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<document id="84C6678C9D1A6DD084706A10AF8B9326" ID-DOI="10.5852/ejt.2024.958.2659" ID-ISSN="2118-9773" ID-Zenodo-Dep="13758948" ID-ZooBank="6FFE3969-D8EF-4082-9107-F98187116C28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726227020474" checkinUser="plazi" docAuthor="Colombo, Wesley D. &amp; Azevedo, Celso O." docDate="2024" docId="7E5987A7FFA6B158E530FDE0F5A1FDC1" docLanguage="en" docName="EJT.2024.958.1-47.pdf" docOrigin="European Journal of Taxonomy 958 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2659/12299" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Megaprosternum nuaulu Colombo &amp; Azevedo, 2024, sp. nov." docType="treatment" docUuid="9A0F3D1D-5268-422F-AE26-A6363740DE37" docUuidSource="ZooBank" docVersion="2" lastPageNumber="33" masterDocId="8260FFDFFFB8B178E761FFABF44DFFE4" masterDocTitle="Revision of the strongly flattened Megaprosternum Azevedo (Hymenoptera, Bethylidae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="31" updateTime="1726227497903" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A56FE2B1E88A6B6771EB9ABBD00A7B3D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0043CDD0DBF0FFAE8B0721B2B4E9193">
<mods:title id="D9610A55834ABAD3944002F29AFE7E0D">Revision of the strongly flattened Megaprosternum Azevedo (Hymenoptera, Bethylidae)</mods:title>
</mods:titleInfo>
<mods:name id="64A4E100EAE6959B121AC14EE72A4AAF" type="personal">
<mods:role id="0BA3B3CC3B5FFD3B7229B57BB2B1BE8B">
<mods:roleTerm id="CD80D5B0FABEE8588FF67CA192F6719C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BAAE1AB0BF96EB773E26BD2D258881A3">Colombo, Wesley D.</mods:namePart>
<mods:nameIdentifier id="6F5F9F94E6EC2CF9C8B513043DD1FA35" type="LSID">B9D7FDE3-BDD8-4714-A53F-640A0994E5F6</mods:nameIdentifier>
<mods:affiliation id="6E5F497BEF5DFD43A7AA43A7BC084885">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Av. Fernando Ferrari 514, Goiabeiras, 29.075 - 910, Vitória, ES, Brazil.</mods:affiliation>
<mods:nameIdentifier id="D7BF5E8A956231B7F9449EBABDFBE1AB" type="email">wesleycolombo@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="47EEB15B9FF3D36C7FD0CBA5A180986B" type="personal">
<mods:role id="E511B8F047DE2CB36CACE942BF7DFDBC">
<mods:roleTerm id="C1BB97424786C2DF3728D0C2DDF78D8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C55980BEEBEF53DF1B468146168D579C">Azevedo, Celso O.</mods:namePart>
<mods:nameIdentifier id="3BF22F41F34B819EA19035D156434A3A" type="LSID">CD98B489-7611-4A33-AF53-3016B9DA1617</mods:nameIdentifier>
<mods:affiliation id="08120B63A05F6F3DBE806B92AA9F4377">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Av. Fernando Ferrari 514, Goiabeiras, 29.075 - 910, Vitória, ES, Brazil.</mods:affiliation>
<mods:nameIdentifier id="32FA7F262D853692AD2C2AB047589301" type="email">bethylidae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4070755EB516C689A63F380A8F32A59B">text</mods:typeOfResource>
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<mods:title id="29E641084395AE9E7B0D342D6A7CB3EC">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D94C4CD4F5182040664330663097D521">
<mods:date id="E78668817EEABF259A7BBB78C59E4A97">2024</mods:date>
<mods:detail id="8F984FFA8B27DF37A8E0DC71B36FBD42" type="pubDate">
<mods:number id="FF27FAEF336FC4AA78E2D83710B6EED9">2024-09-13</mods:number>
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<mods:detail id="6DF0676C7FF73E434A7361025453AAFB" type="volume">
<mods:number id="D13EF242439AD3704997C00FD0FF89EA">958</mods:number>
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<mods:number id="E79FBA63BA2FA462EC859E87FAE93C90">1</mods:number>
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<mods:start id="FCBE42B04BEE752836A13B7797D71352">1</mods:start>
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<mods:url id="47F6A12EED033BF79969D9433993BC8A">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2659/12299</mods:url>
</mods:location>
<mods:classification id="45778F337CE05222FB14EA03C6CEB84B">journal article</mods:classification>
<mods:identifier id="C5CFA048A5FE373AA760781A0747CA7E" type="DOI">10.5852/ejt.2024.958.2659</mods:identifier>
<mods:identifier id="09F98C5B545170723E2BA22112BA1E28" type="ISSN">2118-9773</mods:identifier>
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</mods:mods>
<treatment id="7E5987A7FFA6B158E530FDE0F5A1FDC1" LSID="urn:lsid:zoobank.org:act:9A0F3D1D-5268-422F-AE26-A6363740DE37" httpUri="http://treatment.plazi.org/id/7E5987A7FFA6B158E530FDE0F5A1FDC1" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="BEEA653AFFA6B166E530FDE0F08EFD6C" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="F64F36B1FFA6B166E530FDE0F7AEFD81" blockId="30.[368,1219,586,682]" box="[593,995,586,613]" pageId="30" pageNumber="31">
<heading id="AD0781DDFFA6B166E530FDE0F7AEFD81" box="[593,995,586,613]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="31F04D32FFA6B166E530FDE0F7CCFD81" authorityName="Colombo &amp; Azevedo" authorityYear="2024" box="[593,897,586,613]" class="Insecta" family="Bethylidae" genus="Megaprosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nuaulu" status="sp. nov.">
<emphasis id="C484EAA3FFA6B166E530FDE0F7CCFD81" bold="true" box="[593,897,586,613]" italics="true" pageId="30" pageNumber="31">Megaprosternum nuaulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFB757D8FFA6B166E4E9FDE0F7AEFD81" box="[904,995,587,613]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="F64F36B1FFA6B166E611FDC6F08EFD6C" blockId="30.[368,1219,586,682]" box="[368,1219,621,648]" pageId="30" pageNumber="31">
<uri id="82613AB3FFA6B166E611FDC6F08EFD6C" box="[368,1219,621,648]" pageId="30" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid id="82560C64FFA6B166E5ECFDC6F08EFD6C" box="[653,1219,621,648]" pageId="30" pageNumber="31">9A0F3D1D-5268-422F-AE26-A6363740DE37</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFFA6B166E590FD3BF70FFD4E" box="[753,834,656,682]" pageId="30" pageNumber="31" type="description">
<paragraph id="F64F36B1FFA6B166E590FD3BF70FFD4E" blockId="30.[368,1219,586,682]" box="[753,834,656,682]" pageId="30" pageNumber="31">
<figureCitation id="6ECB2A34FFA6B166E590FD3BF70FFD4E" box="[753,834,656,682]" captionStart="Fig" captionStartId="31.[189,232,1928,1954]" captionTargetBox="[195,1395,608,1870]" captionTargetId="figure-79@31.[189,1398,604,1880]" captionTargetPageId="31" captionText="Fig. 11. Megaprosternum nuaulu sp. nov., holotype, ♀ (RMNH). A–B. Habitus. A. Lateral view. B. Dorsal view. C. Head, dorsal view. D–E. Mesosoma. D. Dorsal view. E. Ventral view. Scale bars: A–B = 250 μm; C–E = 125 μm." figureDoi="http://doi.org/10.5281/zenodo.13758973" httpUri="https://zenodo.org/record/13758973/files/figure.png" pageId="30" pageNumber="31">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFFA6B166E7DCFD7AF6EFFCD8" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="F64F36B1FFA6B166E7DCFD7AF584FD0F" blockId="30.[189,457,721,747]" box="[189,457,721,747]" pageId="30" pageNumber="31">
<heading id="AD0781DDFFA6B166E7DCFD7AF584FD0F" bold="true" box="[189,457,721,747]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="C484EAA3FFA6B166E7DCFD7AF584FD0F" bold="true" box="[189,457,721,747]" pageId="30" pageNumber="31">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="F64F36B1FFA6B166E7DCFD54F6EFFCD8" blockId="30.[187,1398,767,828]" pageId="30" pageNumber="31">
The females of this species are morphologically similar to those of
<taxonomicName id="31F04D32FFA6B166E4DFFCABF079FCFD" authorityName="Colombo &amp; Azevedo" authorityYear="2024" box="[958,1076,768,793]" class="Insecta" family="Bethylidae" genus="Megaprosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="navatu" status="sp. nov.">
<emphasis id="C484EAA3FFA6B166E4DFFCABF079FCFD" box="[958,1076,768,793]" italics="true" pageId="30" pageNumber="31">M. navatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFB757D8FFA6B166E35AFCABF0DBFCFE" box="[1083,1174,768,794]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, as aforementioned in its section of the differential diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFFA6B166E7DCFCC8F5E2FC29" pageId="30" pageNumber="31" type="etymology">
<paragraph id="F64F36B1FFA6B166E7DCFCC8F509FC99" blockId="30.[189,324,867,893]" box="[189,324,867,893]" pageId="30" pageNumber="31">
<heading id="AD0781DDFFA6B166E7DCFCC8F509FC99" bold="true" box="[189,324,867,893]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="C484EAA3FFA6B166E7DCFCC8F509FC99" bold="true" box="[189,324,867,893]" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="F64F36B1FFA6B166E7DCFC3AF5E2FC29" blockId="30.[188,1398,913,973]" pageId="30" pageNumber="31">
The epithet
<taxonomicName id="31F04D32FFA6B166E627FC3AF5DAFC4F" authorityName="Colombo &amp; Azevedo" authorityYear="2024" box="[326,407,913,939]" class="Insecta" family="Bethylidae" genus="Megaprosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nuaulu">
<emphasis id="C484EAA3FFA6B166E627FC3AF5DAFC4F" box="[326,407,913,939]" italics="true" pageId="30" pageNumber="31">nuaulu</emphasis>
</taxonomicName>
is derived from the Nuaulu, a Melanesian people who live in the jungles of the island of Seram,
<collectingCountry id="8EE77621FFA6B166E654FC18F5E6FC29" box="[309,427,947,973]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFFA6B166E7DCFC5FF075FB66" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="F64F36B1FFA6B166E7DCFC5FF5E7FBEA" blockId="30.[189,426,1012,1038]" box="[189,426,1012,1038]" pageId="30" pageNumber="31">
<heading id="AD0781DDFFA6B166E7DCFC5FF5E7FBEA" bold="true" box="[189,426,1012,1038]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="C484EAA3FFA6B166E7DCFC5FF5E7FBEA" bold="true" box="[189,426,1012,1038]" pageId="30" pageNumber="31">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="F64F36B1FFA6B166E78BFB89F511FBD8" blockId="30.[189,1398,1058,1154]" box="[234,348,1058,1084]" pageId="30" pageNumber="31">
<emphasis id="C484EAA3FFA6B166E78BFB89F511FBD8" bold="true" box="[234,348,1058,1084]" pageId="30" pageNumber="31">
<typeStatus id="294B8813FFA6B166E78BFB89F511FBD8" box="[234,348,1058,1084]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="F64F36B1FFA6B166E7DCFBEEF075FB66" blockId="30.[189,1398,1058,1154]" pageId="30" pageNumber="31">
<materialsCitation id="46983CECFFA6B166E7DCFBEEF075FB66" collectingDate="1997-02-28" collectingDateMax="1997-03-21" collectingDateMin="1997-02-28" collectionCode="RMNH" collectorName="C. v. Achterberg &amp; R. de Vries" country="Indonesia" location="North Ceram" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="8EE77621FFA6B166E7DCFBEEF516FBBB" box="[189,347,1093,1119]" name="Indonesia" pageId="30" pageNumber="31">INDONESIA</collectingCountry>
–
<emphasis id="C484EAA3FFA6B166E614FBEEF651FBBB" bold="true" box="[373,540,1093,1119]" pageId="30" pageNumber="31">
<location id="F32F606AFFA6B166E614FBEEF651FBBB" LSID="urn:lsid:plazi:treatment:7E5987A7FFA6B158E530FDE0F5A1FDC1:F32F606AFFA6B166E614FBEEF651FBBB" box="[373,540,1093,1119]" country="Indonesia" name="North Ceram" pageId="30" pageNumber="31">North Ceram</location>
</emphasis>
•
<specimenCount id="E0F6FD38FFA6B166E555FBEEF606FBBB" box="[564,587,1093,1119]" count="1" pageId="30" pageNumber="31" type="female">♀</specimenCount>
; “N[orth] Ceram,
<locationDeviation id="E7FD2F93FFA6B166E47EFBEEF789FBBB" box="[799,964,1093,1119]" location="Wahai" pageId="30" pageNumber="31">
<quantity id="31089B54FFA6B166E47EFBEEF714FBBB" box="[799,857,1093,1119]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="31" unit="km" value="9.0">9 km</quantity>
E Wahai
</locationDeviation>
, nr PHPA-Q, coastal rainforest,
<date id="824E1071FFA6B166E259FBEEF573FB66" pageId="30" pageNumber="31" value="1997-02-28" valueMax="1997-03-21" valueMin="1997-02-28">
<collectingDate id="920AE999FFA6B166E259FBEEF573FB66" pageId="30" pageNumber="31" value="1997-02-28" valueMax="1997-03-21" valueMin="1997-02-28">28.ii- 21.iii.1997</collectingDate>
</date>
, Mal. trap 7,
<collectorName id="5B055367FFA6B166E6BAFBC3F6C3FB66" box="[475,654,1128,1154]" pageId="30" pageNumber="31">C.v.Achterberg</collectorName>
&amp;
<collectorName id="5B055367FFA6B166E5D2FBC3F77CFB66" box="[691,817,1128,1154]" pageId="30" pageNumber="31">R.de Vries</collectorName>
, RMNH’97”;
<collectionCode id="90E1AE74FFA6B166E4B9FBC3F079FB66" box="[984,1076,1128,1154]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="31" type="Museum">RMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFFA6B158E7DCFB03F574FEA1" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="F64F36B1FFA6B166E7DCFB03F503FB26" blockId="30.[189,334,1192,1218]" box="[189,334,1192,1218]" pageId="30" pageNumber="31">
<heading id="AD0781DDFFA6B166E7DCFB03F503FB26" bold="true" box="[189,334,1192,1218]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="C484EAA3FFA6B166E7DCFB03F503FB26" bold="true" box="[189,334,1192,1218]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="F64F36B1FFA6B166E78BFB7CF508FB15" blockId="30.[189,1398,1239,1334]" box="[234,325,1239,1265]" pageId="30" pageNumber="31">
<emphasis id="C484EAA3FFA6B166E78BFB7CF508FB15" bold="true" box="[234,325,1239,1265]" pageId="30" pageNumber="31">Female</emphasis>
</paragraph>
<paragraph id="F64F36B1FFA6B166E7DCFB51F741FAD2" blockId="30.[189,1398,1239,1334]" pageId="30" pageNumber="31">
<smallCapsWord id="F0A9A06DFFA6B166E7DCFB51F523FAF6" baselines="1293,1293" box="[189,366,1274,1300]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="30" pageNumber="31">MEASUREMENTS</smallCapsWord>
(mm). Body length 2.6; LH 0.50; WH 0.40; WF 0.20; WOT 0.06; surface of median clypeal lobe 0.10; HE 0.21; OOL 0.26; LFW 1.36.
</paragraph>
<paragraph id="F64F36B1FFA6B166E7DCFAF6F789FA93" blockId="30.[189,964,1373,1399]" box="[189,964,1373,1399]" pageId="30" pageNumber="31">
<smallCapsWord id="F0A9A06DFFA6B166E7DCFAF6F544FA91" baselines="1392,1392" box="[189,265,1373,1399]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="30" pageNumber="31">COLOR</smallCapsWord>
(
<figureCitation id="6ECB2A34FFA6B166E67AFAF6F5E4FA93" box="[283,425,1373,1399]" captionStart="Fig" captionStartId="31.[189,232,1928,1954]" captionTargetBox="[195,1395,608,1870]" captionTargetId="figure-79@31.[189,1398,604,1880]" captionTargetPageId="31" captionText="Fig. 11. Megaprosternum nuaulu sp. nov., holotype, ♀ (RMNH). A–B. Habitus. A. Lateral view. B. Dorsal view. C. Head, dorsal view. D–E. Mesosoma. D. Dorsal view. E. Ventral view. Scale bars: A–B = 250 μm; C–E = 125 μm." figureDoi="http://doi.org/10.5281/zenodo.13758973" httpUri="https://zenodo.org/record/13758973/files/figure.png" pageId="30" pageNumber="31">Fig. 11A–B</figureCitation>
). Head, mesosoma and metasoma castaneous.
</paragraph>
<paragraph id="F64F36B1FFA6B166E7DCFA35F084F9A6" blockId="30.[188,1399,1438,1603]" pageId="30" pageNumber="31">
<smallCapsWord id="F0A9A06DFFA6B166E7DCFA35F4B1FA52" baselines="1457,1457" box="[189,252,1438,1464]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="30" pageNumber="31">HEAD</smallCapsWord>
(
<figureCitation id="6ECB2A34FFA6B166E66FFA35F53BFA5C" box="[270,374,1438,1464]" captionStart="Fig" captionStartId="31.[189,232,1928,1954]" captionTargetBox="[195,1395,608,1870]" captionTargetId="figure-79@31.[189,1398,604,1880]" captionTargetPageId="31" captionText="Fig. 11. Megaprosternum nuaulu sp. nov., holotype, ♀ (RMNH). A–B. Habitus. A. Lateral view. B. Dorsal view. C. Head, dorsal view. D–E. Mesosoma. D. Dorsal view. E. Ventral view. Scale bars: A–B = 250 μm; C–E = 125 μm." figureDoi="http://doi.org/10.5281/zenodo.13758973" httpUri="https://zenodo.org/record/13758973/files/figure.png" pageId="30" pageNumber="31">Fig. 11C</figureCitation>
). Oval, in lateral view; sides almost parallel, in dorsal view; malar space absent; median clypeal lobe incurved, as long as lateral ones, with pair of lateral carinae, converging posteriorly, lobe delimitation indistinct; mandible with three apical teeth; hypostoma rounded medially; antenna with 11 flagellomeres, pedicel longer than flagellomere I; eye glabrous, contour not protruding; frons smooth coriaceous; ocellar triangle with anterior angle acute, postocellar line shorter than DAO.
</paragraph>
<paragraph id="F64F36B1FFA6B167E7DCF9C1F55BFE8C" blockId="30.[187,1400,1641,2014]" lastBlockId="31.[189,1399,265,360]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<smallCapsWord id="F0A9A06DFFA6B166E7DCF9C1F577F966" baselines="1661,1661" box="[189,314,1642,1668]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesosoma" pageId="30" pageNumber="31">MESOSOMA</smallCapsWord>
(
<figureCitation id="6ECB2A34FFA6B166E633F9C2F5ABF967" box="[338,486,1641,1667]" captionStart="Fig" captionStartId="31.[189,232,1928,1954]" captionTargetBox="[195,1395,608,1870]" captionTargetId="figure-79@31.[189,1398,604,1880]" captionTargetPageId="31" captionText="Fig. 11. Megaprosternum nuaulu sp. nov., holotype, ♀ (RMNH). A–B. Habitus. A. Lateral view. B. Dorsal view. C. Head, dorsal view. D–E. Mesosoma. D. Dorsal view. E. Ventral view. Scale bars: A–B = 250 μm; C–E = 125 μm." figureDoi="http://doi.org/10.5281/zenodo.13758973" httpUri="https://zenodo.org/record/13758973/files/figure.png" pageId="30" pageNumber="31">Fig. 11D–E</figureCitation>
). Pronotal flange polished and coriaceous, with posterior margin at most 1.5× as wide as anterior one; dorsal pronotal area polished, posterior margin outcurved medially, overlapping anterior area of mesoscutum; probasisternum very large, anterior margin strongly angulated, posterior margin curved; epicnemium with anterior margin weakly incurved; mesoscutum longer than mesoscutellum medially; parapsidal signum absent; transscutal fissure conspicuous; mesoscuto-scutellar suture absent; mesopleuron with mesepimeral lobe not evident, anterior mesopleural fovea present, mesopleural epicoxal lobe not evident, mesopleural pit present, upper mesopleural fovea absent; metapectal-propodeal disc almost as long as wide medially, without evident constriction at propodeal spiracle; transverse anterior carina present; lateral marginal carina absent; paraspiracular sulcus absent; paraspiracular carina absent; metapostnotal median carina absent; propodeal spiracle circular, on lateral surface of the metapectal-propodeal complex; metadiscrimen absent; metafurcal pit absent; forewing with prestigmal abscissa of radial 1 triangular; 4.0× as long as pterostigma; 3.0×as wide as Sc+R vein; prestigmal flexion line present; pterostigma small and oval; hind wing with three non-equidistant distal hamuli.
</paragraph>
<paragraph id="F64F36B1FFA7B167E7DCFE3BF5C5FE0A" blockId="31.[189,1400,399,494]" pageId="31" pageNumber="32">
<smallCapsWord id="F0A9A06DFFA7B167E7DCFE3BF574FE4C" baselines="419,419" box="[189,313,400,426]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metasoma" pageId="31" pageNumber="32">METASOMA</smallCapsWord>
. S2aa with anterior margin incurved; S2pa longer than wide medially, evenly wide; second abdominal spiracle oval; third abdominal sternum with anterior margin outcurved; abdominal tergum widening apicad.
</paragraph>
<caption id="A28F6639FFA7B167E7DCF823F679F80D" ID-DOI="http://doi.org/10.5281/zenodo.13758973" ID-Zenodo-Dep="13758973" httpUri="https://zenodo.org/record/13758973/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1928,1954]" targetBox="[195,1395,608,1870]" targetPageId="31">
<paragraph id="F64F36B1FFA7B167E7DCF823F679F80D" blockId="31.[189,1399,1928,2025]" pageId="31" pageNumber="32">
<emphasis id="C484EAA3FFA7B167E7DCF823F552F846" bold="true" box="[189,287,1928,1954]" pageId="31" pageNumber="32">Fig. 11.</emphasis>
<taxonomicName id="31F04D32FFA7B167E64AF822F61EF846" authorityName="Colombo &amp; Azevedo" authorityYear="2024" box="[299,595,1928,1954]" class="Insecta" family="Bethylidae" genus="Megaprosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nuaulu" status="sp. nov.">
<emphasis id="C484EAA3FFA7B167E64AF822F61EF846" box="[299,595,1928,1954]" italics="true" pageId="31" pageNumber="32">Megaprosternum nuaulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFB757D8FFA7B167E501F823F68EF846" box="[608,707,1928,1954]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (RMNH).
<emphasis id="C484EAA3FFA7B167E492F823F067F846" bold="true" box="[1011,1066,1928,1954]" pageId="31" pageNumber="32">A–B</emphasis>
. Habitus.
<emphasis id="C484EAA3FFA7B167E3CFF823F08EF846" bold="true" box="[1198,1219,1928,1954]" pageId="31" pageNumber="32">A</emphasis>
. Lateral view.
<emphasis id="C484EAA3FFA7B167E7DCF800F49CF821" bold="true" box="[189,209,1963,1989]" pageId="31" pageNumber="32">B</emphasis>
. Dorsal view.
<emphasis id="C484EAA3FFA7B167E6E2F800F5DBF821" bold="true" box="[387,406,1963,1989]" pageId="31" pageNumber="32">C</emphasis>
. Head, dorsal view.
<emphasis id="C484EAA3FFA7B167E5F1F800F68BF821" bold="true" box="[656,710,1963,1989]" pageId="31" pageNumber="32">D–E</emphasis>
. Mesosoma.
<emphasis id="C484EAA3FFA7B167E407F800F736F821" bold="true" box="[870,891,1963,1989]" pageId="31" pageNumber="32">D</emphasis>
. Dorsal view.
<emphasis id="C484EAA3FFA7B167E34CF800F072F821" bold="true" box="[1069,1087,1963,1989]" pageId="31" pageNumber="32">E</emphasis>
. Ventral view. Scale bars: A–B = 250 μm; C–E = 125 μm.
</paragraph>
</caption>
<paragraph id="F64F36B1FF98B158E78BFEA2F567FEC7" blockId="32.[189,313,265,326]" box="[234,298,265,291]" pageId="32" pageNumber="33">
<emphasis id="C484EAA3FF98B158E78BFEA2F567FEC7" bold="true" box="[234,298,265,291]" pageId="32" pageNumber="33">Male</emphasis>
</paragraph>
<paragraph id="F64F36B1FF98B158E7DCFE87F574FEA1" blockId="32.[189,313,265,326]" box="[189,313,299,326]" pageId="32" pageNumber="33">Unknown.</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFF98B158E7DCFEC6F574FE51" box="[189,313,365,437]" pageId="32" pageNumber="33" type="biology_ecology">
<paragraph id="F64F36B1FF98B158E7DCFEC6F4B5FE63" blockId="32.[189,248,365,391]" box="[189,248,365,391]" pageId="32" pageNumber="33">
<heading id="AD0781DDFF98B158E7DCFEC6F4B5FE63" bold="true" box="[189,248,365,391]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="C484EAA3FF98B158E7DCFEC6F4B5FE63" bold="true" box="[189,248,365,391]" pageId="32" pageNumber="33">Host</emphasis>
</heading>
</paragraph>
<paragraph id="F64F36B1FF98B158E7DCFE30F574FE51" blockId="32.[189,313,411,437]" box="[189,313,411,437]" pageId="32" pageNumber="33">Unknown.</paragraph>
</subSubSection>
<subSubSection id="BEEA653AFF98B158E7DCFE77F5A1FDC1" box="[189,492,476,549]" pageId="32" pageNumber="33" type="distribution">
<paragraph id="F64F36B1FF98B158E7DCFE77F51BFE12" blockId="32.[189,342,476,502]" box="[189,342,476,502]" pageId="32" pageNumber="33">
<heading id="AD0781DDFF98B158E7DCFE77F51BFE12" bold="true" box="[189,342,476,502]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="C484EAA3FF98B158E7DCFE77F51BFE12" bold="true" box="[189,342,476,502]" pageId="32" pageNumber="33">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="F64F36B1FF98B158E7DCFDA0F5A1FDC1" blockId="32.[189,492,523,549]" box="[189,492,523,549]" pageId="32" pageNumber="33">
<collectingCountry id="8EE77621FF98B158E7DCFDA0F562FDC1" box="[189,303,523,549]" name="Indonesia" pageId="32" pageNumber="33">Indonesia</collectingCountry>
(North Ceram).
</paragraph>
</subSubSection>
</treatment>
</document>
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