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<document ID-CLB-Dataset="6732" ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.60.33075" ID-GBIF-Dataset="dcaff623-aab6-4ab4-bb25-5e494185030d" ID-Pensoft-Pub="2305-2562-1-67" ID-Pensoft-UUID="8DAF8AC953485F52AB974AD46D3F5DB1" ID-Zenodo-Dep="2901066" ID-ZooBank="E51FFDAE5B9B433ABE813EED75F0D070" ModsDocID="2305-2562-60-1-67" checkinTime="1621371389653" checkinUser="pensoft" docAuthor="Londt, Jason G. H." docDate="2019" docId="7E6E86DB161E7A84FAF63B02EFCCF3FF" docLanguage="en" docName="AfrInvert 60(1): 67-82" docOrigin="African Invertebrates 60 (1)" docPubDate="2019-05-10" docSource="http://dx.doi.org/10.3897/afrinvertebr.60.33075" docTitle="Empodiodes greatheadi Oldroyd 1972" docType="treatment" docVersion="5" id="8DAF8AC953485F52AB974AD46D3F5DB1" lastPageNumber="67" masterDocId="8DAF8AC953485F52AB974AD46D3F5DB1" masterDocTitle="A revision of Empodiodes Oldroyd, 1972 with the descriptions of two new species from South Africa (Diptera, Asilidae, Stenopogoninae)" masterLastPageNumber="82" masterPageNumber="67" pageNumber="67" updateTime="1732751666901" updateUser="ExternalLinkService">
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<mods:title id="39271029B998CCE662965D0A95BD8499">A revision of Empodiodes Oldroyd, 1972 with the descriptions of two new species from South Africa (Diptera, Asilidae, Stenopogoninae)</mods:title>
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<mods:namePart id="50BA85FE56B02212E57185B89B5A3460">Londt, Jason G. H.</mods:namePart>
<mods:affiliation id="CCA2F22CFEBCA8A6A48C671F19925F49">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200, South Africa &amp; School of Biological &amp; Conservation Sciences, University of KwaZulu-Natal, Pietermaritzburg, South Africa</mods:affiliation>
<mods:nameIdentifier id="FE96C3F7A18814460B438F410C07DF0B" type="email">londtja@telkomsa.net</mods:nameIdentifier>
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<mods:title id="5824A194AE0E120AD0EBF4888342E9AF">African Invertebrates</mods:title>
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<mods:date id="AFF1D608C3DD8D88B763BAE73CD5DFD7">2019</mods:date>
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<mods:number id="1F43CAD29617571E612E5602C2406B5C">2019-05-10</mods:number>
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<mods:number id="FBE23A7A0DD8340CA3F4E0B346778899">60</mods:number>
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<mods:start id="89BBE32CD927C6FE933935263083B450">67</mods:start>
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<mods:url id="5F6584BFA9236660A0FD7C03BD9211C7">http://dx.doi.org/10.3897/afrinvertebr.60.33075</mods:url>
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<mods:classification id="AD8882C1B294F005B643621E2624B52F">journal article</mods:classification>
<mods:identifier id="EFFA88B47B9E9E0952E2449E0AFDADFD" type="DOI">http://dx.doi.org/10.3897/afrinvertebr.60.33075</mods:identifier>
<mods:identifier id="340F214321AD01E432B4360023FF1169" type="Pensoft-Pub">2305-2562-1-67</mods:identifier>
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<treatment id="7E6E86DB161E7A84FAF63B02EFCCF3FF" ID-GBIF-Taxon="182223236" LSID="urn:lsid:plazi:treatment:7E6E86DB161E7A84FAF63B02EFCCF3FF" httpUri="http://treatment.plazi.org/id/7E6E86DB161E7A84FAF63B02EFCCF3FF" lastPageNumber="67" pageId="0" pageNumber="67" scope_family="Asilidae" scope_order="Diptera" scope_subFamily="Stenopogoninae">
<subSubSection id="120EB1B1AD3EE19C0EAEF12789A1464D" pageId="0" pageNumber="67" type="nomenclature">
<paragraph id="6C9453E465E9AAAF901C8BB8E6577AB7" pageId="0" pageNumber="67">
