239 lines
35 KiB
XML
239 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.923.47104" ID-GBIF-Dataset="1e711d27-f2ea-4e62-9b52-19f33fa223b9" ID-PMC="PMC7142161" ID-Pensoft-Pub="1313-2970-923-65" ID-Pensoft-UUID="5A90240DCD655B319081ECB2F39B90B3" ID-PubMed="32292271" ID-ZooBank="B10E9E718A434560BBAC47F52E401045" ModsDocID="1313-2970-923-65" checkinTime="1585826883453" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2020" docId="7E6813EDDB865FA68860961EAF78AB06" docLanguage="en" docName="ZooKeys 923: 65-77" docOrigin="ZooKeys 923" docSource="http://dx.doi.org/10.3897/zookeys.923.47104" docTitle="Desmopachria mortimer Miller 2020, sp. nov." docType="treatment" docUuid="F8656CD2-3D92-4B2B-94DA-5B04F8F029EC" docUuidSource="ZooBank" docVersion="5" id="5A90240DCD655B319081ECB2F39B90B3" lastPageNumber="65" masterDocId="5A90240DCD655B319081ECB2F39B90B3" masterDocTitle="Two new species of Desmopachria Babington, 1841 in the D. convexa species group (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini)" masterLastPageNumber="77" masterPageNumber="65" pageNumber="65" updateTime="1701286543009" updateUser="admin">
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<mods:title id="9B46CBE5D9BB828A6DDA2BF0997D2081">Two new species of Desmopachria Babington, 1841 in the D. convexa species group (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini)</mods:title>
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<mods:namePart id="D643E7B219A422637C68EB47D56B5888">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="BBBEB2AD1598209938AD778AA4047BDE">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001 USA</mods:affiliation>
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<mods:nameIdentifier id="469D68AC092772C54CDDD874DAB12749" type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:title id="7C237AA56B809CEFCC1CFCBCF6D133F0">ZooKeys</mods:title>
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<mods:part id="7849C9685BB603695B9E451EA09CADF1">
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<mods:date id="1171FD58E57B15A0EB12C489CE9159F6">2020</mods:date>
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<mods:detail id="DDAA8F17CAF379EB0550B6947140453D" type="volume">
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<mods:number id="A211296CE7378321E61AA0879B898EC7">923</mods:number>
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<mods:start id="D8CB66B97706EC2D6198E857F6446D29">65</mods:start>
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<mods:url id="F8974A8C9F5AE9F78BCF093A10056B6D">http://dx.doi.org/10.3897/zookeys.923.47104</mods:url>
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<mods:identifier id="4A324E416B62ACF907AE367B72AFC8DF" type="DOI">http://dx.doi.org/10.3897/zookeys.923.47104</mods:identifier>
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<treatment id="7E6813EDDB865FA68860961EAF78AB06" ID-GBIF-Taxon="163421241" LSID="urn:lsid:zoobank.org:act:F8656CD2-3D92-4B2B-94DA-5B04F8F029EC" httpUri="http://treatment.plazi.org/id/7E6813EDDB865FA68860961EAF78AB06" lastPageNumber="65" pageId="0" pageNumber="65">
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<subSubSection id="AB9A89EBBD4BE3652B277233277D083F" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="4FE658BAE27AEF2209A21CCB8E73E450" pageId="0" pageNumber="65">
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<taxonomicName id="C0482102875F369323E6AEC7078E6E1D" ID-CoL="8LZTC" LSID="7E6813ED-DB86-5FA6-8860-961EAF78AB06" authority="Miller, 2020" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria mortimer" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mortimer" status="sp. nov.">Desmopachria mortimer</taxonomicName>
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<taxonomicNameLabel id="BE69D5AB4A265D5A1478CE20E8ADA805" pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
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<figureCitation id="7D10C3125532B9400E1BEEC8938CC412" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">Figures 6-10</figureCitation>
