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<document id="E67A568AA4DA719CFE5D9AF358EFA056" ID-CLB-Dataset="46003" ID-DOI="10.5281/zenodo.202569" ID-GBIF-Dataset="228bdc70-56b4-4d95-a7f2-beff5f8f1f53" ID-ISSN="1175-5326" ID-Zenodo-Dep="202569" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460408671378" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Yang, Chien-Hui, Okuno, Junji &amp; Chan, Tin-Yam" docDate="2011" docId="7E7187938579FFCDFF56FA93FB6FF99A" docLanguage="en" docName="zt03096p052.pdf" docOrigin="Zootaxa 3096" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Boninpagurus pilosipes Stimpson 1858, n. comb." docType="treatment" docVersion="13" lastPageNumber="45" masterDocId="8248FFEB857BFFC9FFC1FFFCFFD8FF9C" masterDocTitle="Revisiting Pagurus pilosipes (Stimpson, 1858) (Crustacea: Decapoda: Anomura: Paguridae)" masterLastPageNumber="52" masterPageNumber="41" pageNumber="43" updateTime="1732792644779" updateUser="ExternalLinkService">
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<taxonomicName id="31D84D068579FFCBFF56FA93FD55FA15" ID-CoL="d3abfdf7-6976-4017-91fc-a2645316a07c" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[151,653,1391,1417]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="pilosipes" status="comb. nov.">
<emphasis id="C4ACEA978579FFCBFF56FA93FE6DFA15" bold="true" box="[151,437,1391,1417]" italics="true" pageId="2" pageNumber="43">Boninpagurus pilosipes</emphasis>
(Stimpson, 1858)
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,
<taxonomicNameLabel id="DF9F57EC8579FFCBFD5AFA93FCDDFA15" box="[667,773,1391,1417]" pageId="2" pageNumber="43">n. comb.</taxonomicNameLabel>
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(
<figureCitation id="6EE32A008579FFCBFF5EFA6EFEDAFA36" box="[159,258,1426,1450]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="5.[151,250,1825,1848]" captionStartId-1="6.[151,250,1824,1847]" captionStartId-2="7.[151,250,1937,1960]" captionStartId-3="8.[151,250,1867,1890]" captionTargetBox-0="[151,1436,195,1776]" captionTargetBox-1="[248,1353,208,1793]" captionTargetBox-2="[201,1366,215,1937]" captionTargetBox-3="[245,1339,210,1854]" captionTargetId-0="figure@5.[151,1436,195,1776]" captionTargetId-1="figure@6.[233,1376,193,1806]" captionTargetId-2="figure@7.[201,1391,193,1937]" captionTargetId-3="figure@8.[224,1366,193,1854]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1. Boninpagurus pilopsipes (Stimpson, 1858), n. comb. A, D, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands; B, male (SL 1.7 mm; CBM-ZC 10207), Maehama, Chichi-jima Island, Ogasawara Islands; C, E, male with rhizocephalan parasite attached to pleon (SL 3.0 mm; CBM-ZC 10000), Hayama, Sagami Bay. A, B, C, entire animal, dorsal view; D, E, close up of cephalic appendages, showing differences in color pattern of antennular peduncle." captionText-1="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." captionText-2="FIGURE 3. Boninpagurus pilosipes (Stimpson, 1858), n. comb., male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay. A, right cheliped, mesial view (setae omitted); B, same, lateral view; C, left cheliped, mesial view (setae omitted); D, same, lateral view. Scale bar: 1 mm." captionText-3="FIGURE 4. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A, C G, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; B, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, chela of right cheliped, dorsal view (setae omitted); C, chela and carpus of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view; E, left third pereopod, lateral view; F, dactylus of right second pereopod, mesial view; G, dactylus of left third pereopod, mesial view. Scale bars: 1 mm." httpUri-0="https://zenodo.org/record/202570/files/figure.png" httpUri-1="https://zenodo.org/record/202571/files/figure.png" httpUri-2="https://zenodo.org/record/202572/files/figure.png" httpUri-3="https://zenodo.org/record/202573/files/figure.png" pageId="2" pageNumber="43">Figs 14</figureCitation>
)
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<emphasis id="C4ACEA978579FFCBFF56FA27FEB1FA6C" box="[151,361,1498,1520]" italics="true" pageId="2" pageNumber="43">Eupagurus pilosipes</emphasis>
Stimpson, 1858: 249
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(87) [
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locality: Loo Choo (= Okinawa Island)];
<treatmentCitation id="777910948579FFCBFBB3FA26FAF7FA6C" author="Alcock" box="[1138,1327,1498,1520]" page="177" pageId="2" pageNumber="43" year="1905">
<bibRefCitation id="92494B748579FFCBFBB3FA26FB22FA6C" author="Alcock" box="[1138,1274,1498,1520]" pageId="2" pageNumber="43" refString="Alcock, A. (1905) Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum. Part II. Anomura. Fasc. 1. Pagurides. Indian Museum, Calcutta, xi + 197 pp., 16 pls." type="book" year="1905">Alcock 1905</bibRefCitation>
: 177
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;
<treatmentCitation id="777910948579FFCBFAFCFA26FEE8F993" author="Stimpson" page="133" pageId="2" pageNumber="43" year="1907">
<bibRefCitation id="92494B748579FFCBFAFCFA26FF25F993" author="Stimpson" pageId="2" pageNumber="43" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections, 49, 1 - 240, pls. 1 - 26." type="journal article" year="1907">Stimpson 1907</bibRefCitation>
: 133
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;
<treatmentCitation id="777910948579FFCBFEFBFA05FD8EF993" author="Manning" box="[314,598,1529,1551]" page="285" pageId="2" pageNumber="43" year="2006">
<bibRefCitation id="92494B748579FFCBFEFBFA05FDFEF993" author="Manning" box="[314,550,1529,1551]" pageId="2" pageNumber="43" refString="Manning, R. B &amp; Reed, K. J. (2006) Decapod crustaceans deposited in the Zoological Museum of Copenhagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54, 283 - 293." type="journal article" year="2006">Manning &amp; Reed 2006</bibRefCitation>
: 285
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(list).
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</paragraph>
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<taxonomicName id="31D84D068579FFCBFF56F9E5FE94F9B2" ID-CoL="c565937c-1f33-4ded-afe4-ad9f932cc253" box="[151,332,1560,1582]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA978579FFCBFF56F9E5FE94F9B2" box="[151,332,1560,1582]" italics="true" pageId="2" pageNumber="43">Pagurus pilosipes</emphasis>
</taxonomicName>
. —
<treatmentCitation id="777910948579FFCBFEB9F9E4FDDBF9B2" author="Oh" box="[376,515,1560,1582]" page="106" pageId="2" pageNumber="43" year="1983">
<bibRefCitation id="92494B748579FFCBFEB9F9E4FE0CF9B1" author="Oh" box="[376,468,1560,1582]" pageId="2" pageNumber="43" refString="Oh, S. C. (1983) Taxonomical studies on the hermit crabs (Decapoda: Anomura) from Mara Islet, Cheju Island. Journal of the Science and Education Center, Cheju Teachers College, 7, 97 - 124. (in Korean)" type="journal article" year="1983">Oh 1983</bibRefCitation>
: 106
</treatmentCitation>
(in part), pl. 1, figs 3, 4, pl. 2, figs 15;
<treatmentCitation id="777910948579FFCBFC59F9E4FB91F9B1" author="Komai" box="[920,1097,1560,1582]" page="117" pageId="2" pageNumber="43" year="2003">
<bibRefCitation id="92494B748579FFCBFC59F9E4FBC1F9B1" author="Komai" box="[920,1049,1560,1582]" pageId="2" pageNumber="43" refString="Komai, T. (2003) Reassessment of Pagurus pilosipes (Stimpson), supplemental description of P. insulae Asakura, and descriptions of three new species of Pagurus from East Asian waters (Crustacea: Decapoda: Anomura: Paguridae). Natural History Research, 7, 115 - 166." type="journal article" year="2003">Komai 2003</bibRefCitation>
: 117
</treatmentCitation>
, figs 15, 24A;
<treatmentCitation id="777910948579FFCBFB2AF9E5FF28F9D0" author="Hong" page="254" pageId="2" pageNumber="43" year="2006">