<taxonomicName id="0B292AC3815B3288DE6A9BB0F6D01A40" ID-CoL="39LJW" authority="Oldroyd, 1972" authorityName="Oldroyd" authorityYear="1972" class="Insecta" family="Asilidae" genus="Empodiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Empodiodes greatheadi" order="Diptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="greatheadi">Empodiodes greatheadi Oldroyd, 1972</taxonomicName>
<figureCitation id="08C01394DE2A6891E092C2A5E93BB8D5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Empodiodes greatheadi Oldroyd, 1972 holotype. Scale bars: in mm. (Photo BMNH)." figureDoi="10.3897/afrinvertebr.60.33075.figure1" httpUri="https://binary.pensoft.net/fig/300268" pageId="0" pageNumber="67">Figs 1</figureCitation>
<figureCitation id="EEC1389ABCF2EF91C7F4F7D36FFFFE0D" captionStart="Figure 16" captionStartId="F10" captionText="Figure 16. The distribution of Empodiodes species: E. greatheadi Oldroyd, 1972 (●), E. melanoscopaeus Londt, 1992 (○), E. namibiensis Londt, 2012 (■), E. pusillipes sp. nov. (□), E. torridus sp. nov. (▲), E. whittingtoni Londt, 1992 (♦) (Map Zamisa)." figureDoi="10.3897/afrinvertebr.60.33075.figure16" httpUri="https://binary.pensoft.net/fig/300277" pageId="0" pageNumber="67">, 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D71BF29C35867D9954D6740DF4984EF8" pageId="0" pageNumber="67" type="reference_group">
<paragraph id="B0225B68BB1981FBD3AC6EFB4C395693" pageId="0" pageNumber="67">
<taxonomicName id="5B9D832859909E192A85C40DA4DD8070" authorityName="Oldroyd" authorityYear="1972" class="Insecta" family="Asilidae" genus="Empodiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Empodiodes greatheadi" order="Diptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="greatheadi">Empodiodes greatheadi</taxonomicName>
Oldroyd, 1972: 635-9 (figs 1 head lateral, 2 head anterior, 4 metathoracic tarsomere 1, 5 prothoracic tarsus); 1974: 166-8 (figs 162 metathoracic tarsomere 1, 163 prothoracic tarsus, 164 head lateral, 165 head anterior); 1980: 361 (catalogue);
<bibRefCitation id="55F130B39602DBCF911946379E375904" author="Londt, JGH" journalOrPublisher="Journal of African Zoology" pageId="0" pageNumber="67" pagination="55 - 79" refId="B2" refString="Londt, JGH, 1992. Revision of three small Afrotropical asilid genera, Empodiodes Oldroyd, Hynirhynchus Lindner and Lycostommyia Oldroyd (Diptera: Asilidae: Stenopogoninae). Journal of African Zoology 106 (1): 55 - 79" title="Revision of three small Afrotropical asilid genera, Empodiodes Oldroyd, Hynirhynchus Lindner and Lycostommyia Oldroyd (Diptera: Asilidae: Stenopogoninae)." volume="106" year="1992">Londt 1992</bibRefCitation>
: 57-59 (figs 1 antenna, 2 prosternum, 3 wing, 4 metathoracic tarsus, 5 empodium, 6 metathoracic pulvilli, 7-9 ♂ terminalia, 10 metathoracic tarsus 5, 11-13 ♀ terminalia).
</paragraph>
</subSubSection>
<subSubSection id="5D9AB7D78438AC5378DB3D242C4275E5" pageId="0" pageNumber="67" type="materials_examined">
<paragraph id="2409A5F4646CB24360BAD04E40B496A6" pageId="0" pageNumber="67">Material.</paragraph>
<paragraph id="F67DE57663878559067E80DEB7546EBD" pageId="0" pageNumber="67">
<materialsCitation id="64CCDAE9D3CD20AA82F8BCEE82B8E765" collectingDate="1981-10-13" collectorName="F. W. &amp; S. K. Gess" country="SOUTH AFRICA" elevation="650" latitude="-33.183056" location="Rockhurst" longLatPrecision="20" longitude="26.316946" municipality="Grahamstown" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F34C9A7C9A94DCAB4FB8A782D40F8507" name="South Africa">SOUTH AFRICA</collectingCountry>
:
<specimenCount id="897CF4D4BF95AD7FAD6CBF920A3D881D" type="male">2♂</specimenCount>
<collectingMunicipality id="0AA13B8F670D5E20D663C61794265871">Grahamstown</collectingMunicipality>
[= Grahamstown],
<location id="E8A927837D1EDE5D30314FC4F5C2EAED" LSID="urn:lsid:plazi:treatment:7E6E86DB161E7A84FAF63B02EFCCF3FF:E8A927837D1EDE5D30314FC4F5C2EAED" country="SOUTH AFRICA" latitude="-33.183056" longLatPrecision="20" longitude="26.316946" municipality="Grahamstown" name="Rockhurst">Rockhurst</location>
[farm
<geoCoordinate id="F17A151125D79F0834B7DFCDC517FCC5" degrees="33" direction="south" minutes="10" orientation="latitude" precision="15" seconds="59" value="-33.183056">33°10'59&quot;S</geoCoordinate>
,
<geoCoordinate id="42BA316365212C1A969C72575160778B" degrees="26" direction="east" minutes="19" orientation="longitude" precision="15" seconds="01" value="26.316946">26°19'01&quot;E</geoCoordinate>
<emphasis id="1B63A7C295588AA3CC63F71102CC444E" italics="true" pageId="0" pageNumber="67">c.</emphasis>
<quantity id="5968F15806D6689E125546F5D2857E27" metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">
<elevation id="E682C2E59D065103507C4B9D17791625" metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">650m</elevation>
</quantity>
],
<collectingDate id="C8FCDAB0F3FCD26BE61168DB0F5BA495" value="1981-10-13">13.x.1981</collectingDate>
,
<collectorName id="5F13C8CC5435FDADAF49C2BE549802C5">F.W. &amp; S.K. Gess</collectorName>
, 81/82/2, AMGS 101476, AMGS 101477 (AMGS)
</materialsCitation>
;
<materialsCitation id="B97492D24D8503B1A42CF9A7703020E3" collectingDate="1967-10-22" collectorName="C. Jacot-Guillarmod" country="South Africa" elevation="490" latitude="-33.25" location="Hilton" longLatPrecision="20" longitude="26.353334" municipality="Grahamstown" specimenCount="1" specimenCount-female="1">
<specimenCount id="C84EFEA0AA1C88CDEDEFA482FB4D9735" type="female">1♀</specimenCount>
<collectingMunicipality id="EFC342996F1A7D0D33A66F5A34AE1CF6">Grahamstown</collectingMunicipality>
,
<location id="C45A6ADCD8DEA3264AB2F69CCD133DC3" LSID="urn:lsid:plazi:treatment:7E6E86DB161E7A84FAF63B02EFCCF3FF:C45A6ADCD8DEA3264AB2F69CCD133DC3" country="South Africa" latitude="-33.25" longLatPrecision="20" longitude="26.353334" municipality="Grahamstown" name="Hilton">Hilton</location>
[farm
<geoCoordinate id="62C0D4F67309848D269E60EC26D07D98" degrees="33" direction="south" minutes="15" orientation="latitude" precision="15" seconds="00" value="-33.25">33°15'00&quot;S</geoCoordinate>
,
<geoCoordinate id="9C80CF3BA4D4EC766A857AF036AFE062" degrees="26" direction="east" minutes="21" orientation="longitude" precision="15" seconds="12" value="26.353334">26°21'12&quot;E</geoCoordinate>
<emphasis id="CAEBE03A0D31D000F1FCC6EFF0470E66" italics="true" pageId="0" pageNumber="67">c.</emphasis>
<quantity id="11CDF6CAC9B3690DA7CE7616AFC14FD0" metricMagnitude="2" metricUnit="m" metricValue="4.9" unit="m" value="490.0">
<elevation id="D785548113D261EE263EA986052A386A" metricMagnitude="2" metricUnit="m" metricValue="4.9" unit="m" value="490.0">490m</elevation>
</quantity>
],
<collectingDate id="ED58FBAB4913EF080F5F86A4ACC343C2" value="1967-10-22">22.x.1967</collectingDate>
,
<collectorName id="8B96E56D7C049A02889BE0D9D34EA141">C. Jacot-Guillarmod</collectorName>
AMGS 101478 (AMGS)
</materialsCitation>
;
<materialsCitation id="5F165857D9F3D844B236FFF5C2E362B7" collectingDate="2019-01-01" collectingDateMax="2019-12-31" collectingDateMin="2019-01-01" collectorName="D. &amp; A. Greathead'" country="South Africa" elevation="285" latitude="-33.316666" location="Alicedale" longLatPrecision="1205" longitude="26.083334" municipality="South Africa" specimenCount="♂" stateProvince="Cape Province" typeStatus="Holotype">
<specimenCount id="77C154BA1B88488F359C73539C6C391A">♂</specimenCount>
<typeStatus id="5A1961A506F4EA02F5A7EF05A7031899">Holotype</typeStatus>
'
<collectingMunicipality id="7EB45A49FFBD5A002DB6F95ECE758652">South Africa</collectingMunicipality>
/ C.P. [
<collectingRegion id="373F71DB561486F65498D87F5937E55E" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
]
<location id="5ECD757A1C3183CD2700F78BFEC7DAA6" LSID="urn:lsid:plazi:treatment:7E6E86DB161E7A84FAF63B02EFCCF3FF:5ECD757A1C3183CD2700F78BFEC7DAA6" country="South Africa" latitude="-33.316666" longLatPrecision="1205" longitude="26.083334" municipality="South Africa" name="Alicedale" stateProvince="Cape Province">Alicedale</location>
[
<geoCoordinate id="620E5D486356E335320E5373637E6310" degrees="33" direction="south" minutes="19" orientation="latitude" precision="925" value="-33.316666">33°19'S</geoCoordinate>
,
<geoCoordinate id="D7EA6806E997635C67B7DDE7806326A5" degrees="26" direction="east" minutes="05" orientation="longitude" precision="925" value="26.083334">26°05'E</geoCoordinate>
<emphasis id="910B7BF292F8B5EECC28DE0E98A24E80" italics="true" pageId="0" pageNumber="67">c.</emphasis>
<quantity id="FF14D1C2F746A5DE5AF80A614058BF9D" metricMagnitude="2" metricUnit="m" metricValue="2.85" unit="m" value="285.0">
<elevation id="07940086F5CC1A31E5D71FEE2B6EF528" metricMagnitude="2" metricUnit="m" metricValue="2.85" unit="m" value="285.0">285m</elevation>
</quantity>
] / 16.xii.71 /
<collectorName id="2EB1877EBF2F4C395E21CAB5C6674D9D">
D. &amp; A.
<normalizedToken id="41C61429575635D2D2394F7B3BCF1C81" originalValue="Greathead’">Greathead'</normalizedToken>
</collectorName>
NHMUK010628725 (BMNH)
</materialsCitation>
;
<materialsCitation id="69B6C403A504886AAB8CB061E400C1FB" collectingDate="1978-10-30" collectingDateMax="1978-10-31" collectingDateMin="1978-10-30" collectorName="J Londt, R Miller, Karoo, Note" country="South Africa" elevation="935" latitude="-33.333332" location="Prov" longLatPrecision="1205" longitude="23.45" municipality="So. Africa" specimenCount="3324" specimenCount-adult="3323" specimenCount-male="1" stateProvince="Cape" typeStatus="Holotype">
<specimenCount id="3B660C7922833DA50AA633D759C7571C" type="male">1♂</specimenCount>
'
<collectingMunicipality id="C908F3C29978DDD6CA90359C2E1B9974">So. Africa</collectingMunicipality>
:
<collectingRegion id="49D5E56BA5E6EA7CB2837F975F109ECE" country="South Africa" name="Western Cape">Cape</collectingRegion>
<location id="291DBBD1952507EE66127EE2215589BA" LSID="urn:lsid:plazi:treatment:7E6E86DB161E7A84FAF63B02EFCCF3FF:291DBBD1952507EE66127EE2215589BA" country="South Africa" latitude="-33.333332" longLatPrecision="1205" longitude="23.45" municipality="So. Africa" name="Prov" stateProvince="Cape">Prov</location>
/
<locationDeviation id="4A4448812EBDE10141FDB4632240A763" location="Willowmore">
<quantity id="ACF08476898666598E40C6D46F874262" metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
SW Willowmore
</locationDeviation>
[
<emphasis id="27DD54D4B8EBD24CCF33E35B0206F013" italics="true" pageId="0" pageNumber="67">c.</emphasis>
<geoCoordinate id="D4118CF0512ACB468FB73D86009194AB" degrees="33" direction="south" minutes="20" orientation="latitude" precision="925" value="-33.333332">33°20'S</geoCoordinate>
,
<geoCoordinate id="5BCA0667DD6B73817D4AE30EDA28C54B" degrees="23" direction="east" minutes="27" orientation="longitude" precision="925" value="23.45">23°27'E</geoCoordinate>
<quantity id="530AECCE1A4749221A08CF4BA3A82AB7" metricMagnitude="2" metricUnit="m" metricValue="9.35" unit="m" value="935.0">
<elevation id="9FC00D92EEE8852DDE762AA376D6A6A6" metricMagnitude="2" metricUnit="m" metricValue="9.35" unit="m" value="935.0">935m</elevation>
</quantity>
] /
<collectingDate id="E90903F5E09C9E269F2120A3937B7C98" value="1978-10-30" valueMax="1978-10-31" valueMin="1978-10-30">30-31.x.1978</collectingDate>
<specimenCount id="13271844BE6CB4EEA4A7AD4EF2645ECF" type="adult">3323AD</specimenCount>
/
<collectorName id="3E7812D2137D2C7A729D3B2CC21F44BF">J Londt</collectorName>
&amp;
<collectorName id="87AC85963BB8C6FA9F7B7AADE007F3F2">R Miller</collectorName>
/ open
<collectorName id="3C7489A31CBFAFE1FEE2EA174F25D474">Karoo</collectorName>
<normalizedToken id="017A46BCC9A2CF9558166437EBDCB326" originalValue="scrub’">scrub'</normalizedToken>
, NMSA-DIP 81676 (NMSA).
<emphasis id="51160AB26396C3067E6F00E86807C04E" italics="true" pageId="0" pageNumber="67">
<collectorName id="293C50E54B28B772E72701D6B5F5B31E">Note</collectorName>
</emphasis>
:
<typeStatus id="1ED6A80125B9F8BF97A80815DE463508">Holotype</typeStatus>
not personally handled, but excellent photographs supplied confirm identification
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8C013066723738AC947F5817ED07DEC4" pageId="0" pageNumber="67" type="distribution">
<paragraph id="59E328948AC3585E8AF34045F693B21A" pageId="0" pageNumber="67">Distribution, phenology and biology.</paragraph>
<paragraph id="FAF8A27992852F40D89158D64C72D5E6" pageId="0" pageNumber="67">
Entire holotype as in Fig.
<figureCitation id="34631A7409D1C28C7C1169C65A525BBE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Empodiodes greatheadi Oldroyd, 1972 holotype. Scale bars: in mm. (Photo BMNH)." figureDoi="10.3897/afrinvertebr.60.33075.figure1" httpUri="https://binary.pensoft.net/fig/300268" pageId="0" pageNumber="67">1</figureCitation>
. Known from 5 specimens collected at 4 localities in the Eastern Cape Province of South Africa (Fig.
<figureCitation id="6E03011B808FC80A771A388D09C4BB93" captionStart="Figure 16" captionStartId="F10" captionText="Figure 16. The distribution of Empodiodes species: E. greatheadi Oldroyd, 1972 (●), E. melanoscopaeus Londt, 1992 (○), E. namibiensis Londt, 2012 (■), E. pusillipes sp. nov. (□), E. torridus sp. nov. (▲), E. whittingtoni Londt, 1992 (♦) (Map Zamisa)." figureDoi="10.3897/afrinvertebr.60.33075.figure16" httpUri="https://binary.pensoft.net/fig/300277" pageId="0" pageNumber="67">16</figureCitation>
). They were collected in October and November (Table
<tableCitation id="28AF5E6C1FA590194BAC885C1C0937A8" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Phenology of Empodiodes species indicating the number of specimens collected per month (abbreviated and starting with July)." httpUri="http://table.plazi.org/id/4F3B50E21D04B46A64AA0D115C778F5B" pageId="0" pageNumber="67" tableUuid="4F3B50E21D04B46A64AA0D115C778F5B">1</tableCitation>
) in an area located between summer and winter rainfall regions. The only useful label data relating to habitat are the words 'open Karoo
<normalizedToken id="F04031F63B5571890D2C685429F61D7C" originalValue="scrub’">scrub'</normalizedToken>
which describes much of the area. No other biological data are available.
</paragraph>
<caption id="875913CCF3EA51F44FAD2FEDC5617F43" ID-Zenodo-Dep="2901106" doi="10.3897/afrinvertebr.60.33075.figure1" httpUri="https://binary.pensoft.net/fig/300268" pageId="0" pageNumber="67" start="Figure 1" startId="F1">
<paragraph id="110BE67033B2A0B49153059EF6538F1B" pageId="0" pageNumber="67">
<emphasis id="510B2414961475FC2C1CD0EAD787A14C" bold="true" pageId="0" pageNumber="67">Figure 1.</emphasis>
<taxonomicName id="6F7CB8224439E26A62B20AF0555BD5CA" authorityName="Oldroyd" authorityYear="1972" class="Insecta" family="Asilidae" genus="Empodiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Empodiodes greatheadi" order="Diptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="4CFBC0E91EE4B762A0B4318C0E9CA6E2" italics="true" pageId="0" pageNumber="67">Empodiodes greatheadi</emphasis>
</taxonomicName>
Oldroyd, 1972 holotype. Scale bars: in mm. (Photo BMNH).