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<figureCitation id="FE885AA128A2FD7B0B5221FC0A523D49" captionStart="Figures 58, 59" captionStartId="F4" captionText="Figures 58, 59. Desmopachria new species, distribution. 58 D. manco (South America). 59 D. mortimer (Central America)." figureDoi="10.3897/zookeys.923.47104.figures58-59" httpUri="https://binary.pensoft.net/fig/396458" pageId="0" pageNumber="65">, 59</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="BCC47D7674E529A3729B55D9F366E2D2" pageId="0" pageNumber="65" type="locality">
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<paragraph id="3A0A7996CB2A9E3C7FA2C5767DDDA086" pageId="0" pageNumber="65">Type locality.</paragraph>
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<paragraph id="D4CB908496C3302D653516F97A14E129" pageId="0" pageNumber="65">Costa Rica, Cartago Province, Tapanti National Park, pasture by Rio Orosi, ca. 1200 m.</paragraph>
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</subSubSection>
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<subSubSection id="C8A71001563188A08914D93BB08A1A34" pageId="0" pageNumber="65" type="type material">
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<paragraph id="B010F06BC1AE18E4B22FC307FEEC6866" pageId="0" pageNumber="65">Type material.</paragraph>
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<paragraph id="163513A3C3258329C22971791D711191" pageId="0" pageNumber="65">
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Holotype in KUNHM, male labeled, "Costa Rica: Cartago Province Tapanti National Park: 24.v.2006 pasture by Rio Orosi: c. 1200 m leg. A.E.Z. Short, AS-06-043/ SEMC0895195 KUNHM-ENT/ Holotype
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<taxonomicName id="88E4DD810A9783482345B071B818B434" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria mortimer" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mortimer">
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<emphasis id="2D3B0E67E1E6DA2E1B25E61050E81567" italics="true" pageId="0" pageNumber="65">Desmopachria mortimer</emphasis>
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</taxonomicName>
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Miller, 2020 [red label with black line border]." Paratypes, 41, labeled same as holotype except with different specimen barcode labels (Table
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<tableCitation id="6801A073AF3C72889932151C67688A46" captionStart="Table 1" captionStartId="T1" captionText="Table 1. SEMC museum numbers for D. mortimer paratypes." httpUri="http://table.plazi.org/id/78245BEE927AD3828E75A8D8BB37A2F0" pageId="0" pageNumber="65" tableUuid="78245BEE927AD3828E75A8D8BB37A2F0">1</tableCitation>
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) and each with "Paratype
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<taxonomicName id="CA4D2910EFBB0C987A477CF0F9A190EB" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria mortimer" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mortimer">
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<emphasis id="708A34CF8BDBCBA0D543A698AA4BE59C" italics="true" pageId="0" pageNumber="65">Desmopachria mortimer</emphasis>
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</taxonomicName>
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Miller, 2020 [blue label with black line border]."
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</paragraph>
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<caption id="78245BEE927AD3828E75A8D8BB37A2F0" ID-Table-UUID="78245BEE927AD3828E75A8D8BB37A2F0" httpUri="http://table.plazi.org/id/78245BEE927AD3828E75A8D8BB37A2F0" pageId="0" pageNumber="65" start="Table 1" startId="T1">
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<paragraph id="BC6627256F15C701654D4FB8A0828FFB" pageId="0" pageNumber="65">
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<emphasis id="14ABA14F6604EF0BACE14A44D7573D85" bold="true" pageId="0" pageNumber="65">Table 1.</emphasis>
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SEMC museum numbers for
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<taxonomicName id="973B3BD9ED39C486BC194A09E30D4E4C" lsidName="D. mortimer" pageId="0" pageNumber="65" rank="species" species="mortimer">
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<emphasis id="467B13F83B03A8B10E46F5B25FCB4FFD" italics="true" pageId="0" pageNumber="65">D. mortimer</emphasis>
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</taxonomicName>
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paratypes.