<bibRefCitation id="92494B748579FFCBFB2AF9E5FA41F9B2" author="Hong" box="[1259,1433,1560,1582]" pageId="2" pageNumber="43" refString="Hong, B. K., Kim, M. H., Kim, J. N. &amp; Jeon, K. A. (2006) Decapod crustaceans of Dokdo Island, Korea. Journal of Fisheries Science and Technology, 39 (Special Issue), 252 - 258." type="journal article" year="2006">
Hong
<emphasis id="C4ACEA978579FFCBFAEBF9E5FA8DF9B2" box="[1322,1365,1560,1582]" italics="true" pageId="2" pageNumber="43">et al</emphasis>
. 2006
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: 254
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;
<treatmentCitation id="777910948579FFCBFF3BF9CAFDCAF9D0" author="McLaughlin" box="[250,530,1590,1613]" page="34" pageId="2" pageNumber="43" year="2010">
<bibRefCitation id="92494B748579FFCBFF3BF9CAFE35F9D0" author="McLaughlin" box="[250,493,1590,1613]" pageId="2" pageNumber="43" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. and Rahayu D. L. (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galethidae of the Galatheoidea) Part 1 - Lithodoidea, Lomissoidea and Paguroidea. Raffles Bulletin of Zoology, Supplement 23, 5 - 107." type="book chapter" year="2010">
McLaughlin
<emphasis id="C4ACEA978579FFCBFEBCF9C4FE72F9D1" box="[381,426,1591,1613]" italics="true" pageId="2" pageNumber="43">et al</emphasis>
. 2010
</bibRefCitation>
: 34
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. [See Remarks]
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</paragraph>
<paragraph id="F66736858579FFCBFF56F9AAFDADF915" blockId="2.[151,1437,1498,1858]" pageId="2" pageNumber="43">
<treatmentCitationGroup id="D6C811AB8579FFCBFF56F9AAFDADF915" pageId="2" pageNumber="43">
<taxonomicName id="31D84D068579FFCBFF56F9AAFCF1F9F7" ID-CoL="436f9f4e-023d-4ad6-b9d1-0727d019ceb1" authority="Asakura &amp; Tachikawa, 2004: 158" authorityName="Asakura &amp; Tachikawa" authorityPageNumber="158" authorityYear="2004" box="[151,809,1621,1643]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="acanthocheles">
<emphasis id="C4ACEA978579FFCBFF56F9AAFE18F9F7" box="[151,448,1621,1643]" italics="true" pageId="2" pageNumber="43">Boninpagurus acanthocheles</emphasis>
<treatmentCitation id="777910948579FFCBFE08F9A9FCF1F9F7" author="Asakura" box="[457,809,1621,1643]" page="158" pageId="2" pageNumber="43" year="2004">
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: 158
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</taxonomicName>
, figs 17 [
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locality: Ogasawara Islands];
<treatmentCitation id="777910948579FFCBFAC4F9A9FF37F915" author="Asakura" page="145" pageId="2" pageNumber="43" year="2006">
<bibRefCitation id="92494B748579FFCBFAC4F9A9FA40F9F7" author="Asakura" box="[1285,1432,1621,1643]" pageId="2" pageNumber="43" refString="Asakura, A. (2006) Descriptions of color patterns in selected species of pagurid hermit crabs with sexual tubes (Decapoda: Anomura: Paguridae) from Japan. In: Asakura, A. (Ed.), Biology of Anomura II. Crustacean Research, Special Number, 6, 133 - 149." type="journal article" year="2006">Asakura 2006</bibRefCitation>
: 145
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, figs 8, 9.
<treatmentCitation id="777910948579FFCBFE99F98FFDA9F915" author="McLaughlin" box="[344,625,1651,1674]" page="27" pageId="2" pageNumber="43" year="2010">
<bibRefCitation id="92494B748579FFCBFE99F98FFD93F915" author="McLaughlin" box="[344,587,1651,1674]" pageId="2" pageNumber="43" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. and Rahayu D. L. (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galethidae of the Galatheoidea) Part 1 - Lithodoidea, Lomissoidea and Paguroidea. Raffles Bulletin of Zoology, Supplement 23, 5 - 107." type="book chapter" year="2010">
McLaughlin
<emphasis id="C4ACEA978579FFCBFE1DF989FDD0F916" box="[476,520,1652,1674]" italics="true" pageId="2" pageNumber="43">et al</emphasis>
. 2010
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: 27
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.
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</paragraph>
<paragraph id="F66736858579FFCBFF56F96FFBD1F934" blockId="2.[151,1437,1498,1858]" box="[151,1033,1682,1704]" pageId="2" pageNumber="43">
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<emphasis id="C4ACEA978579FFCBFF56F96FFEFEF934" box="[151,294,1683,1704]" italics="true" pageId="2" pageNumber="43">Boninpagurus</emphasis>
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sp. —
<treatmentCitation id="777910948579FFCBFEAEF96EFD85F934" author="Okuno" box="[367,605,1682,1704]" page="153" pageId="2" pageNumber="43" year="2006">
<bibRefCitation id="92494B748579FFCBFEAEF96EFDF2F934" author="Okuno" box="[367,554,1682,1704]" pageId="2" pageNumber="43" refString="Okuno, J., Takeda, M. &amp; Yokota, M. (2006) Inshore hermit crabs (Crustacea: Decapoda: Anomura) collected from the Izu Oceanic Park, eastern coast of Izu Peninsula, Japan. Memoirs of the National Science Museum, 41, 145 - 171." type="journal article" year="2006">
Okuno
<emphasis id="C4ACEA978579FFCBFE7AF96FFE3EF934" box="[443,486,1682,1704]" italics="true" pageId="2" pageNumber="43">et al</emphasis>
. 2006
</bibRefCitation>
: 153
</treatmentCitation>
;
<treatmentCitation id="777910948579FFCBFDA6F96EFCA6F934" author="McLaughlin" box="[615,894,1682,1704]" page="88" pageId="2" pageNumber="43" year="2010">
<bibRefCitation id="92494B748579FFCBFDA6F96EFC81F934" author="McLaughlin" box="[615,857,1682,1704]" pageId="2" pageNumber="43" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. and Rahayu D. L. (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galethidae of the Galatheoidea) Part 1 - Lithodoidea, Lomissoidea and Paguroidea. Raffles Bulletin of Zoology, Supplement 23, 5 - 107." type="book chapter" year="2010">
McLaughlin
<emphasis id="C4ACEA978579FFCBFD2BF96FFCCEF934" box="[746,790,1682,1704]" italics="true" pageId="2" pageNumber="43">et al</emphasis>
. 2010
</bibRefCitation>
: 88
</treatmentCitation>
, 89, fig. 12C.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F66736858579FFCBFF56F94EFD7FF8DE" blockId="2.[151,1437,1498,1858]" pageId="2" pageNumber="43">
<treatmentCitationGroup id="D6C811AB8579FFCBFF56F94EFD7FF8DE" pageId="2" pageNumber="43">
Not
<taxonomicName id="31D84D068579FFCBFF05F94EFEA2F95B" ID-CoL="c565937c-1f33-4ded-afe4-ad9f932cc253" box="[196,378,1713,1735]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA978579FFCBFF05F94EFEA2F95B" box="[196,378,1713,1735]" italics="true" pageId="2" pageNumber="43">Pagurus pilosipes</emphasis>
</taxonomicName>
. —
<treatmentCitation id="777910948579FFCBFE69F94DFD8FF95B" author="Miyake" box="[424,599,1713,1735]" page="91" pageId="2" pageNumber="43" year="1978">
<bibRefCitation id="92494B748579FFCBFE69F94DFDEAF95B" author="Miyake" box="[424,562,1713,1735]" pageId="2" pageNumber="43" refString="Miyake, S. (1978) The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 pp. (English), 161 pp. (Japanese), 4 pls." type="book" year="1978">Miyake 1978</bibRefCitation>
: 91
</treatmentCitation>
, fig. 34 (= mixture of seven species; see
<bibRefCitation id="92494B748579FFCBFC3AF94DFB5CF954" author="Komai" box="[1019,1156,1713,1736]" pageId="2" pageNumber="43" refString="Komai, T. (2003) Reassessment of Pagurus pilosipes (Stimpson), supplemental description of P. insulae Asakura, and descriptions of three new species of Pagurus from East Asian waters (Crustacea: Decapoda: Anomura: Paguridae). Natural History Research, 7, 115 - 166." type="journal article" year="2003">Komai, 2003</bibRefCitation>
);
<treatmentCitation id="777910948579FFCBFB56F94DFF3BF97A" author="Miyake" page="60" pageId="2" pageNumber="43" year="1980">
<bibRefCitation id="92494B748579FFCBFB56F94DFA40F95B" author="Miyake" box="[1175,1432,1713,1735]" pageId="2" pageNumber="43" refString="Miyake, S. &amp; Imafuku, M. (1980) Hermit crabs from Kii Peninsula II. Nankiseibutu: Nanki Biological Society, 22 (2), 59 - 64, pls 1, 2." type="journal article" year="1980">Miyake &amp; Imafuku 1980</bibRefCitation>
: 60
</treatmentCitation>
, pl. 2, fig. 5 (=
<taxonomicName id="31D84D068579FFCBFEBFF92DFD3CF97A" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" authority="Komai, 2003" authorityName="Komai" authorityYear="2003" box="[382,740,1744,1766]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFEBFF92DFD8EF97A" box="[382,598,1745,1766]" italics="true" pageId="2" pageNumber="43">Pagurus nigrivittatus</emphasis>
<bibRefCitation id="92494B748579FFCBFD9DF92CFD3CF97A" author="Komai" box="[604,740,1744,1766]" pageId="2" pageNumber="43" refString="Komai, T. (2003) Reassessment of Pagurus pilosipes (Stimpson), supplemental description of P. insulae Asakura, and descriptions of three new species of Pagurus from East Asian waters (Crustacea: Decapoda: Anomura: Paguridae). Natural History Research, 7, 115 - 166." type="journal article" year="2003">Komai, 2003</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="777910948579FFCBFD37F92CFC69F979" author="Miyake" box="[758,945,1744,1766]" page="132" pageId="2" pageNumber="43" year="1982">
<bibRefCitation id="92494B748579FFCBFD37F92CFCA7F979" author="Miyake" box="[758,895,1744,1766]" pageId="2" pageNumber="43" refString="Miyake, S. (1982) Japanese Crustacean Decapods and Stomatopods in Color, Volume 1: Macrura, Anomura and Stomatopoda. Hoikusha, Osaka, viii + 261 pp., 56 pls. (In Japanese)" type="book" year="1982">Miyake 1982</bibRefCitation>
: 132
</treatmentCitation>
, pl. 44, fig. 5 (=
<taxonomicName id="31D84D068579FFCBFB9BF92DFB37F97A" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" box="[1114,1263,1745,1766]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFB9BF92DFB37F97A" box="[1114,1263,1745,1766]" italics="true" pageId="2" pageNumber="43">P. nigrivittatus</emphasis>
</taxonomicName>
);
<bibRefCitation id="92494B748579FFCBFAC5F92CFA87F979" author="Oh" box="[1284,1375,1744,1766]" pageId="2" pageNumber="43" refString="Oh, S. C. (1983) Taxonomical studies on the hermit crabs (Decapoda: Anomura) from Mara Islet, Cheju Island. Journal of the Science and Education Center, Cheju Teachers College, 7, 97 - 124. (in Korean)" type="journal article" year="1983">Oh 1983</bibRefCitation>
: pl. 1, figs. 3, 4, pl. 3, figs. 15 (=
<taxonomicName id="31D84D068579FFCBFE21F90CFDADF899" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" box="[480,629,1776,1797]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFE21F90CFDADF899" box="[480,629,1776,1797]" italics="true" pageId="2" pageNumber="43">P. nigrivittatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="777910948579FFCBFD4BF912FC99F898" author="Takeda" box="[650,833,1774,1796]" page="124" pageId="2" pageNumber="43" year="1986">
<bibRefCitation id="92494B748579FFCBFD4BF912FCD7F898" author="Takeda" box="[650,783,1774,1796]" pageId="2" pageNumber="43" refString="Takeda, M. (1986) [Anomura]. In: Masuda, H., Hayashi, K., Nakamura, K. &amp; Kobayashi, Y. (Eds.) Marine Invertebrates. Tokai University Press, Tokyo, pp. 104 - 209. (In Japanese)" type="book chapter" year="1986">Takeda 1986</bibRefCitation>
: 124
</treatmentCitation>
, unnumbered fig. (=
<taxonomicName id="31D84D068579FFCBFBD4F90CFB71F899" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" box="[1045,1193,1776,1797]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFBD4F90CFB71F899" box="[1045,1193,1776,1797]" italics="true" pageId="2" pageNumber="43">P. nigrivittatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="777910948579FFCBFB7EF912FAA7F898" author="Asakura" box="[1215,1407,1774,1796]" page="798" pageId="2" pageNumber="43" year="1991">
<bibRefCitation id="92494B748579FFCBFB7EF912FA97F898" author="Asakura" box="[1215,1359,1774,1796]" pageId="2" pageNumber="43" refString="Asakura, A. (1991) A new hermit crab (Decapoda: Paguridea) from rocky shores of the Ogasawara (Bonin) Islands. Proceedings of the Biological Society of Washington, 104, 793 - 799." type="journal article" year="1991">Asakura 1991</bibRefCitation>
: 798
</treatmentCitation>
(=
<taxonomicName id="31D84D068579FFCBFF06F8F2FE86F8BF" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" box="[199,350,1806,1827]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFF06F8F2FE86F8BF" box="[199,350,1806,1827]" italics="true" pageId="2" pageNumber="43">P. nigrivittatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="777910948579FFCBFEB4F8F2FDCBF8BF" author="Wang" box="[373,531,1805,1827]" page="61" pageId="2" pageNumber="43" year="1992">
<bibRefCitation id="92494B748579FFCBFEB4F8F2FE36F8BF" author="Wang" box="[373,494,1806,1827]" pageId="2" pageNumber="43" refString="Wang, F. - Z. (1992) Studies on the hermit crabs fauna of China (Crustacea, Anomura). Donghai Marine Science, 10, 59 - 63. (In Chinese, with English summary)" type="journal article" year="1992">Wang 1992</bibRefCitation>
: 61
</treatmentCitation>
(list); 1994: 570 (list);
<treatmentCitation id="777910948579FFCBFCC8F8F1FC08F8BF" author="Asakura" box="[777,976,1805,1827]" page="363" pageId="2" pageNumber="43" year="1995">
<bibRefCitation id="92494B748579FFCBFCC8F8F1FC44F8BF" author="Asakura" box="[777,924,1805,1827]" pageId="2" pageNumber="43" refString="Asakura, A. (1995) Anomura. In: Nishimura, S. (Ed.), Guide to Sea Shore Animals of Japan with Color Pictures and Keys, Vol. II. Hoikusha, Osaka, pp. 347 - 378, pls. 93 - 100. (In Japanese)." type="book chapter" year="1995">Asakura 1995</bibRefCitation>
: 363
</treatmentCitation>
, pl. 97, fig. 9 (=
<taxonomicName id="31D84D068579FFCBFB44F8F2FAC4F8BF" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" box="[1157,1308,1806,1827]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFB44F8F2FAC4F8BF" box="[1157,1308,1806,1827]" italics="true" pageId="2" pageNumber="43">P. nigrivittatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="777910948579FFCBFAF5F8F1FEF7F8DD" author="Minemizu" page="148" pageId="2" pageNumber="43" year="2000">
<bibRefCitation id="92494B748579FFCBFAF5F8F1FF25F8DD" author="Minemizu" pageId="2" pageNumber="43" refString="Minemizu, R. (2000) Marine Decapod and Stomatopod Crustaceans mainly from Japan. Bun-ichi Sogo Publishing Co., Tokyo, 344 pp. (In Japanese)" type="book" year="2000">Minemizu 2000</bibRefCitation>
: 148
</treatmentCitation>
, unnumbered fig. (=
<taxonomicName id="31D84D068579FFCBFDC5F8D1FD40F8DE" ID-CoL="937738c5-dbbd-46e3-ac7a-f979fd4d4580" box="[516,664,1837,1858]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="nigrivittatus">
<emphasis id="C4ACEA978579FFCBFDC5F8D1FD40F8DE" box="[516,664,1837,1858]" italics="true" pageId="2" pageNumber="43">P. nigrivittatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BEC2650E8579FFCAFF56F892FB88FBD6" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43" type="materials_examined">
<paragraph id="F66736858579FFCBFF56F892FBEBF836" blockId="2.[151,1436,1902,2034]" pageId="2" pageNumber="43">
<emphasis id="C4ACEA978579FFCBFF56F892FE5CF81B" bold="true" box="[151,388,1902,1927]" pageId="2" pageNumber="43">Material examined.</emphasis>
<typeStatus id="296388278579FFCBFE4EF893FE2AF81B" box="[399,498,1903,1927]" pageId="2" pageNumber="43" type="neotype">Neotype</typeStatus>
: female (SL
<quantity id="31209B608579FFCBFD4FF892FD34F81A" box="[654,748,1902,1926]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="43" unit="mm" value="2.5">2.5 mm</quantity>
), CBM-ZC 6383, Heshikiya, Katsuren Peninsula, Okinawa Island, intertidal, on small colony of
<emphasis id="C4ACEA978579FFCBFDF5F86FFD6AF837" box="[564,690,1939,1963]" italics="true" pageId="2" pageNumber="43">Pocilopora</emphasis>
sp.,
<date id="826610458579FFCBFD26F86EFCA7F836" box="[743,895,1938,1962]" pageId="2" pageNumber="43" value="1994-06-23">23 June 1994</date>
, coll. T. Komai.
</paragraph>
<paragraph id="F66736858579FFCAFF04F84BFD80FCB2" blockId="2.[151,1436,1902,2034]" lastBlockId="3.[151,1436,151,2020]" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
Japanese mainland:
<specimenCount id="E0DEFD0C8579FFCBFE68F84AFDD3F852" box="[425,523,1974,1998]" pageId="2" pageNumber="43" type="female">1 female</specimenCount>
(SL
<quantity id="31209B608579FFCBFD84F84AFD78F852" box="[581,672,1974,1998]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="43" unit="mm" value="2.7">2.7 mm</quantity>
), CMNH-ZC 629, Igai-jima Islet, Kamogawa,
<quantity id="31209B608579FFCBFB75F84AFB29F852" box="[1204,1265,1974,1998]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="43" unit="m" value="16.0">16 m</quantity>
,
<date id="826610458579FFCBFB3CF84AFA40F852" box="[1277,1432,1974,1998]" pageId="2" pageNumber="43" value="2001-06-29">29 June 2001</date>
,
<collectionCode id="90C9AE408579FFCBFF56F826FF2DF86E" box="[151,245,2010,2034]" pageId="2" pageNumber="43">SCUBA</collectionCode>
diving, coll. J. Okuno;
<specimenCount id="E0DEFD0C8579FFCBFDC6F826FD8CF86E" box="[519,596,2010,2034]" pageId="2" pageNumber="43" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608579FFCBFD50F826FD35F86E" box="[657,749,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="43" unit="mm" value="3.7">3.7 mm</quantity>
), 1 ovigerous female (SL
<quantity id="31209B608579FFCBFBD8F826FBADF86E" box="[1049,1141,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="43" unit="mm" value="3.2">3.2 mm</quantity>
), CMNH-ZC 630, similar locality,
<quantity id="31209B608578FFCAFF39FF6BFEECFF33" box="[248,308,151,175]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="44" unit="m" value="20.0">20 m</quantity>
,
<date id="826610458578FFCAFE80FF6BFDC7FF33" box="[321,543,151,175]" pageId="3" pageNumber="44" value="2001-11-16">16 November 2001</date>
,
<collectionCode id="90C9AE408578FFCAFDEAFF6BFD51FF33" box="[555,649,151,175]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno; 1 ovigerous female (SL
<quantity id="31209B608578FFCAFB6DFF6BFADFFF33" box="[1196,1287,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="44" unit="mm" value="2.7">2.7 mm</quantity>
), CMNH-ZC 631, same data;
<specimenCount id="E0DEFD0C8578FFCAFE9DFF47FE76FF4F" box="[348,430,187,211]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFE31FF47FD97FF4F" box="[496,591,187,211]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="44" unit="mm" value="2.3">2.3 mm</quantity>
), CMNH-ZC, Ubara Islet, Katsuura, Boso Peninsula,
<date id="826610458578FFCAFB0FFF47FA40FF4F" box="[1230,1432,187,211]" pageId="3" pageNumber="44" value="2001-10-24">24 October 2001</date>
,
<collectionCode id="90C9AE408578FFCAFF56FF23FF2DFF6B" box="[151,245,223,247]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno;
<specimenCount id="E0DEFD0C8578FFCAFDCBFF23FD80FF6B" box="[522,600,223,247]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFD57FF23FD2BFF6B" box="[662,755,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="44" unit="mm" value="2.7">2.7 mm</quantity>
), CMNH-ZC 1330, Awatone, off Kamogawa,
<quantity id="31209B608578FFCAFAC8FF23FAA9FF6B" box="[1289,1393,223,247]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="44" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">1520 m</quantity>
,
<date id="826610458578FFCAFA41FF23FE95FE86" pageId="3" pageNumber="44" value="2002-12-18">18 December 2002</date>
,
<collectionCode id="90C9AE408578FFCAFE98FEFEFE6EFE86" box="[345,438,258,282]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. H. Tachikawa;
<specimenCount id="E0DEFD0C8578FFCAFD3BFEFEFC91FE86" box="[762,841,258,282]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
, CMNH-ZC 1331, same data as CMNH-ZC 1330;
<specimenCount id="E0DEFD0C8578FFCAFA4FFEFEFF15FEA2" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFECFFEDAFEB5FEA2" box="[270,365,294,318]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="44" unit="mm" value="3.1">3.1 mm</quantity>
), CMNH-ZC 1346, Ashika-ne, off Kamogawa, Boso Peninsula,
<quantity id="31209B608578FFCAFBA5FEDAFB7BFEA2" box="[1124,1187,294,318]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="44" unit="m" value="26.0">26 m</quantity>
,
<date id="826610458578FFCAFB75FEDAFA40FEA2" box="[1204,1432,294,318]" pageId="3" pageNumber="44" value="2000-11-26">26 November 2000</date>
,
<collectionCode id="90C9AE408578FFCAFF56FEB5FF2DFEFD" box="[151,245,329,353]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno;
<specimenCount id="E0DEFD0C8578FFCAFDC4FEB5FDBFFEFD" box="[517,615,329,353]" pageId="3" pageNumber="44" type="female">1 female</specimenCount>
(SL
<quantity id="31209B608578FFCAFD63FEB5FD26FEFD" box="[674,766,329,353]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="44" unit="mm" value="2.5">2.5 mm</quantity>
), CMNH-ZC 1347, same data as CMNH-ZC 1346;
<specimenCount id="E0DEFD0C8578FFCAFA8EFEB5FA44FEFD" box="[1359,1436,329,353]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFF0DFE90FEF2FE18" box="[204,298,364,388]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="44" unit="mm" value="3.4">3.4 mm</quantity>
), CMNH-ZC 1378, Hatto-ne, off Kamogawa,
<quantity id="31209B608578FFCAFC88FE90FC51FE18" box="[841,905,364,388]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="44" unit="m" value="15.0">15 m</quantity>
,
<date id="826610458578FFCAFC58FE90FBA3FE18" box="[921,1147,364,388]" pageId="3" pageNumber="44" value="2000-11-26">26 November 2000</date>
,
<collectionCode id="90C9AE408578FFCAFB4AFE90FB30FE18" box="[1163,1256,364,388]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno; 1 ovigerous female (SL
<quantity id="31209B608578FFCAFDC6FE6CFDBBFE34" box="[519,611,400,424]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="44" unit="mm" value="2.1">2.1 mm</quantity>
), CMNH-ZC 1615, Mankuro-ne, off Kamogawa,
<quantity id="31209B608578FFCAFB5BFE6CFB0FFE34" box="[1178,1239,400,424]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="44" unit="m" value="20.0">20 m</quantity>
,
<date id="826610458578FFCAFB25FE6CFA40FE34" box="[1252,1432,400,424]" pageId="3" pageNumber="44" value="2003-10-01">1 October 2003</date>
,
<collectionCode id="90C9AE408578FFCAFF56FE4FFF2CFE57" box="[151,244,435,459]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. H. Tachikawa
<emphasis id="C4ACEA978578FFCAFDFCFE49FDAAFE50" box="[573,626,436,460]" italics="true" pageId="3" pageNumber="44">et al</emphasis>
.;
<specimenCount id="E0DEFD0C8578FFCAFD4DFE4FFD29FE57" box="[652,753,435,459]" pageId="3" pageNumber="44" type="female">1 female</specimenCount>
(SL
<quantity id="31209B608578FFCAFCF1FE4FFC56FE57" box="[816,910,435,459]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="44" unit="mm" value="2.9">2.9 mm</quantity>
), CMNH-ZC 1729, Akino-hama, Izu-Oshima
<collectingCountry id="8ECF76158578FFCAFF56FE2BFF06FE73" box="[151,222,471,495]" name="Iceland" pageId="3" pageNumber="44">Island</collectingCountry>
, Izu Islands,
<quantity id="31209B608578FFCAFEB8FE2BFE6FFE73" box="[377,439,471,495]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="44" unit="m" value="15.0">15 m</quantity>
,
<date id="826610458578FFCAFE04FE2BFD4CFE73" box="[453,660,471,495]" pageId="3" pageNumber="44" value="2004-02-23">23 February 2004</date>
,
<collectionCode id="90C9AE408578FFCAFD63FE2BFD27FE73" box="[674,767,471,495]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. H. Arima; 1 ovigerous female (SL
<quantity id="31209B608578FFCAFAF2FE2BFA48FE73" box="[1331,1424,471,495]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="44" unit="mm" value="1.8">1.8 mm</quantity>
), CMNH-ZC 1740, same locality,
<quantity id="31209B608578FFCAFDD9FE06FD8FFD8E" box="[536,599,506,530]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="44" unit="m" value="15.0">15 m</quantity>
,
<date id="826610458578FFCAFDA9FE06FCC8FD8E" box="[616,784,506,530]" pageId="3" pageNumber="44" value="2004-03-06">6 March 2004</date>
,
<collectionCode id="90C9AE408578FFCAFCE1FE06FCA6FD8E" box="[800,894,506,530]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. H. Arima;
<specimenCount id="E0DEFD0C8578FFCAFB60FE06FB28FD8E" box="[1185,1264,506,530]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFAF3FE06FA48FD8E" box="[1330,1424,506,530]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="44" unit="mm" value="1.7">1.7 mm</quantity>
), CMNH-ZC 1743, same locality,
<quantity id="31209B608578FFCAFDD2FDE2FD89FDAA" box="[531,593,542,566]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="44" unit="m" value="25.0">25 m</quantity>
,
<date id="826610458578FFCAFDA1FDE2FD0BFDAA" box="[608,723,542,566]" pageId="3" pageNumber="44" value="1913-03">13 March</date>
, 2004,
<collectionCode id="90C9AE408578FFCAFCEBFDE2FC50FDAA" box="[810,904,542,566]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. H. Arima;
<specimenCount id="E0DEFD0C8578FFCAFB67FDE2FB2CFDAA" box="[1190,1268,542,566]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFAF2FDE2FA48FDAA" box="[1331,1424,542,566]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="44" unit="mm" value="2.8">2.8 mm</quantity>
), CMNH-ZC 1837, Masaki, Shimizu, Suruga Bay,
<quantity id="31209B608578FFCAFD00FDBDFD36FDC5" box="[705,750,577,601]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="44" unit="m" value="8.0">8 m</quantity>
,
<date id="826610458578FFCAFD3AFDBDFC10FDC5" box="[763,968,577,601]" pageId="3" pageNumber="44" value="2005-11-04">4 November 2005</date>
,
<collectionCode id="90C9AE408578FFCAFC12FDBDFBE8FDC5" box="[979,1072,577,601]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno;
<specimenCount id="E0DEFD0C8578FFCAFAFAFDBDFA44FDC5" box="[1339,1436,577,601]" pageId="3" pageNumber="44" type="female">1 female</specimenCount>
(SL 2.0 mm), CMNH-ZC 1838, same data as CMNH-ZC 1837;
<specimenCount id="E0DEFD0C8578FFCAFCA9FD99FC12FDE1" box="[872,970,613,637]" pageId="3" pageNumber="44" type="female">1 female</specimenCount>
(SL 2.0 mm), CMNH-ZC, Tatsunokuchi, Nagasaki, Kyushu,
<quantity id="31209B608578FFCAFEB9FD74FE70FD3C" box="[376,424,648,672]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="44" unit="m" value="2.0">2 m</quantity>
,
<date id="826610458578FFCAFE76FD74FD41FD3C" box="[439,665,648,672]" pageId="3" pageNumber="44" value="2003-09-13">13 September 2003</date>
,
<collectionCode id="90C9AE408578FFCAFD69FD74FCDDFD3C" box="[680,773,648,672]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno;
<specimenCount id="E0DEFD0C8578FFCAFBDDFD74FBB2FD3C" box="[1052,1130,648,672]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL 2.0 mm), CMNH-ZC 1982, Mankuro-ne, off Kamogawa, Boso Peninsula,
<quantity id="31209B608578FFCAFD3DFD50FCE3FD58" box="[764,827,684,708]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="44" unit="m" value="18.0">18 m</quantity>
,
<date id="826610458578FFCAFC8BFD50FBDFFD58" box="[842,1031,684,708]" pageId="3" pageNumber="44" value="2005-08-11">11 August 2005</date>
,
<collectionCode id="90C9AE408578FFCAFBD4FD50FBAAFD58" box="[1045,1138,684,708]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. J. Okuno;
<specimenCount id="E0DEFD0C8578FFCAFA4FFD50FF00FD7B" pageId="3" pageNumber="44" type="male">2 males</specimenCount>
(SL 2.8, 3.0 mm), CBM-ZC 10000, Sagami Bay, Hayama, Miura Peninsula, ca.
<quantity id="31209B608578FFCAFBB1FD33FB74FD7B" box="[1136,1196,719,743]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="44" unit="m" value="10.0">10 m</quantity>
,
<date id="826610458578FFCAFB78FD33FA40FD7B" box="[1209,1432,719,743]" pageId="3" pageNumber="44" value="2009-09-23">23 September 2009</date>
,
<collectionCode id="90C9AE408578FFCAFF56FD0FFF2CFC97" box="[151,244,755,779]" pageId="3" pageNumber="44">SCUBA</collectionCode>
diving, coll. I. Itou and S. Komai;
<specimenCount id="E0DEFD0C8578FFCAFD51FD0FFD05FC97" box="[656,733,755,779]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFCDBFD0FFCAEFC97" box="[794,886,755,779]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="44" unit="mm" value="2.2">2.2 mm</quantity>
),
<collectionCode id="90C9AE408578FFCAFC4DFD0FFC01FC97" box="[908,985,755,779]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="3" pageNumber="44">NTOU</collectionCode>
<accessionNumber id="E98BAB668578FFCAFC22FD0FFB9CFC97" box="[995,1092,755,779]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M01037" pageId="3" pageNumber="44" type="EnaNcbi">M01037</accessionNumber>
,
<specimenCount id="E0DEFD0C8578FFCAFB93FD0FFB47FC97" box="[1106,1183,755,779]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFB1CFD0FFAE1FC97" box="[1245,1337,755,779]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="44" unit="mm" value="2.1">2.1 mm</quantity>
),
<collectionCode id="90C9AE408578FFCAFA8EFD0FFA44FC97" box="[1359,1436,755,779]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="3" pageNumber="44">NTOU</collectionCode>
<accessionNumber id="E98BAB668578FFCAFF56FCEAFF20FCB2" box="[151,248,790,814]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M01038" pageId="3" pageNumber="44" type="EnaNcbi">M01038</accessionNumber>
, same data as CBM-ZC 10000.
</paragraph>
<paragraph id="F66736858578FFCAFF04FCC5FB0DFC7C" blockId="3.[151,1436,151,2020]" pageId="3" pageNumber="44">
Okinawa Islands,
<collectingCountry id="8ECF76158578FFCAFE51FCC6FE0CFCCD" box="[400,468,826,849]" name="Japan" pageId="3" pageNumber="44">Japan</collectingCountry>
:
<specimenCount id="E0DEFD0C8578FFCAFE1EFCC5FDF3FCCD" box="[479,555,825,849]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFDA4FCC5FD67FCCD" box="[613,703,825,849]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="44" unit="mm" value="1.9">1.9 mm</quantity>
),
<specimenCount id="E0DEFD0C8578FFCAFD12FCC5FC98FCCD" box="[723,832,825,849]" pageId="3" pageNumber="44" type="female">2 females</specimenCount>
(SL 2.2,
<quantity id="31209B608578FFCAFC6AFCC5FBDEFCCD" box="[939,1030,825,849]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="44" unit="mm" value="2.4">2.4 mm</quantity>
), CBM-ZC 6384, same data as
<typeStatus id="296388278578FFCAFAABFCC6FF13FCEA" pageId="3" pageNumber="44" type="neotype">neotype</typeStatus>
;
<specimenCount id="E0DEFD0C8578FFCAFF17FCA1FEF5FCE9" box="[214,301,861,885]" pageId="3" pageNumber="44" type="male">2 males</specimenCount>
(SL 2.0, 2.0 mm),
<specimenCount id="E0DEFD0C8578FFCAFDC2FCA1FDBCFCE9" box="[515,612,861,885]" pageId="3" pageNumber="44" type="female">1 female</specimenCount>
(SL
<quantity id="31209B608578FFCAFD5DFCA1FD2EFCE9" box="[668,758,861,885]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="44" unit="mm" value="1.7">1.7 mm</quantity>
), CBM-ZC 10209, Heshikiya, Katsuren Peninsula, Okinawa Island, subtidal, dead coral,
<date id="826610458578FFCAFE18FC7CFDA8FC04" box="[473,624,896,920]" pageId="3" pageNumber="44" value="2003-07-17">17 July 2003</date>
, coll. M. Osawa;
<specimenCount id="E0DEFD0C8578FFCAFCFDFC7CFC50FC04" box="[828,904,896,920]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFC05FC7CFBF8FC04" box="[964,1056,896,920]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="44" unit="mm" value="2.8">2.8 mm</quantity>
), CBM-ZC 10206, Kouri Fishing Port, Kouri Island, 2.0
<quantity id="31209B608578FFCAFE63FC58FE3BFC20" box="[418,483,932,956]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="44" unit="m" value="2.5">2.5 m</quantity>
,
<date id="826610458578FFCAFE31FC58FD63FC20" box="[496,699,932,956]" pageId="3" pageNumber="44" value="2007-02-25">25 February 2007</date>
, trap, coll. T. Komai; 1 ovigerous female (SL
<quantity id="31209B608578FFCAFB02FC58FAC5FC20" box="[1219,1309,932,956]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="44" unit="mm" value="3.3">3.3 mm</quantity>
), CBM-ZC 10275, Motobu Fishing Port, Motobu, Okinawa Island,
<quantity id="31209B608578FFCAFCC6FC34FC88FC7C" box="[775,848,968,992]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="44" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">34 m</quantity>
,
<date id="826610458578FFCAFC9DFC34FC3FFC7C" box="[860,999,968,992]" pageId="3" pageNumber="44" value="2010-03">March 2010</date>
, trap, coll. T. Komai.
</paragraph>
<paragraph id="F66736858578FFCAFF04FC17FB88FBD6" blockId="3.[151,1436,151,2020]" pageId="3" pageNumber="44">
Ogasawara Islands,
<collectingCountry id="8ECF76158578FFCAFE6AFC10FE37FB9F" box="[427,495,1004,1027]" name="Japan" pageId="3" pageNumber="44">Japan</collectingCountry>
:
<specimenCount id="E0DEFD0C8578FFCAFE3CFC17FD8DFB9F" box="[509,597,1003,1027]" pageId="3" pageNumber="44" type="male">2 males</specimenCount>
(SL 1.4,
<quantity id="31209B608578FFCAFD06FC17FCFCFB9F" box="[711,804,1003,1027]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="44" unit="mm" value="1.7">1.7 mm</quantity>
),
<specimenCount id="E0DEFD0C8578FFCAFCF8FC17FC70FB9F" box="[825,936,1003,1027]" pageId="3" pageNumber="44" type="female">2 females</specimenCount>
(SL 1.5,
<quantity id="31209B608578FFCAFBD8FC17FBAEFB9F" box="[1049,1142,1003,1027]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="44" unit="mm" value="1.5">1.5 mm</quantity>
),
<specimenCount id="E0DEFD0C8578FFCAFB4AFC17FAD0FB9F" box="[1163,1288,1003,1027]" pageId="3" pageNumber="44" type="juvenile">2 juveniles</specimenCount>
(SL 1.2,
<quantity id="31209B608578FFCAFAB8FC17FF1FFBBA" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="44" unit="mm" value="1.3">1.3 mm</quantity>
), CBM-ZC 10207, same locality, subtidal, coral rubble,
<date id="826610458578FFCAFCFAFBF2FC16FBBA" box="[827,974,1038,1062]" pageId="3" pageNumber="44" value="2009-07-17">17 July 2009</date>
, coll. T. Komai;
<collectionCode id="90C9AE408578FFCAFB48FBF2FB0EFBBA" box="[1161,1238,1038,1062]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="3" pageNumber="44">NTOU</collectionCode>
M 0 1039,
<specimenCount id="E0DEFD0C8578FFCAFA90FBF2FA43FBBA" box="[1361,1435,1038,1062]" pageId="3" pageNumber="44" type="male">1 male</specimenCount>
(SL
<quantity id="31209B608578FFCAFF09FBCEFEFAFBD6" box="[200,290,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="44" unit="mm" value="1.4">1.4 mm</quantity>
),
<collectionCode id="90C9AE408578FFCAFEF4FBCEFE5AFBD6" box="[309,386,1074,1098]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="3" pageNumber="44">NTOU</collectionCode>
<accessionNumber id="E98BAB668578FFCAFE48FBCEFE32FBD6" box="[393,490,1074,1098]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M01040" pageId="3" pageNumber="44" type="EnaNcbi">M01040</accessionNumber>
,
<specimenCount id="E0DEFD0C8578FFCAFE37FBCEFD8FFBD6" box="[502,599,1074,1098]" pageId="3" pageNumber="44" type="female">1 female</specimenCount>
(SL
<quantity id="31209B608578FFCAFD4EFBCEFD31FBD6" box="[655,745,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="44" unit="mm" value="1.4">1.4 mm</quantity>
), same data as CBM-ZC 10207.
</paragraph>
</subSubSection>
<subSubSection id="BEC2650E8578FFCDFF04FBA9FECAFD56" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44" type="diagnosis">
<paragraph id="F66736858578FFCDFF04FBA9FDBBFF6B" blockId="3.[151,1436,151,2020]" lastBlockId="4.[151,1436,151,1542]" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
<emphasis id="C4ACEA978578FFCAFF04FBA9FEE7FBF2" bold="true" box="[197,319,1109,1134]" pageId="3" pageNumber="44">Diagnosis.</emphasis>
Shield (
<figureCitation id="6EE32A008578FFCAFE5FFBA9FE39FBF1" box="[414,481,1109,1133]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="3" pageNumber="44">Fig. 2</figureCitation>
A) as long as wide or slightly longer than wide; rostrum moderately broadly triangular, overreaching obtuse anterolateral projections; dorsal surface with distinct paragastric grooves. Ocular peduncles (
<figureCitation id="6EE32A008578FFCAFF12FB60FECFFB28" box="[211,279,1180,1204]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="3" pageNumber="44">Fig. 2</figureCitation>
A) moderately slender, 0.70.8 times as long as shield; corneas not dilated; ocular acicles each with 1 submarginal terminal spine. Antennular peduncles (
<figureCitation id="6EE32A008578FFCAFD18FB3CFCC4FB44" box="[729,796,1216,1240]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="3" pageNumber="44">Fig. 2</figureCitation>
A) overreaching distal corneal margins by 0.20.3 length of ultimate segment. Antennal peduncles (
<figureCitation id="6EE32A008578FFCAFDB0FB1FFD6CFB67" box="[625,692,1251,1275]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="3" pageNumber="44">Fig. 2</figureCitation>
A) reaching nearly to distal corneal margins; second segment with distolateral angle produced, reaching midlength of fourth segment, dorsomesial distal angle with small spine; antennal acicle slightly arcuate, reaching base of cornea. Third maxilliped (
<figureCitation id="6EE32A008578FFCAFC27FAD6FBF2FADE" box="[998,1066,1322,1346]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="3" pageNumber="44">Fig. 2</figureCitation>
C, D) with crista dentata consisting of closely spaced corneous teeth increasing in size proximally; carpus with small dorsodistal and ventrodistal spines. Chelipeds with scattered moderately long setae on dorsal surfaces of chela and carpus. Right cheliped (
<figureCitation id="6EE32A008578FFCAFAABFA8DFF7CFA30" captionStart="FIGURE 3" captionStartId="7.[151,250,1937,1960]" captionTargetBox="[201,1366,215,1937]" captionTargetId="figure@7.[201,1391,193,1937]" captionTargetPageId="7" captionText="FIGURE 3. Boninpagurus pilosipes (Stimpson, 1858), n. comb., male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay. A, right cheliped, mesial view (setae omitted); B, same, lateral view; C, left cheliped, mesial view (setae omitted); D, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/202572/files/figure.png" pageId="3" pageNumber="44">Figs 3</figureCitation>
A, B, 4A, B) moderately stout; dactylus subequal in length to palm in males, distinctly longer than in females, dorsal surface with longitudinal row of small spines or spinulose tubercles adjacent to dorsomesial margin delimited by row of small spines; palm with 57 irregular rows of small spines or spiniform tubercles on dorsal surface, dorsolateral and dorsomesial margins each delimited by row of small spines; carpus with row of strong spines on dorsomesial margin and additional row of spines adjacent to dorsomesial margin, dorsolateral face rounded; merus with row of spines on ventrolateral and ventrodistal margins. Left cheliped (
<figureCitation id="6EE32A008578FFCAFC3DF9BAFB9CF9C2" box="[1020,1092,1606,1630]" captionStart="FIGURE 3" captionStartId="7.[151,250,1937,1960]" captionTargetBox="[201,1366,215,1937]" captionTargetId="figure@7.[201,1391,193,1937]" captionTargetPageId="7" captionText="FIGURE 3. Boninpagurus pilosipes (Stimpson, 1858), n. comb., male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay. A, right cheliped, mesial view (setae omitted); B, same, lateral view; C, left cheliped, mesial view (setae omitted); D, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/202572/files/figure.png" pageId="3" pageNumber="44">Figs 3</figureCitation>
C, D, 4C) moderately slender; dactylus about twice as long as palm, without distinct spines on surfaces; palm with 2 or 3 rows of small to moderately small spines extending onto fixed finger on dorsal surface, dorsal midline weakly elevated, dorsolateral margin delimited by row of small spines, dorsomesial margin with tiny blunt tubercles; carpus weakly widened distally, with row of small spines on dorsolateral and dorsomesial margins; merus with row of small spines on ventrolateral and ventromesial margins. Ambulatory legs (
<figureCitation id="6EE32A008578FFCAFCCDF90BFC89F893" box="[780,849,1783,1807]" captionStart="FIGURE 4" captionStartId="8.[151,250,1867,1890]" captionTargetBox="[245,1339,210,1854]" captionTargetId="figure@8.[224,1366,193,1854]" captionTargetPageId="8" captionText="FIGURE 4. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A, C G, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; B, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, chela of right cheliped, dorsal view (setae omitted); C, chela and carpus of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view; E, left third pereopod, lateral view; F, dactylus of right second pereopod, mesial view; G, dactylus of left third pereopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/202573/files/figure.png" pageId="3" pageNumber="44">Fig. 4</figureCitation>
D, E) with tufts of stiff setae on dorsal and ventral margins; dactyli 1.01.1 times longer than propodi, each with 611 corneous spines on ventral margin, mesial face unarmed (second) (
<figureCitation id="6EE32A008578FFCAFEAEF8C2FE68F8CA" box="[367,432,1854,1878]" captionStart="FIGURE 4" captionStartId="8.[151,250,1867,1890]" captionTargetBox="[245,1339,210,1854]" captionTargetId="figure@8.[224,1366,193,1854]" captionTargetPageId="8" captionText="FIGURE 4. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A, C G, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; B, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, chela of right cheliped, dorsal view (setae omitted); C, chela and carpus of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view; E, left third pereopod, lateral view; F, dactylus of right second pereopod, mesial view; G, dactylus of left third pereopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/202573/files/figure.png" pageId="3" pageNumber="44">Fig. 4</figureCitation>
F) or armed with row of spinules adjacent to dorsal margin (third) (
<figureCitation id="6EE32A008578FFCAFB5CF8C2FB38F8CA" box="[1181,1248,1854,1878]" captionStart="FIGURE 4" captionStartId="8.[151,250,1867,1890]" captionTargetBox="[245,1339,210,1854]" captionTargetId="figure@8.[224,1366,193,1854]" captionTargetPageId="8" captionText="FIGURE 4. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A, C G, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; B, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, chela of right cheliped, dorsal view (setae omitted); C, chela and carpus of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view; E, left third pereopod, lateral view; F, dactylus of right second pereopod, mesial view; G, dactylus of left third pereopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/202573/files/figure.png" pageId="3" pageNumber="44">Fig. 4</figureCitation>
G); propodi each with 1 or 2 corneous spinules on ventrodistal margin and 1 or 2 spinules on ventral margin; carpi only with dorsodistal spine; meri each with few small spines on ventral margin (second) or unarmed (third). Coxa of right fifth pereopod of male with sexual tube of medium length (24 times coxal length), straight or slightly curved, directed laterally or posterolaterally, gradually tapering distally to truncate tip, proximal half opaque, and remaining distal portion transparent. Sixth thoracic sternite (
<figureCitation id="6EE32A00857FFFCDFD48FF6BFD16FF33" box="[649,718,151,175]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="4" pageNumber="45">Fig. 2</figureCitation>
F, J) with subrectangular anterior lobe. Eighth thoracic sternite (
<figureCitation id="6EE32A00857FFFCDFF5EFF47FF3AFF4F" box="[159,226,187,211]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="4" pageNumber="45">Fig. 2</figureCitation>
G, L) consisting of 2 rounded lobes. Telson (
<figureCitation id="6EE32A00857FFFCDFD21FF47FCFCFF4F" box="[736,804,187,211]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="4" pageNumber="45">Fig. 2</figureCitation>
K) with terminal margins slightly to somewhat oblique, each with 4 or 5 moderately large spines.
</paragraph>
<paragraph id="F6673685857FFFCDFF04FEFEFECAFD56" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="45">
<emphasis id="C4ACEA97857FFFCDFF04FEFEFE49FE87" bold="true" box="[197,401,258,283]" pageId="4" pageNumber="45">Coloration in life</emphasis>
(
<figureCitation id="6EE32A00857FFFCDFE60FEFEFE39FE86" box="[417,481,258,282]" captionStart="FIGURE 1" captionStartId="5.[151,250,1825,1848]" captionTargetBox="[151,1436,195,1776]" captionTargetId="figure@5.[151,1436,195,1776]" captionTargetPageId="5" captionText="FIGURE 1. Boninpagurus pilopsipes (Stimpson, 1858), n. comb. A, D, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands; B, male (SL 1.7 mm; CBM-ZC 10207), Maehama, Chichi-jima Island, Ogasawara Islands; C, E, male with rhizocephalan parasite attached to pleon (SL 3.0 mm; CBM-ZC 10000), Hayama, Sagami Bay. A, B, C, entire animal, dorsal view; D, E, close up of cephalic appendages, showing differences in color pattern of antennular peduncle." httpUri="https://zenodo.org/record/202570/files/figure.png" pageId="4" pageNumber="45">Fig. 1</figureCitation>
A, B, D). General background color gray- or green-brown. Shield mottled grayish or greenish brown, paler laterally, with 2 or 3 paired spots. Posterior carapace also mottled brown, with conspicuous pair of spots on either side of posteromedian plate. Ocular peduncles generally light brown, proximally with dark brown transverse band; proximal half of dorsal surface grayish. Antennular peduncles generally grayish or greenish brown; ultimate segment with 2 longitudinal dark brown stripes on dorsal surface; median part of dorsal surface of penultimate segment paler; articulation between segments whitish; dorsal flagellum pale brown. Antennal peduncles generally greenish or grayish brown; fifth segment with brown longitudinal stripes laterally and mesially; antennal flagellum entirely light brown, without particular pattern. Right cheliped with tips of fingers whitish; palm generally brown or grayish brown, with rather obscure longitudinal stripes laterally and mesially; carpus apparently with longitudinal stripes of brown and light gray on dorsal surface, latter consisting of longitudinal row of light gray spots; merus with subdistal, narrow, transverse brown band and irregular dark brown patches or blotches. Left cheliped with similar pattern to right cheliped, but longitudinal stripes on palm, carpus and merus more clearly delineated.
</paragraph>
</subSubSection>
<subSubSection id="BEC2650E857FFFCDFF04FD2AFF05FCC6" pageId="4" pageNumber="45" type="distribution">
<paragraph id="F6673685857FFFCDFF04FD2AFF05FCC6" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="45">
<emphasis id="C4ACEA97857FFFCDFF04FD2AFE86FD73" bold="true" box="[197,350,726,751]" pageId="4" pageNumber="45">Distribution.</emphasis>
Okinawa Islands (Katsuren and Motobu Peninsulas on Okinawa Island, and Kouri Port on Kouri Island), Chichi-jima Island of the Ogasawara Islands, Izu-oshima Island of the Izu Islands, mainland of
<collectingCountry id="8ECF7615857FFFCDFA9DFD07FA44FC8E" box="[1372,1436,763,786]" name="Japan" pageId="4" pageNumber="45">Japan</collectingCountry>
(Boso Peninsula on Honshu southward to Nagasaki on Kyushu), and southern part of
<collectingCountry id="8ECF7615857FFFCDFBB4FCE2FB66FCAA" box="[1141,1214,798,822]" name="South Korea" pageId="4" pageNumber="45">Korea</collectingCountry>
); intertidal to
<quantity id="31209B60857FFFCDFAA0FCE2FA44FCAA" box="[1377,1436,798,822]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="45" unit="m" value="25.0">25 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="BEC2650E857FFFCDFF04FC9AFB6FF99A" pageId="4" pageNumber="45" type="discussion">
<paragraph id="F6673685857FFFCDFF04FC9AFAE6FACE" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="45">
<emphasis id="C4ACEA97857FFFCDFF04FC9AFEE1FCE2" bold="true" box="[197,313,870,894]" pageId="4" pageNumber="45">Remarks.</emphasis>
<bibRefCitation id="92494B74857FFFCDFEFEFC9AFD53FCE2" author="Asakura" box="[319,651,870,894]" pageId="4" pageNumber="45" refString="Asakura, A. &amp; Tachikawa, H. (2004) Boninpagurus acanthocheles, a new genus and new species of hermit crab (Decapoda: Anomura: Paguridae) from shallow waters of the Ogasawara (Bonin) Islands, Japan. Journal of Crustacean Biology, 24, 157 - 167." type="journal article" year="2004">Asakura &amp; Tachikawa (2004)</bibRefCitation>
did not compare their new taxon
<taxonomicName id="31D84D06857FFFCDFBC3FC9BFB19FCE2" box="[1026,1217,870,894]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="acanthocheles">
<emphasis id="C4ACEA97857FFFCDFBC3FC9BFB19FCE2" box="[1026,1217,870,894]" italics="true" pageId="4" pageNumber="45">B. acanthocheles</emphasis>
</taxonomicName>
with
<emphasis id="C4ACEA97857FFFCDFAC1FC9BFAA4FCE2" box="[1280,1404,870,894]" italics="true" pageId="4" pageNumber="45">P. pi lo s ip e s</emphasis>
in spite of close similarities, perhaps because the latter species was considered to lack sexual tube development at the time. The present newly collected male specimens from Okinawa Islands show sexual tube development, closely agreeing with that seen in
<taxonomicName id="31D84D06857FFFCDFE7DFC2FFDA4FC76" box="[444,636,978,1002]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="acanthocheles">
<emphasis id="C4ACEA97857FFFCDFE7DFC2FFDA4FC76" box="[444,636,978,1002]" italics="true" pageId="4" pageNumber="45">B. acanthocheles</emphasis>
</taxonomicName>
. In this study, we examined specimens from various localities in
<collectingCountry id="8ECF7615857FFFCDFA94FC2FFA4FFC76" box="[1365,1431,979,1002]" name="Japan" pageId="4" pageNumber="45">Japan</collectingCountry>
, including those from Okinawa and Ogasawara Islands, representing the
<typeStatus id="29638827857FFFCDFC12FC0BFBDBFB93" box="[979,1027,1015,1039]" pageId="4" pageNumber="45">type</typeStatus>
localities of the two taxa. As mentioned before, two morphs separated by the color of the antenular peduncle are recognized. Nevertheless, we could not find a single morphological character or a suite of characters that will separate one species from another. Comparison of partial sequence of the barcoding gene COI (657 bp) shows that there are only 0.61.5% genetic divergences amongst the specimens from the Japanese mainland, and Okinawa and Ogasawara Islands, while specimens from the same locality have genetic divergences of 0.51.4% (
<tableCitation id="BB5A033E857FFFCDFC14FB56FBF6FB5E" box="[981,1070,1194,1218]" captionStart="TABLE 2" captionStartId="4.[151,235,1590,1611]" captionTargetBox="[151,1436,1670,2011]" captionTargetPageId="4" captionText="TABLE 2. The partial COI gene nucleotide divergence (657 bp) of Boninpagurus pilosipes (Stimpson, 1858) from different localities." httpUri="http://table.plazi.org/id/A2A7660D857FFFCDFF56F9CAFF22F9F7" pageId="4" pageNumber="45" tableUuid="A2A7660D857FFFCDFF56F9CAFF22F9F7">Table 2</tableCitation>
). Such a low genetic divergence is generally considered as an intraspecific variation in decapod crustaceans (e.g.
<bibRefCitation id="92494B74857FFFCDFBDEFB32FA9FFB7A" author="Khamnamtong" box="[1055,1351,1230,1254]" pageId="4" pageNumber="45" refString="Khamnamtong, B., Klinbunga, S. &amp; Menasveta, P. (2009) Genetic diversity and geographic differentiation of the giant tiger shrimp (Penaeus monodon) in Thailand analyzed by mitochondrial COI sequences. Biochemical Genetics, 47, 42 - 55." type="journal article" year="2009">
Khamnamtong
<emphasis id="C4ACEA97857FFFCDFB0DFB33FB26FB7A" box="[1228,1278,1230,1254]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
. 2009
</bibRefCitation>
;
<bibRefCitation id="92494B74857FFFCDFA92FB32FEC8FA96" author="Rhyne" pageId="4" pageNumber="45" refString="Rhyne, A. L., Zhang, D., Lin, J. &amp; Schizas, N. V. (2009) Not any two will do: DNA divergence and interpopulation reproductive compatibility in the simultaneous hermaphroditic shrimp Lysmata wurdemanni. Marine Ecology Progress Series, 388, 185 - 195." type="journal article" year="2009">
Rhyne
<emphasis id="C4ACEA97857FFFCDFF56FB0FFF10FA96" box="[151,200,1266,1290]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
. 2009
</bibRefCitation>
;
<bibRefCitation id="92494B74857FFFCDFEDDFB0EFE0EFA96" author="Silva" box="[284,470,1266,1290]" pageId="4" pageNumber="45" refString="Silva, I. C., Mesquita, N. &amp; Paula, J. (2010) Genetic and morphological differentiation of the mangrove crab Perisesarma guttatum (Brachyura: Sesarmidae) along an East African latitudinal gradient. Biological Journal of the Linnean Society, 99, 28 - 46." type="journal article" year="2010">
Silva
<emphasis id="C4ACEA97857FFFCDFE9DFB0FFE56FA96" box="[348,398,1266,1290]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
. 2010
</bibRefCitation>
;
<bibRefCitation id="92494B74857FFFCDFE23FB0FFD46FA96" author="Yang" box="[482,670,1266,1291]" pageId="4" pageNumber="45" refString="Yang, C. - H., Chan, T. Y. &amp; Chu, K. H. (2010) Two new species of the &quot; Heterocarpus gibbosus Bate, 1888 &quot; species group (Crustacea: Decapoda: Pandalidae) from the western Pacific and NW Australia. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds,) Contributions to shrimp taxonomy. Zootaxa, 2372, 206 - 220." type="journal article" year="2010">
Yang
<emphasis id="C4ACEA97857FFFCDFDE2FB0FFD8CFA96" box="[547,596,1266,1290]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
. 2010
</bibRefCitation>
). Thus, our initial suspicion that two species might be represented in the present collections was not supported by either morphological or molecular comparison. Consequently, we propose to synonymize
<taxonomicName id="31D84D06857FFFCDFEBDFAC7FDE4FACE" box="[380,572,1338,1362]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="acanthocheles">
<emphasis id="C4ACEA97857FFFCDFEBDFAC7FDE4FACE" box="[380,572,1338,1362]" italics="true" pageId="4" pageNumber="45">B. acanthocheles</emphasis>
</taxonomicName>
under
<taxonomicName id="31D84D06857FFFCDFD48FAC7FCD4FACE" box="[649,780,1338,1362]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA97857FFFCDFD48FAC7FCD4FACE" box="[649,780,1338,1362]" italics="true" pageId="4" pageNumber="45">B. pilosipes</emphasis>
</taxonomicName>
, and refer all examined specimens to
<taxonomicName id="31D84D06857FFFCDFB75FAC7FAEFFACE" box="[1204,1335,1338,1362]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA97857FFFCDFB75FAC7FAEFFACE" box="[1204,1335,1338,1362]" italics="true" pageId="4" pageNumber="45">B. pilosipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F6673685857FFFCDFF04FAA2FB6FF99A" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="45">
<bibRefCitation id="92494B74857FFFCDFF04FAA2FEBBFAEA" author="Komai" box="[197,355,1374,1398]" pageId="4" pageNumber="45" refString="Komai, T. (2003) Reassessment of Pagurus pilosipes (Stimpson), supplemental description of P. insulae Asakura, and descriptions of three new species of Pagurus from East Asian waters (Crustacea: Decapoda: Anomura: Paguridae). Natural History Research, 7, 115 - 166." type="journal article" year="2003">Komai (2003)</bibRefCitation>
designated a
<typeStatus id="29638827857FFFCDFE3CFAA3FD8EFAEB" box="[509,598,1375,1399]" pageId="4" pageNumber="45" type="neotype">neotype</typeStatus>
for
<taxonomicName id="31D84D06857FFFCDFD45FAA3FC96FAEA" box="[644,846,1374,1398]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA97857FFFCDFD45FAA3FC96FAEA" box="[644,846,1374,1398]" italics="true" pageId="4" pageNumber="45">Pagurus pilosipes</emphasis>
</taxonomicName>
in the interests of nomenclatural stability, because it was considered that the
<typeStatus id="29638827857FFFCDFE68FA7FFE01FA07" box="[425,473,1411,1435]" pageId="4" pageNumber="45">type</typeStatus>
was destroyed in the Great Chicago Fire of 1871 (
<bibRefCitation id="92494B74857FFFCDFBDDFA7EFB7EFA06" author="Evans" box="[1052,1190,1410,1434]" pageId="4" pageNumber="45" refString="Evans, A. C. (1967) Syntypes of Decapoda described by William Stimspon and James Dana in the collections of the British Museum (Natural History). Journal of Natural History, 1, 399 - 411." type="journal article" year="1967">Evans 1967</bibRefCitation>
). Subsequently,
<bibRefCitation id="92494B74857FFFCDFA9EFA7FFEA6FA22" author="Manning" pageId="4" pageNumber="45" refString="Manning, R. B &amp; Reed, K. J. (2006) Decapod crustaceans deposited in the Zoological Museum of Copenhagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54, 283 - 293." type="journal article" year="2006">Manning &amp; Reed (2006)</bibRefCitation>
published a list of crustaceans that were shipped by W. Stimpson in 1859 to the Zoological Museum in Copenhagen (ZMC). In the list,
<taxonomicName id="31D84D06857FFFCDFD4BFA37FC8DFA7E" box="[650,853,1482,1506]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA97857FFFCDFD4BFA37FC8DFA7E" box="[650,853,1482,1506]" italics="true" pageId="4" pageNumber="45">Pagurus pilosipes</emphasis>
</taxonomicName>
is mentioned (as
<taxonomicName id="31D84D06857FFFCDFBDDFA37FB4FFA7E" box="[1052,1175,1483,1506]" genus="Eupagurus" kingdom="Animalia" pageId="4" pageNumber="45" rank="genus">
<emphasis id="C4ACEA97857FFFCDFBDDFA37FB4FFA7E" box="[1052,1175,1483,1506]" italics="true" pageId="4" pageNumber="45">Eupagurus</emphasis>
</taxonomicName>
), but no specimen was located in the collection of the ZMC. Therefore, the action by
<bibRefCitation id="92494B74857FFFCDFC91FA12FC36F99A" author="Komai" box="[848,1006,1518,1542]" pageId="4" pageNumber="45" refString="Komai, T. (2003) Reassessment of Pagurus pilosipes (Stimpson), supplemental description of P. insulae Asakura, and descriptions of three new species of Pagurus from East Asian waters (Crustacea: Decapoda: Anomura: Paguridae). Natural History Research, 7, 115 - 166." type="journal article" year="2003">Komai (2003)</bibRefCitation>
is still warranted.
</paragraph>
</subSubSection>
</treatment>
</document>