</paragraph>
</caption>
<caption id="4F3B50E21D04B46A64AA0D115C778F5B" ID-Table-UUID="4F3B50E21D04B46A64AA0D115C778F5B" httpUri="http://table.plazi.org/id/4F3B50E21D04B46A64AA0D115C778F5B" pageId="0" pageNumber="67" start="Table 1" startId="T1">
<paragraph id="5011287516E8D80D37115C71E3098506" pageId="0" pageNumber="67">
<emphasis id="44FF48A8C763166BA5565778B34DB908" bold="true" pageId="0" pageNumber="67">Table 1.</emphasis>
Phenology of
<taxonomicName id="FE0806A247B0CD41FCC78A3FD7347766" authorityName="Oldroyd" authorityYear="1972" class="Insecta" family="Asilidae" genus="Empodiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Empodiodes" order="Diptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="9A07924B9884850A2B781489EC9963E3" italics="true" pageId="0" pageNumber="67">Empodiodes</emphasis>
</taxonomicName>
species indicating the number of specimens collected per month (abbreviated and starting with July).
</paragraph>
</caption>
<paragraph id="52A4BB609BA3365A9E822E001A5E7168" pageId="0" pageNumber="67">
<table id="5EB0EC12ACEE42F4CDE60513B52F5495" pageId="0" pageNumber="67">
<tr id="6A45A866CAE1BA9BDC54E7F906A21485" pageId="0" pageNumber="67">
<th id="1C4316AA1084D09BE489D1F135DC2211" colspan="1" pageId="0" pageNumber="67" rowspan="1">Species</th>
<th id="D0AF4447FB0A50426C082466491BAC4B" colspan="1" pageId="0" pageNumber="67" rowspan="1">J</th>
<th id="0CFBA99BE98C4B6FDE4144ED56F9F753" colspan="1" pageId="0" pageNumber="67" rowspan="1">A</th>
<th id="62A638FEBFECB7E0BAB2A7FB74365555" colspan="1" pageId="0" pageNumber="67" rowspan="1">S</th>
<th id="F20EDEAA840AE5CD9507B9306C6A6C98" colspan="1" pageId="0" pageNumber="67" rowspan="1">O</th>
<th id="016B20AC7CCD5D67113C2B4E884465D8" colspan="1" pageId="0" pageNumber="67" rowspan="1">N</th>
<th id="65CB2AE42F26B75F46F07272FFDE7EAB" colspan="1" pageId="0" pageNumber="67" rowspan="1">D</th>
<th id="88644A5D1BABADC88728169161AB78EF" colspan="1" pageId="0" pageNumber="67" rowspan="1">J</th>
<th id="26903A8E2D7633F9D8169F50E8E9C571" colspan="1" pageId="0" pageNumber="67" rowspan="1">F</th>
<th id="8733AE466158DD3E9ABA15A778BE154D" colspan="1" pageId="0" pageNumber="67" rowspan="1">M</th>
<th id="C4842D594A943B77B6767B00B36032C5" colspan="1" pageId="0" pageNumber="67" rowspan="1">A</th>
<th id="F553B73FC0BA27FC56D420B06EE59F7E" colspan="1" pageId="0" pageNumber="67" rowspan="1">M</th>
<th id="75599A5730491A6AC0227DF99BA07B66" colspan="1" pageId="0" pageNumber="67" rowspan="1">J</th>
</tr>
<tr id="4838966BBE31CA23FFAAFE078A35A4BE" pageId="0" pageNumber="67">
<td id="970FB0A0AC6958A444A5428461B2D6F7" colspan="1" pageId="0" pageNumber="67" rowspan="1">
<taxonomicName id="AAB909A6584772887ABA025474590416" lsidName="E. greatheadi" pageId="0" pageNumber="67" rank="species" species="greatheadi">
<emphasis id="94F91614341E2C127F2A82B8806ACB69" italics="true" pageId="0" pageNumber="67">E. greatheadi</emphasis>
</taxonomicName>
</td>
<td id="310771862C094E28B7873FA3384A658D" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="0C69AF7FFE79DDB05263FF7755840D68" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="8CAD2262DD8BABA4018C3DC761E6E930" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="E13BC3FAF3444E60183A387291022B9A" colspan="1" pageId="0" pageNumber="67" rowspan="1">4</td>
<td id="9EC80E09A7991A7D6BC3F52D87FA6675" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="0342FB1C972D081101665DCA405BD40C" colspan="1" pageId="0" pageNumber="67" rowspan="1">1</td>
<td id="DC4F39BD847A4E67C511B2EB53C648D7" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="3339DEEF0CA8B343692A28CA8BBC4246" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="8D91BC600F9C540FBFC18CFFDB3AA498" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="85EEFDB068FA6084D5A907753A023EF1" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="BC74D7C5CE18D7B9D85263EE33AE1A3A" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="A32AE4106332D997577CD4C35EFFEC77" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
</tr>
<tr id="76393929E055882EC404CAD65BF7C78D" pageId="0" pageNumber="67">
<td id="C4987133ECC2C568257A51C969A93FBE" colspan="1" pageId="0" pageNumber="67" rowspan="1">
<taxonomicName id="7DAF595464BBFBDE0DD384DC532151EE" lsidName="E. melanoscopaeus" pageId="0" pageNumber="67" rank="species" species="melanoscopaeus">
<emphasis id="49CA95EAADD85300E02A5AF91EE6D349" italics="true" pageId="0" pageNumber="67">E. melanoscopaeus</emphasis>
</taxonomicName>
</td>
<td id="278A59252223AAC110C9C7EBFD95F54F" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="7C557406F6460AC4DE596D39A2954F40" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="8B9BC410C7ABD1AB50E9E9FB8AC1A65C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="2354C281C8471BA9AF0418F4F5C72531" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="7168C205E49400B70CCD2701B5228CA9" colspan="1" pageId="0" pageNumber="67" rowspan="1">4</td>
<td id="43CCC0122A882E7E320E3F8ED68F72CB" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="D277F88BADD7CBC520CAAAAD04755A23" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="5A8F1333AB4E710348D5E2F178104D13" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="8E8369B563451AEEB551048233961586" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="E31334496167FD1E891FFA0F1CA9FEB3" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="0D5D6248AE55041AEF94F78579B94C07" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="9A7D8337B5E5A77A3F0625B74743354B" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
</tr>
<tr id="655C6C09830765DE6B00DD68D4034599" pageId="0" pageNumber="67">
<td id="D3AC33B1507CE5F00C32735215806FDB" colspan="1" pageId="0" pageNumber="67" rowspan="1">
<taxonomicName id="2548253617816AC8FD4B9299C3CD1B46" lsidName="E. namibiensis" pageId="0" pageNumber="67" rank="species" species="namibiensis">
<emphasis id="2F46B94353FD40B2346F6225309308B9" italics="true" pageId="0" pageNumber="67">E. namibiensis</emphasis>
</taxonomicName>
</td>
<td id="4656EF8E74A580859390CC43D1822A66" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="D1C60E5C4EEAFC4A59B0FF8DD1F6330B" colspan="1" pageId="0" pageNumber="67" rowspan="1">4</td>
<td id="35CFD46C97379F769BC237C8D1698F40" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="2BA7FFDF6C36F895E254A8F607C8E809" colspan="1" pageId="0" pageNumber="67" rowspan="1">36</td>
<td id="524FEBDB99E5E21869C500D15A61CEF7" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="7E2E1820294392D440673B9986FDEAAF" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="9AC70FC67941E010CD9BE2B06B0ED172" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="AA64B146BB22EB4ADB008DBC63451358" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="3BFF510AF2D2DF15890F2A272A44B346" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="C3CC4F0E6A09AF92B0A2603A0F4272D2" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="9F1E7F68DB1A0C75134C682ED7C4E28C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="309034B376D0D3AA648CC4E321D81D6C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
</tr>
<tr id="6DB7D2AC4ECA5CE8A62CA52DA65F735C" pageId="0" pageNumber="67">
<td id="6BAF8E929C14E7DB8A5A1FD24D5F0C15" colspan="1" pageId="0" pageNumber="67" rowspan="1">
<taxonomicName id="F230847D5BFAE86648D968B5C9BBC088" lsidName="E. pusillipes" pageId="0" pageNumber="67" rank="species" species="pusillipes">
<emphasis id="F45D6A05D0164FBC8E5B19B0A075B8D2" italics="true" pageId="0" pageNumber="67">E. pusillipes</emphasis>
</taxonomicName>
sp. nov.
</td>
<td id="F8192632ECD3C0FFAB4BDE576257076D" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="23F87FEACD1716864731345D18F8048A" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="D3A5235F8E25C216830B7EB762EDD73E" colspan="1" pageId="0" pageNumber="67" rowspan="1">1</td>
<td id="F8838126E93A537BE825F751CE9EF879" colspan="1" pageId="0" pageNumber="67" rowspan="1">2</td>
<td id="8EE7BF728878787C79AADE9BA2584B26" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="827D677839DFDFC19C963AD3230D68E3" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="8DFF5BF88066CA71A2301656B6505707" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="06ADE58595E695173A9208211C639C3C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="6047E841C6F539AB75274CF8F977BC1C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="5A6A46286FCF6DD81D195AC4E78C9395" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="3487082BC7EE89D63012A32F5490EE07" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="F9F027AD1800D57CE5D92D8E828EF05A" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
</tr>
<tr id="F39EB91BEC4932593FA3CB961A2CA481" pageId="0" pageNumber="67">
<td id="91AEC7C753A85AEF5E27EF7DE1C20529" colspan="1" pageId="0" pageNumber="67" rowspan="1">
<taxonomicName id="C3A1753BB94CE1B4D043459C10F2FEB0" lsidName="E. torridus" pageId="0" pageNumber="67" rank="species" species="torridus">
<emphasis id="AABD6A2B41CB1876EDC0957355B3668F" italics="true" pageId="0" pageNumber="67">E. torridus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td id="1D4D611908DBD6DF957823DC422CF478" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="1607226DCDF9EF0FB8D75EAE756DDAF7" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="B1CDC82D8E38A2A97B7B5D9710DC2A1A" colspan="1" pageId="0" pageNumber="67" rowspan="1">1</td>
<td id="F28087C2648391BCB35A2412FBDF5EA8" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="017E9893AF8B51941F34A9DCE6BB7861" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="200980E0A49C9806BC22F0EFAA0B5348" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="9D5422C21E626779A8310CA9D98B4898" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="470DC05112F432D7241377E027A4261D" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="B5AB453D829FF9A8B63F62C5C14BE7E5" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="09E6E3659F3C4E0DAA1114BE5C24658A" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="2D14773362040F5725BE19AB4B00518D" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="6A5037C8C432656FDA20B52D8F207E7C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
</tr>
<tr id="42AFEE4854F0325ADFC601F286DA5AD6" pageId="0" pageNumber="67">
<td id="B2E14EC5F549C073ACC036178C84F9F5" colspan="1" pageId="0" pageNumber="67" rowspan="1">
<taxonomicName id="ED8433490AF871E0FF945B1C8DB267A1" lsidName="E. whittingtoni" pageId="0" pageNumber="67" rank="species" species="whittingtoni">
<emphasis id="BD03149C1939ED3E2F07C11F92C5CD6E" italics="true" pageId="0" pageNumber="67">E. whittingtoni</emphasis>
</taxonomicName>
</td>
<td id="ABA7FB5914B7F21D4690F22FE526A81C" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="47548F7E87519C07E2C98BE7C53D5CCD" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="CEE4034F66A241F9FEDDB4BA8C8427BA" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="8B91ACD5E7ADE8ED7BA062DB5C3E1F85" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="9E03370D880EFEEB878CFF5871964AA3" colspan="1" pageId="0" pageNumber="67" rowspan="1">1</td>
<td id="E41191E37F1DDE9997B868D37CCA2BA0" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="CF5AE65291F6BA9EC956F6BA5CDBD971" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="4632D0102E233682AA540D075D9268E8" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="A48E05ABA4EB4BC8E7EFEC14300D19EE" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="36E02711DF8F5B7EAA46AB12F829D012" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="1514A1DBEAEB75847444B71DF36DDB90" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
<td id="182D79D14748F02C3773C4107063FCE4" colspan="1" pageId="0" pageNumber="67" rowspan="1">-</td>
</tr>
<tr id="C1204B85FF5B640EEB81D785076EDA87" pageId="0" pageNumber="67">
<td id="4BEE0E0827B6012D6198F9AF4893D56F" colspan="1" pageId="0" pageNumber="67" rowspan="1">Total</td>
<td id="D629EFFC8E27FBC03BD122E75BC74664" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
<td id="74582B940ED6803AE951303563ADB35C" colspan="1" pageId="0" pageNumber="67" rowspan="1">4</td>
<td id="4DE7D4513B0A377447EF041FCCE6794E" colspan="1" pageId="0" pageNumber="67" rowspan="1">2</td>
<td id="5BA37077BBF4E50514667C7A164EBC4E" colspan="1" pageId="0" pageNumber="67" rowspan="1">42</td>
<td id="30DCA79221F0132FA272912408CF2CCF" colspan="1" pageId="0" pageNumber="67" rowspan="1">5</td>
<td id="4BB76E8F9F0010F26E5DC2D2C7A8565A" colspan="1" pageId="0" pageNumber="67" rowspan="1">1</td>
<td id="C558E1378188C023F9DEFC4F9C50B932" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
<td id="DED16808228CD2A50E4C2BF1E34EE58F" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
<td id="D40FE0CF60EF1F0CEE7F261109AB2C7D" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
<td id="92373F75B174A6633AF3D4C11CEA3F91" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
<td id="6B2283D64189BDD14AFD5428099545EE" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
<td id="A3EAF0AAA91C9047861B5E2846D48251" colspan="1" pageId="0" pageNumber="67" rowspan="1">0</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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