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</paragraph>
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</caption>
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<paragraph id="147A93D02F3DB36BE25F7A6CEE35D3E7" pageId="0" pageNumber="65">
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<table id="1D3153E3C0D4B4A86D609025A112CC3A" pageId="0" pageNumber="65">
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<tr id="6441E775BC753AA6C21A3F1437B0E640" pageId="0" pageNumber="65">
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<td id="B8C71A255943C80AEA5A0055599B2787" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895144</td>
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<td id="00481A660CCEB5DAF176282D416C4253" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895156</td>
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<td id="F212D4C2B7AE12FA27A3E3FCA35EE3F0" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895167</td>
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<td id="15F328692E82C9658453A520C4D5861E" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895182</td>
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</tr>
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<tr id="5A3802574E4B5D52AEEE920ACBEFCF50" pageId="0" pageNumber="65">
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<td id="4DCC823ECC2EF61F0FD9AFF2CC8F9745" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895145</td>
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<td id="38EC77AE2D664B931965324FC9D0FA46" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895157</td>
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<td id="6B2D0D4E6A660B4E987C18721D023278" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895168</td>
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<td id="79AEA7325E622B9245152500F58EA408" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895184</td>
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</tr>
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<tr id="D4C3D0884AD869A8FAC34625EF68A617" pageId="0" pageNumber="65">
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<td id="1072944164C4E38CF9FCB8B70FEA17B7" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895146</td>
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<td id="778DC365978E55B252D48BA6D346BD67" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895158</td>
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<td id="05417B2E67F007042256AC7056DEB8C0" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895169</td>
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<td id="AD63B42B2756D66A9BBE507917412962" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895185</td>
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</tr>
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<tr id="EFDEC2FA767D2E82EF2A62772A872B4C" pageId="0" pageNumber="65">
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<td id="0FFC7A5D6C4EDD52FF9196545A5DF414" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895147</td>
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<td id="EC760DC09438EF1F8220E94302F6C5D1" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895159</td>
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<td id="6181EA635D93B4BC91329ED626A1CB90" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895170</td>
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<td id="524C7CD583C20EE3F59C3F55B7E0B8AD" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895190</td>
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</tr>
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<tr id="F39A3F636D1BF17F68F5B6BAB6153175" pageId="0" pageNumber="65">
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<td id="B5707DA07CDAD2FBDC1676F85B3998D5" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895148</td>
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<td id="3606F04DDAA264EE5CFC6B7A4331DC0E" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895160</td>
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<td id="FC72B41B404384C38FC09F0427AC3AA2" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895171</td>
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<td id="55B32523F4D2CC30CE5BB06485A8D601" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895191</td>
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</tr>
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<tr id="9321D8153A568AA1EB64958F43FD09A2" pageId="0" pageNumber="65">
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<td id="155D416B197C39E443FDB8293C9A298E" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895149</td>
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<td id="034B9A23A530B1B94594298E5C494A8C" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895161</td>
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<td id="48B038DB85FC771DCC4D87D8E728246F" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895172</td>
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<td id="7C7DE540DB8C856DC7103075522EF440" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895192</td>
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</tr>
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<tr id="9AEAA33548C4D8B15EAA88550C6C813B" pageId="0" pageNumber="65">
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<td id="13D012274D3AD1B9757A74BD84D70CDC" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895150</td>
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<td id="8A09C45FE50845DAB5939B3EB3D9036E" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895162</td>
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<td id="C4ADE162C641AD618B4BA26C71EBB7E7" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895173</td>
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<td id="BF76FDA0798B6D2E5AA47848EEFA52F6" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895193</td>
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</tr>
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<tr id="7BA55105F99AA8EFF6099686B05E628F" pageId="0" pageNumber="65">
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<td id="2F375D257B9564EDDD49580D95972460" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895151</td>
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<td id="B43EA329809FFF8713C5B676862BBAF9" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895163</td>
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<td id="A6A640A7570D339A6D4431509541FA90" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895175</td>
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<td id="72CE2A0DC00CE69408DD347199ED5FA9" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895194</td>
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</tr>
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<tr id="B3A8405184B1FDF051EB0DB375390F13" pageId="0" pageNumber="65">
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<td id="A355AEFA63B6594246FE65C0815E73FA" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895152</td>
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<td id="0D6CD6D5E778D72BF7313AE4EFC72E02" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895165</td>
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<td id="265F88CC61D6FCCF1E6E90082DBE86AD" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895177</td>
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<td id="E7AE7BB9821870A2896F5D3DC1EE927A" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895196</td>
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</tr>
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<tr id="14E3B73BB703FAB07297D2877713778A" pageId="0" pageNumber="65">
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<td id="D97824D2D5AB25B258F1A8D4128010A9" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895153</td>
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<td id="042E97D24BC8ED8AEADC103C47594917" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895166</td>
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<td id="407263F77A7D21067E4EC60511A41A6B" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895181</td>
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<td id="52D0099BB4E93DFF38A7000A26920B61" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895197</td>
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</tr>
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||
<tr id="49A995E9C3FA982C7878266EB1AD6B80" pageId="0" pageNumber="65">
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<td id="C7881481BF3E8C4E4BC6C7CF28B44CD9" colspan="1" pageId="0" pageNumber="65" rowspan="1">SEMC0895154</td>
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<td id="08E6D76C80C45A7EBD9DCF5A9A676A54" colspan="1" isEmpty="true" pageId="0" pageNumber="65" rowspan="1">-</td>
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<td id="0B386960F83D03C88E08C76C7E86BE48" colspan="1" isEmpty="true" pageId="0" pageNumber="65" rowspan="1">-</td>
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<td id="4781B59627EC306377BFDEABB7BE8A59" colspan="1" isEmpty="true" pageId="0" pageNumber="65" rowspan="1">-</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection id="C6DE4FD2A423B51C7F1A78C32A56F6D1" pageId="0" pageNumber="65" type="diagnosis">
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<paragraph id="F3B9CE4F044BC7E0E61C4F2B04C73CF8" pageId="0" pageNumber="65">Diagnosis.</paragraph>
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<paragraph id="6E91D6AAD2C1546C49AF89A5999A8511" pageId="0" pageNumber="65">
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This is a moderately sized, somewhat elongate species of
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<taxonomicName id="027C5C2483F2F8482AE694BE2476BA46" authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis id="5996D60C2F042B831C53C15A74363E15" italics="true" pageId="0" pageNumber="65">Desmopachria</emphasis>
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||
</taxonomicName>
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(Fig.
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<figureCitation id="4EE7C2C70CAAB66AC4F91E67B41EDC95" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
|
||
; TL = 1.7-1.9 mm). The dorsal coloration is characteristic with diffuse darker regions medially and extending laterally onto the surface of the elytron (Fig.
|
||
<figureCitation id="B705C3C32C7B09B867E5A546E38E83E5" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">6</figureCitation>
|
||
). The shape of the male genitalia is diagnostic (Figs
|
||
<figureCitation id="7438E8012DB83B53B62CBA791124AD60" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">7-10</figureCitation>
|
||
) and place the species in the
|
||
<taxonomicName id="C00731728D1FB9B90D01800A9AA670AB" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria convexa-convexa" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="convexa-convexa">
|
||
<emphasis id="148723976F68AAB4D773060FBA622B7F" italics="true" pageId="0" pageNumber="65">Desmopachria convexa-convexa</emphasis>
|
||
</taxonomicName>
|
||
subgroup (see above). The median lobe is short (Figs
|
||
<figureCitation id="260739E6D45DB212DBC58414D6B2B518" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">7</figureCitation>
|
||
,
|
||
<figureCitation id="B8B547276BBD382724EDB47411AEC823" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">8</figureCitation>
|
||
), and in lateral aspect it is irregularly broad basally with the apical portion slender, short and curved dorsally to a narrowly rounded apex (Fig.
|
||
<figureCitation id="FEAE2C5765A23F9D120226F58BC9064A" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">7</figureCitation>
|
||
). In ventral aspect it is very broad, basally broadly U-shaped, apically deeply bifid, with each ramus slender and strongly curved medially (Fig.
|
||
<figureCitation id="6643212AEE4B790417D6DADAAF500767" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">8</figureCitation>
|
||
). The lateral lobe in ventral aspect is irregularly shaped, broad, with the apex narrowly truncate and with a subapical articulable lobe that extends well beyond the apex of the lateral lobe and is apically acuminate and sharply pointed (Fig.
|
||
<figureCitation id="DBD77E8BD0D0146955635ECD8A881F4C" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">9</figureCitation>
|
||
). The lateral lobe in lateral aspect is very broad basally with the apex slender, elongate, directed dorsad, and with the subapical articulable lobe elongate, curved and apically sharply pointed (Fig.
|
||
<figureCitation id="34DDC466EB17CAADD9477308B63E75BE" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46C3A94B5E3AD105ABD53EC2035201BF" pageId="0" pageNumber="65" type="description">
|
||
<paragraph id="15365F89011FF6330E5E32FFCF43C759" pageId="0" pageNumber="65">Description.</paragraph>
|
||
<paragraph id="BF7A6276D150C2682075103A39B7A469" pageId="0" pageNumber="65">
|
||
<emphasis id="58C5EE00DC3DF3CCA447078AD7864E2D" bold="true" italics="true" pageId="0" pageNumber="65">Measurements.</emphasis>
|
||
TL = 1.7-1.9 mm, GW = 1.1-1.2 mm, PW = 0.9-1.0 mm, HW = 0.5-0.6 mm, EW = 0.3-0.4 mm, TL/GW = 1.5-1.6, HW/EW = 1.2-1.4. Body broad but slightly elongate posteriorly, lateral margins continuous between pronotum and elytron (Fig.
|
||
<figureCitation id="BDF1B02FBD026D0A0D4DB5B091F174EA" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">6</figureCitation>
|
||
), dorsoventrally broad.
|
||
</paragraph>
|
||
<paragraph id="22DB21A7FD6B15D56A3969F6F5DEFD6F" pageId="0" pageNumber="65">
|
||
<emphasis id="153C625FEB367EE8232CF20EC9852305" bold="true" italics="true" pageId="0" pageNumber="65">Coloration</emphasis>
|
||
(Fig.
|
||
<figureCitation id="C870475F03F958AD87B05330E58C76D3" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">6</figureCitation>
|
||
). Head orange, diffusely darker orange posteriorly. Pronotum orange, with darker orange region along posterior margin. Elytron orange, with diffuse, weakly margined darker regions along anteromedial margin, along elytral suture and expanded somewhat onto disc medially; surface not iridescent. Mesoventrite, metacoxa and abdominal ventrites dark orange, other ventral surfaces and appendages lighter orange.
|
||
</paragraph>
|
||
<paragraph id="593F6E3706F85CA14719A6E2BFC944C7" pageId="0" pageNumber="65">
|
||
<emphasis id="CCD35BD5548A949F9C8AC3DD30AC82BE" bold="true" italics="true" pageId="0" pageNumber="65">Sculpture and structure.</emphasis>
|
||
Head (Fig.
|
||
<figureCitation id="C9624749B2AC678B6B755A4D53E7FA89" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">6</figureCitation>
|
||
) broad, anteriorly produced in broadly rounded margin; anterior margin of clypeus margined with conspicuous, continuous narrow bead; surface of head shiny, very finely and sparsely punctate, punctation slightly denser posteriorly; eyes large (HW/EW = 2.0); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, lobe at anterodorsal angle, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, slightly curved with narrow bead, bead wider medially, slightly angulate medially; surface shiny, nearly impunctate medially, more but sparsely punctate along anterior and posterior margins, punctation variable, fine to course. Elytron moderately broad, laterally broadly curved; surface shiny, more coarsely and evenly punctate than pronotum, punctation shallow and indistinct, of variable sizes. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process slender anteriorly, with distinctive, small medial tubercle, apically short and broad, concave, apically broadly pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, with very few indistinct punctures, anteromedially with curved transverse carina between posterior margins of mesocoxal cavities; metaventrite wings extremely slender. Metacoxa with medial portion short, less than 1/2 length of metaventrite medially, metacoxal lines distinctly divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, evenly but shallowly and indistinctly punctate, punctures evenly distributed. Metatrochanter large, longer than ventral margin of metafemur anterior to metatrochanter apex; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, nearly impunctate.
