">17</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFFE8E67FF0FFAB516B5FA3B" bold="true" box="[151,179,1404,1429]" pageId="7" pageNumber="421">19</emphasis>
|
||
(
|
||
<figureCitation id="6F9E2A52FFFE8E67FF5AFAB51701FA3B" box="[194,263,1404,1429]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5</figureCitation>
|
||
); vEry difficult to Extract, and with a dimorphiSm charactErizEd by diffErEnt SizE and ornamEntation. ThE largEr
|
||
<emphasis id="C5D1EAC5FFFE8E67FF78FA6816E8FA16" box="[224,238,1441,1464]" italics="true" pageId="7" pageNumber="421">a</emphasis>
|
||
SEta (lEngth: 391, 415, 416, 420 µm; width: 11, 17, 17, 29 µm) almoSt Straight or Slightly brackEt-ShapEd (
|
||
<figureCitation id="6F9E2A52FFFE8E67FF07FA0D16FFFA73" box="[159,249,1476,1501]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5A</figureCitation>
|
||
) with diStal half undulatEd (
|
||
<figureCitation id="6F9E2A52FFFE8E67FDDAFA0D149CFA73" box="[578,666,1476,1501]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5B</figureCitation>
|
||
) and with a hook- or claw-ShapEd apEx (
|
||
<figureCitation id="6F9E2A52FFFE8E67FBEFFA0D12E5FA73" box="[1143,1251,1476,1501]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig 5C,E</figureCitation>
|
||
); ornamEntation charactErizEd by irrEgular and ScarcE thornS or notchES juSt bEforE thE apEx (
|
||
<figureCitation id="6F9E2A52FFFE8E67FB98FA2E1273F9AE" box="[1024,1141,1511,1536]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5C,D</figureCitation>
|
||
); in SomE individualS, thE SEta apparEntly within a covEr, but apEx frEE (
|
||
<figureCitation id="6F9E2A52FFFE8E67FD0AF9C514ECF98B" box="[658,746,1548,1573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5C</figureCitation>
|
||
). ThE ShortEr
|
||
<emphasis id="C5D1EAC5FFFE8E67FC1CF9C51594F98A" box="[900,914,1548,1572]" italics="true" pageId="7" pageNumber="421">b</emphasis>
|
||
SEta (lEngth: 315 µm; width: 12.3 µm) Slightly curvEd (
|
||
<figureCitation id="6F9E2A52FFFE8E67FF6EF9E6174BF9E6" box="[246,333,1583,1608]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5F</figureCitation>
|
||
) and with an acutE apEx curvEd 90° or morE in thE SamE dirEction of curvaturE (
|
||
<figureCitation id="6F9E2A52FFFE8E67FA92F9E6136EF9E6" box="[1290,1384,1583,1608]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5H</figureCitation>
|
||
); no undulationS arE prESEnt and ornamEntation charactErizEd by irrEgular notchES (
|
||
<figureCitation id="6F9E2A52FFFE8E67FB81F99D1268F9C3" box="[1049,1134,1620,1645]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig 5G</figureCitation>
|
||
), limitEd to diStal End juSt bEforE apEx (
|
||
<figureCitation id="6F9E2A52FFFE8E67FEA9F9BE1789F93E" box="[305,399,1655,1680]" captionStart="FIGURE 5" captionStartId="10.[151,250,1875,1897]" captionTargetBox="[226,1335,194,1828]" captionTargetId="figure@10.[224,1336,193,1828]" captionTargetPageId="10" captionText="FIGURE 5. Lavellodrilus sheylae sp. nov. SEM photographs. Large right penial setae a of segment 19 from Paratype IEOL 4830: A. Complete setae (scale 100 µm), B. Undulate region (scale 25 µm), C. Distal region and apex (scale 10 µm); from Paratype IEOL 4833: D. Ornamentation from distal region (scale 10 µm), E. Apex (scale 5 µm). Smaller right penial setae b of segment 19 (Paratype IEOL 4833): F. Complete setae (scale 100 µm), G. Distal ornamentation (scale 5 µm), H. Distal region and apex (scale 5 µm)." httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="7" pageNumber="421">Fig. 5H</figureCitation>
|
||
). GEnital SEtaE abSEnt. ClitEllum dark or light orangE, SaddlE-ShapEd, in
|
||
<emphasis id="C5D1EAC5FFFE8E67FB79F9BE1321F93E" bold="true" box="[1249,1319,1655,1680]" pageId="7" pageNumber="421">13–17</emphasis>
|
||
, vEntrally rEaching
|
||
<emphasis id="C5D1EAC5FFFE8E67FE9CF955172EF91D" box="[260,296,1692,1715]" italics="true" pageId="7" pageNumber="421">BC</emphasis>
|
||
(cloSE to
|
||
<emphasis id="C5D1EAC5FFFE8E67FE30F95417BFF91A" box="[424,441,1693,1716]" italics="true" pageId="7" pageNumber="421">B</emphasis>
|
||
) or
|
||
<emphasis id="C5D1EAC5FFFE8E67FE6CF9551403F91D" box="[500,517,1692,1715]" italics="true" pageId="7" pageNumber="421">A</emphasis>
|
||
(
|
||
<figureCitation id="6F9E2A52FFFE8E67FD82F9551495F91A" box="[538,659,1692,1717]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig. 3A,D</figureCitation>
|
||
). DorSal porES prESEnt all along thE body, firSt porE in 10/11 (7 individualS) or 11/12 (onE individual). SpErmathEcal porES mESial in 7/8 and 8/9, vEry Small, only SEEn in two individualS (
|
||
<figureCitation id="6F9E2A52FFFE8E67FEBBF92D1778F952" box="[291,382,1764,1789]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig. 3D</figureCitation>
|
||
). FEmalE porES in
|
||
<emphasis id="C5D1EAC5FFFE8E67FDD6F92D146CF953" bold="true" box="[590,618,1764,1789]" pageId="7" pageNumber="421">14</emphasis>
|
||
, prESEtal, mEdian to
|
||
<emphasis id="C5D1EAC5FFFE8E67FCCFF92C1563F952" box="[855,869,1765,1788]" italics="true" pageId="7" pageNumber="421">a</emphasis>
|
||
; in SomE individualS within a bilobular or ovoidal papilla (
|
||
<figureCitation id="6F9E2A52FFFE8E67FF6FF8CE1752F88E" box="[247,340,1799,1824]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig. 3C</figureCitation>
|
||
). Two pairS of mESial proStatic porES in
|
||
<emphasis id="C5D1EAC5FFFE8E67FCA3F8CE1551F88E" bold="true" box="[827,855,1799,1824]" pageId="7" pageNumber="421">17</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFFE8E67FC0FF8CE15B5F88E" bold="true" box="[919,947,1799,1824]" pageId="7" pageNumber="421">19</emphasis>
|
||
, ovEr Elliptical prominEncES tranSvErSally oriEntEd; from thE prominEncES, SEminal groovES curvE 90° to continuE ovEr
|
||
<emphasis id="C5D1EAC5FFFE8E67FB86F8E51246F8ED" box="[1054,1088,1836,1859]" italics="true" pageId="7" pageNumber="421">AB</emphasis>
|
||
in SEgmEnt
|
||
<emphasis id="C5D1EAC5FFFE8E67FB4CF8E512F6F8EB" bold="true" box="[1236,1264,1836,1861]" pageId="7" pageNumber="421">18</emphasis>
|
||
(
|
||
<figureCitation id="6F9E2A52FFFE8E67FA9BF8E51392F8EA" box="[1283,1428,1836,1861]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig. 3A,B,D</figureCitation>
|
||
) Surrounding gEnital markS. MalE porES not SEEn, probably within SEminal groovES and opEning SomEwhErE in
|
||
<emphasis id="C5D1EAC5FFFE8E67FAE1F8861393F8C6" bold="true" box="[1401,1429,1871,1896]" pageId="7" pageNumber="421">18</emphasis>
|
||
. GEnital markS aS pairEd papilla locatEd in
|
||
<emphasis id="C5D1EAC5FFFE8E67FDEAF8BD1488F823" bold="true" box="[626,654,1908,1933]" pageId="7" pageNumber="421">16</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFFE8E67FD5EF8BD14E4F823" bold="true" box="[710,738,1908,1933]" pageId="7" pageNumber="421">18</emphasis>
|
||
, Each onE ExtEnding from midvEntral linE to
|
||
<emphasis id="C5D1EAC5FFFE8E67FB47F8BC12F6F822" box="[1247,1264,1909,1932]" italics="true" pageId="7" pageNumber="421">B</emphasis>
|
||
and occupying all or ¾ of thE rESpEctivE SEgmEnt. ThoSE of
|
||
<emphasis id="C5D1EAC5FFFE8E67FD09F85114ABF81F" bold="true" box="[657,685,1944,1969]" pageId="7" pageNumber="421">16</emphasis>
|
||
ovoidal and longitudinally oriEntEd; thoSE of
|
||
<emphasis id="C5D1EAC5FFFE8E67FB2CF85112D6F81F" bold="true" box="[1204,1232,1944,1969]" pageId="7" pageNumber="421">18</emphasis>
|
||
largEr and almoSt circular (
|
||
<figureCitation id="6F9E2A52FFFE8E67FF64F875178CF87A" box="[252,394,1980,2005]" captionStart="FIGURE 3" captionStartId="8.[151,250,1690,1712]" captionTargetBox="[171,1395,230,1639]" captionTargetId="figure@8.[151,1436,195,1668]" captionTargetPageId="8" captionText="FIGURE 3. Lavellodrilus sheylae sp. nov. External. A. Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm). B. Latero-ventral view of segments 16–20 of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm) C. Ventral view of segments 14 and 15 showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm). D. Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm)." httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="7" pageNumber="421">Fig. 3A,B,D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A3DA665FFFF18E68FF0FF9531339F884" httpUri="https://zenodo.org/record/1446943/files/figure.png" pageId="8" pageNumber="422" startId="8.[151,250,1690,1712]" targetBox="[171,1395,230,1639]" targetPageId="8">
|
||
<paragraph id="F71A36D7FFF18E68FF0FF9531339F884" blockId="8.[151,1436,1689,1834]" pageId="8" pageNumber="422">
|
||
<emphasis id="C5D1EAC5FFF18E68FF0FF9531468F901" bold="true" box="[151,622,1689,1712]" pageId="8" pageNumber="422">
|
||
FIGURE 3.
|
||
<taxonomicName id="30A54D54FFF18E68FEBEF950140CF901" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[294,522,1689,1711]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="422" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFF18E68FEBEF950140CF901" bold="true" box="[294,522,1689,1711]" italics="true" pageId="8" pageNumber="422">Lavellodrilus sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFF18E68FD8FF9531468F901" box="[535,622,1690,1711]" pageId="8" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
External.
|
||
<emphasis id="C5D1EAC5FFF18E68FD7DF95314F8F901" bold="true" box="[741,766,1690,1711]" pageId="8" pageNumber="422">A.</emphasis>
|
||
Ventral view of anterior region of an individual with single diverticulum spermathecae (IEOL 4832; scale 1 mm).
|
||
<emphasis id="C5D1EAC5FFF18E68FD49F97014EEF960" bold="true" box="[721,744,1721,1742]" pageId="8" pageNumber="422">B.</emphasis>
|
||
Latero-ventral view of segments
|
||
<emphasis id="C5D1EAC5FFF18E68FBD2F971128CF963" bold="true" box="[1098,1162,1720,1742]" pageId="8" pageNumber="422">16–20</emphasis>
|
||
of an individual with two diverticula spermathecae (Holotype IEOL 4831) showing genital papillae (Gp) and prostatic pores (Pp) (scale 500 µm)
|
||
<emphasis id="C5D1EAC5FFF18E68FA1BF91E139AF943" bold="true" box="[1411,1436,1751,1773]" pageId="8" pageNumber="422">C.</emphasis>
|
||
Ventral view of segments
|
||
<emphasis id="C5D1EAC5FFF18E68FE3AF93F17BAF8A5" bold="true" box="[418,444,1782,1803]" pageId="8" pageNumber="422">14</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF18E68FE76F93F140EF8A5" bold="true" box="[494,520,1782,1803]" pageId="8" pageNumber="422">15</emphasis>
|
||
showing the papilla (P) that contains female pores (Fp) (Holotype IEOL 4831; scale 500 µm).
|
||
<emphasis id="C5D1EAC5FFF18E68FF56F8DC16E1F884" bold="true" box="[206,231,1813,1834]" pageId="8" pageNumber="422">D.</emphasis>
|
||
Ventral view of anterior region of an individual with two diverticula spermathecae (IEOL 4836; scale 1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A3DA665FFFF08E69FF0FF8D31274F846" httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="9" pageNumber="423" startId="9.[151,250,1818,1840]" targetBox="[182,1376,205,1764]" targetPageId="9">
|
||
<paragraph id="F71A36D7FFF08E69FF0FF8D31274F846" blockId="9.[151,1436,1818,2024]" pageId="9" pageNumber="423">
|
||
<emphasis id="C5D1EAC5FFF08E69FF0FF8D3144AF89E" bold="true" box="[151,588,1818,1840]" pageId="9" pageNumber="423">
|
||
FIGURE 4.
|
||
<taxonomicName id="30A54D54FFF08E69FE80F8D317F3F89E" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[280,501,1818,1840]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="423" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFF08E69FE80F8D317F3F89E" bold="true" box="[280,501,1818,1840]" italics="true" pageId="9" pageNumber="423">Lavellodrilus sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFF08E69FE64F8D2144AF89E" box="[508,588,1819,1840]" pageId="9" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Internal.
|
||
<emphasis id="C5D1EAC5FFF08E69FD36F8D314C0F881" bold="true" box="[686,710,1818,1839]" pageId="9" pageNumber="423">A.</emphasis>
|
||
Dorsal view of segments
|
||
<emphasis id="C5D1EAC5FFF08E69FC55F8D3121FF89F" bold="true" box="[973,1049,1818,1841]" pageId="9" pageNumber="423">11–18 (</emphasis>
|
||
Holotype IEOL 4831; scale 1 mm).
|
||
<emphasis id="C5D1EAC5FFF08E69FA1CF8D2139DF89E" bold="true" box="[1412,1435,1819,1840]" pageId="9" pageNumber="423">B.</emphasis>
|
||
Dorsal view of segments
|
||
<emphasis id="C5D1EAC5FFF08E69FE04F8F017DDF8E0" bold="true" box="[412,475,1849,1871]" pageId="9" pageNumber="423">15–17</emphasis>
|
||
from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment
|
||
<emphasis id="C5D1EAC5FFF08E69FDA7F89E145EF8C2" bold="true" box="[575,600,1879,1900]" pageId="9" pageNumber="423">16</emphasis>
|
||
where intestine was cut down to look for typhlosole.
|
||
<emphasis id="C5D1EAC5FFF08E69FBE6F89E1291F8C3" bold="true" box="[1150,1175,1879,1901]" pageId="9" pageNumber="423">C.</emphasis>
|
||
Parietes of segments
|
||
<emphasis id="C5D1EAC5FFF08E69FAEEF89E16B6F825" bold="true" pageId="9" pageNumber="423">16– 19</emphasis>
|
||
, showing parietal lateral-ventral vessels and prostates of
|
||
<emphasis id="C5D1EAC5FFF08E69FD66F8BF151EF825" bold="true" box="[766,792,1910,1931]" pageId="9" pageNumber="423">17</emphasis>
|
||
(P17) and 1
|
||
<emphasis id="C5D1EAC5FFF08E69FC00F8BF15A3F825" bold="true" box="[920,933,1910,1931]" pageId="9" pageNumber="423">9</emphasis>
|
||
(P19) (Paratype IEOL 4837; scale 0.86 mm).
|
||
<emphasis id="C5D1EAC5FFF08E69FA1BF8BE139AF822" bold="true" box="[1411,1436,1911,1932]" pageId="9" pageNumber="423">D.</emphasis>
|
||
Dorso-lateral view of single diverticulum spermatheca of segment
|
||
<emphasis id="C5D1EAC5FFF08E69FCDDF85C1554F804" bold="true" box="[837,850,1941,1962]" pageId="9" pageNumber="423">8</emphasis>
|
||
(Paratype IEOL 4830; scale 200 µm).
|
||
<emphasis id="C5D1EAC5FFF08E69FB7CF85C12FAF804" bold="true" box="[1252,1276,1941,1962]" pageId="9" pageNumber="423">E.</emphasis>
|
||
Dorsal view of two diverticula spermatheca of segment
|
||
<emphasis id="C5D1EAC5FFF08E69FDB7F87D143AF867" bold="true" box="[559,572,1972,1993]" pageId="9" pageNumber="423">8</emphasis>
|
||
(Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A3DA665FFFF38E6AFF0FF89A1778F840" httpUri="https://zenodo.org/record/1446947/files/figure.png" pageId="10" pageNumber="424" startId="10.[151,250,1875,1897]" targetBox="[226,1335,194,1828]" targetPageId="10">
|
||
<paragraph id="F71A36D7FFF38E6AFF0FF89A1778F840" blockId="10.[151,1436,1875,2030]" pageId="10" pageNumber="424">
|
||
<emphasis id="C5D1EAC5FFF38E6AFF0FF89A1455F8C6" bold="true" box="[151,595,1875,1897]" pageId="10" pageNumber="424">
|
||
FIGURE 5.
|
||
<taxonomicName id="30A54D54FFF38E6AFE83F89A17FCF8C7" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[283,506,1875,1897]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="424" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFF38E6AFE83F89A17FCF8C7" bold="true" box="[283,506,1875,1897]" italics="true" pageId="10" pageNumber="424">Lavellodrilus sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFF38E6AFD9AF89A1455F8C6" box="[514,595,1875,1896]" pageId="10" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
SEM photographs. Large right penial setae
|
||
<emphasis id="C5D1EAC5FFF38E6AFB84F89D122FF8C7" box="[1052,1065,1876,1897]" italics="true" pageId="10" pageNumber="424">a</emphasis>
|
||
of segment
|
||
<emphasis id="C5D1EAC5FFF38E6AFB30F89A12C7F8C6" bold="true" box="[1192,1217,1875,1896]" pageId="10" pageNumber="424">19</emphasis>
|
||
from Paratype IEOL 4830:
|
||
<emphasis id="C5D1EAC5FFF38E6AFF41F8BD16F7F827" bold="true" box="[217,241,1908,1929]" pageId="10" pageNumber="424">A.</emphasis>
|
||
Complete setae (scale 100 µm),
|
||
<emphasis id="C5D1EAC5FFF38E6AFDD3F8BC1464F824" bold="true" box="[587,610,1909,1930]" pageId="10" pageNumber="424">B.</emphasis>
|
||
Undulate region (scale 25 µm),
|
||
<emphasis id="C5D1EAC5FFF38E6AFC2FF8BD15D6F824" bold="true" box="[951,976,1908,1930]" pageId="10" pageNumber="424">C.</emphasis>
|
||
Distal region and apex (scale 10 µm); from Paratype IEOL 4833:
|
||
<emphasis id="C5D1EAC5FFF38E6AFEEDF85F1788F805" bold="true" box="[373,398,1942,1963]" pageId="10" pageNumber="424">D.</emphasis>
|
||
Ornamentation from distal region (scale 10 µm),
|
||
<emphasis id="C5D1EAC5FFF38E6AFC1EF85F159BF805" bold="true" box="[902,925,1942,1963]" pageId="10" pageNumber="424">E.</emphasis>
|
||
Apex (scale 5 µm). Smaller right penial setae
|
||
<emphasis id="C5D1EAC5FFF38E6AFAECF85C1387F805" box="[1396,1409,1941,1963]" italics="true" pageId="10" pageNumber="424">b</emphasis>
|
||
of segment 19 (Paratype IEOL 4833):
|
||
<emphasis id="C5D1EAC5FFF38E6AFD9EF87E1411F862" bold="true" box="[518,535,1975,1996]" pageId="10" pageNumber="424">F.</emphasis>
|
||
Complete setae (scale 100 µm),
|
||
<emphasis id="C5D1EAC5FFF38E6AFCF4F87E1586F863" bold="true" box="[876,896,1975,1997]" pageId="10" pageNumber="424">G.</emphasis>
|
||
Distal ornamentation (scale 5 µm),
|
||
<emphasis id="C5D1EAC5FFF38E6AFB6EF87E1317F862" bold="true" box="[1270,1297,1975,1996]" pageId="10" pageNumber="424">H.</emphasis>
|
||
Distal region and apex (scale 5 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A3DA665FFFF28E6BFF0BFD9E16D5FD26" pageId="11" pageNumber="425">
|
||
<paragraph id="F71A36D7FFF28E6BFF0BFD9E16D5FD26" blockId="11.[147,1818,598,648]" pageId="11" pageNumber="425">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFD9E16FAFDC2" bold="true" box="[147,252,599,620]" pageId="11" pageNumber="425">TABLE 2.</emphasis>
|
||
Morphological comparisons among the currently recognized
|
||
<taxonomicName id="30A54D54FFF28E6BFCD5FD9F15CBFDC2" authorityName="Fragoso" authorityYear="1988" box="[845,973,598,620]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="genus">
|
||
<emphasis id="C5D1EAC5FFF28E6BFCD5FD9F15CBFDC2" box="[845,973,598,620]" italics="true" pageId="11" pageNumber="425">Lavellodrilus</emphasis>
|
||
</taxonomicName>
|
||
species from Mexico. Data obtained from Fragoso (1988, 1991) and Fragoso and Rojas (2016)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F71A36D7FFF28E6BFD8BFD6910D8FA15" pageId="11" pageNumber="425">
|
||
<table id="85A5C477FFF2719FFF0BFD691153F9A0" box="[147,1877,672,1550]" gridcols="6" gridrows="8" pageId="11" pageNumber="425" rowsContinueIn="12.[147,1943,735,1560]">
|
||
<tr id="49953495FFF2719FFF0BFD691153FD7E" box="[147,1877,672,720]" gridrow="0" pageId="11" pageNumber="425" rowspan-0="1">
|
||
<th id="0A445DE9FFF2719FFD8BFD691484FD7E" box="[531,642,672,720]" gridcol="1" gridrow="0" pageId="11" pageNumber="425">Size (mm)</th>
|
||
<th id="0A445DE9FFF2719FFD5BFD691530FD7E" box="[707,822,672,720]" gridcol="2" gridrow="0" pageId="11" pageNumber="425">
|
||
1
|
||
<taxonomicName id="30A54D54FFF28E6BFD54FD691526FD1B" box="[716,800,672,693]" pageId="11" pageNumber="425" rank="subSpecies" subSpecies="dorsal">st dorsal</taxonomicName>
|
||
pore
|
||
</th>
|
||
<th id="0A445DE9FFF2719FFCCBFD69125AFD7E" box="[851,1116,672,720]" gridcol="3" gridrow="0" pageId="11" pageNumber="425">Genital marks</th>
|
||
<th id="0A445DE9FFF2719FFBE7FD691076FD7E" box="[1151,1648,672,720]" gridcol="4" gridrow="0" pageId="11" pageNumber="425">Genital setae/Penial setae</th>
|
||
<th id="0A445DE9FFF2719FF909FD691153FD7E" box="[1681,1877,672,720]" gridcol="5" gridrow="0" pageId="11" pageNumber="425">Gizzard</th>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFD1E1153FC8B" box="[147,1877,727,805]" gridrow="1" pageId="11" pageNumber="425">
|
||
<th id="0A445DE9FFF2719FFF0BFD1E17D6FC8B" box="[147,464,727,805]" gridcol="0" gridrow="1" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFD11173CFCA7" authority="Fragoso, 1991 Bonampak, Chis." authorityName="Fragoso" authorityYear="1991" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="bonampakensis">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFD111746FD43" box="[147,320,727,749]" italics="true" pageId="11" pageNumber="425">L. bonampakensis</emphasis>
|
||
Fragoso, 1991 Bonampak, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFD1E1484FC8B" box="[531,642,727,805]" gridcol="1" gridrow="1" pageId="11" pageNumber="425">
|
||
122
|
||
<emphasis id="C5D1EAC5FFF28E6BFDAFFD111445FD43" box="[567,579,728,749]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
130 L 3.4
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FD3D143BFCA7" box="[561,573,756,777]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
3.7 W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFD5BFD1E1530FC8B" box="[707,822,727,805]" gridcol="2" gridrow="1" pageId="11" pageNumber="425">9/10</td>
|
||
<td id="0A445DE9FFF2719FFCCBFD1E125AFC8B" box="[851,1116,727,805]" gridcol="3" gridrow="1" pageId="11" pageNumber="425">
|
||
bilobulated midventral papillae in
|
||
<emphasis id="C5D1EAC5FFF28E6BFC26FD3D15CCFCA7" bold="true" box="[958,970,756,777]" pageId="11" pageNumber="425">9</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF28E6BFC61FD3D1217FCA7" bold="true" box="[1017,1041,756,777]" pageId="11" pageNumber="425">17</emphasis>
|
||
(2); ovoidal in 19/20 (1)
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FD1E1076FC8B" box="[1151,1648,727,805]" gridcol="4" gridrow="1" pageId="11" pageNumber="425">
|
||
present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFD1112E9FD43" bold="true" box="[1251,1263,728,749]" pageId="11" pageNumber="425">9</emphasis>
|
||
, bracket shaped and lanceolate apex/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFD3D1319FCA7" bold="true" box="[1251,1311,756,777]" pageId="11" pageNumber="425">17, 19</emphasis>
|
||
bracket shaped and scarcely ornamented at distal end
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FD1E1153FC8B" box="[1681,1877,727,805]" gridcol="5" gridrow="1" pageId="11" pageNumber="425">
|
||
one esophageal in
|
||
<emphasis id="C5D1EAC5FFF28E6BF8DBFD111153FD43" bold="true" box="[1859,1877,728,749]" pageId="11" pageNumber="425">5,</emphasis>
|
||
small; one intestinal in
|
||
<emphasis id="C5D1EAC5FFF28E6BF932FCD910CEFC8B" bold="true" box="[1706,1736,784,805]" pageId="11" pageNumber="425">15,</emphasis>
|
||
larger
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFC8F1153FC01" box="[147,1877,838,943]" gridrow="2" pageId="11" pageNumber="425" rowspan-2="1">
|
||
<th id="0A445DE9FFF2719FFF0BFC8F17D6FC01" box="[147,464,838,943]" gridcol="0" gridrow="2" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFC8E1714FCD9" authority="Fragoso, 1991 Chajul, Chis." authorityName="Fragoso" authorityYear="1991" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="ilkus">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFC8E16DCFCF2" box="[147,218,838,860]" italics="true" pageId="11" pageNumber="425">L. ilkus</emphasis>
|
||
Fragoso, 1991 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFC8F1484FC01" box="[531,642,838,943]" gridcol="1" gridrow="2" pageId="11" pageNumber="425">
|
||
230
|
||
<emphasis id="C5D1EAC5FFF28E6BFDAFFC8E1445FCF2" box="[567,579,839,860]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
335 L 4.3
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FCAA143BFCD6" box="[561,573,867,888]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
5.0 W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFCCBFC8F125AFC01" box="[851,1116,838,943]" gridcol="3" gridrow="2" pageId="11" pageNumber="425">
|
||
bilobulated midventral papillae in
|
||
<emphasis id="C5D1EAC5FFF28E6BFC26FCAB15CCFCD9" bold="true" box="[958,970,866,887]" pageId="11" pageNumber="425">7</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFC4EFCAB15E4FCD9" bold="true" box="[982,994,866,887]" pageId="11" pageNumber="425">8</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFC76FCAB15FCFCD9" bold="true" box="[1006,1018,866,887]" pageId="11" pageNumber="425">9</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF28E6BFBB1FCAB1247FCD9" bold="true" box="[1065,1089,866,887]" pageId="11" pageNumber="425">17</emphasis>
|
||
(4); ovoidal in 19/20, 20/21 (2)
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FC8F1076FC01" box="[1151,1648,838,943]" gridcol="4" gridrow="2" pageId="11" pageNumber="425">
|
||
absent/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFAB5FC8E1343FCF2" bold="true" box="[1325,1349,839,860]" pageId="11" pageNumber="425">17</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFAD3FC8E1365FCF2" bold="true" box="[1355,1379,839,860]" pageId="11" pageNumber="425">19</emphasis>
|
||
; all bracket shaped and scarcely ornamented at distal end; spoon bowl- shaped apex
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FC8F1153FC01" box="[1681,1877,838,943]" gridcol="5" gridrow="2" pageId="11" pageNumber="425">
|
||
one esophageal in
|
||
<emphasis id="C5D1EAC5FFF28E6BF8DBFC8E1153FCF2" bold="true" box="[1859,1877,839,860]" pageId="11" pageNumber="425">5,</emphasis>
|
||
small; one intestinal in
|
||
<emphasis id="C5D1EAC5FFF28E6BF932FCB710CEFC3D" bold="true" box="[1706,1736,894,915]" pageId="11" pageNumber="425">15,</emphasis>
|
||
larger
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFC181153FBB3" box="[147,1877,977,1053]" gridrow="3" pageId="11" pageNumber="425">
|
||
<th id="0A445DE9FFF2719FFF0BFC1817D6FBB3" box="[147,464,977,1053]" gridcol="0" gridrow="3" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFC181714FBAF" authority="Fragoso, 1988 Chajul, Chis." authorityName="Fragoso" authorityYear="1988" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="maya">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFC1816E6FC48" box="[147,224,977,998]" italics="true" pageId="11" pageNumber="425">L. maya</emphasis>
|
||
Fragoso, 1988 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFC181484FBB3" box="[531,642,977,1053]" gridcol="1" gridrow="3" pageId="11" pageNumber="425">
|
||
81
|
||
<emphasis id="C5D1EAC5FFF28E6BFDB3FC181431FC48" box="[555,567,977,998]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
114 L 1.7
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FC24143BFBAC" box="[561,573,1005,1026]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
2.4 W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFD5BFC181530FBB3" box="[707,822,977,1053]" gridcol="2" gridrow="3" pageId="11" pageNumber="425">8/9, 9/10</td>
|
||
<td id="0A445DE9FFF2719FFCCBFC18125AFBB3" box="[851,1116,977,1053]" gridcol="3" gridrow="3" pageId="11" pageNumber="425">
|
||
midventral papillae in 16/17, 21/22 (2) and sweallings in
|
||
<emphasis id="C5D1EAC5FFF28E6BFC4FFBC115E5FBB3" bold="true" box="[983,995,1032,1053]" pageId="11" pageNumber="425">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF28E6BFB8AFBC11218FBB3" bold="true" box="[1042,1054,1032,1053]" pageId="11" pageNumber="425">9</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FC181076FBB3" box="[1151,1648,977,1053]" gridcol="4" gridrow="3" pageId="11" pageNumber="425">
|
||
present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFC1812E9FC48" bold="true" box="[1251,1263,977,998]" pageId="11" pageNumber="425">8</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFB6DFC181307FC48" bold="true" box="[1269,1281,977,998]" pageId="11" pageNumber="425">9</emphasis>
|
||
/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFAEFFC181389FC48" bold="true" box="[1399,1423,977,998]" pageId="11" pageNumber="425">17</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFA0EFC1813A8FC48" bold="true" box="[1430,1454,977,998]" pageId="11" pageNumber="425">19</emphasis>
|
||
; all bracket shaped and ornamented at distal end
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FC181153FBB3" box="[1681,1877,977,1053]" gridcol="5" gridrow="3" pageId="11" pageNumber="425">
|
||
one in
|
||
<emphasis id="C5D1EAC5FFF28E6BF94AFC1810D8FC48" bold="true" box="[1746,1758,977,998]" pageId="11" pageNumber="425">5</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFBEA1153FB25" box="[147,1877,1059,1163]" gridrow="4" pageId="11" pageNumber="425">
|
||
<th id="0A445DE9FFF2719FFF0BFBEA17D6FB25" box="[147,464,1059,1163]" gridcol="0" gridrow="4" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFBED1714FBDE" authority="Fragoso & Rojas, 2016 Chajul, Chis." authorityName="Fragoso & Rojas" authorityYear="2016" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="notosetosus">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFBED171AFB97" box="[147,284,1060,1081]" italics="true" pageId="11" pageNumber="425">L. notosetosus</emphasis>
|
||
Fragoso & Rojas, 2016 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFBEA1484FB25" box="[531,642,1059,1163]" gridcol="1" gridrow="4" pageId="11" pageNumber="425">
|
||
135 L 1.2
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FB89143BFBFB" box="[561,573,1088,1109]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
1.5 W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFD5BFBEA1530FB25" box="[707,822,1059,1163]" gridcol="2" gridrow="4" pageId="11" pageNumber="425">13/14</td>
|
||
<td id="0A445DE9FFF2719FFCCBFBEA125AFB25" box="[851,1116,1059,1163]" gridcol="3" gridrow="4" pageId="11" pageNumber="425">
|
||
midventral papillae in 16/17,19/20, 20/21,21/22, 22/23 (5) and butterfly shaped swelling in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB94FBBF1222FB25" bold="true" box="[1036,1060,1142,1163]" pageId="11" pageNumber="425">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FBEA1076FB25" box="[1151,1648,1059,1163]" gridcol="4" gridrow="4" pageId="11" pageNumber="425">
|
||
present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFBEA12FDFB96" bold="true" box="[1251,1275,1059,1080]" pageId="11" pageNumber="425">12</emphasis>
|
||
, almost straight, highly ornamented with lanceolate apex/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFA2DFBF613F1FBFA" bold="true" box="[1461,1527,1087,1108]" pageId="11" pageNumber="425">17, 19,</emphasis>
|
||
very large, highly curved and with undulated distal end ending in an arrow shaped apex
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FBEA1153FB25" box="[1681,1877,1059,1163]" gridcol="5" gridrow="4" pageId="11" pageNumber="425">
|
||
one in
|
||
<emphasis id="C5D1EAC5FFF28E6BF94AFBEA10D8FB96" bold="true" box="[1746,1758,1059,1080]" pageId="11" pageNumber="425">5</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFB671153FABB" box="[147,1877,1198,1301]" gridrow="5" pageId="11" pageNumber="425">
|
||
<th id="0A445DE9FFF2719FFF0BFB6717D6FABB" box="[147,464,1198,1301]" gridcol="0" gridrow="5" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFB671714FB70" authority="Fragoso, 1988 Chajul, Chis." authorityName="Fragoso" authorityYear="1988" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="parvus">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFB6716EBFB6D" box="[147,237,1198,1219]" italics="true" pageId="11" pageNumber="425">L. parvus</emphasis>
|
||
Fragoso, 1988 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFB671484FABB" box="[531,642,1198,1301]" gridcol="1" gridrow="5" pageId="11" pageNumber="425">
|
||
25
|
||
<emphasis id="C5D1EAC5FFF28E6BFDB3FB671431FB6D" box="[555,567,1198,1219]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
32 L 0.8
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FB03143BFB71" box="[561,573,1226,1247]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
1.5 W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFD5BFB671530FABB" box="[707,822,1198,1301]" gridcol="2" gridrow="5" pageId="11" pageNumber="425">10/11,11/12</td>
|
||
<td id="0A445DE9FFF2719FFCCBFB67125AFABB" box="[851,1116,1198,1301]" gridcol="3" gridrow="5" pageId="11" pageNumber="425">
|
||
paired papillae in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB98FB67121EFB6D" bold="true" box="[1024,1048,1198,1219]" pageId="11" pageNumber="425">17</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFBBCFB67123AFB6D" bold="true" box="[1060,1084,1198,1219]" pageId="11" pageNumber="425">19</emphasis>
|
||
; midventral unpaired in
|
||
<emphasis id="C5D1EAC5FFF28E6BFBACFB00124AFB70" bold="true" box="[1076,1100,1225,1246]" pageId="11" pageNumber="425">20</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFCCBFB2C156DFB54" bold="true" box="[851,875,1253,1274]" pageId="11" pageNumber="425">21</emphasis>
|
||
; midventral sweallings in
|
||
<emphasis id="C5D1EAC5FFF28E6BFCF4FAC9157EFABB" bold="true" box="[876,888,1280,1301]" pageId="11" pageNumber="425">9</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF28E6BFCE6FAC91590FABB" bold="true" box="[894,918,1280,1301]" pageId="11" pageNumber="425">10</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FB671076FABB" box="[1151,1648,1198,1301]" gridcol="4" gridrow="5" pageId="11" pageNumber="425">
|
||
present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFB6712E9FB6D" bold="true" box="[1251,1263,1198,1219]" pageId="11" pageNumber="425">9</emphasis>
|
||
, bracket shaped and lanceolate apex/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFB001319FB70" bold="true" box="[1251,1311,1225,1246]" pageId="11" pageNumber="425">17, 19</emphasis>
|
||
with two curvatures; all ornamented at distal en.
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FB671153FABB" box="[1681,1877,1198,1301]" gridcol="5" gridrow="5" pageId="11" pageNumber="425">
|
||
one in
|
||
<emphasis id="C5D1EAC5FFF28E6BF94AFB6710D8FB6D" bold="true" box="[1746,1758,1198,1219]" pageId="11" pageNumber="425">5</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFAF11153FA2A" box="[147,1877,1336,1412]" gridrow="6" pageId="11" pageNumber="425">
|
||
<th id="0A445DE9FFF2719FFF0BFAF117D6FA2A" box="[147,464,1336,1412]" gridcol="0" gridrow="6" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFAF11714FAC6" authority="Fragoso, 1988 Chajul, Chis." authorityName="Fragoso" authorityYear="1988" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="riparius">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFAF116FFFAE3" box="[147,249,1336,1357]" italics="true" pageId="11" pageNumber="425">L. riparius</emphasis>
|
||
Fragoso, 1988 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFAF11484FA2A" box="[531,642,1336,1412]" gridcol="1" gridrow="6" pageId="11" pageNumber="425">
|
||
110
|
||
<emphasis id="C5D1EAC5FFF28E6BFDAFFAF11445FAE3" box="[567,579,1336,1357]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
133 L 2.1
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FA9A143BFAC6" box="[561,573,1363,1384]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
3.5W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFD5BFAF11530FA2A" box="[707,822,1336,1412]" gridcol="2" gridrow="6" pageId="11" pageNumber="425">8/9, 9/10</td>
|
||
<td id="0A445DE9FFF2719FFCCBFAF1125AFA2A" box="[851,1116,1336,1412]" gridcol="3" gridrow="6" pageId="11" pageNumber="425">
|
||
midventral papillae in 16/17, 20/21
|
||
<emphasis id="C5D1EAC5FFF28E6BFC55FA9A15DFFAC6" box="[973,985,1363,1384]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
24/25 (6) and a swealling in
|
||
<emphasis id="C5D1EAC5FFF28E6BFC47FAA615EDFA2A" bold="true" box="[991,1003,1391,1412]" pageId="11" pageNumber="425">9</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FAF11076FA2A" box="[1151,1648,1336,1412]" gridcol="4" gridrow="6" pageId="11" pageNumber="425">
|
||
present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFB7BFAF112E9FAE3" bold="true" box="[1251,1263,1336,1357]" pageId="11" pageNumber="425">9</emphasis>
|
||
/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFAC7FAF1139AFAE3" bold="true" box="[1375,1436,1336,1357]" pageId="11" pageNumber="425">17, 19</emphasis>
|
||
; all bracket shaped and ornamented at distal end
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FAF11153FA2A" box="[1681,1877,1336,1412]" gridcol="5" gridrow="6" pageId="11" pageNumber="425">
|
||
one in
|
||
<emphasis id="C5D1EAC5FFF28E6BF94AFAF110D8FAE3" bold="true" box="[1746,1758,1336,1357]" pageId="11" pageNumber="425">5</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF2719FFF0BFA6C1153F9A0" box="[147,1877,1445,1550]" gridrow="7" pageId="11" pageNumber="425">
|
||
<th id="0A445DE9FFF2719FFF0BFA6C17D6F9A0" box="[147,464,1445,1550]" gridcol="0" gridrow="7" pageId="11" pageNumber="425">
|
||
<taxonomicName id="30A54D54FFF28E6BFF0BFA6F16F2FA15" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[147,244,1445,1467]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="425" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF0BFA6F16F2FA15" box="[147,244,1445,1467]" italics="true" pageId="11" pageNumber="425">L. sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFF28E6BFF62FA6F1741FA15" box="[250,327,1446,1467]" pageId="11" pageNumber="425" rank="species">
|
||
<emphasis id="C5D1EAC5FFF28E6BFF62FA6F1741FA15" bold="true" box="[250,327,1446,1467]" pageId="11" pageNumber="425">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Comalcalco and Teapa, Tab. MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF2719FFD8BFA6C1484F9A0" box="[531,642,1445,1550]" gridcol="1" gridrow="7" pageId="11" pageNumber="425">
|
||
18
|
||
<emphasis id="C5D1EAC5FFF28E6BFDB3FA6F1431FA15" box="[555,567,1446,1467]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
30 L 0.9
|
||
<emphasis id="C5D1EAC5FFF28E6BFDA9FA0B143BFA79" box="[561,573,1474,1495]" italics="true" pageId="11" pageNumber="425">–</emphasis>
|
||
1.1 W
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFD5BFA6C1530F9A0" box="[707,822,1445,1550]" gridcol="2" gridrow="7" pageId="11" pageNumber="425">10/11</td>
|
||
<td id="0A445DE9FFF2719FFCCBFA6C125AF9A0" box="[851,1116,1445,1550]" gridcol="3" gridrow="7" pageId="11" pageNumber="425">
|
||
paired papilla located in
|
||
<emphasis id="C5D1EAC5FFF28E6BFBD9FA6F125FFA15" bold="true" box="[1089,1113,1446,1467]" pageId="11" pageNumber="425">16</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF28E6BFCE4FA0B1592FA79" bold="true" box="[892,916,1474,1495]" pageId="11" pageNumber="425">18</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF2719FFBE7FA6C1076F9A0" box="[1151,1648,1445,1550]" gridcol="4" gridrow="7" pageId="11" pageNumber="425">
|
||
absent/ present in
|
||
<emphasis id="C5D1EAC5FFF28E6BFAB5FA6F136FFA15" bold="true" box="[1325,1385,1446,1467]" pageId="11" pageNumber="425">17, 19</emphasis>
|
||
, dimorphic; setae
|
||
<emphasis id="C5D1EAC5FFF28E6BF982FA6F1020FA15" box="[1562,1574,1446,1467]" italics="true" pageId="11" pageNumber="425">a</emphasis>
|
||
longer, undulated at distal end; setae
|
||
<emphasis id="C5D1EAC5FFF28E6BFA02FA0813A0FA79" box="[1434,1446,1473,1495]" italics="true" pageId="11" pageNumber="425">b</emphasis>
|
||
smaller and non- undulated; all slightly curved, with scarce distal ornamentation and with hook-claw shaped apex
|
||
</td>
|
||
<td id="0A445DE9FFF2719FF909FA6C1153F9A0" box="[1681,1877,1445,1550]" gridcol="5" gridrow="7" pageId="11" pageNumber="425">one in 5</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFF28E6BF9B3F9DC115FF985" blockId="11.[1579,1881,1557,1579]" box="[1579,1881,1557,1579]" pageId="11" pageNumber="425">
|
||
<emphasis id="C5D1EAC5FFF28E6BF9B3F9DC115FF985" box="[1579,1881,1557,1579]" italics="true" pageId="11" pageNumber="425">……continued on the next page</emphasis>
|
||
</paragraph>
|
||
<caption id="A3DA665FFFF58E6CFF0BFD611769FD12" box="[147,367,680,700]" pageId="12" pageNumber="426">
|
||
<paragraph id="F71A36D7FFF58E6CFF0BFD611769FD12" blockId="12.[147,367,680,700]" box="[147,367,680,700]" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFD6116FFFD12" bold="true" box="[147,249,680,700]" pageId="12" pageNumber="426">TABLE 2.</emphasis>
|
||
(Continued)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F71A36D7FFF58E6CFDBAFD16103FFA53" pageId="12" pageNumber="426">
|
||
<table id="85A5C477FFF5719FFF0BFD161191F9B6" box="[147,1943,735,1560]" gridcols="6" gridrows="8" pageId="12" pageNumber="426" rowsContinueFrom="11.[147,1877,672,1550]">
|
||
<tr id="49953495FFF5719FFF0BFD161191FCA0" box="[147,1943,735,782]" gridrow="0" pageId="12" pageNumber="426" rowspan-0="1">
|
||
<th id="0A445DE9FFF5719FFDBAFD161510FCA0" box="[546,790,735,782]" gridcol="1" gridrow="0" pageId="12" pageNumber="426">Esophagus</th>
|
||
<th id="0A445DE9FFF5719FFCDFFD1615BBFCA0" box="[839,957,735,782]" gridcol="2" gridrow="0" pageId="12" pageNumber="426">
|
||
1
|
||
<taxonomicName id="30A54D54FFF58E6CFCC8FD1615BBFD5D" box="[848,957,735,755]" pageId="12" pageNumber="426" rank="subSpecies" subSpecies="intestinal">st intestinal</taxonomicName>
|
||
segment
|
||
</th>
|
||
<th id="0A445DE9FFF5719FFC74FD1612E0FCA0" box="[1004,1254,735,782]" gridcol="3" gridrow="0" pageId="12" pageNumber="426">Typhlosole</th>
|
||
<th id="0A445DE9FFF5719FFA8AFD161358FCA0" box="[1298,1374,735,782]" gridcol="4" gridrow="0" pageId="12" pageNumber="426">Seminal vesicles</th>
|
||
<th id="0A445DE9FFF5719FFA2FFD161191FCA0" box="[1463,1943,735,782]" gridcol="5" gridrow="0" pageId="12" pageNumber="426">Spermathecae</th>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFCDC1191FCCE" box="[147,1943,789,864]" gridrow="1" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFCDC17F9FCCE" box="[147,511,789,864]" gridcol="0" gridrow="1" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFCDC1733FCEB" authority="Fragoso, 1991 Bonampak, Chis." authorityName="Fragoso" authorityYear="1991" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="bonampakensis">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFCDC173DFC84" box="[147,315,789,810]" italics="true" pageId="12" pageNumber="426">L. bonampakensis</emphasis>
|
||
Fragoso, 1991 Bonampak, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFCDC1510FCCE" box="[546,790,789,864]" gridcol="1" gridrow="1" pageId="12" pageNumber="426">
|
||
esophageal pouches in
|
||
<emphasis id="C5D1EAC5FFF58E6CFD61FCDF1516FC84" bold="true" box="[761,784,790,810]" pageId="12" pageNumber="426">14</emphasis>
|
||
, highly vascularized
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFCDFFCDC15BBFCCE" box="[839,957,789,864]" gridcol="2" gridrow="1" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFCDF1559FC84" bold="true" box="[839,863,790,810]" pageId="12" pageNumber="426">16</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FCDC12E0FCCE" box="[1004,1254,789,864]" gridcol="3" gridrow="1" pageId="12" pageNumber="426">
|
||
present, laminar, starting from
|
||
<emphasis id="C5D1EAC5FFF58E6CFB87FCF81231FCEB" bold="true" box="[1055,1079,817,837]" pageId="12" pageNumber="426">16</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFCDC1358FCCE" box="[1298,1374,789,864]" gridcol="4" gridrow="1" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFCDF132CFC84" bold="true" box="[1298,1322,790,810]" pageId="12" pageNumber="426">11</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CFAADFCDF134BFC84" bold="true" box="[1333,1357,790,810]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFCDC1191FCCE" box="[1463,1943,789,864]" gridcol="5" gridrow="1" pageId="12" pageNumber="426">
|
||
two pairs in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9B3FCDF1031FC84" bold="true" box="[1579,1591,790,810]" pageId="12" pageNumber="426">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF58E6CF9FCFCDF1076FC84" bold="true" box="[1636,1648,790,810]" pageId="12" pageNumber="426">9</emphasis>
|
||
; short duct, ovoid ampulla; diverticulum smaller than ampulla, and lateral to ental part of duct
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFC491191FC65" box="[147,1943,896,971]" gridrow="2" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFC4917F9FC65" box="[147,511,896,971]" gridcol="0" gridrow="2" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFC491708FC1E" authority="Fragoso, 1991 Chajul, Chis." authorityName="Fragoso" authorityYear="1991" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="ilkus">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFC4916D1FC3B" box="[147,215,896,917]" italics="true" pageId="12" pageNumber="426">L. ilkus</emphasis>
|
||
Fragoso, 1991 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFC491510FC65" box="[546,790,896,971]" gridcol="1" gridrow="2" pageId="12" pageNumber="426">
|
||
esophageal pouches in
|
||
<emphasis id="C5D1EAC5FFF58E6CFD61FC481516FC3B" bold="true" box="[761,784,897,917]" pageId="12" pageNumber="426">14</emphasis>
|
||
, highly vascularized
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFCDFFC4915BBFC65" box="[839,957,896,971]" gridcol="2" gridrow="2" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFC4B1559FC38" bold="true" box="[839,863,898,918]" pageId="12" pageNumber="426">16</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FC4912E0FC65" box="[1004,1254,896,971]" gridcol="3" gridrow="2" pageId="12" pageNumber="426">
|
||
present, laminar, starting from
|
||
<emphasis id="C5D1EAC5FFF58E6CFB87FC551231FC1E" bold="true" box="[1055,1079,924,944]" pageId="12" pageNumber="426">16</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFC491358FC65" box="[1298,1374,896,971]" gridcol="4" gridrow="2" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFC48132CFC3B" bold="true" box="[1298,1322,897,917]" pageId="12" pageNumber="426">11</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CFAADFC48134BFC3B" bold="true" box="[1333,1357,897,917]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFC491191FC65" box="[1463,1943,896,971]" gridcol="5" gridrow="2" pageId="12" pageNumber="426">
|
||
two pairs in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9B3FC481031FC3B" bold="true" box="[1579,1591,897,917]" pageId="12" pageNumber="426">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF58E6CF9FCFC481076FC3B" bold="true" box="[1636,1648,897,917]" pageId="12" pageNumber="426">9</emphasis>
|
||
; short duct, ovoid ampulla; diverticulum smaller than ampulla, and lateral to ental part of duct
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFC221191FB98" box="[147,1943,1003,1078]" gridrow="3" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFC2217F9FB98" box="[147,511,1003,1078]" gridcol="0" gridrow="3" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFC221708FBB5" authority="Fragoso, 1988 Chajul, Chis." authorityName="Fragoso" authorityYear="1988" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="maya">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFC2216DBFBAE" box="[147,221,1003,1024]" italics="true" pageId="12" pageNumber="426">L. maya</emphasis>
|
||
Fragoso, 1988 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFC221510FB98" box="[546,790,1003,1078]" gridcol="1" gridrow="3" pageId="12" pageNumber="426">without glands; in some cases distended</td>
|
||
<td id="0A445DE9FFF5719FFCDFFC2215BBFB98" box="[839,957,1003,1078]" gridcol="2" gridrow="3" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFC241559FBAF" bold="true" box="[839,863,1005,1025]" pageId="12" pageNumber="426">14</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FC2212E0FB98" box="[1004,1254,1003,1078]" gridcol="3" gridrow="3" pageId="12" pageNumber="426">
|
||
present, laminar, starting small in
|
||
<emphasis id="C5D1EAC5FFF58E6CFBA5FBCE1277FBB5" bold="true" box="[1085,1137,1031,1051]" pageId="12" pageNumber="426">16,17</emphasis>
|
||
, and gradually increasing size
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFC221358FB98" box="[1298,1374,1003,1078]" gridcol="4" gridrow="3" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFC25132CFBAE" bold="true" box="[1298,1322,1004,1024]" pageId="12" pageNumber="426">11</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CFAADFC25134BFBAE" bold="true" box="[1333,1357,1004,1024]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFC221191FB98" box="[1463,1943,1003,1078]" gridcol="5" gridrow="3" pageId="12" pageNumber="426">
|
||
two pairs in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9B3FC251031FBAE" bold="true" box="[1579,1591,1004,1024]" pageId="12" pageNumber="426">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF58E6CF9FCFC251076FBAE" bold="true" box="[1636,1648,1004,1024]" pageId="12" pageNumber="426">9</emphasis>
|
||
; short duct, ovoidal ampulla; diverticulum smaller than ampulla, and lateral to ental part of duct
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFB9F1191FB12" box="[147,1943,1110,1212]" gridrow="4" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFB9F17F9FB12" box="[147,511,1110,1212]" gridcol="0" gridrow="4" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFB9F1708FB28" authority="Fragoso & Rojas, 2016 Chajul, Chis." authorityName="Fragoso & Rojas" authorityYear="2016" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="notosetosus">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFB9F171EFBC5" box="[147,280,1110,1131]" italics="true" pageId="12" pageNumber="426">L. notosetosus</emphasis>
|
||
Fragoso & Rojas, 2016 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFB9F1510FB12" box="[546,790,1110,1212]" gridcol="1" gridrow="4" pageId="12" pageNumber="426">
|
||
without glands; in
|
||
<emphasis id="C5D1EAC5FFF58E6CFD48FB9E1500FBC5" bold="true" box="[720,774,1111,1131]" pageId="12" pageNumber="426">13-18</emphasis>
|
||
vascularized and in some cases distended
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFCDFFB9F15BBFB12" box="[839,957,1110,1212]" gridcol="2" gridrow="4" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFB9E1584FBC5" bold="true" box="[839,898,1111,1132]" pageId="12" pageNumber="426">
|
||
18
|
||
<emphasis id="C5D1EAC5FFF58E6CFCC7FB9E156CFBC2" bold="true" box="[863,874,1111,1132]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
20
|
||
</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FB9F12E0FB12" box="[1004,1254,1110,1212]" gridcol="3" gridrow="4" pageId="12" pageNumber="426">
|
||
present, laminar, small, starting in
|
||
<emphasis id="C5D1EAC5FFF58E6CFBC8FBBB1296FB28" bold="true" box="[1104,1168,1137,1158]" pageId="12" pageNumber="426">
|
||
20
|
||
<emphasis id="C5D1EAC5FFF58E6CFBFFFBB81275FB28" bold="true" box="[1127,1139,1137,1158]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
22
|
||
</emphasis>
|
||
reaching maximal size two, three segments backwards
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFB9F1358FB12" box="[1298,1374,1110,1212]" gridcol="4" gridrow="4" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFB9E132CFBC5" bold="true" box="[1298,1322,1111,1131]" pageId="12" pageNumber="426">11</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CFAADFB9E134BFBC5" bold="true" box="[1333,1357,1111,1131]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFB9F1191FB12" box="[1463,1943,1110,1212]" gridcol="5" gridrow="4" pageId="12" pageNumber="426">
|
||
two pairs in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9B3FB9E1031FBC5" bold="true" box="[1579,1591,1111,1131]" pageId="12" pageNumber="426">8</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CF9DAFB9E1048FBC5" bold="true" box="[1602,1614,1111,1131]" pageId="12" pageNumber="426">9</emphasis>
|
||
; diverticulum size ½ than ampulla, and lateral to ectal part of duct
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFB151191FA89" box="[147,1943,1244,1319]" gridrow="5" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFB1517F9FA89" box="[147,511,1244,1319]" gridcol="0" gridrow="5" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFB151708FAA2" authority="Fragoso, 1988 Chajul, Chis." authorityName="Fragoso" authorityYear="1988" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="parvus">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFB1516ECFB5F" box="[147,234,1244,1265]" italics="true" pageId="12" pageNumber="426">L. parvus</emphasis>
|
||
Fragoso, 1988 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFB151510FA89" box="[546,790,1244,1319]" gridcol="1" gridrow="5" pageId="12" pageNumber="426">
|
||
without glands; in
|
||
<emphasis id="C5D1EAC5FFF58E6CFD48FB141502FB5F" bold="true" box="[720,772,1245,1266]" pageId="12" pageNumber="426">
|
||
7
|
||
<emphasis id="C5D1EAC5FFF58E6CFD43FB1414E1FB5C" bold="true" box="[731,743,1245,1266]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
14
|
||
</emphasis>
|
||
highly vascularized
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFCDFFB1515BBFA89" box="[839,957,1244,1319]" gridcol="2" gridrow="5" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFB171559FB5C" bold="true" box="[839,863,1246,1266]" pageId="12" pageNumber="426">14</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FB1512E0FA89" box="[1004,1254,1244,1319]" gridcol="3" gridrow="5" pageId="12" pageNumber="426">
|
||
present, starting large and bifid in
|
||
<emphasis id="C5D1EAC5FFF58E6CFBAEFB31124BFAA2" bold="true" box="[1078,1101,1272,1292]" pageId="12" pageNumber="426">15</emphasis>
|
||
; from
|
||
<emphasis id="C5D1EAC5FFF58E6CFB14FB3112CBFAA2" bold="true" box="[1164,1229,1271,1292]" pageId="12" pageNumber="426">
|
||
20
|
||
<emphasis id="C5D1EAC5FFF58E6CFB3CFB3E12B6FAA2" bold="true" box="[1188,1200,1271,1292]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
22
|
||
</emphasis>
|
||
laminar
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFB151358FA89" box="[1298,1374,1244,1319]" gridcol="4" gridrow="5" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFB14132CFB5F" bold="true" box="[1298,1322,1245,1265]" pageId="12" pageNumber="426">11</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CFAADFB14134BFB5F" bold="true" box="[1333,1357,1245,1265]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFB151191FA89" box="[1463,1943,1244,1319]" gridcol="5" gridrow="5" pageId="12" pageNumber="426">
|
||
two pairs in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9B3FB141031FB5F" bold="true" box="[1579,1591,1245,1265]" pageId="12" pageNumber="426">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF58E6CF9FCFB141076FB5F" bold="true" box="[1636,1648,1245,1265]" pageId="12" pageNumber="426">9</emphasis>
|
||
; short duct and one fold ampulla; diverticulum of the same size than ampulla and lateral to ental part of duct
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFA8E1191FA3C" box="[147,1943,1351,1426]" gridrow="6" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFA8E17F9FA3C" box="[147,511,1351,1426]" gridcol="0" gridrow="6" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFA8E1708FAD9" authority="Fragoso, 1988 Chajul, Chis." authorityName="Fragoso" authorityYear="1988" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="riparius">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFA8E16F0FAF2" box="[147,246,1351,1372]" italics="true" pageId="12" pageNumber="426">L. riparius</emphasis>
|
||
Fragoso, 1988 Chajul, Chis.
|
||
</taxonomicName>
|
||
MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFA8E1510FA3C" box="[546,790,1351,1426]" gridcol="1" gridrow="6" pageId="12" pageNumber="426">without glands; in some cases distended</td>
|
||
<td id="0A445DE9FFF5719FFCDFFA8E15BBFA3C" box="[839,957,1351,1426]" gridcol="2" gridrow="6" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFA801558FAF3" bold="true" box="[839,862,1353,1373]" pageId="12" pageNumber="426">14</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FA8E12E0FA3C" box="[1004,1254,1351,1426]" gridcol="3" gridrow="6" pageId="12" pageNumber="426">
|
||
present, laminar, starting small in
|
||
<emphasis id="C5D1EAC5FFF58E6CFBA5FAAA1271FAD9" bold="true" box="[1085,1143,1378,1399]" pageId="12" pageNumber="426">
|
||
15
|
||
<emphasis id="C5D1EAC5FFF58E6CFBCCFAAB1259FAD9" bold="true" box="[1108,1119,1378,1399]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
18
|
||
</emphasis>
|
||
, and gradually increasing size
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFA8E1358FA3C" box="[1298,1374,1351,1426]" gridcol="4" gridrow="6" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFA81132CFAF2" bold="true" box="[1298,1322,1352,1372]" pageId="12" pageNumber="426">11</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF58E6CFAADFA81134BFAF2" bold="true" box="[1333,1357,1352,1372]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFA8E1191FA3C" box="[1463,1943,1351,1426]" gridcol="5" gridrow="6" pageId="12" pageNumber="426">
|
||
two pairs in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9B3FA811031FAF2" bold="true" box="[1579,1591,1352,1372]" pageId="12" pageNumber="426">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF58E6CF9FCFA811076FAF2" bold="true" box="[1636,1648,1352,1372]" pageId="12" pageNumber="426">9</emphasis>
|
||
; short duct, ovoid ampulla; diverticulum smaller than ampulla, and lateral to ental part of duct
|
||
</td>
|
||
</tr>
|
||
<tr id="49953495FFF5719FFF0BFA7B1191F9B6" box="[147,1943,1458,1560]" gridrow="7" pageId="12" pageNumber="426">
|
||
<th id="0A445DE9FFF5719FFF0BFA7B17F9F9B6" box="[147,511,1458,1560]" gridcol="0" gridrow="7" pageId="12" pageNumber="426">
|
||
<taxonomicName id="30A54D54FFF58E6CFF0BFA7B16F7FA69" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[147,241,1458,1479]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="426" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF0BFA7B16F7FA69" box="[147,241,1458,1479]" italics="true" pageId="12" pageNumber="426">L. sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFF58E6CFF6FFA7A1744FA69" box="[247,322,1459,1479]" pageId="12" pageNumber="426" rank="species">
|
||
<emphasis id="C5D1EAC5FFF58E6CFF6FFA7A1744FA69" bold="true" box="[247,322,1459,1479]" pageId="12" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Comalcalco and Teapa, Tab. MEXICO
|
||
</th>
|
||
<td id="0A445DE9FFF5719FFDBAFA7B1510F9B6" box="[546,790,1458,1560]" gridcol="1" gridrow="7" pageId="12" pageNumber="426">
|
||
without glands; in
|
||
<emphasis id="C5D1EAC5FFF58E6CFD48FA7A1502FA69" bold="true" box="[720,772,1459,1480]" pageId="12" pageNumber="426">
|
||
8
|
||
<emphasis id="C5D1EAC5FFF58E6CFD43FA7A14E1FA66" bold="true" box="[731,743,1459,1480]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
12
|
||
</emphasis>
|
||
vascularized; in
|
||
<emphasis id="C5D1EAC5FFF58E6CFD22FA0714FDFA4C" bold="true" box="[698,763,1485,1506]" pageId="12" pageNumber="426">
|
||
10
|
||
<emphasis id="C5D1EAC5FFF58E6CFD4AFA0414D8FA4C" bold="true" box="[722,734,1485,1506]" italics="true" pageId="12" pageNumber="426">–</emphasis>
|
||
11
|
||
</emphasis>
|
||
dilated
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFCDFFA7B15BBF9B6" box="[839,957,1458,1560]" gridcol="2" gridrow="7" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFCDFFA7D1559FA66" bold="true" box="[839,863,1460,1480]" pageId="12" pageNumber="426">14</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFC74FA7B12E0F9B6" box="[1004,1254,1458,1560]" gridcol="3" gridrow="7" pageId="12" pageNumber="426">
|
||
present, laminar, starting with maximal size in
|
||
<emphasis id="C5D1EAC5FFF58E6CFB2DFA0712CBFA4C" bold="true" box="[1205,1229,1486,1506]" pageId="12" pageNumber="426">15</emphasis>
|
||
; two-four segments after diminishing to normal size
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA8AFA7B1358F9B6" box="[1298,1374,1458,1560]" gridcol="4" gridrow="7" pageId="12" pageNumber="426">
|
||
<emphasis id="C5D1EAC5FFF58E6CFA8AFA7D132CFA66" bold="true" box="[1298,1322,1460,1480]" pageId="12" pageNumber="426">12</emphasis>
|
||
</td>
|
||
<td id="0A445DE9FFF5719FFA2FFA7B1191F9B6" box="[1463,1943,1458,1560]" gridcol="5" gridrow="7" pageId="12" pageNumber="426">
|
||
unpaired; anterior in
|
||
<emphasis id="C5D1EAC5FFF58E6CF9E3FA7A1081FA69" bold="true" box="[1659,1671,1459,1479]" pageId="12" pageNumber="426">8</emphasis>
|
||
or
|
||
<emphasis id="C5D1EAC5FFF58E6CF93DFA7A10B7FA69" bold="true" box="[1701,1713,1459,1479]" pageId="12" pageNumber="426">7</emphasis>
|
||
and posterior in
|
||
<emphasis id="C5D1EAC5FFF58E6CF8CDFA7A1167FA69" bold="true" box="[1877,1889,1459,1479]" pageId="12" pageNumber="426">8</emphasis>
|
||
or
|
||
<emphasis id="C5D1EAC5FFF58E6CF81DFA7A1197FA69" bold="true" box="[1925,1937,1459,1479]" pageId="12" pageNumber="426">9</emphasis>
|
||
two or one lateral diverticula emerging from ental region of duct
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFF48E6DFF5FFF5E17C5FD90" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="427">
|
||
<emphasis id="C5D1EAC5FFF48E6DFF5FFF5E1736FF1E" bold="true" box="[199,304,151,176]" pageId="13" pageNumber="427">Internal.</emphasis>
|
||
SEptum 5/6 thin, Slightly muScular, in SomE individualS almoSt mEmbranouS; SEpta
|
||
<date id="831B1017FFF48E6DFAB4FF5E139AFF1E" box="[1324,1436,151,176]" pageId="13" pageNumber="427" value="2012-07-06" valueMax="2012-11-06" valueMin="2012-07-06">6/7–11/12</date>
|
||
muScular; SEptum 12/13 thin but Still muScular; SEptum 13/14 and backwardS mEmbranouS and thin; SEpta
|
||
<date id="831B1017FFF48E6DFADFFF7516B2FF56" pageId="13" pageNumber="427" value="2011-06-05" valueMax="2011-10-05" valueMin="2011-06-05">5/6–10/ 11</date>
|
||
, and in SomE individualS alSo 11/12, funnEl-ShapEd and Slightly imbricatEd. OnE largE rEctangular gizzard in
|
||
<emphasis id="C5D1EAC5FFF48E6DFA1FFF161393FF56" bold="true" box="[1415,1429,223,248]" pageId="13" pageNumber="427">5</emphasis>
|
||
. Gizzard dimEnSionS (lEngth by width): 0.36 by
|
||
<quantity id="305D9B32FFF48E6DFD5DFECD152AFEB2" box="[709,812,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="427" unit="mm" value="0.64">0.64 mm</quantity>
|
||
(
|
||
<typeStatus id="281E8875FFF48E6DFCA7FECD15AEFEB3" box="[831,936,260,285]" pageId="13" pageNumber="427" type="holotype">holotypE</typeStatus>
|
||
), 0.54 by
|
||
<quantity id="305D9B32FFF48E6DFBB9FECD128EFEB2" box="[1057,1160,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="13" pageNumber="427" unit="mm" value="0.65">0.65 mm</quantity>
|
||
(
|
||
<typeStatus id="281E8875FFF48E6DFB02FECD12F8FEB2" box="[1178,1278,260,284]" pageId="13" pageNumber="427" type="paratype">ParatypE</typeStatus>
|
||
IEOL 4836). ESophaguS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFEA3FEEE1775FEEE" bold="true" box="[315,371,295,320]" pageId="13" pageNumber="427">9–12</emphasis>
|
||
without intErnal lamEllaE; vaScularization in ESophagEal wallS variablE, in SomE individualS prESEnt in SEgmEntS
|
||
<quantity id="305D9B32FFF48E6DFEE6FE8517D2FECB" box="[382,468,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="13" pageNumber="427" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">
|
||
<emphasis id="C5D1EAC5FFF48E6DFEE6FE8517B0FECB" bold="true" box="[382,438,332,357]" pageId="13" pageNumber="427">8–10</emphasis>
|
||
in
|
||
</quantity>
|
||
othErS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFDDFFE851479FECB" bold="true" box="[583,639,332,357]" pageId="13" pageNumber="427">9–12</emphasis>
|
||
; dilatationS rESEmbling pouchES fairly conStant in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFAB4FE85134EFECB" bold="true" box="[1324,1352,332,357]" pageId="13" pageNumber="427">10</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFA19FE85139CFECB" bold="true" box="[1409,1434,332,357]" pageId="13" pageNumber="427">11</emphasis>
|
||
(obSErvEd in 7 individualS, including
|
||
<typeStatus id="281E8875FFF48E6DFDDFFEA614A8FE26" box="[583,686,367,392]" pageId="13" pageNumber="427" type="holotype">holotypE</typeStatus>
|
||
); thEy could corrESpond to ESophagEal glandS, mainly bEcauSE in SomE individualS thEy arE highly vaScularizEd. IntEStinE Starting in 13/14 (6 ind.) (
|
||
<figureCitation id="6F9E2A52FFF48E6DFBF9FE5D12C6FE02" box="[1121,1216,404,429]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4A</figureCitation>
|
||
). A laminar dorSal typhloSolE StartS abruptly with maximal SizE in SEgmEnt
|
||
<emphasis id="C5D1EAC5FFF48E6DFC88FE7E152AFE7E" bold="true" box="[784,812,439,464]" pageId="13" pageNumber="427">15</emphasis>
|
||
, diminiShing 2–3 SEgmEntS poStEriorly to a normal SizE (not morE than ½ of lumEn) until SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFD19FE15149BFE5B" bold="true" box="[641,669,476,501]" pageId="13" pageNumber="427">20</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF48E6DFD34FE1514CEFE5B" bold="true" box="[684,712,476,501]" pageId="13" pageNumber="427">24</emphasis>
|
||
(thrEE ind.) or
|
||
<emphasis id="C5D1EAC5FFF48E6DFCEAFE151588FE5B" bold="true" box="[882,910,476,501]" pageId="13" pageNumber="427">31</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF48E6DFC05FE1515BFFE5B" bold="true" box="[925,953,476,501]" pageId="13" pageNumber="427">34</emphasis>
|
||
(thrEE ind.), whErE it abruptly dEcrEaSES in SizE to a wEak ribbon-likE StructurE; thiS ribbon gradually diminiShES in SizE, Ending 20 or 39 SEgmEntS bEforE anuS. LatEral typhloSolES abSEnt.
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFF48E6DFF5FFD801373FBF4" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="427">
|
||
SinglE dorSal vESSEl viSiblE throughout; in
|
||
<emphasis id="C5D1EAC5FFF48E6DFD3BFD8014B9FDCC" bold="true" box="[675,703,585,610]" pageId="13" pageNumber="427">15</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFD6DFD801517FDCC" bold="true" box="[757,785,585,610]" pageId="13" pageNumber="427">16</emphasis>
|
||
conSpicuouSly EnlargEd, and coincidEnt with largEr dorSal typhloSolE (
|
||
<figureCitation id="6F9E2A52FFF48E6DFE84FDA4177EFD28" box="[284,376,621,646]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4A</figureCitation>
|
||
); in two individualS (IEOL 4832, 4837) doublE in
|
||
<emphasis id="C5D1EAC5FFF48E6DFC21FDA415D3FD28" bold="true" box="[953,981,621,646]" pageId="13" pageNumber="427">16</emphasis>
|
||
and with latEral vESSElS full of blood in 15/16 and 16/17 (
|
||
<figureCitation id="6F9E2A52FFF48E6DFEF8FD5817BFFD04" box="[352,441,657,682]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4B</figureCitation>
|
||
). Supra-ESophagEal vESSEl clEarly viSiblE in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFBB8FD58123FFD04" bold="true" box="[1056,1081,657,682]" pageId="13" pageNumber="427">11</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFBEBFD581289FD04" bold="true" box="[1139,1167,657,682]" pageId="13" pageNumber="427">12</emphasis>
|
||
; in
|
||
<emphasis id="C5D1EAC5FFF48E6DFB25FD5812DFFD04" bold="true" box="[1213,1241,657,682]" pageId="13" pageNumber="427">10</emphasis>
|
||
SEEn only in onE individual. LatEral hEartS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFE4FFD7C17E3FD60" bold="true" box="[471,485,693,718]" pageId="13" pageNumber="427">9</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFD86FD7C143CFD60" bold="true" box="[542,570,693,718]" pageId="13" pageNumber="427">10</emphasis>
|
||
; latEro-ESophagEal hEartS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFC18FD7C159DFD60" bold="true" box="[896,923,693,718]" pageId="13" pageNumber="427">11</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFC4BFD7C15E9FD60" bold="true" box="[979,1007,693,718]" pageId="13" pageNumber="427">12</emphasis>
|
||
. VEntral vESSEl prESEnt. Extra pariEtal latEral-vEntral vESSElS prESEnt, viSiblE EvEn from thE ExtErior; thEy run ovEr
|
||
<emphasis id="C5D1EAC5FFF48E6DFC7DFD13120FFD5F" box="[997,1033,730,753]" italics="true" pageId="13" pageNumber="427">BC</emphasis>
|
||
, Starting in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFB65FD10131FFD5C" bold="true" box="[1277,1305,729,754]" pageId="13" pageNumber="427">14</emphasis>
|
||
(1 ind.),
|
||
<emphasis id="C5D1EAC5FFF48E6DFA18FD10139AFD5C" bold="true" box="[1408,1436,729,754]" pageId="13" pageNumber="427">16</emphasis>
|
||
(1) (
|
||
<figureCitation id="6F9E2A52FFF48E6DFF5FFD341718FCB8" box="[199,286,765,790]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4C</figureCitation>
|
||
),
|
||
<emphasis id="C5D1EAC5FFF48E6DFEABFD341749FCB8" bold="true" box="[307,335,765,790]" pageId="13" pageNumber="427">17</emphasis>
|
||
(1),
|
||
<emphasis id="C5D1EAC5FFF48E6DFE1CFD3417A6FCB8" bold="true" box="[388,416,765,790]" pageId="13" pageNumber="427">18</emphasis>
|
||
(3) or
|
||
<emphasis id="C5D1EAC5FFF48E6DFE75FD34140FFCB8" bold="true" box="[493,521,765,790]" pageId="13" pageNumber="427">19</emphasis>
|
||
(1), bEing Still conSpicuouS and viSiblE in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFBCBFD341269FCB8" bold="true" box="[1107,1135,765,790]" pageId="13" pageNumber="427">25</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF48E6DFBE5FD34129FFCB8" bold="true" box="[1149,1177,765,790]" pageId="13" pageNumber="427">27</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF48E6DFB3FFD3412C5FCB8" bold="true" box="[1191,1219,765,790]" pageId="13" pageNumber="427">28</emphasis>
|
||
; in two individualS Still viSiblE in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFE3CFCE817C6FC94" bold="true" box="[420,448,801,826]" pageId="13" pageNumber="427">40</emphasis>
|
||
,
|
||
<emphasis id="C5D1EAC5FFF48E6DFE56FCE817ECFC94" bold="true" box="[462,490,801,826]" pageId="13" pageNumber="427">41</emphasis>
|
||
. A pair of infra-ESophagEal vESSElS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFC0FFCE815B5FC94" bold="true" box="[919,947,801,826]" pageId="13" pageNumber="427">10</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFC71FCE81204FC94" bold="true" box="[1001,1026,801,826]" pageId="13" pageNumber="427">11</emphasis>
|
||
, viSiblE only in thE
|
||
<typeStatus id="281E8875FFF48E6DFB78FCE81340FC94" box="[1248,1350,801,826]" pageId="13" pageNumber="427" type="holotype">holotypE</typeStatus>
|
||
. Holoic without vESiclES. ThE ExonEphric, and apparEntly StomatE holonEphridia arE vEntro-pariEtal from SEgmEnt
|
||
<emphasis id="C5D1EAC5FFF48E6DFAD0FC8C1350FCF0" bold="true" box="[1352,1366,837,862]" pageId="13" pageNumber="427">7</emphasis>
|
||
; from thiS SEgmEnt nEphridia gradually movE towardS latEral-pariEtES, bEing complEtEly pariEtal from
|
||
<emphasis id="C5D1EAC5FFF48E6DFB5DFCA012E7FC2C" bold="true" box="[1221,1249,873,898]" pageId="13" pageNumber="427">13</emphasis>
|
||
backwardS (
|
||
<figureCitation id="6F9E2A52FFF48E6DFAF6FCA016BBFC08" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4A</figureCitation>
|
||
). NEphroStomES and nEphroporES not SEEn. Holandric. IridiScEnt malE funnElS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFBC3FC441271FC08" bold="true" box="[1115,1143,909,934]" pageId="13" pageNumber="427">10</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFB29FC4412CCFC08" bold="true" box="[1201,1226,909,934]" pageId="13" pageNumber="427">11</emphasis>
|
||
; tEStES difficult to SEE; rEcordEd only in two individualS, in SEgmEnt
|
||
<emphasis id="C5D1EAC5FFF48E6DFD55FC7814E0FC64" bold="true" box="[717,742,945,970]" pageId="13" pageNumber="427">11</emphasis>
|
||
. MalE gonoduct zigzagging, doublE, Supra-pariEtal, obSErvEd only in two individualS in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFDA4FC1C1484FC40" bold="true" box="[572,642,981,1006]" pageId="13" pageNumber="427">13–16</emphasis>
|
||
. OnE pair of SEminal vESiclES in
|
||
<emphasis id="C5D1EAC5FFF48E6DFB9BFC1C1220FC40" bold="true" box="[1027,1062,981,1006]" pageId="13" pageNumber="427">12,</emphasis>
|
||
Small and partially covEring thE ESophagouS. Two pairS of Small tubular proStatES in
|
||
<emphasis id="C5D1EAC5FFF48E6DFD4FFC3014F5FBBC" bold="true" box="[727,755,1017,1042]" pageId="13" pageNumber="427">17</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFCB0FC301542FBBC" bold="true" box="[808,836,1017,1042]" pageId="13" pageNumber="427">19</emphasis>
|
||
, with onE fold, limitEd to itS SEgmEntS or ExtEnding to adjacEnt antErior or poStErior SEgmEnt (
|
||
<figureCitation id="6F9E2A52FFF48E6DFDC9FBD414ADFB98" box="[593,683,1053,1078]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4C</figureCitation>
|
||
); in two individualS Spirally coilEd; glandular part 2–4 timES largEr than muScular rEgion. PEnial SEtaE folliclES fixEd to floor and latEral wallS, limitEd to thEir rESpEctivE SEgmEntS.
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFF48E6DFF5FFBAC15DAFA6F" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="427">
|
||
PairEd ovariES and fEmalE funnElS in
|
||
<emphasis id="C5D1EAC5FFF48E6DFDFCFBAC1486FBD0" bold="true" box="[612,640,1125,1150]" pageId="13" pageNumber="427">13</emphasis>
|
||
. OvariES fan or (mainly) buSh-ShapEd, vEry largE and with conSpicuouS EggS not arrangEd in rowS; fEmalE funnElS fixEd to floor and vEry cloSE to mid-vEntral linE. Only two SpErmathEcaE, onE opEning in 7/8 and thE othEr onE in 8/9. AntErior and poStErior SpErmathEcaE locatEd in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF48E6DFAA4FB64134CFB68" bold="true" box="[1340,1354,1197,1222]" pageId="13" pageNumber="427">8</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFA1FFB641393FB68" bold="true" box="[1415,1429,1197,1222]" pageId="13" pageNumber="427">9</emphasis>
|
||
, rESpEctivEly (6 ind.); or locatEd in
|
||
<emphasis id="C5D1EAC5FFF48E6DFDAAFB181446FB44" bold="true" box="[562,576,1233,1258]" pageId="13" pageNumber="427">7</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF48E6DFDE6FB18148AFB44" bold="true" box="[638,652,1233,1258]" pageId="13" pageNumber="427">9</emphasis>
|
||
(1 ind.); or both locatEd in
|
||
<emphasis id="C5D1EAC5FFF48E6DFC45FB1815EDFB44" bold="true" box="[989,1003,1233,1258]" pageId="13" pageNumber="427">8</emphasis>
|
||
(1 ind.). Two
|
||
<typeStatus id="281E8875FFF48E6DFB03FB1B12D0FB44" box="[1179,1238,1234,1258]" pageId="13" pageNumber="427">typES</typeStatus>
|
||
of SpErmathEcaE rEcordEd: in two individualS (
|
||
<typeStatus id="281E8875FFF48E6DFE7BFB3F1457FAA0" box="[483,593,1270,1294]" pageId="13" pageNumber="427" type="paratype">paratypES</typeStatus>
|
||
4830 and 4832) Each SpErmathEca with a SinglE, latEral divErticulum (
|
||
<figureCitation id="6F9E2A52FFF48E6DFAF7FB3C16BDFA9C" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4C</figureCitation>
|
||
);
|
||
<typeStatus id="281E8875FFF48E6DFF4BFAD01733FA9C" box="[211,309,1305,1330]" pageId="13" pageNumber="427" type="holotype">holotypE</typeStatus>
|
||
and othEr
|
||
<typeStatus id="281E8875FFF48E6DFE2DFAD31426FA9C" box="[437,544,1306,1330]" pageId="13" pageNumber="427" type="paratype">paratypES</typeStatus>
|
||
with two latEral and oppoSitE divErticula pEr SpErmathEca (
|
||
<figureCitation id="6F9E2A52FFF48E6DFB4DFAD01335FA9C" box="[1237,1331,1305,1330]" captionStart="FIGURE 4" captionStartId="9.[151,250,1818,1840]" captionTargetBox="[182,1376,205,1764]" captionTargetId="figure@9.[164,1400,193,1781]" captionTargetPageId="9" captionText="FIGURE 4. Lavellodrilus sheylae sp. nov. Internal. A. Dorsal view of segments 11–18 (Holotype IEOL 4831; scale 1 mm). B. Dorsal view of segments 15–17 from an individual with two diverticula spermathecae (Paratype IEOL 4837; scale 0.76 mm), showing double dorsal vessel in segment 16 where intestine was cut down to look for typhlosole. C. Parietes of segments 16– 19, showing parietal lateral-ventral vessels and prostates of 17 (P17) and 19 (P19) (Paratype IEOL 4837; scale 0.86 mm). D. Dorso-lateral view of single diverticulum spermatheca of segment 8 (Paratype IEOL 4830; scale 200 µm). E. Dorsal view of two diverticula spermatheca of segment 8 (Paratype IEOL 4833; scale 200 µm). A= ampulla, Di= diverticulum, Du= duct, Dv= dorsal vessel, E=esophagus, H= holonephridium, I= intestine, Lv= parietal lateral ventral vessels." httpUri="https://zenodo.org/record/1446945/files/figure.png" pageId="13" pageNumber="427">Fig. 4D</figureCitation>
|
||
). In both
|
||
<typeStatus id="281E8875FFF48E6DFF0FFAF716D4FAF8" box="[151,210,1342,1366]" pageId="13" pageNumber="427">typES</typeStatus>
|
||
of SpErmathEcaE, divErticula EmErgE from Ental rEgion of duct; duct aS widE aS ampulla or Slightly narrowEr, oriEntEd along thE SamE axiS. DivErticula with SpErm inSidE in all but
|
||
<typeStatus id="281E8875FFF48E6DFC3EFAAB1200FAD4" box="[934,1030,1378,1402]" pageId="13" pageNumber="427" type="paratype">paratypE</typeStatus>
|
||
4832, half or lESS aS half aS widE aS duct or ampulla. MEaSurEd lEngthS of SpErmathEcaE, onE divErticulum
|
||
<typeStatus id="281E8875FFF48E6DFC34FA4F15E7FA30" box="[940,993,1414,1438]" pageId="13" pageNumber="427">typE</typeStatus>
|
||
: antErior 0.34,
|
||
<quantity id="305D9B32FFF48E6DFB10FA4F12E8FA33" box="[1160,1262,1414,1438]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="427" unit="mm" value="0.38">0.38 mm</quantity>
|
||
, poStErior 0.41,
|
||
<quantity id="305D9B32FFF48E6DFF0FFA6316F8FA6F" box="[151,254,1450,1474]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="427" unit="mm" value="0.44">0.44 mm</quantity>
|
||
; two divErticula
|
||
<typeStatus id="281E8875FFF48E6DFE21FA6317E8FA6C" box="[441,494,1450,1474]" pageId="13" pageNumber="427">typE</typeStatus>
|
||
: antErior
|
||
<quantity id="305D9B32FFF48E6DFDCEFA6014BAFA6F" box="[598,700,1449,1474]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="13" pageNumber="427" unit="mm" value="0.53">0.53 mm</quantity>
|
||
, poStErior 0.50,
|
||
<quantity id="305D9B32FFF48E6DFCE9FA6015DEFA6F" box="[881,984,1449,1474]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="13" pageNumber="427" unit="mm" value="0.56">0.56 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFBF655CFFF48E6DFF5FFA0414B1F9A4" pageId="13" pageNumber="427" type="etymology">
|
||
<paragraph id="F71A36D7FFF48E6DFF5FFA0414B1F9A4" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="427">
|
||
<emphasis id="C5D1EAC5FFF48E6DFF5FFA041740FA48" bold="true" box="[199,326,1485,1510]" pageId="13" pageNumber="427">Etymology</emphasis>
|
||
. ThE namE of thE SpEciES iS dEdicatEd to Dr. ShEyla UribE, in rEcognition of hEr EnthuSiaSm and EffortS to Study thE EarthwormS of
|
||
<collectingRegion id="3561F835FFF48E6DFD82FA381473F9A4" box="[538,629,1521,1546]" country="Mexico" name="Tabasco" pageId="13" pageNumber="427">TabaSco</collectingRegion>
|
||
StatE.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFBF655CFFF48E6EFF5FF9DC12E3FD06" lastPageId="14" lastPageNumber="428" pageId="13" pageNumber="427" type="discussion">
|
||
<paragraph id="F71A36D7FFF48E6EFF5FF9DC1347FF59" blockId="13.[151,1437,151,2014]" lastBlockId="14.[151,1437,151,680]" lastPageId="14" lastPageNumber="428" pageId="13" pageNumber="427">
|
||
<emphasis id="C5D1EAC5FFF48E6DFF5FF9DC173DF980" bold="true" box="[199,315,1557,1582]" pageId="13" pageNumber="427">Remarks.</emphasis>
|
||
<taxonomicName id="30A54D54FFF48E6DFEDAF9DF17B2F983" authority="Fragoso & Rojas, 2018" authorityName="Fragoso & Rojas" authorityYear="2018" box="[322,436,1557,1581]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="13" pageNumber="427" phylum="Annelida" rank="species" species="sheylae" status="sp. nov.">
|
||
<emphasis id="C5D1EAC5FFF48E6DFEDAF9DF17B2F983" box="[322,436,1557,1581]" italics="true" pageId="13" pageNumber="427">L. sheylae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEE257BEFFF48E6DFE23F9DC1413F980" box="[443,533,1557,1582]" pageId="13" pageNumber="427" rank="species">
|
||
<emphasis id="C5D1EAC5FFF48E6DFE23F9DC1413F980" bold="true" box="[443,533,1557,1582]" pageId="13" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
iS uniquE in thE gEnuS by thE unpairEd condition of antErior (diScharging in 7/8) and poStErior (diScharging in 8/9) SpErmathEcaE (
|
||
<tableCitation id="BA27036CFFF48E6DFD72F9F0154CF9FC" box="[746,842,1593,1618]" captionStart="TABLE 2" captionStartId="11.[147,229,599,620]" captionText="TABLE 2. Morphological comparisons among the currently recognized Lavellodrilus species from Mexico. Data obtained from Fragoso (1988, 1991) and Fragoso and Rojas (2016)" pageId="13" pageNumber="427">TablE 2</tableCitation>
|
||
), a condition not obSErvEd in any of thE holoic acanthodrilinES prESEnt in
|
||
<collectingRegion id="3561F835FFF48E6DFE56F9941421F9D8" box="[462,551,1629,1654]" country="Mexico" name="Mexico" pageId="13" pageNumber="427">
|
||
<collectingCountry id="8FB27647FFF48E6DFE56F9941421F9D8" box="[462,551,1629,1654]" name="Mexico" pageId="13" pageNumber="427">MExico</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="8FB27647FFF48E6DFDAFF9941474F9D8" box="[567,626,1629,1654]" name="Cuba" pageId="13" pageNumber="427">Cuba</collectingCountry>
|
||
and CEntral AmErican or thE holoic and mEroic diplocardinES from northErn
|
||
<collectingRegion id="3561F835FFF48E6DFE98F948175FF934" box="[256,345,1665,1690]" country="Mexico" name="Mexico" pageId="13" pageNumber="427">
|
||
<collectingCountry id="8FB27647FFF48E6DFE98F948175FF934" box="[256,345,1665,1690]" name="Mexico" pageId="13" pageNumber="427">MExico</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="8FB27647FFF48E6DFEFFF94B1799F934" box="[359,415,1666,1690]" name="United States of America" pageId="13" pageNumber="427">USA</collectingCountry>
|
||
and thE CaribbEan (
|
||
<bibRefCitation id="93344B26FFF48E6DFD17F94B159BF934" box="[655,925,1665,1690]" pageId="13" pageNumber="427" refId="ref6603" refString="Fragoso, C. & Rojas, P. (2016) Lavellodrilus notosetosus sp. nov. (Annelida, Crassiclitellata, Acanthodrilidae): a new Mexican earthworm with uncommon characters, revealed by a preliminary revision of subfamily Acanthodrilinae. Zootaxa, 4154 (2), 101 - 138. http: // doi. org / 10.11646 / zootaxa. 4154.2.1" type="journal article">FragoSo & RojaS 2016</bibRefCitation>
|
||
). Although unpairEd SpErmathEcaE arE quitE frEquEnt in SEvEral BEnhamiinaE acanthodrilid gEnEra (
|
||
<bibRefCitation id="93344B26FFF48E6DFC92F96C1599F910" box="[778,927,1701,1726]" pageId="13" pageNumber="427" refId="ref6354" refString="Csuzdi, C. S. (2010) A monograph of the Paleotropical Benhamiinae earthworms. Hungarian Natural History Museum & Hungarian Academy of Sciences, Budapest, 348 pp." type="book">CSuzdi 2010</bibRefCitation>
|
||
), thiS unpairEd condition of SpErmathEcaE in SuccESSivE SEgmEntS iS not obSErvEd in any of thE world acanthodrilinES rEcEntly rEviSEd by
|
||
<bibRefCitation id="93344B26FFF48E6DFB5EF90316E3F8A8" pageId="13" pageNumber="427" refId="ref6603" refString="Fragoso, C. & Rojas, P. (2016) Lavellodrilus notosetosus sp. nov. (Annelida, Crassiclitellata, Acanthodrilidae): a new Mexican earthworm with uncommon characters, revealed by a preliminary revision of subfamily Acanthodrilinae. Zootaxa, 4154 (2), 101 - 138. http: // doi. org / 10.11646 / zootaxa. 4154.2.1" type="journal article">FragoSo and RojaS (2016)</bibRefCitation>
|
||
. Only in thE caSE of thE South African SpEciES
|
||
<taxonomicName id="30A54D54FFF48E6DFCD2F92412FEF8A8" authority="Pickford 1937" authorityName="Pickford" authorityYear="1937" box="[842,1272,1773,1798]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="13" pageNumber="427" phylum="Annelida" rank="species" species="bavenda">
|
||
<emphasis id="C5D1EAC5FFF48E6DFCD2F9241238F8AB" box="[842,1086,1773,1797]" italics="true" pageId="13" pageNumber="427">Parachilota bavenda</emphasis>
|
||
<bibRefCitation id="93344B26FFF48E6DFBD7F92412FEF8A8" box="[1103,1272,1773,1798]" pageId="13" pageNumber="427" refId="ref7053" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. W. Heffer & Sons Ltd., Cambridge, 612 pp." type="book">Pickford 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
(with mESial SpErmathEcal porES) thE poStErior pairEd SpErmathEcaE arE SubStitutEd by a SinglE onE, whErEaS thE antErior pair iS conSErvEd (
|
||
<bibRefCitation id="93344B26FFF48E6DFE82F8FC17C5F8E0" box="[282,451,1845,1870]" pageId="13" pageNumber="427" refId="ref7053" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. W. Heffer & Sons Ltd., Cambridge, 612 pp." type="book">Pickford 1937</bibRefCitation>
|
||
). In thE caSE of thE mEroic acanthodrilidS from
|
||
<collectingRegion id="3561F835FFF48E6DFC6EF8FC124DF8E0" box="[1014,1099,1845,1870]" country="Mexico" name="Mexico" pageId="13" pageNumber="427">
|
||
<collectingCountry id="8FB27647FFF48E6DFC6EF8FC124DF8E0" box="[1014,1099,1845,1870]" name="Mexico" pageId="13" pageNumber="427">MExico</collectingCountry>
|
||
</collectingRegion>
|
||
and CEntral AmErica (gEnuS
|
||
<taxonomicName id="30A54D54FFF48E6DFF0FF890171EF8DF" authorityName="Michaelsen" authorityYear="1935" box="[151,280,1881,1905]" class="Clitellata" family="Acanthodrilidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="13" pageNumber="427" phylum="Annelida" rank="genus">
|
||
<emphasis id="C5D1EAC5FFF48E6DFF0FF890171EF8DF" box="[151,280,1881,1905]" italics="true" pageId="13" pageNumber="427">Ramiellona</emphasis>
|
||
</taxonomicName>
|
||
) thE loSS of thE pairEd condition of SpErmathEcaE haS bEEn obSErvEd in at lEaSt two SpEciES:
|
||
<taxonomicName id="30A54D54FFF48E6DFA83F8901705F83B" authorityName="Gates" authorityYear="1962" class="Clitellata" family="Acanthodrilidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="13" pageNumber="427" phylum="Annelida" rank="species" species="mexicana">
|
||
<emphasis id="C5D1EAC5FFF48E6DFA83F8901705F83B" italics="true" pageId="13" pageNumber="427">Ramiellona mexicana</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="93344B26FFF48E6DFE91F8B71789F838" box="[265,399,1917,1942]" pageId="13" pageNumber="427" refId="ref6730" refString="Gates, G. (1962) On some earthworms of Eisen's collections. Proceedings of the California Academy of Sciences, 4 (31), 185 - 225." type="journal article">GatES, 1962</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="93344B26FFF48E6DFE05F8B71419F838" box="[413,543,1917,1942]" pageId="13" pageNumber="427" refId="ref6730" refString="Gates, G. (1962) On some earthworms of Eisen's collections. Proceedings of the California Academy of Sciences, 4 (31), 185 - 225." type="journal article">GatES 1962</bibRefCitation>
|
||
) and anothEr tiny undEScribEd SpEciES from SouthErn
|
||
<collectingRegion id="3561F835FFF48E6DFBEFF8B412CAF838" box="[1143,1228,1917,1942]" country="Mexico" name="Mexico" pageId="13" pageNumber="427">
|
||
<collectingCountry id="8FB27647FFF48E6DFBEFF8B412CAF838" box="[1143,1228,1917,1942]" name="Mexico" pageId="13" pageNumber="427">MExico</collectingCountry>
|
||
</collectingRegion>
|
||
(
|
||
<bibRefCitation id="93344B26FFF48E6DFB43F8B717B3F814" pageId="13" pageNumber="427" refId="ref6573" refString="Fragoso, C. (1993) Les Peuplements de Uers de Terre dans L'Est et Sud-Est du Mexique. These, Universite Paris, Paris, 228 pp." type="book">
|
||
FragoSo 1993, aS "GEn.
|
||
<date id="831B1017FFF48E6DFF47F86B1722F814" box="[223,292,1954,1978]" pageId="13" pageNumber="427">
|
||
nov.
|
||
<specimenCount id="E1A3FD5EFFF48E6DFE8EF86B1722F814" box="[278,292,1954,1978]" pageId="13" pageNumber="427" type="generic">3</specimenCount>
|
||
</date>
|
||
Sp.
|
||
<date id="831B1017FFF48E6DFECBF86B17ACF814" box="[339,426,1953,1978]" pageId="13" pageNumber="427" value="2016-11">nov. 16</date>
|
||
"
|
||
</bibRefCitation>
|
||
). In thE firSt caSE (Similar but invErSE to thE pattErn obSErvEd in
|
||
<taxonomicName id="30A54D54FFF48E6DFB1FF86B1304F817" authorityName="Pickford" authorityYear="1937" box="[1159,1282,1953,1977]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="13" pageNumber="427" phylum="Annelida" rank="species" species="bavenda">
|
||
<emphasis id="C5D1EAC5FFF48E6DFB1FF86B1304F817" box="[1159,1282,1953,1977]" italics="true" pageId="13" pageNumber="427">P. bavenda</emphasis>
|
||
</taxonomicName>
|
||
), thErE iS onE antErior SpErmathEca with two divErticula and thErE arE two poStErior SpErmathEcaE, Each onE with onE divErticulum; in thE SEcond caSE thErE iS onE antErior SpErmathEca and onE poStErior, Each onE with two divErticula. It SEEmS that in SouthErn
|
||
<collectingRegion id="3561F835FFF78E6EFE36FF751405FF7B" box="[430,515,188,213]" country="Mexico" name="Mexico" pageId="14" pageNumber="428">
|
||
<collectingCountry id="8FB27647FFF78E6EFE36FF751405FF7B" box="[430,515,188,213]" name="Mexico" pageId="14" pageNumber="428">MExico</collectingCountry>
|
||
</collectingRegion>
|
||
thE pattErn of unpairEd SpErmathEcaE opEning to thE ExtErior through mESial porES haS bEEn dEvElopEd indEpEndEntly in SEvEral diffErEnt linEagES. WE will go dEEpEr into thiS aSpEct latEr on.
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFF78E6EFF5FFECD1202FE02" blockId="14.[151,1437,151,680]" pageId="14" pageNumber="428">
|
||
FurthEr diStinguiShing fEaturES arE a dorSal thyploSolE which StartS with maximal SizE (in thE othEr
|
||
<taxonomicName id="30A54D54FFF78E6EFF0FFEE1172BFEEE" authorityName="Fragoso" authorityYear="1988" box="[151,301,296,320]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="14" pageNumber="428" phylum="Annelida" rank="genus">
|
||
<emphasis id="C5D1EAC5FFF78E6EFF0FFEE1172BFEEE" box="[151,301,296,320]" italics="true" pageId="14" pageNumber="428">Lavellodrilus</emphasis>
|
||
</taxonomicName>
|
||
SpEciES it StartS Small, gradually incrEaSing itS SizE), thE highly dEvElopEd dorSal vESSEl in SEgmEntS
|
||
<emphasis id="C5D1EAC5FFF78E6EFF0FFE8516B5FECB" bold="true" box="[151,179,332,357]" pageId="14" pageNumber="428">15</emphasis>
|
||
and
|
||
<emphasis id="C5D1EAC5FFF78E6EFF71FE851703FECB" bold="true" box="[233,261,332,357]" pageId="14" pageNumber="428">16</emphasis>
|
||
(EvEn doublE in SomE individualS) in coincidEncE with a biggEr typhloSolE and a pair of SEminal vESiclES in
|
||
<emphasis id="C5D1EAC5FFF78E6EFF2CFEA616D6FE26" bold="true" box="[180,208,367,392]" pageId="14" pageNumber="428">12</emphasis>
|
||
(
|
||
<tableCitation id="BA27036CFFF78E6EFF47FEA61731FE26" box="[223,311,367,392]" captionStart="TABLE 2" captionStartId="11.[147,229,599,620]" captionText="TABLE 2. Morphological comparisons among the currently recognized Lavellodrilus species from Mexico. Data obtained from Fragoso (1988, 1991) and Fragoso and Rojas (2016)" pageId="14" pageNumber="428">TablE 2</tableCitation>
|
||
). By thE poSition of thE firSt dorSal porE, thE Small body SizE and thE pairEd naturE of gEnital markS, thE nEw SpEciES would bE morE rElatEd to
|
||
<taxonomicName id="30A54D54FFF78E6EFDF4FE5D15F9FE02" authority="FragoSo, 1988" authorityName="Fragoso" authorityYear="1988" box="[620,1023,404,428]" class="Clitellata" family="Acanthodrilidae" genus="Lavellodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="14" pageNumber="428" phylum="Annelida" rank="species" species="parvus">
|
||
<emphasis id="C5D1EAC5FFF78E6EFDF4FE5D1553FE02" box="[620,853,404,428]" italics="true" pageId="14" pageNumber="428">Lavellodrilus parvus</emphasis>
|
||
<bibRefCitation id="93344B26FFF78E6EFCC4FE5D15F9FE02" box="[860,1023,404,428]" pageId="14" pageNumber="428" refId="ref6476" refString="Fragoso, C. (1988) Sistematica y ecologia de un genero nuevo de lombriz de tierra (Acanthodrilini, Oligochaeta) de la Selva Lacandona, Chiapas. Mexico. Acta Zoologica Mexicana, Nueva Serie, 25, 1 - 34." type="journal article">FragoSo, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F71A36D7FFF78E6EFF5FFE7E12E3FD06" blockId="14.[151,1437,151,680]" pageId="14" pageNumber="428">
|
||
Variation in thE numbEr of divErticula (onE or two) for a SinglE SpEciES iS nEw for MExican acanthodrilinES, holoic or mEroic. In othEr rEgionS, howEvEr, SomE variation rElatEd to thE numbEr of SpErmathEcaE haS bEEn obSErvEd. For ExamplE in
|
||
<taxonomicName id="30A54D54FFF78E6EFE50FDC8150EFDB6" authority="(Pickford 1937)" authorityName="Pickford" authorityYear="1937" baseAuthorityName="Pickford" baseAuthorityYear="1937" box="[456,776,511,536]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="14" pageNumber="428" phylum="Annelida" rank="species" species="bavenda">
|
||
<emphasis id="C5D1EAC5FFF78E6EFE50FDC81440FDB6" box="[456,582,512,536]" italics="true" pageId="14" pageNumber="428">P. bavenda</emphasis>
|
||
(
|
||
<bibRefCitation id="93344B26FFF78E6EFDCFFE3614F9FDB6" box="[599,767,511,536]" pageId="14" pageNumber="428" refId="ref7053" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. W. Heffer & Sons Ltd., Cambridge, 612 pp." type="book">Pickford 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, two or thrEE SpErmathEcaE wErE rEcordEd in individualS bElonging to thE SamE or diffErEnt population; intErEStingly, whEn only onE (antErior or poStErior) SpErmathEca waS prESEnt, it waS alwayS on onE SidE and nEvEr midvEntral. WE arE currEntly looking for morE matErial of thiS SpEciES in
|
||
<collectingRegion id="3561F835FFF78E6EFF2FFDA516F4FD2B" box="[183,242,620,645]" country="American Samoa" name="order" pageId="14" pageNumber="428">ordEr</collectingRegion>
|
||
to undErtakE COI analySiS of thE two morphotypES. ThE rESulting gEnEtic diStancES (gd) will pErmit to diScriminatE bEtwEEn a SpErmathEcal polymorphiSm (gd<15%) or two diffErEnt SpEciES (gd>15%).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |