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<document id="FA5C5D6D110A08DB2735F12B10A339D4" ID-DOI="10.3897/zitteliana.98.e138605" ID-ZooBank="9FB6F185-9EA1-48EB-A5C9-632BD52A8B6F" ID-publisher-id="138605" URI-arpha="88AF8F00-FD88-5C61-809C-0C929B9B2F5A" URI-zoobank="http://zoobank.org/9FB6F185-9EA1-48EB-A5C9-632BD52A8B6F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1733930160955" checkinUser="pensoft" docAuthor="Gründel, Joachim &amp; Nützel, Alexander" docDate="2024" docId="7E5A4F7ACD125E71A551A40896937D10" docLanguage="en" docName="Zitteliana 98: 19-72" docOrigin="Zitteliana 98" docSource="https://zitteliana.pensoft.net/article/138605/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Columbellaria corallina" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="72" masterDocId="88AF8F00FD885C61809C0C929B9B2F5A" masterDocTitle="Late Jurassic (Upper Kimmeridgian) gastropods from Saal near Kelheim (Germany, Bavaria)" masterLastPageNumber="72" masterPageNumber="19" pageNumber="19" updateTime="1733940029114" updateUser="admin">
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<mods:title id="602B109671A8DFD8DCAF5F6975F40C2F">Late Jurassic (Upper Kimmeridgian) gastropods from Saal near Kelheim (Germany, Bavaria)</mods:title>
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<mods:namePart id="B20FA94D0F6FAB9E0C94B7818F4D1F8B">Gründel, Joachim</mods:namePart>
<mods:affiliation id="4004CC8CF44192C1069608EE112D8751">Institut für Geowissenschaften, Fachrichtung Paläontologie, Freie Universität Berlin, Malteserstraße 74 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="C562BEE45BA527FBE01CDC5FC029A560">Nützel, Alexander</mods:namePart>
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<mods:affiliation id="503312B5EDE5979D7F0CB97BFB7080D8">SNSB-Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Str. 10, 80333 München, Germany &amp; Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universität München, Richard-Wagner-Str. 10, 80333 München, Germany</mods:affiliation>
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<mods:title id="025D6F0C8AEFCF1FE65B4F1412E27D06">Zitteliana</mods:title>
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<mods:date id="6EA423A2C9F276CD3CB55414A8C9FEB8">2024</mods:date>
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<mods:number id="53514AD718616E8DE742783814B6D207">98</mods:number>
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<treatment id="7E5A4F7ACD125E71A551A40896937D10" ID-arpha="7E5A4F7A-CD12-5E71-A551-A40896937D10" LSID="urn:lsid:plazi:treatment:7E5A4F7ACD125E71A551A40896937D10" httpUri="http://treatment.plazi.org/id/7E5A4F7ACD125E71A551A40896937D10" scope_family="Colombellinidae" scope_subClass="Caenogastropoda" scope_superOrder="Latrogastropoda">
<subSubSection id="FF15614F635438B2CCA3D59BE2CAB514" type="nomenclature">
<paragraph id="54BF155BA5EA0E5716AD5F65E655C4D5">
<taxonomicName id="0C02D1E92107A56ADF3F9D0B7D28638E" ID-CoL="CD49J" authority="(Quenstedt, 1852)" baseAuthorityName="Quenstedt" baseAuthorityYear="1852" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="corallina" subClass="Caenogastropoda">
<emphasis id="BB9235834826AAC021979F0D7D50AF13" italics="true">Columbellaria corallina</emphasis>
(
<bibRefCitation id="6C500C1C5B46E36704954215C782B11E" author="Quenstedt FA" refId="B148" refString="Quenstedt FA (1852) Handbuch der Petrefaktenkunde. 1. Auflage, 792 pp., Tubingen (Laupp)." year="1852">Quenstedt, 1852</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="52EC7540F234349C8E56B5EA2839B6B6" type="description">
<paragraph id="5A244ED42B2449E15660703119326D08">
<figureCitation id="A0E713C6C908556B3D56A8B04D830F12" captionStart="Plate 10" captionStartId="F10" captionText="Plate 10. (1) Neuburgensia angulata sp. nov., holotype, SNSB-BSPG 2016 XXI 1849, collection Lang, lateral view, height 24 mm. (23) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1850, collection Lang, (2) earliest preserved whorls in lateral view, height 1 mm, (3) lateral view, height 6 mm. (4) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1851, collection Lang, lateral view, height 10 mm. (5) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1852, collection Schäfer ¸ basal view, width 6 mm. (6) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1853, collection Lang, oblique apical, width 6.5 mm. (79) Neuburgensia rara sp. nov., holotype, SNSB-BSPG 2016 XXI 1856, (7) lateral view, height 30 mm, (8) early whorls in lateral view, height ca. 7 mm, (9) middle whorls in lateral view, height 12 mm. (10, 11) Purpuroidea lapierrea (Buvignier, 1843), SNSB-BSPG 2016 XXI 1857, lateral and oblique views, height 30 mm. (12, 13) Columbellaria corallina (Quenstedt, 1852), SNSB-BSPG 2016 XXI 1859, (12) apex in lateral view, height 10 mm, (13) lateral view, height 19 mm. (14, 15) Columbellaria corallina (Quenstedt, 1852), SNSB 1860, lateral and slightly oblique lateral views, height 21 mm. (16) Diarthema aspera sp. nov., paratype, SNSB-BSPG 2016 XXI 1862, lateral view, height 44 mm." figureDoi="10.3897/zitteliana.98.e138605.plate10" httpUri="https://binary.pensoft.net/fig/1196366">Plate 10: figs 1215</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3EFD65465CBBA0F56B57222CB20814ED" type="reference_group">
<paragraph id="D2A753722D2010C7EB3AA7F2A5801F21">
<treatmentCitationGroup id="F989EC31CB7C0D17BE6E3E8AE81EF7FE">
2017
<treatmentCitation id="01424923FD28FE3562645169FFDAFE7F">
<taxonomicName id="3B63586797547FB93102A85DE39D126B" ID-CoL="CD49J" authority="(Quenstedt, 1852)" baseAuthorityName="Quenstedt" baseAuthorityYear="1852" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" isUncertain="true" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="corallina">
<emphasis id="BE19334C530B33C6859228C9D9E6F882" italics="true">Columbellaria</emphasis>
cf.
<emphasis id="0236EB54AD6D3AE4BDDE5358C43E6C0F" italics="true">corallina</emphasis>
(
<bibRefCitation id="570B2D5859E63FEBCAD9517C0025A65A" author="Quenstedt FA" refId="B148" refString="Quenstedt FA (1852) Handbuch der Petrefaktenkunde. 1. Auflage, 792 pp., Tubingen (Laupp)." year="1852">Quenstedt, 1852</bibRefCitation>
)
</taxonomicName>
</treatmentCitation>
Werner, Nützel and Nose: 32, fig. 3 A C.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B1EF3768674DD70C4676C123276F2D1B">
<treatmentCitationGroup id="7E48C641E1F77789653884EB2CF085EE">
2019
<treatmentCitation id="F23BCBD2BE8DA5F356A966446F6D0EDF">
<taxonomicName id="B80D6FD55CD2BF6387C104BF8D0537CD" ID-CoL="CD49J" authority="(Quenstedt, 1852)" baseAuthorityName="Quenstedt" baseAuthorityYear="1852" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="corallina">
<emphasis id="591B8368ED5804105914FFCED05B394A" italics="true">Columbellaria corallina</emphasis>
(
<bibRefCitation id="C435A06EE4DD58869AFF5EE9C8147478" author="Quenstedt FA" refId="B148" refString="Quenstedt FA (1852) Handbuch der Petrefaktenkunde. 1. Auflage, 792 pp., Tubingen (Laupp)." year="1852">Quenstedt, 1852</bibRefCitation>
)
</taxonomicName>
</treatmentCitation>
Gründel, Keupp and Lang: 133, pl. 9, figs 1117 (here more chresonymy and synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2AF851D598567ABD9465C6769F423961">
<treatmentCitationGroup id="67343A57FA40134C0F2A3B91F37183D2">
<treatmentCitation id="3EC30107E1C1674B4DB59467C7DD88D2">
2019
<taxonomicName id="71FB6F208C6A9CE16F1B2723F5860297" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="undefined-1">
<emphasis id="60B012A42D4C9BBA7045F71068DC5AF4" italics="true">Columbellaria</emphasis>
sp. 1
</taxonomicName>
Gründel, Keupp and Lang: 136, pl. 9, fig. 18, pl. 10, fig. 1.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENFFK" type="material">
<paragraph id="F73D37A015797EB0BC73302AEE6E1CA7">
<heading id="8BD8B825C9AEE87FBFA25B8C2ED0C35F" reason="title">Material.</heading>
</paragraph>
<paragraph id="66520AB4063AE509494501746DF73F9B">
<specimenCount id="21E90102654A3D4ECB7695BACEA47214" count="3" type="generic">3 specimens</specimenCount>
,
<specimenCode id="DBE208972FA92D98E91FAD1E1884AA9E">SNSB-BSPG 2016 XXI 1859</specimenCode>
1861.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESFFK" type="description">
<paragraph id="289DC50D27B9BD7A6826C98F84E58201">
<heading id="A807D907F7AC7D15DA0C804F0EAFED6E" reason="title">Description.</heading>
</paragraph>
<paragraph id="708E6027C6BE20484149A48FA43FE32F">
Shell egg-shaped, higher than wide, with distinctly elevated, slender, gradate spire, consisting of more than 5 whorls; illustrated specimen (Plate
<figureCitation id="A23555FCF952B7BC39D35766EF38333E" captionStart="Plate 10" captionStartId="F10" captionText="Plate 10. (1) Neuburgensia angulata sp. nov., holotype, SNSB-BSPG 2016 XXI 1849, collection Lang, lateral view, height 24 mm. (23) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1850, collection Lang, (2) earliest preserved whorls in lateral view, height 1 mm, (3) lateral view, height 6 mm. (4) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1851, collection Lang, lateral view, height 10 mm. (5) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1852, collection Schäfer ¸ basal view, width 6 mm. (6) Neuburgensia angulata sp. nov., paratype, SNSB-BSPG 2016 XXI 1853, collection Lang, oblique apical, width 6.5 mm. (79) Neuburgensia rara sp. nov., holotype, SNSB-BSPG 2016 XXI 1856, (7) lateral view, height 30 mm, (8) early whorls in lateral view, height ca. 7 mm, (9) middle whorls in lateral view, height 12 mm. (10, 11) Purpuroidea lapierrea (Buvignier, 1843), SNSB-BSPG 2016 XXI 1857, lateral and oblique views, height 30 mm. (12, 13) Columbellaria corallina (Quenstedt, 1852), SNSB-BSPG 2016 XXI 1859, (12) apex in lateral view, height 10 mm, (13) lateral view, height 19 mm. (14, 15) Columbellaria corallina (Quenstedt, 1852), SNSB 1860, lateral and slightly oblique lateral views, height 21 mm. (16) Diarthema aspera sp. nov., paratype, SNSB-BSPG 2016 XXI 1862, lateral view, height 44 mm." figureDoi="10.3897/zitteliana.98.e138605.plate10" httpUri="https://binary.pensoft.net/fig/1196366">10</figureCitation>
: figs 14, 15)
<quantity id="1CE8AFEA5F4C0C76ED27199199044D42" metricMagnitude="-2" metricUnit="m" metricValue="2.1" unit="mm" value="21.0">21 mm</quantity>
high; already first preserved whorls angulated forming edge that later becomes a keel; keel separates oblique ramp and vertical abapical whorl face; ramp ornamented with several very weak spiral cords over several whorls; spiral cords no longer visible on last two spire whorls; instead, formation of subsutural spiral cord covered with small tubercles and a stronger spiral cord, also with nodes, between subsutural cord and keel; whorl face of early teleoconch with axial ribs (approximately 10 on last spire whorl); intersections of axial ribs and spiral cords nodular; all mentioned spiral cords continue onto body whorl without interruption; axial ribs disappear at transition from spire whorls to body whorl and keel is weakened into a spiral cord of equal strength as in other spiral cords; transition from convex whorl face to strongly convex base evenly rounded; body whorl covered with approximately 15 nodular spiral cords of approximately same strength; spiral cords on whorl face slightly more distant from each other than those on base; aperture with widened outer lip and spine-like protrusions where spiral cords are situated; aperture elongated with anterior and posterior siphonal canals (see
<bibRefCitation id="864E401B1FE5F8F3DB3D56BBF2A1503B" author="Hagele" firstAuthor="Hagele" journalOrPublisher="Fossilien, Sonderband" pagination="1-144" refId="B91" refString="Hägele G (1997) Juraschnecken. Fossilien, Sonderband 11: 1144." title="Juraschnecken." volume="11" year="1997">Hägele 1997</bibRefCitation>
;
<bibRefCitation id="C79F66C11CCB58E1AF0391027957035A" author="Werner" etAl="et al." firstAuthor="Werner" refId="B175" refString="Werner W, Nützel A, Nose M (2017) Fossilien aus dem Oberjura von Saal Die Sammlung Jürgen Sylla. Freunde der Bayerischen Staatssammlung für Paläontologie und Historische Geologie München e. V., Jahresbericht 2016 und Mitteilungen, 2837." year="2017">Werner et al. 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEGFK" type="remarks">
<paragraph id="BD0BBE38FAD6D32ACA130B28F62739F1">
<heading id="FA72F5FCF25CCC956A19BF4FCDD0EFEB" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="8E1C527B0D8A18BF646AD65CE0A69192">
The spire of the present specimens combines characteristics of
<taxonomicName id="AC1E4BF06DFC9F5537084FC30D389746" authority="sensu Grundel et al. (2019)" authorityName="Grundel" authorityYear="2019" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="corallina">
<emphasis id="D2AA4EEA63EB702E993FD5F3E0570FD2" italics="true">Columbellaria corallina</emphasis>
sensu
<bibRefCitation id="9B4C8FB067307F8B43F88A8E66E19EEB" author="Gründel J &amp; Keupp H &amp; Lang F" etAl="et al." firstAuthor="Grundel" journalOrPublisher="Zitteliana" pagination="97-142" refId="B86" refString="Gründel J, Keupp H, Lang F (2019) Arten der Unterklasse Caenogastropoda aus der Korallenfazies des oberen Kimmeridgiums (Ober-Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland). Zitteliana 93: 97142." title="Arten der Unterklasse Caenogastropoda aus der Korallenfazies des oberen Kimmeridgiums (Ober-Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland)." volume="93" year="2019">Gründel et al. (2019)</bibRefCitation>
</taxonomicName>
(the last whorl is approximately
<quantity id="06C1E53644992621588E3C5C11F2E077" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
wide; distinct spiral cord between adapical suture and keel) with that of
<taxonomicName id="990F50569FD106FC37F560DF9AF1836D" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="undefined-1">
<emphasis id="12788D3A43F02AC3D87748D444AB80AE" italics="true">Columbellaria</emphasis>
sp. 1
</taxonomicName>
sensu
<bibRefCitation id="FA87CAA5AA4C1A7C07DA6B222EA6B680" author="Gründel J &amp; Keupp H &amp; Lang F" etAl="et al." firstAuthor="Grundel" journalOrPublisher="Zitteliana" pagination="97-142" refId="B86" refString="Gründel J, Keupp H, Lang F (2019) Arten der Unterklasse Caenogastropoda aus der Korallenfazies des oberen Kimmeridgiums (Ober-Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland). Zitteliana 93: 97142." title="Arten der Unterklasse Caenogastropoda aus der Korallenfazies des oberen Kimmeridgiums (Ober-Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland)." volume="93" year="2019">Gründel et al. (2019)</bibRefCitation>
(strong keel on whorls until end of spire, stronger nodes on keel). It is possible that all forms described
<bibRefCitation id="BA04CC181AADB579865648057235B693" author="Gründel J &amp; Keupp H &amp; Lang F" etAl="et al." firstAuthor="Grundel" journalOrPublisher="Zitteliana" pagination="97-142" refId="B86" refString="Gründel J, Keupp H, Lang F (2019) Arten der Unterklasse Caenogastropoda aus der Korallenfazies des oberen Kimmeridgiums (Ober-Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland). Zitteliana 93: 97142." title="Arten der Unterklasse Caenogastropoda aus der Korallenfazies des oberen Kimmeridgiums (Ober-Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland)." volume="93" year="2019">Gründel et al. (2019)</bibRefCitation>
and described here are variants of a single species. Due to the lack of more extensive and better-preserved material, these forms are summarized herein under the name
<taxonomicName id="C91612BD8977EC167F4A39A49C3F77ED" authority="(Quenstedt, 1852)" baseAuthorityName="Quenstedt" baseAuthorityYear="1852" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="corallina">
<emphasis id="0DD21E72A8B478B2D68D054BEEFBE5C1" italics="true">Columbellaria corallina</emphasis>
(
<bibRefCitation id="F3089ECA656F89592AD661B5E3DF409F" author="Quenstedt FA" refId="B148" refString="Quenstedt FA (1852) Handbuch der Petrefaktenkunde. 1. Auflage, 792 pp., Tubingen (Laupp)." year="1852">Quenstedt, 1852</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="3BE30F17D53770B4708041FB04CBC606">
<taxonomicName id="47A70B911822F18613964D4D6B50857D" authority="Grundel et al. (2022)" authorityName="Grundel" authorityYear="2022" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="rara">
<emphasis id="7D27AF76F0F4772AE47728FB313EA3D8" italics="true">Columbellaria rara</emphasis>
<bibRefCitation id="3B20D299858B651943EF81E208E72FEF" author="Gründel J &amp; Hostettler B &amp; Menkveld-Gfeller U" etAl="et al." firstAuthor="Grundel" journalOrPublisher="Revue de Paléobiologie" pagination="29-84" refId="B83" refString="Gründel J, Hostettler B, Menkveld-Gfeller U (2022) Die Gastropoden aus der Korallenrifffazies der St-Ursanne-Formation (mittleres Oxfordien) des Schweizer Jura. 3. Die Unterklasse Caenogastropoda Cox, 1960. Revue de Paléobiologie 40: 2984." title="Die Gastropoden aus der Korallenrifffazies der St-Ursanne-Formation (mittleres Oxfordien) des Schweizer Jura. 3. Die Unterklasse Caenogastropoda Cox, 1960." volume="40" year="2022">Gründel et al. (2022)</bibRefCitation>
</taxonomicName>
has significantly fewer spiral cords on the body whorl and these are more bulging.
<taxonomicName id="DADCDB3DCBC6F28DB0D979EF4064F808" authority="Gemmellaro, 1870" authorityName="Gemmellaro" authorityYear="1870" class="Gastropoda" family="Colombellinidae" genus="Zittelia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="picteti">
<emphasis id="7B033D7AC0707C14FEB692D4DD773EFE" italics="true">Zittelia picteti</emphasis>
Gemmellaro, 1870
</taxonomicName>
has a wider and more convex body whorl, the spiral cords are at least as wide as the spiral furrows between them and the spire whorls are not keeled.
<taxonomicName id="63C6FE4EDA17BEAECED5DC43F27CEBAB" authority="Guirand &amp; Ogerien, 1865" authorityName="Guirand &amp; Ogerien" authorityYear="1865" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="victoria">
<emphasis id="CF39008D2505CE6F750680D8D0CCADC7" italics="true">Columbellaria victoria</emphasis>
<bibRefCitation id="61A2795B7F862AFA47CA772C2CE4CFF2" author="Guirand M &amp; Ogérien LF" journalOrPublisher="Travaux de la Société dÉmulation du Jura, für" pagination="369-394" refId="B88" refString="Guirand M, Ogerien LF (1865) Quelques fossiles nouveaux du Corallien du Jura. Travaux de la Societe d'Emulation du Jura, fur 1865: 369 - 394." title="Quelques fossiles nouveaux du Corallien du Jura." volume="1865" year="1865">Guirand &amp; Ogérien, 1865</bibRefCitation>
</taxonomicName>
and sensu Loriol in
<bibRefCitation id="D5B2379ABD5ADED4608BB389375C4B09" author="Loriol P de &amp; Bourgeat AE" refId="B111" refString="Loriol P de, Bourgeat AE (18861888) Études sur les mollusques des couches coralligènes de Valfin (Jura). Mémoires de la Société Paléontologique Suisse 1315: 1369.">Loriol and Bourgeat (18861888</bibRefCitation>
) has spiral cords of equal strength on the body whorl, which are at least as wide as the spiral furrows between them. Its body whorl is more convex and it lacks a subsutural row of small nodes.
<taxonomicName id="A64E0C222AF9DEC4F33875A5D44DD5DF" authority="Zittel, 1873" authorityName="Zittel" authorityYear="1873" class="Gastropoda" family="Colombellinidae" genus="Columbellaria" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="denticulata">
<emphasis id="02C5B85F728B0D34F220EC6F10E88195" italics="true">Columbellaria denticulata</emphasis>
<bibRefCitation id="D1623A109EDE1BE8FC77A9AC277F887B" author="Zittel K" refId="B183" refString="Zittel K (1873) Die Gastropoden der Stramberger Schichten. Palaeontographica, Supplement 2 / 1: 194 - 373." year="1873">Zittel, 1873</bibRefCitation>
</taxonomicName>
has more nodes on the last spire whorl and lacks subsutural row of small nodes is missing there; the outer lip of the body whorl lacks furrows.
</paragraph>
</subSubSection>
</treatment>
</document>