|
||
</paragraph>
|
||
<paragraph id="51376864C252F904B9B899CD21D9BDFF" pageId="0" pageNumber="65">
|
||
<emphasis id="7A08DFE083BD0424179BB963C67AFDAA" bold="true" italics="true" pageId="0" pageNumber="65">Male genitalia.</emphasis>
|
||
Male median lobe in lateral aspect irregularly shaped, apical portion slender, short, slightly curved dorsad, apically narrowly rounded (Fig.
|
||
<figureCitation id="98E155CDE360CF34B13B60CC339A6BB8" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">7</figureCitation>
|
||
). Median lobe in ventral aspect very broad, basally U-shaped, apically deeply and broadly bifid, each ramus slender, apically narrowly rounded, gently curved, and convergent medially at apices (Fig.
|
||
<figureCitation id="0F90639B079E9697AB087921A35F0283" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">8</figureCitation>
|
||
). Lateral lobe in ventral aspect irregularly shaped, broad, with subapical articulable lobe slender, undulate, apically sharply pointed and elongate, extending well beyond apex of lateral lobe (Fig.
|
||
<figureCitation id="D67B5B761AD8419C8497911917246FEF" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">9</figureCitation>
|
||
). Lateral lobe in lateral aspect broad basally, apex slender, oblique, directed dorsad, subapical articulable process slender, elongate curved and sharply pointed (Fig.
|
||
<figureCitation id="FA491776470AA2610DB27A48C78B6348" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AA20DF4AD9FF95353D24A8D1121F30C0" pageId="0" pageNumber="65" type="etymology">
|
||
<paragraph id="8B46F87E5D0BF46E535BC78A95021357" pageId="0" pageNumber="65">Etymology.</paragraph>
|
||
<paragraph id="03202C50E2E5997576001F6CB833D083" pageId="0" pageNumber="65">
|
||
This species is named
|
||
<taxonomicName id="2421866619144CF390A7834DD6F82C2C" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="mortimer" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mortimer">
|
||
<emphasis id="74DEFD505922B76722DDA73288113599" italics="true" pageId="0" pageNumber="65">mortimer</emphasis>
|
||
</taxonomicName>
|
||
, after Colonel Douglas Mortimer, the older bounty hunter in the film "For a Few Dollars More".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31729734257418BEEAF41FA9782A0524" pageId="0" pageNumber="65" type="distribution">
|
||
<paragraph id="2FBB8B0AB28D051ED4C4750399CB8ED4" pageId="0" pageNumber="65">Distribution.</paragraph>
|
||
<paragraph id="3B6440D91FD4CA81CD602F651FCD832B" pageId="0" pageNumber="65">
|
||
This species is known only from one locality in Costa Rica, Prov. Cartago (Fig.
|
||
<figureCitation id="2053332E33C91E41C497789533365157" captionStart="Figures 58, 59" captionStartId="F4" captionText="Figures 58, 59. Desmopachria new species, distribution. 58 D. manco (South America). 59 D. mortimer (Central America)." figureDoi="10.3897/zookeys.923.47104.figures58-59" httpUri="https://binary.pensoft.net/fig/396458" pageId="0" pageNumber="65">58</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDA2AB3FEF1CAA86AA85C1E707243B53" pageId="0" pageNumber="65" type="habitat">
|
||
<paragraph id="59251D8E1F8758504E67DC455DF44657" pageId="0" pageNumber="65">Habitat.</paragraph>
|
||
<paragraph id="4C97A136511A4A7F88E7B6DC5596290E" pageId="0" pageNumber="65">
|
||
The only known habitat information is the type series collected in a
|
||
<normalizedToken id="7D3FB871D8BFD7FB0C3E9CEF53AA750B" originalValue="“pasture.”">"pasture."</normalizedToken>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |