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<document ID-CLB-Dataset="6659" ID-DOI="http://dx.doi.org/10.3897/zse.97.59143" ID-GBIF-Dataset="83d5b180-4e0b-44d1-ad83-67ad55ee0355" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="BD006A3AD2C6516AB596CDA05E0F375D" ID-ZooBank="1D5553BA24894F2C9527B7E46B9B432B" ModsDocID="1860-0743-97-1-1" checkinTime="1621423297205" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Pall-Gergely, Barna, Sutcharit, Chirasak &amp; Tongkerd, Piyoros" docDate="2021" docId="7E65B4F0822C58BABD666E44B1A0C99B" docLanguage="en" docName="ZoosystEvol 97(1): 1-20" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/zse.97.59143" docTitle="Dicharax burchi Jirapatrasilp &amp; Pall-Gergely 2021, sp. nov." docType="treatment" docUuid="8797FE78-84C5-4887-AF61-923C1E341734" docUuidSource="ZooBank" docVersion="5" id="BD006A3AD2C6516AB596CDA05E0F375D" lastPageNumber="1" masterDocId="BD006A3AD2C6516AB596CDA05E0F375D" masterDocTitle="The operculate micro land snail genus Dicharax Kobelt &amp; Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species" masterLastPageNumber="20" masterPageNumber="1" pageNumber="1" updateTime="1732748620476" updateUser="ExternalLinkService">
<mods:mods id="B502E155F7F48901589A5DBB51A64761" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="804DDA2CD5E5FD206E0052FDA4932CCB">
<mods:title id="A89A2D9BDCE280B806BEAF16A045EE7B">The operculate micro land snail genus Dicharax Kobelt &amp; Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species</mods:title>
</mods:titleInfo>
<mods:name id="F8AB1E927933B78B79CF8593686E68CD" type="personal">
<mods:role id="DE5C7B25BFE618FAFE016846E9B88B9B">
<mods:roleTerm id="9711739D49ABB5593D0C2B49BAEFE3B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8EB9F49A506B8EF3A3A0D69D26298730">Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier id="BA35E41854A4B489D03C17B1ACFD5656" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation id="5A06105E6DBD6C752ADCA8A4DD3F776C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name id="7E76B717CEE5C8C21FDA6E0BD5CB51D2" type="personal">
<mods:role id="DAB04A36F82EEDD7B08B7F706D58115C">
<mods:roleTerm id="F0757F5281F72E860C6CE9FC8761E883">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="626BC59D6ED9E1ACADF6BFE9C59BDD1E">Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier id="4726F5B3769E7EA90664F2AC6E82E8A5" type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation id="AB6E6DCBCB0C1F4D1C00890713F79A27">Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name id="0E6AD41B6D73FF279EC7FD5E3EFC3585" type="personal">
<mods:role id="86A968724A58F131DF80F40727FF8098">
<mods:roleTerm id="6B2F2555F6A70E4DC589047B4D530C43">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D00AF02B1842102257F4038D8BAFCCB1">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="E20FBF4039E5E530AEC4181536E0412C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name id="31BDEBA1089B0C402965E6BE93104152" type="personal">
<mods:role id="4975DC31BF5B2E7E979D4CD5F93FC2A2">
<mods:roleTerm id="7F75D664D37540DD9A9E754FE3CA8E8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D04C0883CAE62EA1E3FC8E661A36EEA0">Tongkerd, Piyoros</mods:namePart>
<mods:nameIdentifier id="B2B8D07583C7B024209D0983358BC957" type="ORCID">https://orcid.org/0000-0001-9221-9293</mods:nameIdentifier>
<mods:affiliation id="4225D95C1A617A344DFE1A4FC0EF87CB">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="D12EE98B09AA639FF04B1D254C5D613B" type="email">piyorose@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F1ED21AA181C77D02035A842E19F571A">text</mods:typeOfResource>
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<mods:title id="76F1B388FF4EEEF89E2EB1DAD264089D">Zoosystematics and Evolution</mods:title>
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<mods:part id="91923285497DFF7FAA7C23F0319405A2">
<mods:date id="9F3C7F747444AAD8D10A9613D2E08909">2021</mods:date>
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<mods:number id="376DA8DD773725C156331E9AD4E3F676">2021-01-05</mods:number>
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<mods:detail id="8A9DF1FC1C8D0FFE4B3155BA91C897C5" type="volume">
<mods:number id="750CBE574942CC945D4D5A8F3550765D">97</mods:number>
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<mods:classification id="C7A4E23FED9853E7A805EE07305BFA1D">journal article</mods:classification>
<mods:identifier id="3BB213B376745212C24926F681784D21" type="DOI">http://dx.doi.org/10.3897/zse.97.59143</mods:identifier>
<mods:identifier id="42B532B6402BD48A2958D70A23E1659B" type="Pensoft-Pub">1860-0743-1-1</mods:identifier>
<mods:identifier id="0179092AD4FFF57DC535099FF8711C6C" type="ZooBank">1D5553BA24894F2C9527B7E46B9B432B</mods:identifier>
<mods:identifier id="B170F6D899E6C821847508A339237AE6" type="Pensoft-UUID">BD006A3AD2C6516AB596CDA05E0F375D</mods:identifier>
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<treatment id="7E65B4F0822C58BABD666E44B1A0C99B" ID-GBIF-Taxon="182236446" LSID="urn:lsid:zoobank.org:act:8797FE78-84C5-4887-AF61-923C1E341734" httpUri="http://treatment.plazi.org/id/7E65B4F0822C58BABD666E44B1A0C99B" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Alycaeidae" scope_subClass="Caenogastropoda">
<subSubSection id="1AEA35401DBF8CA4397697D9C6BAF391" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="833FBA91116DD512B770B071C2657B29" pageId="0" pageNumber="1">
<taxonomicName id="222335008E21C70C746067AE3F723CF4" LSID="http://zoobank.org/8797FE78-84C5-4887-AF61-923C1E341734" authority="Jirapatrasilp &amp; Páll-Gergely" authorityName="Jirapatrasilp &amp; Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax burchi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burchi" status="sp. nov.">
Dicharax burchi Jirapatrasilp &amp;
<normalizedToken id="1155A2967E6FAC668BBB3BD7B7110F35" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="FF90BD061BC36E1AB0569D9BEC0CDD16" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="8388BE4740A24BA5193026999017B1A6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">Figs 6F</figureCitation>
<figureCitation id="0CCA3114D9F372D92CE6AB4728E34CAA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation id="FE18CE92A384654291055218A7DDE508" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">, 11A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E9B777744D9E84E3F8025D69B831D68A" pageId="0" pageNumber="1" type="type material">
<paragraph id="99576D6D81430A5EFCC4EDD7016434A6" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="7DCBA1BE847CC4061AD39C9B5DC801F7" pageId="0" pageNumber="1">
<emphasis id="CC3EB1AF08D92CC071F4097AECEE6EE9" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
CUMZ 7428/1 (Fig.
<figureCitation id="9B27594BBB30EFA36E4C19E76C25B3D6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10A</figureCitation>
),
<emphasis id="C44635E94F55842B1021E819A32599AA" bold="true" italics="true" pageId="0" pageNumber="1">paratypes</emphasis>
CUMZ 7428/2 (1 shell and 6 specimens in ethanol; Fig.
<figureCitation id="E2EFA462F6E56DA4D71E48EBD613359F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">6F</figureCitation>
), HNHM (1 shell), NHMUK 20200326 (2 shells) and SMF (2 shells) from Tham Phaya Nakharat, Khon Kaen Province, Thailand,
<geoCoordinate id="4039694C3D1F4B863A93453B70DF803C" degrees="16" direction="north" minutes="48" orientation="latitude" precision="1" seconds="32.8" value="16.80911">16°48'32.8&quot;N</geoCoordinate>
,
<geoCoordinate id="53D99EBC7A32F8788BA3EEC6A559E62F" degrees="101" direction="east" minutes="57" orientation="longitude" precision="1" seconds="23.9" value="101.956635">101°57'23.9&quot;E</geoCoordinate>
, 21 July 2020, coll. P. Tongkerd, A. Pholyotha.
</paragraph>
<caption id="25A0F93F1F356F1CA648D436C344BE90" doi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph id="D77D85E0A6528179F99E13EE46FB1206" pageId="0" pageNumber="1">
<emphasis id="CB6EC92CDDF80739A55E5B4F0B5CAFE3" bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName id="A2426B86181FC6F81790673B22EBC657" authorityName="Jirapatrasilp &amp; Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax burchi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burchi">
<emphasis id="0700B17B7986987DBE0F003E01A5B612" italics="true" pageId="0" pageNumber="1">Dicharax burchi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="87794A284D069C2289794C50696F489A" bold="true" pageId="0" pageNumber="1">A.</emphasis>
Holotype CUMZ 7428/1 with operculum;
<emphasis id="B7B0DE1A6748A65F0B8E78877B50FC6D" bold="true" pageId="0" pageNumber="1">B, C.</emphasis>
Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province;
<emphasis id="8354C6C180DE0E7DD1A1100E7BD25897" bold="true" pageId="0" pageNumber="1">D.</emphasis>
Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="35F148177118B1473F48DB5A6AF2E1A4" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="F2B9AE58B6803CA7495B241C27869BC3" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="48D37B074435B6EB374C3924176AB715" pageId="0" pageNumber="1">
<materialsCitation id="3350DDA4587765BCE4CCE6651A4AD0AB" accessionNumber="UF 347279" collectingDate="1987-06-23" collectorName="NW Ban Soppong, F. G. Thompson" country="Thailand" elevation="800" latitude="19.55305" location="Soppong" longLatPrecision="1" longitude="98.192215" specimenCount="1" stateProvince="Mae Hong Son Province">
<accessionNumber id="C9A927B812066BBDD0399D221F7E3C11" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/UF347279">UF 347279</accessionNumber>
from limestone pass 10.0 km
<collectorName id="6EACC48BE75E79D28FCE12D426151CC9">
NW Ban
<location id="755052286A1814E9B41D741532591067" LSID="urn:lsid:plazi:treatment:7E65B4F0822C58BABD666E44B1A0C99B:755052286A1814E9B41D741532591067" country="Thailand" latitude="19.55305" longLatPrecision="1" longitude="98.192215" name="Soppong" stateProvince="Mae Hong Son Province">Soppong</location>
</collectorName>
,
<collectingRegion id="672F7F8BC5C7320208E464A76AF25E1A" country="Thailand" name="Mae Hong Son">Mae Hong Son Province</collectingRegion>
,
<collectingCountry id="E5CFB9D89E52C9288010A6B54D84EEF1" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="9244D17295D284143D7D38BA1D750F0C" degrees="19" direction="north" minutes="33.183" orientation="latitude" precision="1" value="19.55305">19°33.183'N</geoCoordinate>
,
<geoCoordinate id="32CB6FC5970671E5055A08DD82BA0277" degrees="98" direction="east" minutes="11.533" orientation="longitude" precision="1" value="98.192215">98°11.533'E</geoCoordinate>
,
<elevation id="30CB33B67407990A032B6DC90CCE57C0" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<quantity id="97E88955FA6D32B5B5DA14F6C2035FD9" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="56D0AA0C1CEFBA64189A8681EC4D16C2" value="1987-06-23">23 Jun. 1987</collectingDate>
, coll.
<collectorName id="D1B47180930097412EF9F5178CAAA309">F.G. Thompson</collectorName>
(1 shell; Fig.
<figureCitation id="B5C0208014B61333284F23C096036A9A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10D</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="856AB89DB90AE63319F81B902ACF186F" accessionNumber="UF 345622" collectingDate="1988-03-20" collectingDateMax="2018-10-07" collectingDateMin="1988-03-20" collectorName="K. Auffenberg, A. Hunyadi, Hunyadi, K, Okubo, J. U, Doi Chiang Dao, S. Panha, Panha, J. B, Burch, P" country="Thailand" elevation="820" latitude="19.24075" location="Road" longLatPrecision="1" longitude="98.8805" specimenCount="1" stateProvince="Mae Hong Son Province">
<accessionNumber id="DE2D4F306848A60CA6306725601D54C8" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/UF345622">UF 345622</accessionNumber>
from
<locationDeviation id="738737DCD90280E33BDA88EC118694E2" location="Soppong">
<quantity id="95F2E716C359E6C7C6C67EFEE1A74252" metricMagnitude="4" metricUnit="m" metricValue="1.02" unit="km" value="10.2">10.2 km</quantity>
WNW of Soppong
</locationDeviation>
,
<location id="724369B2E036B74DC723D2E4F03734B6" LSID="urn:lsid:plazi:treatment:7E65B4F0822C58BABD666E44B1A0C99B:724369B2E036B74DC723D2E4F03734B6" country="Thailand" latitude="19.24075" longLatPrecision="1" longitude="98.8805" name="Road" stateProvince="Mae Hong Son Province">Road</location>
1095,
<collectingRegion id="EBC0BC92EC6960B75771255594A34ACB" country="Thailand" name="Mae Hong Son">Mae Hong Son Province</collectingRegion>
,
<collectingCountry id="3F59D10D71004B3E5377ECA7691D8A9B" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="047967D0F89D6B92BD9BBD6357F1BB61" degrees="19" direction="north" minutes="33" orientation="latitude" precision="925" value="19.55">19°33'N</geoCoordinate>
,
<geoCoordinate id="36F3F5BF988EBC129DDC06D19F674156" degrees="98" direction="east" minutes="03" orientation="longitude" precision="925" value="98.05">98°03'E</geoCoordinate>
,
<elevation id="9379E6B5C6BFC2DEDB5E7801525D8B81" metricMagnitude="2" metricUnit="m" metricValue="8.2" unit="m" value="820.0">
<quantity id="64B604E11A9B19F5695ED9B5118EE164" metricMagnitude="2" metricUnit="m" metricValue="8.2" unit="m" value="820.0">820 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="126DC79F336B80CFF23F50A069647F16" value="1988-03-20">20 Mar. 1988</collectingDate>
, coll.
<collectorName id="01F8F404365C78715DB00398D2030043">K. Auffenberg</collectorName>
(3 shells). HA (locality code: 2018/41) from Hsihseng centre E ca.
<quantity id="0201036D554A8160E085FD32E45D0EBD" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
, right side of rd. +
<quantity id="30B3F8EB3A1DE8BC589F79E7BCE3B157" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation id="C8D68317FF97BF4478490525E2018B0C" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
on unpaved rd., limestone hill, Shan State,
<collectingCountry id="BC6B678B2006CC5DE311C5ADF3EB3058" name="Myanmar">Myanmar</collectingCountry>
,
<geoCoordinate id="4A5BDB82A7030A23D006A8DF0AA8A88C" degrees="20" direction="north" minutes="8.002" orientation="latitude" precision="1" value="20.133368">20°8.002'N</geoCoordinate>
,
<geoCoordinate id="602419E9CD49922A0952710AEE2BE04E" degrees="97" direction="east" minutes="18.024" orientation="longitude" precision="1" value="97.3004">97°18.024'E</geoCoordinate>
,
<elevation id="1948D21AFF0076A51C369601E8E7B839" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<quantity id="DBBB76C923967B3A0BEB2E96DF2B9E1E" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="46D62304E480D24AE8079DA3341C8615" value="2018-10-07">7 Oct. 2018</collectingDate>
, coll.
<collectorName id="1F1DB4BCC024C9A8EFFA6167CE5C5E0B">
A.
<collectorName id="EA91A45E261E25F1E694DDBF6DB64A37">Hunyadi, K</collectorName>
</collectorName>
.
<collectorName id="232DD393234417BFC5159C2501E07549">Okubo, J.U</collectorName>
. Otani (1 shell; Fig.
<figureCitation id="AA04120C5FB10ABF8B0811BCD5E4AF93" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11A</figureCitation>
). CUMZ 7431 from
<collectorName id="FD03AEC59CCF327AD41BC65978822100">Doi Chiang Dao</collectorName>
,
<collectingRegion id="41AFEF3CF16E302AA8AC74BE2AD4E152" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="184531B1F0C53B25B91B2C945A04D6E2" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="B30D7A89CB2D48DBDEB01CC35D6EB4D6" degrees="19" direction="north" minutes="14" orientation="latitude" precision="1" seconds="26.7" value="19.24075">19°14'26.7&quot;N</geoCoordinate>
,
<geoCoordinate id="D0B118DD5787612964117C58068C4EBF" degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="49.8" value="98.8805">98°52'49.8&quot;E</geoCoordinate>
,
<collectingDate id="5FDE7D364DD2DC271C687865A57C8A30" value="1997-05-24" valueMax="1997-05-28" valueMin="1997-05-24">24-28 May 1997</collectingDate>
, coll.
<collectorName id="7E722AF78F432294112790A9E25BABD5">
S.
<collectorName id="C753BC097913E3AA2230D96700F1D9E2">Panha, J.B</collectorName>
</collectorName>
.
<collectorName id="EBDBCD2173FC7E2123EFE4076A55DE58">Burch, P</collectorName>
. Dumrongrojwattana (4 shells; Fig.
<figureCitation id="C91CAB53FD56B835F63D4B9F04BB5ECA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10B, C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4178D822095FD037ED5E0316E5E14824" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="89B165F9B01C6CA51500616E787FA32E" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="CE06A763A9E389D583220E06CBCFA608" pageId="0" pageNumber="1">
Shell small (SH up to 3.6 mm, SW up to 3.5 mm), conical. Spire ca.
<normalizedToken id="DE837514A8FA51B223F60745F698102D" originalValue="½">1/2</normalizedToken>
of shell height. R1 with regular, strong ribs; R2 as long as R3; R2 with 36-43 ribs. R3 with either sharp and narrow or blunt and more elongated swelling in the middle of R3 or near aperture. Aperture round. Inner peristome blunt, slightly protruding. Outer peristome expanded (most conspicuously at the upper palatal position and near the umbilicus), reflected. Umbilicus partly or entirely covered by expanded outer peristome. Operculum multispiral.
</paragraph>
</subSubSection>
<subSubSection id="FFD3B8C0CA0644EE01900E42F1F68530" pageId="0" pageNumber="1" type="description">
<paragraph id="522A570EBFD6BEC8BF17F1CA2C500166" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="5E3271D7261645D32A62938B1BF79FFE" pageId="0" pageNumber="1">
Shell small (SH up to 3.6 mm, SW up to 3.5 mm), conical, solid, semitransparent, yellowish to pale orange. Shell outline round in apical view, spire ca.
<normalizedToken id="ED4F655C9CF9ADE374FAC723E13D9263" originalValue="½">1/2</normalizedToken>
of shell height. Whorls approaching 5, last whorl large. Protoconch low, ca. two whorls, glossy and smooth. R1 ca. 2 whorls, with regular, strong ribs; with ca. 21 ribs in
<normalizedToken id="0F102719821946312CEAB0EE7BDFB85B" originalValue="¼">1/4</normalizedToken>
whorl adjoining R2, ribs sharper near suture and flatter near umbilicus, most prominent when approaching R2. Boundary between R1 and R2 distinct as R2 contains more close-set ribs than R1; R2 with 36-43 ribs that are curved towards aperture; R2 ca.
<normalizedToken id="A297425C2367A063905779A9B11CF3FA" originalValue="¼">1/4</normalizedToken>
whorl, as long as R3. Boundary between R2 and R3 distinct due to shallow constriction; R3 smooth with indistinct growth lines; with either sharp and narrow or blunt and more elongated swelling in the middle of R3 or near aperture. Aperture round; slightly oblique to shell axis. Peristome double with prominent outer peristome. Inner peristome blunt, slightly protruding. Outer peristome strongly thickened, expanded, reflected, most conspicuously at the upper palatal position and near the umbilicus, multilayered (visible in lateral view). Umbilicus is partly or entirely closed by the reflected outer peristome. Operculum thin; multispiral ridges not significantly elevated.
</paragraph>
</subSubSection>
<subSubSection id="8128085531113F2A759CB62AC8EA2D64" pageId="0" pageNumber="1" type="etymology">
<paragraph id="9259190A51DE701C52282DCC7C79A6BC" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="E35B4A62B38B5AD74BFFDC609DDA8CE4" pageId="0" pageNumber="1">In honor of Prof. John B. Burch, a prominent American malacologist, who, together with Prof. S. Panha, extensively studied the taxonomy and systematics of micro land snails of Thailand.</paragraph>
</subSubSection>
<subSubSection id="99657A248D3F5A04CC8B80D8F9FCF051" pageId="0" pageNumber="1" type="distribution">
<paragraph id="D28D90B5E284086C35053CEF22123A7B" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="B642CC87714B4B08B0B9C07BD65AD6D3" pageId="0" pageNumber="1">The new species is known from the type locality, Tham Phaya Nakharat, Khon Kaen Province, Northeastern Thailand. The other localities are Doi Chiang Dao, Chiang Mai Province and Ban Soppong, Mae Hong Son Province, Northern Thailand, and also from Shan State, Myanmar.</paragraph>
</subSubSection>
<subSubSection id="845939303B5B051457B23C61C51E6286" pageId="0" pageNumber="1" type="remarks">
<paragraph id="F26F2CF54F4516F9FAAC9A9B3FFDF778" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="AB8E76DE3300CA317955FC05EBE87C43" pageId="0" pageNumber="1">
Differs from
<taxonomicName id="7995B66BD51075D243D96B9CED684F0F" lsidName="D. vestitus" pageId="0" pageNumber="1" rank="species" species="vestitus">
<emphasis id="92415E8C98CD3573D43827C8DA77A022" italics="true" pageId="0" pageNumber="1">D. vestitus</emphasis>
</taxonomicName>
(Blanford, 1862) from Moditoung [Modi Taung, Arakan Hills] (Fig.
<figureCitation id="1E98E1F04FB8D62D97690B88B95EACD1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11B</figureCitation>
), and its subspecies
<taxonomicName id="15A48B9B7F831E02AC399C1F37973C03" lsidName="D. v. subsp. akyabensis" pageId="0" pageNumber="1" rank="subSpecies" species="v." subSpecies="akyabensis">
<emphasis id="7957A4B97E5D6EE3A5F91A95C9D32995" italics="true" pageId="0" pageNumber="1">D. v. akyabensis</emphasis>
</taxonomicName>
(Godwin-Austen, 1914) from Baumi, Akyab [Sittwe] (Fig.
<figureCitation id="8CF5BAE1A4BFADB28DD753EC172529B6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11C</figureCitation>
), both from Rhakhine State, Myanmar, by having a smaller shell with a higher spire, a longer last whorl and a less distinct and a narrower constriction between R2 and R3. Differs from
<taxonomicName id="EAD574064AC8DFBE336FB59CB274CABA" lsidName="D. conicus" pageId="0" pageNumber="1" rank="species" species="conicus">
<emphasis id="F44AC8BC66BC34B7A0BD507C2758A8B4" italics="true" pageId="0" pageNumber="1">D. conicus</emphasis>
</taxonomicName>
(Godwin-Austen, 1871) from Samiamri, E of the Kopili R., Assam State, India (Fig.
<figureCitation id="9859B127717C748FCDBC56B40505D3CF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11D</figureCitation>
) in having a longer last whorl, more close-set ribs on R2, and the inner and outer peristomes are narrower and less reflected. Differs from
<taxonomicName id="188A175908F50F98979082449DC6BDE8" lsidName="D. imitator" pageId="0" pageNumber="1" rank="species" species="imitator">
<emphasis id="E514E4DC268B5CF1050315F84C18BDC3" italics="true" pageId="0" pageNumber="1">D. imitator</emphasis>
</taxonomicName>
<normalizedToken id="F353FB15D7F197A2081B6C0DA5E071C9" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi in
<bibRefCitation id="2C40DF6805757D5EA9E6A82D48320B38" DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken id="32B0678D838892706BE9DDFDB0944352" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al., 2017
</bibRefCitation>
from Guangxi, China in having a less smooth R1, a much thicker peristome, a longer R2, a shallower constriction between R2 and R3, the last whorl less oblique compared to the columellar axis and a less triangular aperture (
<bibRefCitation id="4A15FAB1FB7862E6E9C263CDB7886D81" DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken id="66BEDB0470F82EF5E1A630BD4621D54C" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
).
</paragraph>
<caption id="C6DB1A0ABF6355AFF53B29C2EADC0DEB" doi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph id="27BC02AEA5096B5ADCA7332E740FA9DD" pageId="0" pageNumber="1">
<emphasis id="FA1B2F32C55BFB2C067FE2698FF4E51C" bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
<emphasis id="5352FA38D23FE4DD4931AA9B89B18720" bold="true" pageId="0" pageNumber="1">A.</emphasis>
<taxonomicName id="B1B246C4732EF11162F77820F0932F89" authorityName="Jirapatrasilp &amp; Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax burchi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burchi">
<emphasis id="84F28EB19DE52F3886E9C1B50FAB626A" italics="true" pageId="0" pageNumber="1">Dicharax burchi</emphasis>
</taxonomicName>
sp. nov., specimen HA (locality code: 2018/41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar;
<emphasis id="02540D9F0A6449633B2BB2BF74EEFA6E" bold="true" pageId="0" pageNumber="1">B.</emphasis>
<taxonomicName id="C682E6E89714A57E1823A1A4330232A8" baseAuthorityName="W.T.Blanford" baseAuthorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax vestitus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vestitus">
<emphasis id="FA45104FAB5DB32A9C17BA3C83572C4B" italics="true" pageId="0" pageNumber="1">Dicharax vestitus</emphasis>
</taxonomicName>
, holotype NHMUK 1906.4.4.53;
<emphasis id="672F44EAAFA34AB8AD8F317AEC9A3CFB" bold="true" pageId="0" pageNumber="1">C.</emphasis>
<taxonomicName id="9A084FFB0027F997B5074508EBC38D83" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax vestitus subsp. akyabensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="vestitus" subSpecies="akyabensis">
<emphasis id="6588E3CEEA792C140C6F7C73191D5D79" italics="true" pageId="0" pageNumber="1">Dicharax vestitus akyabensis</emphasis>
</taxonomicName>
, syntype NHMUK 1888.12.4.251-252;
<emphasis id="04263987FBE985BEDAD5BBF03AEC06D8" bold="true" pageId="0" pageNumber="1">D.</emphasis>
<taxonomicName id="45415B6FAD0F70090F39D345EE24A817" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1871" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax conicus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="conicus">
<emphasis id="B73F87E648F5C0837D6861564A1AC5D0" italics="true" pageId="0" pageNumber="1">Dicharax conicus</emphasis>
</taxonomicName>
, syntype NHMUK 1903.7.1.2674. Photo: NHM (
<emphasis id="5276B686F983945FE415D133F2CAC698" bold="true" pageId="0" pageNumber="1">B-D</emphasis>
).
</paragraph>
</caption>
<paragraph id="43FED7D67BF3F737622B69B575E8C2C6" pageId="0" pageNumber="1">
This species exhibits the variation of R1 sculpture from nearly smooth (Fig.
<figureCitation id="85B8C44EF8D117FDC7A2087400BE9921" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10A</figureCitation>
) to distinctly ribbed (Fig.
<figureCitation id="AC8AAC3D105821CA5BFDF75B4556CBD1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10C, D</figureCitation>
) and R3 swelling from sharp (Fig.
<figureCitation id="1421DBC3471CEE4B072C972CE0B4E532" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10B, C</figureCitation>
) to blunter and longer by having the anterior slope of swelling steeper than posterior slope (Fig.
<figureCitation id="B19FF9D91A6F5865D49757E1076D6AE0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10D</figureCitation>
). In addition, the umbilicus is open, or covered by the reflected outer peristome. The latter trait even shows variation within the same population (maybe also depends on the age of the examined specimens).
</paragraph>
</subSubSection>
</treatment>
</document>
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