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<document id="C4072D4C245899CBCBE703B0650E11CF" ID-DOI="10.11646/zootaxa.5507.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747694" ID-ZooBank="09AC0223-FC47-4C34-8719-0376CD92C837" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055664737" checkinUser="plazi" docAuthor="Viraktamath, C. A., Webb, M. D. &amp; Yeshwanth, H. M." docDate="2024" docId="7E7D0B1F766E381FFF1EFB88FB9EFD8B" docLanguage="en" docName="zootaxa.5507.1.4.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Libengaia (Libengaia) trifida Viraktamath, Webb &amp; Yeshwanth, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="113" masterDocId="8244736776673816FF89FFB7F83CFB36" masterDocTitle="Opsiini leafhopper genus Libengaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) in Africa and India with description of one new subgenus and six new species" masterLastPageNumber="122" masterPageNumber="104" pageNumber="113" updateTime="1726056483101" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9B0936F4E3A498FBB7DF7CE1A0F9A0E3">Opsiini leafhopper genus Libengaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) in Africa and India with description of one new subgenus and six new species</mods:title>
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<mods:namePart id="126A7256AA675C8E4C479A6694A1B67E">Viraktamath, C. A.</mods:namePart>
<mods:affiliation id="0C90783ECE61D6E1D718BD6B6F4AA39D">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India</mods:affiliation>
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<mods:namePart id="8A3B662ABED11E71A7041E341B900184">Webb, M. D.</mods:namePart>
<mods:affiliation id="A2E1DEF8833D688A727AD5D062242B52">Department of Life Science, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom</mods:affiliation>
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<mods:namePart id="5E4B0F44EB6D0F7C90AF41C760849EF7">Yeshwanth, H. M.</mods:namePart>
<mods:affiliation id="BFF99BD457CB3E6715494367FF9E98A9">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India</mods:affiliation>
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<treatment id="7E7D0B1F766E381FFF1EFB88FB9EFD8B" ID-DOI="http://doi.org/10.5281/zenodo.13747614" ID-Zenodo-Dep="13747614" LSID="urn:lsid:plazi:treatment:7E7D0B1F766E381FFF1EFB88FB9EFD8B" httpUri="http://treatment.plazi.org/id/7E7D0B1F766E381FFF1EFB88FB9EFD8B" lastPageNumber="113" pageId="9" pageNumber="113">
<subSubSection id="BECEE982766E381FFF1EFB88FA62FF6F" box="[151,606,1087,1114]" pageId="9" pageNumber="113" type="nomenclature">
<paragraph id="F66BBA09766E381FFF1EFB88FA62FF6F" blockId="9.[151,606,1087,1150]" box="[151,606,1087,1114]" pageId="9" pageNumber="113">
<heading id="AD230D65766E381FFF1EFB88FA62FF6F" bold="true" box="[151,606,1087,1114]" fontSize="11" level="1" pageId="9" pageNumber="113" reason="1">
<emphasis id="C4A0661B766E381FFF1EFB88FA62FF6F" bold="true" box="[151,606,1087,1114]" pageId="9" pageNumber="113">
<taxonomicName id="31D4C18A766E381FFF1EFB88F9C5FF6C" ID-CoL="8HSDP" authorityName="Viraktamath &amp; Webb &amp; Yeshwanth" authorityYear="2024" box="[151,505,1087,1114]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="113" phylum="Arthropoda" rank="species" species="trifida" status="sp. nov." subGenus="Libengaia">
<emphasis id="C4A0661B766E381FFF1EFB88F92DFF6C" bold="true" box="[151,273,1087,1114]" italics="true" pageId="9" pageNumber="113">Libengaia</emphasis>
(
<emphasis id="C4A0661B766E381FFEAAFB88F9A1FF6C" bold="true" box="[291,413,1087,1114]" italics="true" pageId="9" pageNumber="113">Libengaia</emphasis>
)
<emphasis id="C4A0661B766E381FFE27FB88F9C5FF6C" bold="true" box="[430,505,1087,1114]" italics="true" pageId="9" pageNumber="113">trifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DF93DB60766E381FFD89FB88FA62FF6F" box="[512,606,1087,1113]" pageId="9" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="BECEE982766E381FFF1EFBD4FCD1FD43" pageId="9" pageNumber="113" type="description">
<paragraph id="F66BBA09766E381FFF1EFBD4F944FF48" blockId="9.[151,606,1087,1150]" box="[151,376,1123,1150]" pageId="9" pageNumber="113">
<figureCitation id="6EEFA68C766E381FFF1EFBD4F8D0FF4B" box="[151,236,1123,1149]" captionStart="FIGURE 1" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[156,1430,205,1829]" captionTargetId="figure-18@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 1 AP. Species of Libengaia (Libengaia), habitus dorsal (AF), face (GN) and ant. A, G, L. (L.) brevistylus sp. nov. male; BC, HI, L. (L.) deemingi sp. nov.: male (BH), female (CI); D, JK, L. (L.) immaculata male; E, L, M, L. (L.) longistylus sp. nov.; F, N, L. (L.) trifida sp. nov., holotype male, face; OP, ant, Crematogaster sp. dorsal and lateral view." figureDoi="http://doi.org/10.5281/zenodo.13747696" httpUri="https://zenodo.org/record/13747696/files/figure.png" pageId="9" pageNumber="113">Figs 1F</figureCitation>
, MN, 5AF.
</paragraph>
<paragraph id="F66BBA09766E381FFF1EFB1BF9CEFFDF" blockId="9.[151,1437,1195,1725]" pageId="9" pageNumber="113">
<emphasis id="C4A0661B766E381FFF1EFB1BF935FFF3" box="[151,265,1196,1221]" italics="true" pageId="9" pageNumber="113">Diagnosis</emphasis>
. Coloration similar to that in
<taxonomicName id="31D4C18A766E381FFDE7FB1BFB00FFF3" authorityName="Viraktamath &amp; Webb &amp; Yeshwanth" authorityYear="2024" box="[622,828,1195,1221]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="113" phylum="Arthropoda" rank="species" species="brevistylus" subGenus="Libengaia">
<emphasis id="C4A0661B766E381FFDE7FB1BFAB8FFF3" box="[622,644,1196,1221]" italics="true" pageId="9" pageNumber="113">L.</emphasis>
(
<emphasis id="C4A0661B766E381FFD10FB1BFA93FFF3" box="[665,687,1196,1221]" italics="true" pageId="9" pageNumber="113">L.</emphasis>
)
<emphasis id="C4A0661B766E381FFD4DFB1CFB00FFF3" box="[708,828,1195,1221]" italics="true" pageId="9" pageNumber="113">brevistylus</emphasis>
</taxonomicName>
, but the maculae bolder and denser. Aedeagal shaft appearing trifid in lateral view.
</paragraph>
<paragraph id="F66BBA09766E381FFF4EFB43F8DAFE07" blockId="9.[151,1437,1195,1725]" pageId="9" pageNumber="113">
<emphasis id="C4A0661B766E381FFF4EFB43F977FE3B" box="[199,331,1268,1293]" italics="true" pageId="9" pageNumber="113">Description</emphasis>
. Male
<quantity id="312C17EC766E381FFE17FB44F9CAFE38" box="[414,502,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="113" unit="mm" value="3.6">3.6 mm</quantity>
long. Coloration similar to that in
<taxonomicName id="31D4C18A766E381FFC03FB43FC68FE3B" authorityName="Viraktamath &amp; Webb &amp; Yeshwanth" authorityYear="2024" box="[906,1108,1267,1293]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="113" phylum="Arthropoda" rank="species" species="brevistylus" subGenus="Libengaia">
<emphasis id="C4A0661B766E381FFC03FB43FB9DFE3B" box="[906,929,1268,1293]" italics="true" pageId="9" pageNumber="113">L.</emphasis>
(
<emphasis id="C4A0661B766E381FFC3AFB43FBF6FE3B" box="[947,970,1268,1293]" italics="true" pageId="9" pageNumber="113">L.</emphasis>
)
<emphasis id="C4A0661B766E381FFC55FB44FC68FE3B" box="[988,1108,1267,1293]" italics="true" pageId="9" pageNumber="113">brevistylus</emphasis>
</taxonomicName>
, but the maculae bolder and denser.
</paragraph>
<paragraph id="F66BBA09766E381FFF4EFA8CFAF1FD1B" blockId="9.[151,1437,1195,1725]" pageId="9" pageNumber="113">
<emphasis id="C4A0661B766E381FFF4EFA8CF94CFE63" box="[199,368,1339,1365]" italics="true" pageId="9" pageNumber="113">Male genitalia</emphasis>
. Pygofer about as long as height at base in lateral view, dorsal and ventral margins almost straight, posteroventral angle rounded, macrosetae present in posterior 0.33. Style with well-developed preapical lobe, apophysis slender, slightly laterally curved, with sculptured lateral surface. Connective stem 2× as long as arms, apex widened. Aedeagal shafts about
<geoCoordinate id="93E0DCCE766E381FFDF5FA10FA82FEF4" box="[636,702,1447,1474]" degrees="0.33" direction="south" orientation="latitude" pageId="9" pageNumber="113" precision="555" value="-0.33">0.33 s</geoCoordinate>
long as ventral process, slightly shorter than preatrium, each shaft with lamellate ledge laterally, distally projected as short process slightly beyond gonopore, preatrium with slender elongate process arising at base shafts and extending beyond atrial wall, forked before apex into unequal branches; shaft in dorsal view straight and divergent distally.
</paragraph>
<paragraph id="F66BBA09766E381FFF4EF980FD01FD67" blockId="9.[151,1437,1195,1725]" box="[199,1341,1591,1618]" pageId="9" pageNumber="113">
<emphasis id="C4A0661B766E381FFF4EF980F9A7FD67" box="[199,411,1591,1617]" italics="true" pageId="9" pageNumber="113">Material examined</emphasis>
.
<materialsCitation id="46BCB054766E381FFE20F980FD01FD67" box="[425,1341,1591,1618]" collectingDate="1975-02-02" collectionCode="BMNH" collectorName="Ilaro Forest, W &amp; J. Riley" country="Nigeria" location="Ilaro Forest" pageId="9" pageNumber="113" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="296F04AB766E381FFE20F980FA01FD67" box="[425,573,1591,1617]" pageId="9" pageNumber="113" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="E0D27180766E381FFDCDF98FFA67FD64" box="[580,603,1592,1618]" count="1" pageId="9" pageNumber="113" type="male"></specimenCount>
,
<collectingCountry id="8EC3FA99766E381FFDEEF980FAE3FD67" box="[615,735,1591,1617]" name="Nigeria" pageId="9" pageNumber="113">NIGERIA</collectingCountry>
:
<collectorName id="5B21DFDF766E381FFD60F980FBA8FD64" box="[745,916,1591,1618]" pageId="9" pageNumber="113">
<location id="F30BECD2766E381FFD60F980FB52FD64" LSID="urn:lsid:plazi:treatment:7E7D0B1F766E381FFF1EFB88FB9EFD8B:F30BECD2766E381FFD60F980FB52FD64" box="[745,878,1591,1618]" country="Nigeria" name="Ilaro Forest" pageId="9" pageNumber="113">Ilaro Forest</location>
, W
</collectorName>
. Nigeria,
<date id="826A9CC9766E381FFB89F980FC5AFD64" box="[1024,1126,1591,1618]" pageId="9" pageNumber="113" value="1975-02-02">
<collectingDate id="922E6521766E381FFB89F980FC5AFD64" box="[1024,1126,1591,1618]" pageId="9" pageNumber="113" value="1975-02-02">2.ii.1975</collectingDate>
</date>
,
<collectorName id="5B21DFDF766E381FFBFBF98FFCF4FD67" box="[1138,1224,1591,1618]" pageId="9" pageNumber="113">J. Riley</collectorName>
(
<collectionCode id="90C522CC766E381FFB5FF980FD0CFD67" box="[1238,1328,1591,1617]" pageId="9" pageNumber="113">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F66BBA09766E381FFF4EF9ECFCD1FD43" blockId="9.[151,1437,1195,1725]" box="[199,1261,1627,1654]" pageId="9" pageNumber="113">
<emphasis id="C4A0661B766E381FFF4EF9ECF903FD43" box="[199,319,1627,1653]" italics="true" pageId="9" pageNumber="113">Etymology</emphasis>
. The specific epithet refers to the three pronged apex of each of the aedeagal shafts.
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766E381FFF4EF9C8FB9EFD8B" pageId="9" pageNumber="113" type="discussion">
<paragraph id="F66BBA09766E381FFF4EF9C8FB9EFD8B" blockId="9.[151,1437,1195,1725]" pageId="9" pageNumber="113">
<emphasis id="C4A0661B766E381FFF4EF9C8F908FDAF" bold="true" box="[199,308,1663,1689]" pageId="9" pageNumber="113">Remarks</emphasis>
. This species has a shorter aedeagal shaft as in the other species of the nominate subgenus found in Africa but differs in having each shaft appearing trifid in lateral view.
</paragraph>
</subSubSection>
</treatment>
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