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<document id="5C1BE66672B6DEAF6D9C6B0BE238641A" ID-DOI="10.11646/zootaxa.5507.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747694" ID-ZooBank="09AC0223-FC47-4C34-8719-0376CD92C837" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055664737" checkinUser="plazi" docAuthor="Viraktamath, C. A., Webb, M. D. &amp; Yeshwanth, H. M." docDate="2024" docId="7E7D0B1F766C381DFF1EFDDDFC50FEEC" docLanguage="en" docName="zootaxa.5507.1.4.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Libengaia (Nandiana) bharatiensis Viraktamath &amp; Webb &amp; Yeshwanth 2024, comb. nov." docType="treatment" docVersion="2" lastPageNumber="115" masterDocId="8244736776673816FF89FFB7F83CFB36" masterDocTitle="Opsiini leafhopper genus Libengaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) in Africa and India with description of one new subgenus and six new species" masterLastPageNumber="122" masterPageNumber="104" pageNumber="115" updateTime="1726056483101" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2BAA65CE48123C03FCDC5F9090AF77C8">Opsiini leafhopper genus Libengaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) in Africa and India with description of one new subgenus and six new species</mods:title>
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<mods:namePart id="A107B312E603599154187DF99B497125">Webb, M. D.</mods:namePart>
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<mods:namePart id="EDC1F52851A852DE8450070C0B073D4F">Yeshwanth, H. M.</mods:namePart>
<mods:affiliation id="71B6C2A3629DFA8054C9ADA2F13E5CAA">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India</mods:affiliation>
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<taxonomicName id="31D4C18A766C381DFF1EFDDDFB61F9B2" ID-CoL="BV68B" authority="(Malhotra &amp; Sharma)" authorityName="Viraktamath &amp; Webb &amp; Yeshwanth" authorityYear="2024" baseAuthorityName="Viraktamath &amp; Webb &amp; Yeshwanth" baseAuthorityYear="2024" box="[151,861,618,645]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="bharatiensis" status="comb. nov." subGenus="Nandiana">
<emphasis id="C4A0661B766C381DFF1EFDDDFA7CF9B3" bold="true" box="[151,576,618,645]" italics="true" pageId="11" pageNumber="115">Libengaia (Nandiana) bharatiensis</emphasis>
(Malhotra &amp; Sharma)
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<figureCitation id="6EEFA68C766C381DFF1EFD38F930F99F" box="[151,268,655,681]" captionStart="FIGURE 6" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[156,1417,186,1857]" captionTargetId="figure-18@12.[151,1435,181,1862]" captionTargetPageId="12" captionText="FIGURE 6 A–P. Species of Libengaia (Nandiana), habitus dorsal and lateral views.A– J, color variation in L. (N.) bharatiensis (Malhotra &amp; Shrma): specimens from Bandipur (AB), Khandala (CD), Mudigere (EF), Thirthahalli (GH) and Waghai (IJ); K–N, L. (N.) lamerallis Sunil &amp; Meshram, male (KL), female (MN); O–P, L. (N.) ramamurthyi sp. nov., holotype male." figureDoi="http://doi.org/10.5281/zenodo.13747706" httpUri="https://zenodo.org/record/13747706/files/figure.png" pageId="11" pageNumber="115">Figs 6A–J</figureCitation>
,
<figureCitation id="6EEFA68C766C381DFE91FD38F9A5F99F" box="[280,409,655,681]" captionStart="FIGURE 7" captionStartId="13.[151,250,1866,1890]" captionTargetBox="[164,1414,195,1837]" captionTargetId="figure-28@13.[151,1436,181,1842]" captionTargetPageId="13" captionText="FIGURE 7 A–R. Species of Libengaia (Nandiana), faces and female genitalia.A–E, L. (N.) bharatiensis (Malhotra &amp; Sharma), color variation in face: specimens from Bandipur (A), Khandala (B), Mudigere (C), Thirthahalli (D) and Waghai (E); F–G, L. (N.) lamerallis Sunil &amp; Meshram, male and female; H, L. (N.) ramamurthyi sp. nov., holotype male face; I–M, L. (N.) bharatiensis: I, female sternite VII; JK, valvula I; LM, valvula II; N–R, L. (N.) lamerallis: N, female sternite VII; OP, valvula I; QR, valvula II." figureDoi="http://doi.org/10.5281/zenodo.13747708" httpUri="https://zenodo.org/record/13747708/files/figure.png" pageId="11" pageNumber="115">7A–E, I–M</figureCitation>
,
<figureCitation id="6EEFA68C766C381DFE2CFD38F9D1F99F" box="[421,493,655,681]" captionStart="FIGURE 8" captionStartId="14.[151,250,1891,1915]" captionTargetBox="[152,1406,194,1841]" captionTargetId="figure-18@14.[151,1436,181,1867]" captionTargetPageId="14" captionText="FIGURE 8 A–G. Libengaia (Nandiana) bharatiensis (Malhotra &amp; Sharma, male genitalia: A, pygofer and anal tube, lateral view; B, valve, ventral view; C, styles and subgenital plates dorsal view; D, style, dorsal view; E, connective and aedeagus, dorsal view; F, aedeagus, ventral view; G, connective and aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13747712" httpUri="https://zenodo.org/record/13747712/files/figure.png" pageId="11" pageNumber="115">8A–G</figureCitation>
.
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<taxonomicName id="31D4C18A766C381DFF1EFD63FA8FF9DD" ID-CoL="3VNV2" authority="Malhotra &amp; Sharma 1977: 27 - 29" authorityName="Malhotra &amp; Sharma" authorityPageNumber="27 - 29" authorityYear="1977" box="[151,691,723,747]" class="Insecta" family="Cicadellidae" genus="Litura" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="bharatiensis">
<emphasis id="C4A0661B766C381DFF1EFD63F966F9DD" box="[151,346,723,747]" italics="true" pageId="11" pageNumber="115">Litura bharatiensis</emphasis>
<bibRefCitation id="9245C7F8766C381DFEE9FD64FA8FF9DD" author="Malhotra, Y. R. &amp; Sharma, B." box="[352,691,723,747]" pageId="11" pageNumber="115" pagination="27 - 29" refId="ref8738" refString="Malhotra, Y. R. &amp; Sharma, B. (1977) A new species of Litura from Jammu, India (Homoptera: Cicadellidae). Oriental Insects, 11, 27 - 29. https: // doi. org / 10.1080 / 00305316.1977.10432079" type="journal article" year="1977">Malhotra &amp; Sharma 1977: 27–29</bibRefCitation>
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<figureCitation id="6EEFA68C766C381DFD34FD64FB24F9DD" box="[701,792,723,747]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStartId-0="4.[151,250,1858,1882]" captionStartId-1="5.[151,250,1943,1967]" captionStartId-2="7.[151,250,1943,1967]" captionStartId-3="8.[151,250,1938,1962]" captionStartId-4="10.[151,250,1940,1964]" captionStartId-5="12.[151,250,1885,1909]" captionStartId-6="13.[151,250,1866,1890]" captionStartId-7="14.[151,250,1891,1915]" captionStartId-8="15.[151,250,1908,1932]" captionTargetBox-0="[156,1430,205,1829]" captionTargetBox-1="[180,1416,190,1912]" captionTargetBox-2="[174,1409,186,1908]" captionTargetBox-3="[153,1396,191,1869]" captionTargetBox-4="[170,1402,213,1887]" captionTargetBox-5="[156,1417,186,1857]" captionTargetBox-6="[164,1414,195,1837]" captionTargetBox-7="[152,1406,194,1841]" captionTargetBox-8="[160,1435,194,1877]" captionTargetId-0="figure-18@4.[151,1435,181,1834]" captionTargetId-1="figure-28@5.[170,1417,181,1920]" captionTargetId-2="figure-28@7.[169,1419,181,1920]" captionTargetId-3="figure-18@8.[151,1436,181,1915]" captionTargetId-4="figure-18@10.[158,1430,181,1917]" captionTargetId-5="figure-18@12.[151,1435,181,1862]" captionTargetId-6="figure-28@13.[151,1436,181,1842]" captionTargetId-7="figure-18@14.[151,1436,181,1867]" captionTargetId-8="figure-28@15.[151,1436,181,1885]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="10" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="15" captionText-0="FIGURE 1 A–P. Species of Libengaia (Libengaia), habitus dorsal (A–F), face (G–N) and ant. A, G, L. (L.) brevistylus sp. nov. male; BC, HI, L. (L.) deemingi sp. nov.: male (BH), female (CI); D, JK, L. (L.) immaculata male; E, L, M, L. (L.) longistylus sp. nov.; F, N, L. (L.) trifida sp. nov., holotype male, face; OP, ant, Crematogaster sp. dorsal and lateral view." captionText-1="FIGURE 2 A–E. Libengaia (Libengaia) brevistylus sp. nov., male genitalia. A, pygofer, lateral view; B, style, dorsal view; C, connective, dorsal view; D, aedeagus, dorsal view; E, aedeagus, lateral view." captionText-2="FIGURE 3 A–E. Libengaia (Libengaia) immaculata sp. nov., male genitalia. A, genital capsule, lateral view; B, style, dorsal view; C, connective, dorsal view; D, aedeagus, ventral view; E, aedeagus, lateral view." captionText-3="FIGURE 4 A–G. Species of Libengaia (Libengaia) male genitalia. A–D, L. (L.) deemingi sp. nov.: A, genital capsule, lateral view; B, style, dorsal view; C, aedeagus, ventral view; D, aedeagus, lateral view. E–G, L. (L.) longistylus sp. nov.: E, aedeagus, lateral view; F, aedeagus, dorsal view; G, style, dorsal view." captionText-4="FIGURE 5 A–F. Libengaia (Libengaia) trifida sp. nov., male genitalia. A, pygofer, lateral view; B, subgenital plate, ventral view; C, connective, dorsal view; D, style, dorsal view and apex of style, magnified; E, aedeagus, dorsal view; F, aedeagus, lateral view." captionText-5="FIGURE 6 A–P. Species of Libengaia (Nandiana), habitus dorsal and lateral views.A– J, color variation in L. (N.) bharatiensis (Malhotra &amp; Shrma): specimens from Bandipur (AB), Khandala (CD), Mudigere (EF), Thirthahalli (GH) and Waghai (IJ); K–N, L. (N.) lamerallis Sunil &amp; Meshram, male (KL), female (MN); O–P, L. (N.) ramamurthyi sp. nov., holotype male." captionText-6="FIGURE 7 A–R. Species of Libengaia (Nandiana), faces and female genitalia.A–E, L. (N.) bharatiensis (Malhotra &amp; Sharma), color variation in face: specimens from Bandipur (A), Khandala (B), Mudigere (C), Thirthahalli (D) and Waghai (E); F–G, L. (N.) lamerallis Sunil &amp; Meshram, male and female; H, L. (N.) ramamurthyi sp. nov., holotype male face; I–M, L. (N.) bharatiensis: I, female sternite VII; JK, valvula I; LM, valvula II; N–R, L. (N.) lamerallis: N, female sternite VII; OP, valvula I; QR, valvula II." captionText-7="FIGURE 8 A–G. Libengaia (Nandiana) bharatiensis (Malhotra &amp; Sharma, male genitalia: A, pygofer and anal tube, lateral view; B, valve, ventral view; C, styles and subgenital plates dorsal view; D, style, dorsal view; E, connective and aedeagus, dorsal view; F, aedeagus, ventral view; G, connective and aedeagus, lateral view." captionText-8="FIGURE 9 A–H. Libengaia (Nandiana) lamerallis Sunil &amp; Meshram, metatibia and male genitalia: A, chaetotaxy of apex of metatibia, ventral view; B, pygofer, lateral view; C, subgenital plate, ventral view; D, connective, dorsal view; E, style, dorsal view; F, aedeagus dorsal view; G, aedeagal shaft, dorsolateral view; H, aedeagus, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13747696" figureDoi-1="http://doi.org/10.5281/zenodo.13747698" figureDoi-2="http://doi.org/10.5281/zenodo.13747700" figureDoi-3="http://doi.org/10.5281/zenodo.13747702" figureDoi-4="http://doi.org/10.5281/zenodo.13747704" figureDoi-5="http://doi.org/10.5281/zenodo.13747706" figureDoi-6="http://doi.org/10.5281/zenodo.13747708" figureDoi-7="http://doi.org/10.5281/zenodo.13747712" figureDoi-8="http://doi.org/10.5281/zenodo.13747714" httpUri-0="https://zenodo.org/record/13747696/files/figure.png" httpUri-1="https://zenodo.org/record/13747698/files/figure.png" httpUri-2="https://zenodo.org/record/13747700/files/figure.png" httpUri-3="https://zenodo.org/record/13747702/files/figure.png" httpUri-4="https://zenodo.org/record/13747704/files/figure.png" httpUri-5="https://zenodo.org/record/13747706/files/figure.png" httpUri-6="https://zenodo.org/record/13747708/files/figure.png" httpUri-7="https://zenodo.org/record/13747712/files/figure.png" httpUri-8="https://zenodo.org/record/13747714/files/figure.png" pageId="11" pageNumber="115">Figs 1–9</figureCitation>
.
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<treatmentCitation id="77759C18766C381DFF1EFD45FABDF83C" author="Viraktamath, C. A." box="[151,641,754,778]" page="243" pageId="11" pageNumber="115" year="1978">
<taxonomicName id="31D4C18A766C381DFF1EFD45FABDF83C" ID-CoL="8R82G" authority="Viraktamath 1978: 241 - 243" authorityName="Viraktamath" authorityPageNumber="241 - 243" authorityYear="1978" box="[151,641,754,778]" class="Insecta" family="Cicadellidae" genus="Cestius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="terminaliae">
<emphasis id="C4A0661B766C381DFF1EFD45F961F83C" box="[151,349,754,778]" italics="true" pageId="11" pageNumber="115">Cestius terminaliae</emphasis>
<bibRefCitation id="9245C7F8766C381DFEEAFD45FABDF83C" author="Viraktamath, C. A." box="[355,641,754,778]" pageId="11" pageNumber="115" pagination="241 - 244" refId="ref8925" refString="Viraktamath, C. A. (1978) A new species of Cestius Distant (Homoptera: Cicadellidae) from southern India. Journal of Natural History, 12, 241 - 244. https: // doi. org / 10.1080 / 00222937800770131" type="journal article" year="1978">Viraktamath 1978: 241–243</bibRefCitation>
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</treatmentCitation>
,
<figureCitation id="6EEFA68C766C381DFD05FD45FAFBF83C" box="[652,711,754,778]" captionStart="FIGURE 1" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[156,1430,205,1829]" captionTargetId="figure-18@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 1 A–P. Species of Libengaia (Libengaia), habitus dorsal (A–F), face (G–N) and ant. A, G, L. (L.) brevistylus sp. nov. male; BC, HI, L. (L.) deemingi sp. nov.: male (BH), female (CI); D, JK, L. (L.) immaculata male; E, L, M, L. (L.) longistylus sp. nov.; F, N, L. (L.) trifida sp. nov., holotype male, face; OP, ant, Crematogaster sp. dorsal and lateral view." figureDoi="http://doi.org/10.5281/zenodo.13747696" httpUri="https://zenodo.org/record/13747696/files/figure.png" pageId="11" pageNumber="115">Figs 1</figureCitation>
–11, s
<emphasis id="C4A0661B766C381DFC8FFD45FB65F83C" bold="true" box="[774,857,754,778]" pageId="11" pageNumber="115">yn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766C381DFF1EFC8EFB84FE10" pageId="11" pageNumber="115" type="diagnosis">
<paragraph id="F66BBA09766C381DFF1EFC8EF8DFF8AC" blockId="11.[151,1437,824,1499]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF1EFC8EF935F864" box="[151,265,825,850]" italics="true" pageId="11" pageNumber="115">Diagnosis</emphasis>
. This species can be recognized from other species of the subgenus by the aedeagal shaft having one retrorse apical process that is slightly widened beyond midlength. Coloration especially in males is variable (
<figureCitation id="6EEFA68C766C381DFAE3FCEBF8EBF8AC" captionStart="FIGURE 6" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[156,1417,186,1857]" captionTargetId="figure-18@12.[151,1435,181,1862]" captionTargetPageId="12" captionText="FIGURE 6 A–P. Species of Libengaia (Nandiana), habitus dorsal and lateral views.A– J, color variation in L. (N.) bharatiensis (Malhotra &amp; Shrma): specimens from Bandipur (AB), Khandala (CD), Mudigere (EF), Thirthahalli (GH) and Waghai (IJ); K–N, L. (N.) lamerallis Sunil &amp; Meshram, male (KL), female (MN); O–P, L. (N.) ramamurthyi sp. nov., holotype male." figureDoi="http://doi.org/10.5281/zenodo.13747706" httpUri="https://zenodo.org/record/13747706/files/figure.png" pageId="11" pageNumber="115">Figs 6A–J</figureCitation>
).
</paragraph>
<paragraph id="F66BBA09766C381DFF4EFC12FC93FF78" blockId="11.[151,1437,824,1499]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF4EFC12F977F888" box="[199,331,933,958]" italics="true" pageId="11" pageNumber="115">Description</emphasis>
. Female. Coloration similar to male but less variable. Sternite VII twice as wide as long medially, posterior margin broadly produced medially into bilobed lobe, lateral angles produced and conically rounded. Valvula I curved dorsally, dorsal sculpturing strigate, occupying distal half length, strigae dorsally becoming cellular on dorsal margin. Valvula II curved dorsally, with one tooth almost at midlength dorsally, regular toothed area occupying distal 0.4 length, each tooth slightly broader than height, with secondary dentition.
</paragraph>
<paragraph id="F66BBA09766C381DFF4EFBEFFB84FE10" blockId="11.[151,1437,824,1499]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF4EFBEFF99EFF44" box="[199,418,1112,1138]" italics="true" pageId="11" pageNumber="115">Material examined</emphasis>
.
<materialsCitation id="46BCB054766C381DFE3EFBEFFC54FFA0" collectingDate="1974-06-19" collectionCode="UASB" collectorName="C. A. Viraktamath" country="India" location="Lalbagh" municipality="Bangalore" pageId="11" pageNumber="115" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Karnataka" typeStatus="holotype">
<collectingCountry id="8EC3FA99766C381DFE3EFBEFFA37FF44" box="[439,523,1112,1138]" name="India" pageId="11" pageNumber="115">INDIA</collectingCountry>
:
<collectingRegion id="341074EB766C381DFD95FBEFFAA8FF44" box="[540,660,1112,1138]" country="India" name="Karnataka" pageId="11" pageNumber="115">Karnataka</collectingRegion>
:
<typeStatus id="296F04AB766C381DFD2CFBEFFB05FF44" box="[677,825,1112,1138]" pageId="11" pageNumber="115" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="E0D27180766C381DFCCFFBEEFB61FF45" box="[838,861,1113,1139]" count="1" pageId="11" pageNumber="115" type="male">♂</specimenCount>
, and
<specimenCount id="E0D27180766C381DFC2FFBEFFBF7FF44" box="[934,971,1112,1138]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<specimenCount id="E0D27180766C381DFC57FBEFFC3DFF44" box="[990,1025,1112,1138]" count="1" pageId="11" pageNumber="115" type="female">1♀</specimenCount>
,
<typeStatus id="296F04AB766C381DFB9FFBEEFCBDFF45" box="[1046,1153,1113,1139]" pageId="11" pageNumber="115" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="31D4C18A766C381DFB3DFBEFFDA9FF44" authorityName="Viraktamath" authorityYear="1978" box="[1204,1429,1112,1138]" class="Insecta" family="Cicadellidae" genus="Cestius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="terminaliae">
<emphasis id="C4A0661B766C381DFB3DFBEFFDA9FF44" box="[1204,1429,1112,1138]" italics="true" pageId="11" pageNumber="115">Cestius terminaliae</emphasis>
</taxonomicName>
.
<collectingMunicipality id="160F2073766C381DFF1EFBCBF931FFA0" box="[151,269,1148,1174]" pageId="11" pageNumber="115">Bangalore</collectingMunicipality>
,
<location id="F30BECD2766C381DFE96FBCBF942FFA0" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EFDDDFC50FEEC:F30BECD2766C381DFE96FBCBF942FFA0" box="[287,382,1148,1174]" country="India" municipality="Bangalore" name="Lalbagh" pageId="11" pageNumber="115" stateProvince="Karnataka">Lalbagh</location>
,
<emphasis id="C4A0661B766C381DFE19FBCAFAB7FFA0" box="[400,651,1148,1174]" italics="true" pageId="11" pageNumber="115">
ex
<taxonomicName id="31D4C18A766C381DFE31FBCBFAB7FFA0" box="[440,651,1148,1174]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="115" phylum="Tracheophyta" rank="species" species="arjuna">Terminalia arjuna</taxonomicName>
</emphasis>
,
<date id="826A9CC9766C381DFD29FBCBFB25FFA0" box="[672,793,1148,1174]" pageId="11" pageNumber="115" value="1974-06-19">
<collectingDate id="922E6521766C381DFD29FBCBFB25FFA0" box="[672,793,1148,1174]" pageId="11" pageNumber="115" value="1974-06-19">19.vi.1974</collectingDate>
</date>
,
<collectorName id="5B21DFDF766C381DFCA4FBCBFBC5FFA0" box="[813,1017,1148,1174]" pageId="11" pageNumber="115">C.A. Viraktamath</collectorName>
(
<collectionCode id="90C522CC766C381DFB99FBCBFC5DFFA0" box="[1040,1121,1148,1174]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="11" pageNumber="115">UASB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="46BCB054766C381DFBF3FBCBF9DFFFE8" collectingDate="1976-07-15" collectingDateMax="1978-05-22" collectingDateMin="1976-07-15" collectorName="Thithimathi &amp; Channapatna" country="India" county="Bangalore" location="Nagarahole" municipality="Mudigere" pageId="11" pageNumber="115" specimenCount="13" specimenCount-female="4" specimenCount-male="9">
<typeStatus id="296F04AB766C381DFBF3FBCBFD14FFA0" box="[1146,1320,1148,1174]" isOtherTypeStatus="true" pageId="11" pageNumber="115">Other material</typeStatus>
:
<specimenCount id="E0D27180766C381DFAB3FBCBFD62FFA0" box="[1338,1374,1148,1174]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<specimenCount id="E0D27180766C381DFAFBFBCBFDABFFA0" box="[1394,1431,1148,1174]" count="2" pageId="11" pageNumber="115" type="female">2♀</specimenCount>
,
<collectingCounty id="1F0AC285766C381DFF1EFB17F931FF8C" box="[151,269,1184,1210]" pageId="11" pageNumber="115">Bangalore</collectingCounty>
,
<date id="826A9CC9766C381DFE90FB17F9A8FF8C" box="[281,404,1184,1210]" pageId="11" pageNumber="115" value="1976-09-12">
<collectingDate id="922E6521766C381DFE90FB17F9A8FF8C" box="[281,404,1184,1210]" pageId="11" pageNumber="115" value="1976-09-12">12.ix.1976</collectingDate>
</date>
;
<specimenCount id="E0D27180766C381DFE29FB17F9F8FF8C" box="[416,452,1184,1210]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<collectingMunicipality id="160F2073766C381DFE58FB17FA7DFF8C" box="[465,577,1184,1210]" pageId="11" pageNumber="115">Mudigere</collectingMunicipality>
,
<date id="826A9CC9766C381DFDC4FB17FA82FF8C" box="[589,702,1184,1210]" pageId="11" pageNumber="115" value="1978-05-22">
<collectingDate id="922E6521766C381DFDC4FB17FA82FF8C" box="[589,702,1184,1210]" pageId="11" pageNumber="115" value="1978-05-22">22.v.1978</collectingDate>
</date>
;
<specimenCount id="E0D27180766C381DFD43FB17FAD2FF8C" box="[714,750,1184,1210]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<location id="F30BECD2766C381DFD72FB17FB43FF8C" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EFDDDFC50FEEC:F30BECD2766C381DFD72FB17FB43FF8C" box="[763,895,1184,1210]" country="India" county="Bangalore" municipality="Mudigere" name="Nagarahole" pageId="11" pageNumber="115">Nagarahole</location>
,
<date id="826A9CC9766C381DFC02FB17FBC4FF8C" box="[907,1016,1184,1210]" pageId="11" pageNumber="115" value="1978-01-17">
<collectingDate id="922E6521766C381DFC02FB17FBC4FF8C" box="[907,1016,1184,1210]" pageId="11" pageNumber="115" value="1978-01-17">17.i.1978</collectingDate>
</date>
;
<specimenCount id="E0D27180766C381DFB8DFB17FC15FF8D" box="[1028,1065,1184,1211]" count="5" pageId="11" pageNumber="115" type="male">5♂</specimenCount>
,
<specimenCount id="E0D27180766C381DFBBFFB17FC66FF8C" box="[1078,1114,1184,1210]" count="2" pageId="11" pageNumber="115" type="female">2♀</specimenCount>
,
<collectorName id="5B21DFDF766C381DFBEFFB17FCD1FF8C" box="[1126,1261,1184,1210]" pageId="11" pageNumber="115">Thithimathi</collectorName>
,
<date id="826A9CC9766C381DFB70FB17FD5AFF8C" box="[1273,1382,1184,1210]" pageId="11" pageNumber="115" value="1978-01-20">
<collectingDate id="922E6521766C381DFB70FB17FD5AFF8C" box="[1273,1382,1184,1210]" pageId="11" pageNumber="115" value="1978-01-20">20.i.1978</collectingDate>
</date>
;
<specimenCount id="E0D27180766C381DFAFBFB17FDABFF8C" box="[1394,1431,1184,1210]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<collectorName id="5B21DFDF766C381DFF1EFB73F910FFE8" box="[151,300,1220,1246]" pageId="11" pageNumber="115">Channapatna</collectorName>
,
<date id="826A9CC9766C381DFEB0FB73F987FFE9" box="[313,443,1220,1247]" pageId="11" pageNumber="115" value="1976-07-15">
<collectingDate id="922E6521766C381DFEB0FB73F987FFE9" box="[313,443,1220,1247]" pageId="11" pageNumber="115" value="1976-07-15">15.vii.1976</collectingDate>
</date>
, all
</materialsCitation>
<materialsCitation id="46BCB054766C381DFE62FB73FB57FE34" collectingDate="1975-01-16" collectingDateMax="1985-01-18" collectingDateMin="1975-01-16" collectionCode="UASB" collectorName="C. A. Viraktamath &amp; S. A. Viraktamath &amp; Bandipur &amp; Viraktamath" country="India" location="Biligiri Rangana Hills" pageId="11" pageNumber="115" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
collected by
<collectorName id="5B21DFDF766C381DFDF4FB73FB7BFFE8" box="[637,839,1220,1246]" pageId="11" pageNumber="115">C.A. Viraktamath</collectorName>
;
<specimenCount id="E0D27180766C381DFCDDFB73FB44FFE8" box="[852,888,1220,1246]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<specimenCount id="E0D27180766C381DFC0FFB73FB95FFE8" box="[902,937,1220,1246]" count="1" pageId="11" pageNumber="115" type="female">1♀</specimenCount>
,
<location id="F30BECD2766C381DFC31FB73FC88FFE8" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EFDDDFC50FEEC:F30BECD2766C381DFC31FB73FC88FFE8" box="[952,1204,1220,1247]" country="India" name="Biligiri Rangana Hills" pageId="11" pageNumber="115">Biligiri Rangana Hills</location>
,
<date id="826A9CC9766C381DFB48FB73FD6BFFE9" box="[1217,1367,1220,1247]" pageId="11" pageNumber="115" value="1975-01-16" valueMax="1975-01-17" valueMin="1975-01-16">
<collectingDate id="922E6521766C381DFB48FB73FD6BFFE9" box="[1217,1367,1220,1247]" pageId="11" pageNumber="115" value="1975-01-16" valueMax="1975-01-17" valueMin="1975-01-16">16–17.i.1975</collectingDate>
</date>
,
<collectorName id="5B21DFDF766C381DFAEDFB73F919FE34" pageId="11" pageNumber="115">S. A. Viraktamath</collectorName>
;
<specimenCount id="E0D27180766C381DFEA7FB5FF96DFE34" box="[302,337,1256,1282]" count="1" pageId="11" pageNumber="115" type="female">1♀</specimenCount>
,
<collectorName id="5B21DFDF766C381DFED4FB5FF9FBFE34" box="[349,455,1256,1282]" pageId="11" pageNumber="115">Bandipur</collectorName>
,
<date id="826A9CC9766C381DFE59FB5FFA00FE35" box="[464,572,1256,1283]" pageId="11" pageNumber="115" value="1985-01-18">
<collectingDate id="922E6521766C381DFE59FB5FFA00FE35" box="[464,572,1256,1283]" pageId="11" pageNumber="115" value="1985-01-18">18.i.1985</collectingDate>
</date>
, S.A.
<collectorName id="5B21DFDF766C381DFDF2FB5FFB39FE34" box="[635,773,1256,1282]" pageId="11" pageNumber="115">Viraktamath</collectorName>
(
<collectionCode id="90C522CC766C381DFC9AFB5FFB58FE34" box="[787,868,1256,1282]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="11" pageNumber="115">UASB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="46BCB054766C381DFCFDFB5FF8C4FE10" collectingDate="1981-01-18" collectionCode="UASB" collectorName="C. A. Viraktamath" country="India" location="Waghai" pageId="11" pageNumber="115" specimenCount="1" specimenCount-male="1" stateProvince="Gujarat">
<collectingRegion id="341074EB766C381DFCFDFB5FFBF1FE34" box="[884,973,1256,1282]" country="India" name="Gujarat" pageId="11" pageNumber="115">Gujarat</collectingRegion>
:
<specimenCount id="E0D27180766C381DFC5DFB5FFBC5FE34" box="[980,1017,1256,1282]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<location id="F30BECD2766C381DFB8AFB5FFC67FE34" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EFDDDFC50FEEC:F30BECD2766C381DFB8AFB5FFC67FE34" box="[1027,1115,1256,1282]" country="India" name="Waghai" pageId="11" pageNumber="115" stateProvince="Gujarat">Waghai</location>
,
<date id="826A9CC9766C381DFBEAFB5FFCECFE34" box="[1123,1232,1256,1282]" pageId="11" pageNumber="115" value="1981-01-18">
<collectingDate id="922E6521766C381DFBEAFB5FFCECFE34" box="[1123,1232,1256,1282]" pageId="11" pageNumber="115" value="1981-01-18">18.i.1981</collectingDate>
</date>
,
<collectorName id="5B21DFDF766C381DFB50FB5FFDA0FE34" box="[1241,1436,1256,1282]" pageId="11" pageNumber="115">C.A. Viraktamath</collectorName>
(
<collectionCode id="90C522CC766C381DFF16FABBF8CCFE10" box="[159,240,1292,1318]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="11" pageNumber="115">UASB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="46BCB054766C381DFE8AFABBFB88FE10" box="[259,948,1292,1319]" collectingDate="1977-11-18" collectionCode="T, UASB" collectorName="M. M. Ali" country="India" location="Khandala" pageId="11" pageNumber="115" specimenCount="1" specimenCount-male="1" stateProvince="Maharashtra">
<collectingRegion id="341074EB766C381DFE8AFABBF9AFFE10" box="[259,403,1292,1318]" country="India" name="Maharashtra" pageId="11" pageNumber="115">Maharashtra</collectingRegion>
:
<specimenCount id="E0D27180766C381DFE14FABBF9FEFE10" box="[413,450,1292,1318]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<location id="F30BECD2766C381DFE47FABBFA00FE10" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EFDDDFC50FEEC:F30BECD2766C381DFE47FABBFA00FE10" box="[462,572,1292,1318]" country="India" name="Khandala" pageId="11" pageNumber="115" stateProvince="Maharashtra">Khandala</location>
,
<date id="826A9CC9766C381DFDC1FABBFAFEFE10" box="[584,706,1292,1318]" pageId="11" pageNumber="115" value="1977-11-18">
<collectingDate id="922E6521766C381DFDC1FABBFAFEFE10" box="[584,706,1292,1318]" pageId="11" pageNumber="115" value="1977-11-18">18.xi.1977</collectingDate>
</date>
,
<collectionCode id="90C522CC766C381DFD44FABAFADCFE11" box="[717,736,1293,1319]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="115">T</collectionCode>
.
<collectorName id="5B21DFDF766C381DFD6AFABAFB70FE10" box="[739,844,1292,1319]" pageId="11" pageNumber="115">M.M. Ali</collectorName>
(
<collectionCode id="90C522CC766C381DFCD2FABBFB90FE10" box="[859,940,1292,1318]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="11" pageNumber="115">UASB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766C381DFF4EFA87FC50FEEC" pageId="11" pageNumber="115" type="discussion">
<paragraph id="F66BBA09766C381DFF4EFA87FC50FEEC" blockId="11.[151,1437,824,1499]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF4EFA87F907FE7C" bold="true" box="[199,315,1328,1354]" pageId="11" pageNumber="115">Remarks.</emphasis>
<bibRefCitation id="9245C7F8766C381DFECDFA87FA1FFE7C" author="Viraktamath, C. A." box="[324,547,1328,1354]" pageId="11" pageNumber="115" pagination="241 - 244" refId="ref8925" refString="Viraktamath, C. A. (1978) A new species of Cestius Distant (Homoptera: Cicadellidae) from southern India. Journal of Natural History, 12, 241 - 244. https: // doi. org / 10.1080 / 00222937800770131" type="journal article" year="1978">Viraktamath (1978)</bibRefCitation>
adequately described the species and also illustrated the male genitalia; here only the male genitalia are described and illustrations of both male and female genitalia are given in addition to habitus. The color is very variable from light brownish maculation on the head, thorax and abdomen to jet black overall coloration. Some females are very pale with well-developed ivory markings on the pronotum and wings. It resembles
<taxonomicName id="31D4C18A766C381DFE87FA76F9EDFEEC" authorityName="Sunil &amp; Meshram" authorityYear="2023" box="[270,465,1472,1498]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="lamerallis" subGenus="Nandiana">
<emphasis id="C4A0661B766C381DFE87FA76F9EDFEEC" box="[270,465,1472,1498]" italics="true" pageId="11" pageNumber="115">L. (N.) lamerallis</emphasis>
</taxonomicName>
but can be distinguished by the characters given in the key.
</paragraph>
</subSubSection>
</treatment>
</document>
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